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<document id="E2B8BE49A906C278FAB94B4FE5C3AB60" ID-CLB-Dataset="198291" ID-DOI="10.11646/palaeoentomology.5.5.7" ID-GBIF-Dataset="8de17f19-6f61-45d9-b227-0e9c0a2f53f8" ID-ISSN="2624-2834" ID-Zenodo-Dep="7333739" ID-ZooBank="70EB2402-2833-46F1-9244-E7BFEFA564D8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668767170269" checkinUser="plazi" docAuthor="PEÑALVER, ENRIQUE, ARILLO, ANTONIO &amp; NEL, ANDRÉ" docDate="2022" docId="03EB87B7FFF2DF4051D7A898FB1BF9F9" docLanguage="en" docName="palaeoentomology.5.5.7.pdf" docOrigin="Palaeoentomology 5 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.5.7" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Eltxo grimaldii PEÑALVER &amp; ARILLO &amp; NEL 2022, sp. nov." docType="treatment" docUuid="8A273D67-2B1B-4798-A67B-5D2E8AFE245B" docUuidSource="ZooBank" docVersion="7" lastPageNumber="462" masterDocId="FFD2FFCFFFF3DF41515FAD3DFF8DFF8A" masterDocTitle="A review of the Cretaceous genus Eltxo (Diptera: Cecidomyiidae) with description of the new species Eltxo grimaldii from El Soplao amber" masterLastPageNumber="467" masterPageNumber="461" pageNumber="462" updateTime="1699411768036" updateUser="ExternalLinkService">
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<mods:title id="561C51C50A7F85A089D4145C6726DABA">A review of the Cretaceous genus Eltxo (Diptera: Cecidomyiidae) with description of the new species Eltxo grimaldii from El Soplao amber</mods:title>
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<mods:namePart id="00394988BD44E7E5A466FD52C76B1545">PEÑALVER, ENRIQUE</mods:namePart>
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<mods:namePart id="8F2135DC7BA63F6D2A9F418D5196B9B2">ARILLO, ANTONIO</mods:namePart>
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<mods:namePart id="10BAD77571669D0DBC2072006A11226D">NEL, ANDRÉ</mods:namePart>
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<mods:title id="395FC39407173D0DA93FD7E07DC34F2F">Palaeoentomology</mods:title>
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<mods:date id="B3516D133A3B5F1679788856BF177E05">2022</mods:date>
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<treatment id="03EB87B7FFF2DF4051D7A898FB1BF9F9" ID-DOI="http://doi.org/10.5281/zenodo.7383860" ID-GBIF-Taxon="205276666" ID-Zenodo-Dep="7383860" LSID="urn:lsid:zoobank.org:act:8A273D67-2B1B-4798-A67B-5D2E8AFE245B" httpUri="http://treatment.plazi.org/id/03EB87B7FFF2DF4051D7A898FB1BF9F9" lastPageNumber="462" pageId="1" pageNumber="462">
<subSubSection id="C358652AFFF2DF4051D7A898FE15FA4A" box="[136,408,1445,1472]" pageId="1" pageNumber="462" type="nomenclature">
<paragraph id="8BFD36A1FFF2DF4051D7A898FE15FA4A" blockId="1.[136,778,1445,1582]" box="[136,408,1445,1472]" pageId="1" pageNumber="462">
<heading id="D0B581CDFFF2DF4051D7A898FE15FA4A" bold="true" box="[136,408,1445,1472]" fontSize="10" level="2" pageId="1" pageNumber="462" reason="3">
<emphasis id="B936EAB3FFF2DF4051D7A898FE15FA4A" bold="true" box="[136,408,1445,1472]" pageId="1" pageNumber="462">
<taxonomicName id="4C424D22FFF2DF4051D7A898FEBAFA4A" ID-CoL="9J6LJ" authority="PEÑALVER &amp; ARILLO &amp; NEL, 2022" authorityName="PEÑALVER &amp; ARILLO &amp; NEL" authorityYear="2022" box="[136,311,1445,1472]" class="Insecta" family="Cecidomyiidae" genus="Eltxo" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="462" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
