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<mods:title id="FA72EE39207071BE5D47C4DE6D397847">Revision of the Central Asian scorpion genus Anomalobuthus Kraepelin, 1900, with descriptions of three new species and a generic synonymy (Scorpiones: Buthidae)</mods:title>
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<mods:namePart id="22132C72871F5BEB6FDA951DE90B1A66">Teruel, Rolando</mods:namePart>
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<paragraph id="8BCE36FEFB7EFFACFCEBF03EFA22ED39" blockId="12.[842,1429,1617,1739]" box="[842,1428,1617,1645]" pageId="12" pageNumber="11">
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<taxonomicName id="4C714D7DFB7EFFACFCEBF03EFA22ED39" ID-CoL="6774Q" authority="Teruel, Kovarik et" authorityName="Teruel, Kovarik et Fet" authorityYear="2018" box="[842,1428,1617,1645]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="krivochatskyi" status="sp. nov.">
<emphasis id="B905EAECFB7EFFACFCEBF03EFB03ED38" bold="true" box="[842,1205,1617,1644]" italics="true" pageId="12" pageNumber="11">Anomalobuthus krivochatskyi</emphasis>
Teruel, Kovařík et
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Fet, 
<emphasis id="B905EAECFB7EFFACFBC9F01BFB1CEDDA" bold="true" box="[1128,1194,1652,1678]" pageId="12" pageNumber="11">sp. n.</emphasis>
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<figureCitation id="134A2A7BFB7EFFACFCEBF0FAFC53EDF8" box="[842,997,1685,1708]" captionStart-0="Figures 34–37" captionStart-1="Figures 38–41" captionStart-2="Figures 42–45" captionStart-3="Figures 46–53" captionStart-4="Figures 54–58" captionStart-5="Figures 59–64" captionStartId-0="13.[192,278,1630,1653]" captionStartId-1="14.[192,278,1754,1777]" captionStartId-2="16.[192,278,1584,1607]" captionStartId-3="17.[192,278,1398,1421]" captionStartId-4="18.[192,278,1378,1401]" captionStartId-5="19.[192,278,1542,1565]" captionTargetBox-0="[212,1415,153,1571]" captionTargetBox-1="[201,1418,199,1682]" captionTargetBox-2="[212,1422,194,1547]" captionTargetBox-3="[191,1437,191,1368]" captionTargetBox-4="[204,1423,190,1328]" captionTargetBox-5="[204,1417,183,1488]" captionTargetId-0="figure-6@13.[192,1439,144,1601]" captionTargetId-1="figure-12@14.[192,1439,192,1697]" captionTargetId-2="figure-12@16.[192,1440,192,1554]" captionTargetId-3="figure-6@17.[192,1437,192,1367]" captionTargetId-4="figure-12@18.[192,1437,168,1348]" captionTargetId-5="figure-6@19.[192,1437,168,1512]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="16" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="19" captionText-0="Figures 34–37: Anomalobuthus krivochatskyi sp. n., adult male holotype (34–35) and adult female paratopotype (36–37): fullbody views, dorsal (34, 36) and ventral (35, 37). Scale bar = 10 mm." captionText-1="Figures 38–41: Anomalobuthus krivochatskyi sp. n., adult male (38–39) and adult female paratypes (40–41) from between Bukhara and Gazli, Uzbekistan: full-body views, dorsal (38, 40) and ventral (39, 41). Scale bar = 10 mm." captionText-2="Figures 42–45: Anomalobuthus krivochatskyi sp. n., adult female (42–43) and juvenile male paratypes (44–45) from Chardara, Kazakhstan: full-body views, dorsal (42, 44) and ventral (43, 45)." captionText-3="Figures 46–53: Anomalobuthus krivochatskyi sp. n., adult male holotype (46–47) and adult female paratopotype (48–53): chelicerae, carapace and tergites I–III, dorsal (46, 48), sternopectinal region and sternites III–V, ventral (47, 49), distal part of right legs I–IV, internal (50–53)." captionText-4="Figures 54–58: Anomalobuthus krivochatskyi sp. n., adult female paratopotype: right pedipalp trochanter and femur, dorsal (54), right pedipalp patella, dorsal (55) and external (56), right pedipalp chela, dorsoexternal (57), right pedipalp movable finger, dorsoexternal (58). Black dots depict trichobothria." captionText-5="Figures 59–64: Anomalobuthus krivochatskyi sp. n., adult male holotype (59–61) and adult female paratopotype (62–64): metasoma and telson, lateral (59, 62), dorsal (60, 63) and ventral (61, 64)." pageId="12" pageNumber="11">Figures 34–64</figureCitation>