<emphasis id="B936EAB3FFF2DF4051D7A898FEBAFA4A" bold="true" box="[136,311,1445,1472]" italics="true" pageId="1" pageNumber="462">Eltxo grimaldii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557C8FFF2DF405061A89BFE15FA4A" box="[318,408,1446,1472]" pageId="1" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C358652AFFF2DF4051D7A8F7FE18F9A4" pageId="1" pageNumber="462" type="description">
<paragraph id="8BFD36A1FFF2DF4051D7A8F7FF71FA6E" blockId="1.[136,778,1445,1582]" box="[136,252,1482,1508]" pageId="1" pageNumber="462">
(
<figureCitation id="13792A24FFF2DF4051CFA8F7FF5AFA6E" box="[144,215,1482,1508]" captionStart="FIGURE 1" captionStartId="2.[136,235,1919,1943]" captionTargetBox="[151,1436,189,1896]" captionTargetId="figure-22@2.[151,1436,189,1896]" captionTargetPageId="2" captionText="FIGURE 1. Micrographs of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Habitus in lateral view. B, Detail of costal wing margin. C, Detail of wing margin close to end of CuA. D, Detail of head in dorso-lateral view. E, Palpi. F, Detail of basal part of left wing. G, Detail of pretarsal claws of hind leg. H, Detail of hind tarsus. Abbreviations: p = pedicel, s = scape. Images A, D, F and H composed on photographs taken at different focal planes." figureDoi="http://doi.org/10.5281/zenodo.7333741" httpUri="https://zenodo.org/record/7333741/files/figure.png" pageId="1" pageNumber="462">Figs 1</figureCitation>
,
<figureCitation id="13792A24FFF2DF4051BAA8F7FF79FA6E" box="[229,244,1482,1508]" captionStart="FIGURE 2" captionStartId="3.[136,235,1409,1433]" captionTargetBox="[177,1406,197,1385]" captionTargetId="figure-242@3.[151,1436,189,1385]" captionTargetPageId="3" captionText="FIGURE 2. Camera lucida drawings of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Head in ventro-lateral view. B, Palpi enlarged from A. C, Left wing as preserved (all macrotrichia drawn, microtrichia only drawn in some zones of wing membrane). D, Hind tarsus. Abbreviations: C = costal vein, CuA = cubitus anterior, M = median vein, p = pedicel, R1R5 = radial veins, Rs = radial sector, s = scape, Sc = Subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.7333743" httpUri="https://zenodo.org/record/7333743/files/figure.png" pageId="1" pageNumber="462">2</figureCitation>
)
</paragraph>
<paragraph id="8BFD36A1FFF2DF4051D7A8D2FE18F9A4" blockId="1.[136,778,1445,1582]" pageId="1" pageNumber="462">
LSID:
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C358652AFFF2DF4051D7AB61FF60F869" pageId="1" pageNumber="462" type="materials_examined">
<paragraph id="8BFD36A1FFF2DF4051D7AB61FF60F869" blockId="1.[136,778,1628,2093]" pageId="1" pageNumber="462">
<materialsCitation id="3B2A3CFCFFF2DF4051D7AB61FF60F869" ID-GBIF-Occurrence="3978769301" collectionCode="ES-" country="Spain" county="Cantabria" location="Cantabria" pageId="1" pageNumber="462" specimenCode="07-40" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B936EAB3FFF2DF4051D7AB61FF71F9FC" bold="true" box="[136,252,1628,1654]" pageId="1" pageNumber="462">
<typeStatus id="54F98803FFF2DF4051D7AB61FF7AF9FC" box="[136,247,1628,1654]" pageId="1" pageNumber="462" type="holotype">Holotype</typeStatus>
.