, 
<figureCitation id="134A2A7BFB7EFFACFC51F0FAFBE6EDF8" box="[1008,1104,1685,1708]" captionStart="Figures 120–125" captionStartId="34.[192,278,1734,1757]" captionTargetBox="[191,1440,200,1705]" captionTargetId="figure-12@34.[192,1439,201,1704]" captionTargetPageId="34" captionText="Figures 120–125: Anomalobuthus krivochatskyi sp. n., adult male paratopotype: SEM views of chelicerae, dorsal and ventral (120), metasoma V and telson, lateral (121); anterior half of carapace, dorsal (122); close-up of trichobothria (123); left leg IV basitarsus andtelotarsus, ventrointernal (124); left leg IV basitarsus and telotarsus, external (125). Images courtesy Jan Štundl." pageId="12" pageNumber="11">120–125</figureCitation>
, 
<figureCitation id="134A2A7BFB7EFFACFBFAF0FAFB0DEDF8" box="[1115,1211,1685,1708]" captionStart="Figures 126–133" captionStartId="36.[192,278,1688,1711]" captionTargetBox="[191,1440,167,1659]" captionTargetId="figure-12@36.[192,1439,168,1658]" captionTargetPageId="36" captionText="Figures 126–133: Anomalobuthus spp., SEM views of right pedipalp of adult male A. rickmersi from Repetek (126–127) and adult male paratopotype A. krivochatskyi sp. n. (128–133): chela (126, 128), close-up of manus (127, 129), three ventrointernal magnified views of mid part of both fingers in consecutive figures (130–132, movable finger on top), ventrointernal close-up of fingertips (133, movable finger on top). See trichobothria highlighted in red." pageId="12" pageNumber="11">128–133</figureCitation>
, 
<figureCitation id="134A2A7BFB7EFFACFB67F0FAFA27EDF8" box="[1222,1425,1685,1708]" captionStart-0="Figures 134–140" captionStart-1="Figure 141" captionStartId-0="37.[192,278,1748,1771]" captionStartId-1="40.[192,268,1144,1167]" captionTargetBox-0="[191,1440,191,1719]" captionTargetBox-1="[191,1440,191,1114]" captionTargetId-0="figure-6@37.[192,1439,192,1718]" captionTargetId-1="figure-12@40.[192,1439,192,1114]" captionTargetPageId-0="37" captionTargetPageId-1="40" captionText-0="Figures 134–140: Anomalobuthus spp., SEM views showing size variation of leg IV tibial spur (highlighted in red): large in paratopotype female A. krivochatskyi sp. n. (134), moderately large in paratype female A. krivochatskyi sp. n. from Bukhara (135), small in paratopotype female A. krivochatskyi sp. n. (136), absent in male A. rickmersi from Repetek (137), close-up of a medium-sized leg IV tibial spur of a paratopotype female A. krivochatskyi sp. n. from Zarafshan (138), see spinulate and striate sculpture. Anomalobuthus krivochatskyi sp. n., adult female paratopotype: SEM views of right pecten and adjacent parts of leg coxae and sternite III (139), peg sensilla of tooth from same pecten (140)." captionText-1="Figure 141: Distribution of the genus Anomalobuthus: A. krivochatskyi sp. n. (1), A. lowei sp. n. (2), A. pavlovskyi sp. n. (3), A. rickmersi (4), A. talebii (5), A. zarudnyi comb. n. (6), and literature records whose identification is pending (black squares). Data compiled from specimens studied herein and the following supplementary sources: Rickmer Rickmers (1913), Fet (1989, 1994), Gromov &amp; Kopdykbaev (1994), Gromov (2006), Mityaev et al. (2008), and Graham et al. (2012)." pageId="12" pageNumber="11">134–136, 138–141</figureCitation>
. 
<tableCitation id="C6F30345FB7EFFACFB86F0DBFB32ED9F" box="[1063,1156,1716,1739]" captionStart="Table 2" captionStartId="20.[192,257,1203,1226]" captionText="Table 2: Measurements of four types of Anomalobuthus krivochatskyi sp. n. Abbreviations: length (L), width (W), posterior width (Wp), depth (D)." pageId="12" pageNumber="11">Tables 2</tableCitation>
, 
<tableCitation id="C6F30345FB7EFFACFB31F0DBFB0EED9F" box="[1168,1208,1716,1739]" captionStart-0="Table 4" captionStart-1="Table 5" captionStartId-0="35.[192,257,1030,1053]" captionStartId-1="38.[192,257,680,703]" captionText-0="Table 4: Comparison between five species of Anomalobuthus based upon selected morphometric ratios. Abbreviations: length (L)," captionText-1="Table 5: Pectinal tooth count variation in all known species of Anomalobuthus. Abbreviations: number of pectines (N), average (Ave.), standard deviation (SD). This includes counts taken from sources with precise and reliable geographic data (Birula, 1905, 1910)." pageId="12" pageNumber="11">4–5</tableCitation>
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<emphasis id="B905EAECFB7EFFACFCE5F084FA2CEC56" bold="true" box="[836,1434,1771,1794]" pageId="12" pageNumber="11">http://zoobank.org/urn:lsid:zoobank.org:act:3936A6</emphasis>
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<emphasis id="B905EAECFB7EFFACFC01F165FA88EC75" bold="true" box="[928,1342,1802,1825]" pageId="12" pageNumber="11">8B-96D5-461C-9568-5077BEA67488</emphasis>
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<taxonomicName id="4C714D7DFB7EFFACFC9FF128FAC6EC0A" ID-CoL="5V74N" authority=": Soleglad &amp; Fet, 2003 a: 5" authorityName=": Soleglad &amp; Fet" authorityPageNumber="5" authorityYear="2003" box="[830,1392,1863,1886]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB7EFFACFC9FF128FBE4EC0A" box="[830,1106,1863,1886]" italics="true" pageId="12" pageNumber="11">Anomalobuthus rickmersi</emphasis>
: Soleglad &amp; Fet, 2003a: 5
</taxonomicName>
, fig. 4 (in part; Bukhara); Soleglad &amp; Fet, 2003b: 7, 59, 156, tab. 2 (in part; Bukhara); Fet et al., 2005: 2, 6, 11; fig. 10 (in part; Bukhara); Fet et al., 2006: 2, 10, tab. 3; 
<bibRefCitation id="EFE04B0FFB7FFFADFE9DF09EFDC3EC5C" author="GRAHAM" box="[316,629,1777,1800]" pageId="13" pageNumber="12" pagination="95 - 110" refId="ref24752" refString="GRAHAM, M. R., V. OLAH-HEMMINGS &amp; V. FET. 2012. Phylogeography of co-distributed dune scorpions identifies the Amu Darya River as a longstanding component of Central Asian biogeography (Scorpiones: Buthidae). Zoology in the Middle East, 55: 95 - 110." type="journal article" year="2012">Graham et al., 2012: 95–106</bibRefCitation>
, figs. 1–2, 6 (in part; between Bukhara and Gazli); Loria &amp; Prendini, 2014: S1: 2, S2: 2; 
<bibRefCitation id="EFE04B0FFB7FFFADFBE8F0DBFA9AED9F" author="TERUEL" box="[1097,1324,1716,1739]" pageId="13" pageNumber="12" pagination="1 - 10" refId="ref26171" refString="TERUEL, R., F. KOVARIK, S. NAVIDPOUR &amp; V. FET. 2014. The first record of the genus Anomalobuthus Kraepelin, 1900 from Iran, with description of a new species (Scorpiones: Buthidae). Euscorpius, 192: 1 - 10." type="journal article" year="2014">Teruel et al., 2014: 1</bibRefCitation>
, 4, 6–7, 9, fig. 20, tabs. 1–2 (in part; 
<collectingCountry id="F366766EFB7FFFADFBF5F0BDFB7AEDBD" box="[1108,1228,1746,1769]" name="Uzbekistan" pageId="13" pageNumber="12">Uzbekistan</collectingCountry>
).