</emphasis>
Specimen
<collectionCode id="ED53AE64FFF2DF4050C5AB61FE4BF9FC" box="[410,454,1628,1654]" pageId="1" pageNumber="462">ES-</collectionCode>
<specimenCode id="DBE49EDAFFF2DF40509AAB61FD85F9FC" box="[453,520,1628,1654]" pageId="1" pageNumber="462">07-40</specimenCode>
,
<specimenCount id="9D44FD28FFF2DF40537BAB61FDFEF9FC" box="[548,627,1628,1654]" pageId="1" pageNumber="462" type="female">female</specimenCount>
, housed at the Institutional Collection from the El Soplao Cave (Government of Cantabria), Celis,
<collectingCounty id="629C4E2DFFF2DF40537BAB98FD19F935" box="[548,660,1701,1727]" pageId="1" pageNumber="462">Cantabria</collectingCounty>
, Northern
<collectingCountry id="F3557631FFF2DF4051D7ABF7FF46F96E" box="[136,203,1738,1764]" pageId="1" pageNumber="462">Spain</collectingCountry>
. It is virtually complete but lacking the apices of the two antennae and distal parts of all the legs except one of the metathoracic legs, due to the limits of the amber fragment that contains it. The specimen is in a small fragment of amber, 4 × 3 ×
<quantity id="4CBA9B44FFF2DF40509BAA61FD8BF8FC" box="[452,518,1884,1910]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="462" unit="mm" value="1.0">1 mm</quantity>
in size, and embedded in a regular prism of epoxy resin, 23 × 9 ×
<quantity id="4CBA9B44FFF2DF40532EAABDFD3EF811" box="[625,691,1920,1947]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="462" unit="mm" value="1.0">1 mm</quantity>
in size, for optimal viewing and curation. No syninclusions were detected.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C358652AFFF2DF4051E2AAD3FC84F7A6" pageId="1" pageNumber="462" type="etymology">
<paragraph id="8BFD36A1FFF2DF4051E2AAD3FC84F7A6" blockId="1.[136,778,1628,2093]" pageId="1" pageNumber="462">
<emphasis id="B936EAB3FFF2DF4051E2AAD3FEC9F782" bold="true" box="[189,324,2030,2056]" pageId="1" pageNumber="462">Etymology.</emphasis>
Honoring Dr David Grimaldi (AMNH), for his relevant contributions on fossil insects and amber.
</paragraph>
</subSubSection>
<subSubSection id="C358652AFFF2DF405200ADA3FCFEFDAD" pageId="1" pageNumber="462" type="diagnosis">
<paragraph id="8BFD36A1FFF2DF405200ADA3FC4EFE1F" blockId="1.[810,1452,158,1651]" pageId="1" pageNumber="462">
<emphasis id="B936EAB3FFF2DF405200ADA3FC54FF33" bold="true" box="[863,985,158,185]" pageId="1" pageNumber="462">Diagnosis.</emphasis>
Female. The new species differs from the only species described,
<taxonomicName id="4C424D22FFF2DF40556DADFEFB53FF57" authorityName="Arillo &amp; Nel" authorityYear="2000" box="[1074,1246,195,221]" class="Insecta" family="Cecidomyiidae" genus="Eltxo" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="462" phylum="Arthropoda" rank="species" species="cretaceus">
<emphasis id="B936EAB3FFF2DF40556DADFEFB53FF57" box="[1074,1246,195,221]" italics="true" pageId="1" pageNumber="462">Eltxo cretaceus</emphasis>
</taxonomicName>
(which was based on a unique male), by the following features: distal section of R
<subScript id="17C634E4FFF2DF405203AC22FCE9FEA7" attach="left" box="[860,868,287,301]" fontSize="6" pageId="1" pageNumber="462">1</subScript>
vein elongate, vein M present in a short basal part and evanescent, arculus vein contacting M very basal and CuA vein contacting wing margin distad level of basal section of Rs.