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<emphasis id="B905EAECFB7FFFADFF61F031FEDEED21" bold="true" box="[192,360,1630,1653]" pageId="13" pageNumber="12">Figures 34–37:</emphasis>
<taxonomicName id="4C714D7DFB7FFFADFED1F00FFD3BED21" authority="Teruel &amp; Kovařík &amp; Fet, 2018" authorityName="Teruel &amp; Kovařík &amp; Fet" authorityYear="2018" box="[368,653,1632,1653]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="krivochatskyi" status="sp. nov.">
<emphasis id="B905EAECFB7FFFADFED1F00FFD3BED21" box="[368,653,1632,1653]" italics="true" pageId="13" pageNumber="12">Anomalobuthus krivochatskyi</emphasis>
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<taxonomicNameLabel id="A2365797FB7FFFADFD35F00FFD7DED21" box="[660,715,1632,1653]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
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, adult male holotype (34–35) and adult female paratopotype (36–37): fullbody views, dorsal (34, 36) and ventral (35, 37). Scale bar = 10 mm.
</paragraph>
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(
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, 
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, 
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SSE of
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<paragraph id="8BCE36FEFB7CFFAEFF61F0B5FB02EC5E" blockId="14.[192,1440,1754,1802]" pageId="14" pageNumber="13">
<emphasis id="B905EAECFB7CFFAEFF61F0B5FEDAEDA5" bold="true" box="[192,364,1754,1777]" pageId="14" pageNumber="13">Figures 38–41:</emphasis>
<taxonomicName id="4C714D7DFB7CFFAEFED4F0B3FD23EDA5" authority="Teruel &amp; Kovařík &amp; Fet, 2018" authorityName="Teruel &amp; Kovařík &amp; Fet" authorityYear="2018" box="[373,661,1756,1777]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="krivochatskyi" status="sp. nov.">
<emphasis id="B905EAECFB7CFFAEFED4F0B3FD23EDA5" box="[373,661,1756,1777]" italics="true" pageId="14" pageNumber="13">Anomalobuthus krivochatskyi</emphasis>
</taxonomicName>
<emphasis id="B905EAECFB7CFFAEFD3EF0B3FD6EEDA5" bold="true" box="[671,728,1756,1777]" pageId="14" pageNumber="13">
<taxonomicNameLabel id="A2365797FB7CFFAEFD3EF0B3FD6EEDA5" box="[671,728,1756,1777]" pageId="14" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult male (38–39) and adult female paratypes (40–41) from between Bukhara and Gazli, Uzbekistan: full-body views, dorsal (38, 40) and ventral (39, 41). Scale bar = 10 mm.
</paragraph>
</caption>
<paragraph id="8BCE36FEFB7DFFAFFF61F6A8FEF7EBA9" blockId="15.[192,802,199,253]" pageId="15" pageNumber="14">
<materialsCitation id="3B193CA3FB7DFFAFFF61F6A8FE8BEBA9" collectingDate="1998-04-18" collectingDateMax="1998-04-28" collectingDateMin="1998-04-18" collectionCode="V" latitude="41.533333" longLatPrecision="1155" longitude="64.2" pageId="15" pageNumber="14" specimenCount="1">
Zarafshan, 
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<geoCoordinate id="EE455039FB7DFFAFFE3DF6A8FE47EB8A" box="[412,497,199,222]" degrees="64" direction="east" minutes="12" orientation="longitude" pageId="15" pageNumber="14" precision="925" value="64.2">64º12'E</geoCoordinate>
, 
<date id="FFCF103EFB7DFFAFFDA1F6A8FD73EB8A" box="[512,709,199,222]" pageId="15" pageNumber="14" value="1998-04-18" valueMax="1998-04-28" valueMin="1998-04-18">
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</date>
, leg. A. 
<collectionCode id="ED60AE3BFB7DFFAFFF61F689FF60EBA9" box="[192,214,230,253]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="14" type="Museum">V</collectionCode>
. Gromov
</materialsCitation>
.