</paragraph>
<paragraph id="8BFD36A1FFF2DF405200ACA2FCFEFDAD" blockId="1.[810,1452,158,1651]" pageId="1" pageNumber="462">
<emphasis id="B936EAB3FFF2DF405200ACA2FBD0FE33" bold="true" box="[863,1117,415,441]" pageId="1" pageNumber="462">Locality and horizon.</emphasis>
Middle Albian amber outcrop of Rábago-El Soplao (near Rábago village,
<collectingRegion id="4986F843FFF2DF405468ACFEFA2AFE57" box="[1335,1447,451,477]" country="Spain" name="Cantabria" pageId="1" pageNumber="462">Cantabria</collectingRegion>
, northern
<collectingCountry id="F3557631FFF2DF4052C6ACD5FC50FD88" box="[921,989,488,514]" name="Spain" pageId="1" pageNumber="462">Spain</collectingCountry>
), Peñosas Fm. (
<bibRefCitation id="EFD34B50FFF2DF4055F5ACD5FA2AFD88" author="Najarro, M. &amp; Penalver, E. &amp; Rosales, I. &amp; Perez-de &amp; la Fuente, R. &amp; Daviero-Gomez, V. &amp; Gomez, B. &amp; Delclos, X." box="[1194,1447,488,515]" pageId="1" pageNumber="462" pagination="363 - 387" refId="ref3954" refString="Najarro, M., Penalver, E., Rosales, I., Perez-de la Fuente, R., Daviero-Gomez, V., Gomez, B. &amp; Delclos, X. (2009) Unusual concentration of Early Albian arthropod - bearing amber in the Basque-Cantabrian Basin (El Soplao, Cantabria, northern Spain): Palaeoenvironmental and palaeobiological implications. Geologica Acta, 7, 363 - 387. https: // doi. org / 10.1344 / 105.000001443" type="journal article" year="2009">
Najarro
<emphasis id="B936EAB3FFF2DF40544CACD4FAD8FD88" box="[1299,1365,488,514]" italics="true" pageId="1" pageNumber="462">et al.</emphasis>
, 2009
</bibRefCitation>
,
<bibRefCitation id="EFD34B50FFF2DF405275AF30FCEAFDAD" author="Najarro, M. &amp; Penalver, E. &amp; Rosales, I. &amp; Soriano, C. &amp; Perez-de &amp; la Fuente, R. &amp; Menor-Salvan, C. &amp; Valle, R. &amp; Blanco, J. &amp; Delclos, X." box="[810,871,525,551]" pageId="1" pageNumber="462" pagination="959 - 976" refId="ref4040" refString="Najarro, M., Penalver, E., Rosales, I., Soriano, C., Perez-de la Fuente, R., Menor-Salvan, C., Lopez del Valle, R., Ortega- Blanco, J. &amp; Delclos, X. (2010) A review of the El Soplao amber outcrop, Early Cretaceous of Cantabria (Spain). Acta Geologica Sinica (English Edition), 84, 959 - 976. https: // doi. org / 10.1111 / j. 1755 - 6724.2010.00258. x" type="journal article" year="2010">2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C358652AFFF2DF405200AF0CFB1BF9F9" pageId="1" pageNumber="462" type="description">
<paragraph id="8BFD36A1FFF2DF405200AF0CFB1BF9F9" blockId="1.[810,1452,158,1651]" pageId="1" pageNumber="462">
<emphasis id="B936EAB3FFF2DF405200AF0CFC7CFDC1" bold="true" box="[863,1009,561,587]" pageId="1" pageNumber="462">Description.</emphasis>
Female (
<figureCitation id="13792A24FFF2DF40553BAF0CFB3DFDC6" box="[1124,1200,561,588]" captionStart="FIGURE 1" captionStartId="2.[136,235,1919,1943]" captionTargetBox="[151,1436,189,1896]" captionTargetId="figure-22@2.[151,1436,189,1896]" captionTargetPageId="2" captionText="FIGURE 1. Micrographs of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Habitus in lateral view. B, Detail of costal wing margin. C, Detail of wing margin close to end of CuA. D, Detail of head in dorso-lateral view. E, Palpi. F, Detail of basal part of left wing. G, Detail of pretarsal claws of hind leg. H, Detail of hind tarsus. Abbreviations: p = pedicel, s = scape. Images A, D, F and H composed on photographs taken at different focal planes." figureDoi="http://doi.org/10.5281/zenodo.7333741" httpUri="https://zenodo.org/record/7333741/files/figure.png" pageId="1" pageNumber="462">Figs 1</figureCitation>
,
<figureCitation id="13792A24FFF2DF40559CAF0FFB59FDC6" box="[1219,1236,562,588]" captionStart="FIGURE 2" captionStartId="3.[136,235,1409,1433]" captionTargetBox="[177,1406,197,1385]" captionTargetId="figure-242@3.[151,1436,189,1385]" captionTargetPageId="3" captionText="FIGURE 2. Camera lucida drawings of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Head in ventro-lateral view. B, Palpi enlarged from A. C, Left wing as preserved (all macrotrichia drawn, microtrichia only drawn in some zones of wing membrane). D, Hind tarsus. Abbreviations: C = costal vein, CuA = cubitus anterior, M = median vein, p = pedicel, R1R5 = radial veins, Rs = radial sector, s = scape, Sc = Subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.7333743" httpUri="https://zenodo.org/record/7333743/files/figure.png" pageId="1" pageNumber="462">2</figureCitation>
). Body length
<emphasis id="B936EAB3FFF2DF4054D5AF0FFA29FDC1" box="[1418,1444,562,587]" italics="true" pageId="1" pageNumber="462">ca</emphasis>
.