</paragraph>
<paragraph id="8BCE36FEFB7DFFAFFF51F74CFD6CE91A" blockId="15.[192,802,291,590]" pageId="15" pageNumber="14">
<materialsCitation id="3B193CA3FB7DFFAFFF51F74CFDFFEA0D" collectionCode="FKCP, RTOC" collectorName="Same" country="Uzbekistan" county="District" location="Navoyi Province" municipality="Tamdy" pageId="15" pageNumber="14" specimenCount="8" specimenCount-female="6" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54CA885CFB7DFFAFFF51F74CFEDBEA6E" box="[240,365,291,314]" pageId="15" pageNumber="14" type="paratype">PARATYPES</typeStatus>
. 
<collectorName id="26845328FB7DFFAFFE27F74CFE77EA6E" box="[390,449,291,314]" pageId="15" pageNumber="14">Same</collectorName>
data as holotype, 
<specimenCount id="9D77FD77FB7DFFAFFD13F74CFD65EA6E" box="[690,723,291,314]" pageId="15" pageNumber="14" type="male">1♂</specimenCount>
, 
<specimenCount id="9D77FD77FB7DFFAFFD4DF74CFC94EA6E" box="[748,802,291,314]" pageId="15" pageNumber="14" type="female">5♀♀</specimenCount>
(
<collectionCode id="ED60AE3BFB7DFFAFFF69F72DFEA6EA0D" box="[200,272,322,345]" pageId="15" pageNumber="14">FKCP</collectionCode>
), 
<specimenCount id="9D77FD77FB7DFFAFFE82F72DFEF2EA0D" box="[291,324,322,345]" pageId="15" pageNumber="14" type="male">1♂</specimenCount>
, 
<specimenCount id="9D77FD77FB7DFFAFFEF3F72DFEC2EA0D" box="[338,372,322,345]" pageId="15" pageNumber="14" type="female">1♀</specimenCount>
(
<collectionCode id="ED60AE3BFB7DFFAFFE25F72DFE79EA0D" box="[388,463,322,345]" pageId="15" pageNumber="14">RTOC</collectionCode>
: Sco-0233)
</materialsCitation>
. 
<materialsCitation id="3B193CA3FB7DFFAFFDF9F72DFE55EAA6" collectingDate="1979-04-30" collectingDateMax="2002-05-11" collectingDateMin="1979-04-30" collectionCode="V, NMPC, ZMUH, SOFM, RTOC" collectorName="Fet" country="Uzbekistan" county="Kizylkum Desert" elevation="206" latitude="40.083332" location="Bukhara" longLatPrecision="1165" longitude="64.066666" municipality="Bukhara Province" pageId="15" pageNumber="14" specimenCount="9" specimenCount-female="2" specimenCount-juvenile="6" specimenCount-male="1" stateProvince="Bukhara Province" typeStatus="paratype">
<emphasis id="B905EAECFB7DFFAFFDF9F72DFD61EA0D" bold="true" box="[600,727,322,345]" pageId="15" pageNumber="14">
<collectingCountry id="F366766EFB7DFFAFFDF9F72DFD61EA0D" box="[600,727,322,345]" name="Uzbekistan" pageId="15" pageNumber="14">Uzbekistan</collectingCountry>
</emphasis>
, 
<collectingCounty id="62AF4E72FB7DFFAFFD44F72DFEFDEA23" pageId="15" pageNumber="14">Kizylkum Desert</collectingCounty>
, 
<collectingRegion id="49B5F81CFB7DFFAFFEFBF70FFD94EA23" box="[346,546,352,375]" inferredBySuffix="true" pageId="15" pageNumber="14">
<collectingMunicipality id="6BAAAC84FB7DFFAFFEFBF70FFD94EA23" box="[346,546,352,375]" pageId="15" pageNumber="14">Bukhara Province</collectingMunicipality>
</collectingRegion>
, 
<locationDeviation id="9A7C2FDCFB7DFFAFFD90F70FFF4CEAC2" firstLocation="Bukhara" pageId="15" pageNumber="14" secondLocation="Gazli">between Bukhara and Gazli</locationDeviation>
, 
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<geoCoordinate id="EE455039FB7DFFAFFEC6F710FE0CEAC2" box="[359,442,383,406]" degrees="64" direction="east" minutes="04" orientation="longitude" pageId="15" pageNumber="14" precision="925" value="64.066666">64º04'E</geoCoordinate>
, 
<elevation id="005CD1CDFB7DFFAFFE6BF710FDFFEAC2" box="[458,585,383,406]" metricMagnitude="2" metricUnit="m" metricValue="2.06" pageId="15" pageNumber="14" unit="m" value="206.0">
<quantity id="4C899B1BFB7DFFAFFE6BF710FDB9EAC2" box="[458,527,383,406]" metricMagnitude="2" metricUnit="m" metricValue="2.06" pageId="15" pageNumber="14" unit="m" value="206.0">206 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFCF103EFB7DFFAFFDF8F710FD58EAC2" box="[601,750,383,406]" pageId="15" pageNumber="14" value="2002-05-11">
<collectingDate id="EF8BE9D6FB7DFFAFFDF8F710FD58EAC2" box="[601,750,383,406]" pageId="15" pageNumber="14" value="2002-05-11">11 May 2002</collectingDate>
</date>
, leg. 
<collectionCode id="ED60AE3BFB7DFFAFFF63F7F1FF6EEAE1" box="[194,216,414,437]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="14" type="Museum">V</collectionCode>
. 
<collectorName id="26845328FB7DFFAFFF4EF7F1FEA2EAE1" box="[239,276,414,437]" pageId="15" pageNumber="14">Fet</collectorName>
, 
<specimenCount id="9D77FD77FB7DFFAFFE8AF7F1FEFDEAE1" box="[299,331,414,437]" pageId="15" pageNumber="14" type="male">1♂</specimenCount>
, 
<specimenCount id="9D77FD77FB7DFFAFFEC5F7F1FE30EAE1" box="[356,390,414,437]" pageId="15" pageNumber="14" type="female">1♀</specimenCount>
, 
<specimenCount id="9D77FD77FB7DFFAFFE01F7F1FD96EAE1" box="[416,544,414,437]" pageId="15" pageNumber="14" type="juvenile">4 juveniles</specimenCount>
(
<collectionCode id="ED60AE3BFB7DFFAFFD9AF7F1FD3FEAE1" box="[571,649,414,437]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="15" pageNumber="14">NMPC</collectionCode>
), 
<specimenCount id="9D77FD77FB7DFFAFFD0AF7F1FC94EAE1" box="[683,802,414,437]" pageId="15" pageNumber="14" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="ED60AE3BFB7DFFAFFF66F7D3FEAFEA87" box="[199,281,444,467]" country="Viet Nam" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoological Museum, University of Hanoi" pageId="15" pageNumber="14">ZMUH</collectionCode>
), 
<specimenCount id="9D77FD77FB7DFFAFFE92F7D3FE14EA87" box="[307,418,444,467]" pageId="15" pageNumber="14" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="ED60AE3BFB7DFFAFFE16F7D3FDB6EA87" box="[439,512,444,467]" country="Bulgaria" httpUri="http://grbio.org/cool/dcs4-ffmk" name="National Museum of Natural History, Sofia" pageId="15" pageNumber="14">SOFM</collectionCode>
), 
<specimenCount id="9D77FD77FB7DFFAFFDBAF7D3FD8AEA87" box="[539,572,444,467]" pageId="15" pageNumber="14" type="female">1♀</specimenCount>
(
<collectionCode id="ED60AE3BFB7DFFAFFDF0F7D3FD2AEA87" box="[593,668,444,467]" pageId="15" pageNumber="14">RTOC</collectionCode>
: Sco-0208); 
<location id="8EAE6025FB7DFFAFFF61F7B4FEABEAA6" LSID="urn:lsid:plazi:treatment:03D887E8FB7EFFB5FCEBF03EFDD7EA87:8EAE6025FB7DFFAFFF61F7B4FEABEAA6" box="[192,285,475,498]" country="Uzbekistan" county="Kizylkum Desert" latitude="40.083332" longLatPrecision="1165" longitude="64.066666" municipality="Bukhara Province" name="Bukhara" pageId="15" pageNumber="14" stateProvince="Bukhara Province">Bukhara</location>
, 
<date id="FFCF103EFB7DFFAFFE94F7B4FE55EAA6" box="[309,483,475,498]" pageId="15" pageNumber="14" value="1979-04-30">
<collectingDate id="EF8BE9D6FB7DFFAFFE94F7B4FE55EAA6" box="[309,483,475,498]" pageId="15" pageNumber="14" value="1979-04-30">30 April 1979</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B193CA3FB7DFFAFFE5FF7B4FCA9EAA6" box="[510,799,475,498]" collectionCode="FKCP" collectorName="Antus" country="Uzbekistan" pageId="15" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Bukhara Province" typeStatus="paratype">
leg. 