<quantity id="4CBA9B44FFF2DF405275AF6BFC07FDFB" box="[810,906,598,625]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="462" unit="mm" value="1.1">1.1 mm</quantity>
. Antennae with only scape, pedicel and 45 flagellomeres preserved due to limits of amber fragment (
<figureCitation id="13792A24FFF2DF40526DAFA2FC1CFD30" box="[818,913,671,698]" captionStart="FIGURE 1" captionStartId="2.[136,235,1919,1943]" captionTargetBox="[151,1436,189,1896]" captionTargetId="figure-22@2.[151,1436,189,1896]" captionTargetPageId="2" captionText="FIGURE 1. Micrographs of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Habitus in lateral view. B, Detail of costal wing margin. C, Detail of wing margin close to end of CuA. D, Detail of head in dorso-lateral view. E, Palpi. F, Detail of basal part of left wing. G, Detail of pretarsal claws of hind leg. H, Detail of hind tarsus. Abbreviations: p = pedicel, s = scape. Images A, D, F and H composed on photographs taken at different focal planes." figureDoi="http://doi.org/10.5281/zenodo.7333741" httpUri="https://zenodo.org/record/7333741/files/figure.png" pageId="1" pageNumber="462">Figs 1D</figureCitation>
,
<figureCitation id="13792A24FFF2DF4052FFAF9DFC4AFD30" box="[928,967,672,698]" captionStart="FIGURE 2" captionStartId="3.[136,235,1409,1433]" captionTargetBox="[177,1406,197,1385]" captionTargetId="figure-242@3.[151,1436,189,1385]" captionTargetPageId="3" captionText="FIGURE 2. Camera lucida drawings of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Head in ventro-lateral view. B, Palpi enlarged from A. C, Left wing as preserved (all macrotrichia drawn, microtrichia only drawn in some zones of wing membrane). D, Hind tarsus. Abbreviations: C = costal vein, CuA = cubitus anterior, M = median vein, p = pedicel, R1R5 = radial veins, Rs = radial sector, s = scape, Sc = Subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.7333743" httpUri="https://zenodo.org/record/7333743/files/figure.png" pageId="1" pageNumber="462">2A</figureCitation>
): preservation of flagellomeres preventing identification of potential sensoria. Ocelli absent. Palpi three-segmented and elongate (
<figureCitation id="13792A24FFF2DF4055EEAFD4FA94FC89" box="[1201,1305,745,771]" captionStart="FIGURE 1" captionStartId="2.[136,235,1919,1943]" captionTargetBox="[151,1436,189,1896]" captionTargetId="figure-22@2.[151,1436,189,1896]" captionTargetPageId="2" captionText="FIGURE 1. Micrographs of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Habitus in lateral view. B, Detail of costal wing margin. C, Detail of wing margin close to end of CuA. D, Detail of head in dorso-lateral view. E, Palpi. F, Detail of basal part of left wing. G, Detail of pretarsal claws of hind leg. H, Detail of hind tarsus. Abbreviations: p = pedicel, s = scape. Images A, D, F and H composed on photographs taken at different focal planes." figureDoi="http://doi.org/10.5281/zenodo.7333741" httpUri="https://zenodo.org/record/7333741/files/figure.png" pageId="1" pageNumber="462">Figs 1E</figureCitation>
,
<figureCitation id="13792A24FFF2DF40546CAFD4FADAFC89" box="[1331,1367,745,771]" captionStart="FIGURE 2" captionStartId="3.[136,235,1409,1433]" captionTargetBox="[177,1406,197,1385]" captionTargetId="figure-242@3.[151,1436,189,1385]" captionTargetPageId="3" captionText="FIGURE 2. Camera lucida drawings of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Head in ventro-lateral view. B, Palpi enlarged from A. C, Left wing as preserved (all macrotrichia drawn, microtrichia only drawn in some zones of wing membrane). D, Hind tarsus. Abbreviations: C = costal vein, CuA = cubitus anterior, M = median vein, p = pedicel, R1R5 = radial veins, Rs = radial sector, s = scape, Sc = Subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.7333743" httpUri="https://zenodo.org/record/7333743/files/figure.png" pageId="1" pageNumber="462">2B</figureCitation>