<collectorName id="26845328FB7DFFAFFD99F7B4FDCCEAA6" box="[568,634,475,498]" pageId="15" pageNumber="14">Antuš</collectorName>
, 
<specimenCount id="9D77FD77FB7DFFAFFD33F7B4FD05EAA6" box="[658,691,475,498]" pageId="15" pageNumber="14" type="female">1♀</specimenCount>
(
<collectionCode id="ED60AE3BFB7DFFAFFD6FF7B4FCA1EAA6" box="[718,791,475,498]" pageId="15" pageNumber="14">FKCP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B193CA3FB7DFFAFFF61F795FD60E91A" collectingDate="2016-05-23" collectingDateMax="2016-05-24" collectingDateMin="2016-05-23" collectionCode="FKCP" collectorName="P. Kucera" country="Kazakhstan" elevation="250" latitude="41.270554" location="Chardara" longLatPrecision="19" longitude="67.88389" pageId="15" pageNumber="14" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" stateProvince="Ongtustik Qazaqstan Oblysy" typeStatus="paratype">
<emphasis id="B905EAECFB7DFFAFFF61F795FEFEE945" bold="true" box="[192,328,506,529]" pageId="15" pageNumber="14">
<collectingCountry id="F366766EFB7DFFAFFF61F795FEFEE945" box="[192,328,506,529]" name="Kazakhstan" pageId="15" pageNumber="14">Kazakhstan</collectingCountry>
</emphasis>
, 
<collectingRegion id="49B5F81CFB7DFFAFFEF9F795FD34E945" box="[344,642,506,529]" country="Kazakhstan" name="Ongtustik Qazaqstan Oblysy" pageId="15" pageNumber="14">South Kazakhstan Province</collectingRegion>
, 
<quantity id="4C899B1BFB7DFFAFFD30F795FD7FE945" box="[657,713,506,529]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="14" unit="km" value="5.0">5 km</quantity>
west of 
<location id="8EAE6025FB7DFFAFFF63F477FE94E97B" LSID="urn:lsid:plazi:treatment:03D887E8FB7EFFB5FCEBF03EFDD7EA87:8EAE6025FB7DFFAFFF63F477FE94E97B" box="[194,290,536,559]" country="Kazakhstan" latitude="41.270554" longLatPrecision="19" longitude="67.88389" name="Chardara" pageId="15" pageNumber="14" stateProvince="Ongtustik Qazaqstan Oblysy">Chardara</location>
, 
<geoCoordinate id="EE455039FB7DFFAFFE95F477FE07E97B" box="[308,433,536,559]" degrees="41" direction="north" minutes="16" orientation="latitude" pageId="15" pageNumber="14" precision="15" seconds="14" value="41.270554">41°16'14&quot;N</geoCoordinate>
<geoCoordinate id="EE455039FB7DFFAFFE1DF477FD8FE97B" box="[444,569,536,559]" degrees="67" direction="east" minutes="53" orientation="longitude" pageId="15" pageNumber="14" precision="15" seconds="02" value="67.88389">67°53'02&quot;E</geoCoordinate>
, 
<elevation id="005CD1CDFB7DFFAFFDE8F477FD7BE97B" box="[585,717,536,559]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="14" unit="m" value="250.0">
<quantity id="4C899B1BFB7DFFAFFDE8F477FD27E97B" box="[585,657,536,559]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="14" unit="m" value="250.0">250 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFCF103EFB7DFFAFFD7EF477FE9BE91A" pageId="15" pageNumber="14" value="2016-05-23" valueMax="2016-05-24" valueMin="2016-05-23">
<collectingDate id="EF8BE9D6FB7DFFAFFD7EF477FE9BE91A" pageId="15" pageNumber="14" value="2016-05-23" valueMax="2016-05-24" valueMin="2016-05-23">23–24 May 2016</collectingDate>
</date>
, leg. 