), with palpomeres subequal, distal palpomere slightly longer than others, palpomeres 2.5 times longer than wide; other mouthpart structures obscure. Wing
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long,
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wide. Wing venation reduced (
<figureCitation id="13792A24FFF2DF4055A7AE46FADDFC1C" box="[1272,1360,891,918]" captionStart="FIGURE 1" captionStartId="2.[136,235,1919,1943]" captionTargetBox="[151,1436,189,1896]" captionTargetId="figure-22@2.[151,1436,189,1896]" captionTargetPageId="2" captionText="FIGURE 1. Micrographs of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Habitus in lateral view. B, Detail of costal wing margin. C, Detail of wing margin close to end of CuA. D, Detail of head in dorso-lateral view. E, Palpi. F, Detail of basal part of left wing. G, Detail of pretarsal claws of hind leg. H, Detail of hind tarsus. Abbreviations: p = pedicel, s = scape. Images A, D, F and H composed on photographs taken at different focal planes." figureDoi="http://doi.org/10.5281/zenodo.7333741" httpUri="https://zenodo.org/record/7333741/files/figure.png" pageId="1" pageNumber="462">Figs 1F</figureCitation>
,
<figureCitation id="13792A24FFF2DF405401AE41FA0EFC1C" box="[1374,1411,892,918]" captionStart="FIGURE 2" captionStartId="3.[136,235,1409,1433]" captionTargetBox="[177,1406,197,1385]" captionTargetId="figure-242@3.[151,1436,189,1385]" captionTargetPageId="3" captionText="FIGURE 2. Camera lucida drawings of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Head in ventro-lateral view. B, Palpi enlarged from A. C, Left wing as preserved (all macrotrichia drawn, microtrichia only drawn in some zones of wing membrane). D, Hind tarsus. Abbreviations: C = costal vein, CuA = cubitus anterior, M = median vein, p = pedicel, R1R5 = radial veins, Rs = radial sector, s = scape, Sc = Subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.7333743" httpUri="https://zenodo.org/record/7333743/files/figure.png" pageId="1" pageNumber="462">2C</figureCitation>
). C extending beyond Rs with a clear end. Apex of R1 at half wing lenght.Apex of Rs meeting C just beyond wing apex. Sc preserved. Basal section of Rs slightly evanescent. Vein M present in a short basal part and evanescent. Vein CuA unforked and evanescent at its apex; apex of CuA well distad level of basal section of Rs. Arculus present, with distal half evanescent, contacting M very close to separation of M and CuA. Wings with dense microtrichia and sparse macrotrichia. Wing margin with setae in costal margin and macrotrichia in posterior margin (but only very few of them still visible) (
<figureCitation id="13792A24FFF2DF40558CA832FADCFAA3" box="[1235,1361,1295,1321]" captionStart="FIGURE 1" captionStartId="2.[136,235,1919,1943]" captionTargetBox="[151,1436,189,1896]" captionTargetId="figure-22@2.[151,1436,189,1896]" captionTargetPageId="2" captionText="FIGURE 1. Micrographs of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Habitus in lateral view. B, Detail of costal wing margin. C, Detail of wing margin close to end of CuA. D, Detail of head in dorso-lateral view. E, Palpi. F, Detail of basal part of left wing. G, Detail of pretarsal claws of hind leg. H, Detail of hind tarsus. Abbreviations: p = pedicel, s = scape. Images A, D, F and H composed on photographs taken at different focal planes." figureDoi="http://doi.org/10.5281/zenodo.7333741" httpUri="https://zenodo.org/record/7333741/files/figure.png" pageId="1" pageNumber="462">Fig. 1B, C</figureCitation>
). Distal portions of legs not preserved due to limits of the amber fragment, except to a complete metathoracic leg, which is long and setose, with five tarsomeres; tarsomere 1 longer than 2 (
<figureCitation id="13792A24FFF2DF4052D3A89CFC7DFA31" box="[908,1008,1441,1467]" captionStart="FIGURE 1" captionStartId="2.[136,235,1919,1943]" captionTargetBox="[151,1436,189,1896]" captionTargetId="figure-22@2.[151,1436,189,1896]" captionTargetPageId="2" captionText="FIGURE 1. Micrographs of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Habitus in lateral view. B, Detail of costal wing margin. C, Detail of wing margin close to end of CuA. D, Detail of head in dorso-lateral view. E, Palpi. F, Detail of basal part of left wing. G, Detail of pretarsal claws of hind leg. H, Detail of hind tarsus. Abbreviations: p = pedicel, s = scape. Images A, D, F and H composed on photographs taken at different focal planes." figureDoi="http://doi.org/10.5281/zenodo.7333741" httpUri="https://zenodo.org/record/7333741/files/figure.png" pageId="1" pageNumber="462">Figs 1H</figureCitation>
,
<figureCitation id="13792A24FFF2DF40555CA89CFBA7FA31" box="[1027,1066,1441,1467]" captionStart="FIGURE 2" captionStartId="3.[136,235,1409,1433]" captionTargetBox="[177,1406,197,1385]" captionTargetId="figure-242@3.[151,1436,189,1385]" captionTargetPageId="3" captionText="FIGURE 2. Camera lucida drawings of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Head in ventro-lateral view. B, Palpi enlarged from A. C, Left wing as preserved (all macrotrichia drawn, microtrichia only drawn in some zones of wing membrane). D, Hind tarsus. Abbreviations: C = costal vein, CuA = cubitus anterior, M = median vein, p = pedicel, R1R5 = radial veins, Rs = radial sector, s = scape, Sc = Subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.7333743" httpUri="https://zenodo.org/record/7333743/files/figure.png" pageId="1" pageNumber="462">2D</figureCitation>
). All tibiae lacking tibial spurs. Pretarsal claws short and with a small basal prolongation with rounded apex; empodium insconspicuous (
<figureCitation id="13792A24FFF2DF40541BA8D6FA13F98F" box="[1348,1438,1515,1541]" captionStart="FIGURE 1" captionStartId="2.[136,235,1919,1943]" captionTargetBox="[151,1436,189,1896]" captionTargetId="figure-22@2.[151,1436,189,1896]" captionTargetPageId="2" captionText="FIGURE 1. Micrographs of Eltxo grimaldii sp. nov. (Cecidomyiidae), holotype ES-07-40, El Soplao amber (middle Albian). A, Habitus in lateral view. B, Detail of costal wing margin. C, Detail of wing margin close to end of CuA. D, Detail of head in dorso-lateral view. E, Palpi. F, Detail of basal part of left wing. G, Detail of pretarsal claws of hind leg. H, Detail of hind tarsus. Abbreviations: p = pedicel, s = scape. Images A, D, F and H composed on photographs taken at different focal planes." figureDoi="http://doi.org/10.5281/zenodo.7333741" httpUri="https://zenodo.org/record/7333741/files/figure.png" pageId="1" pageNumber="462">Fig. 1G</figureCitation>
). Abdomen
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long (up to the cerci apices). Genitalia obscure, but long ovipositor (
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); contour of the cerci well visible in lateral view.
</paragraph>
</subSubSection>
</treatment>
</document>