<collectorName id="26845328FB7DFFAFFEC9F458FE67E91A" box="[360,465,567,590]" pageId="15" pageNumber="14">P. Kučera</collectorName>
, 
<specimenCount id="9D77FD77FB7DFFAFFE7FF458FDB6E91A" box="[478,512,567,590]" pageId="15" pageNumber="14" type="female">1♀</specimenCount>
, 
<specimenCount id="9D77FD77FB7DFFAFFDACF458FDC1E91A" box="[525,631,567,590]" pageId="15" pageNumber="14" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="ED60AE3BFB7DFFAFFD24F458FD78E91A" box="[645,718,567,590]" pageId="15" pageNumber="14">FKCP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36B6575FB7DFFAFFF51F41BFC94E836" pageId="15" pageNumber="14" type="etymology">
<paragraph id="8BCE36FEFB7DFFAFFF51F41BFC94E836" blockId="15.[192,802,628,866]" pageId="15" pageNumber="14">
ETYMOLOGY. We are pleased to name this species after our colleague, Viktor Anatolyevich Krivochatsky (b. 1954) (
<collectingRegion id="49B5F81CFB7DFFAFFE9CF4DDFE6CE99D" box="[317,474,690,713]" country="Russia" name="Saint Petersburg" pageId="15" pageNumber="14">St. Petersburg</collectingRegion>
, 
<collectingCountry id="F366766EFB7DFFAFFE4CF4DDFD8EE99D" box="[493,568,690,713]" name="Russia" pageId="15" pageNumber="14">Russia</collectingCountry>
), an entomologist, a specialist in ant-lions (
<taxonomicName id="4C714D7DFB7DFFAFFE12F4BFFD86E9B3" box="[435,560,720,743]" class="Insecta" kingdom="Animalia" order="Neuroptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="order">Neuroptera</taxonomicName>
: 
<taxonomicName id="4C714D7DFB7DFFAFFD9AF4BFFD5DE9B3" authorityName="Latreille" authorityYear="1802" box="[571,747,720,743]" class="Insecta" family="Myrmeleontidae" kingdom="Animalia" order="Neuroptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="family">Myrmeleontidae</taxonomicName>
) and an old friend of V.F. from the time of their work in the natural reserves of 
<collectingCountry id="F366766EFB7DFFAFFE01F561FD82E871" box="[416,564,782,805]" name="Turkmenistan" pageId="15" pageNumber="14">Turkmenistan</collectingCountry>
in the 1970s-1980s. Since 1991, Dr. Krivochatsky works in ZISP and is in charge of the famed scorpion collection of Alexei Birula.
</paragraph>
</subSubSection>
<subSubSection id="C36B6575FB7DFFAFFF51F5E7FD55EEDE" pageId="15" pageNumber="14" type="diagnosis">
<paragraph id="8BCE36FEFB7DFFAFFF51F5E7FD55EEDE" blockId="15.[192,802,904,1418]" pageId="15" pageNumber="14">
DIAGNOSIS. Adult size 
<quantity id="4C899B1BFB7DFFAFFE50F5E7FDDEE8CB" box="[497,616,904,927]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.2" metricValueMin="2.3" pageId="15" pageNumber="14" unit="mm" value="27.5" valueMax="32.0" valueMin="23.0">23–32 mm</quantity>
in males, 
<quantity id="4C899B1BFB7DFFAFFD7EF5E7FF5CE8EA" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.7" metricValueMin="2.3" pageId="15" pageNumber="14" unit="mm" value="30.0" valueMax="37.0" valueMin="23.0">23–37 mm</quantity>
in females. Coloration orange yellow, essentially immaculate, only with some regular blackish spots on pedipalps and metasoma (slightly darker and more extended in female); metasomal segment V and telson blackish. Pedipalp fingers with 9–11 principal rows of denticles and 6–7 internal accessory denticles. Pectines with 22–25 teeth in males and 
<quantity id="4C899B1BFB7DFFAFFDB7F230FDC1EF22" box="[534,631,1119,1142]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.588" metricValueMin="4.826" pageId="15" pageNumber="14" unit="in" value="20.5" valueMax="22.0" valueMin="19.0">19–22 in</quantity>
females. Tibial spur in males absent, in females present in legs III–IV. Metasoma slightly attenuate, with most carinae weak; ventral lateral carinae of segment V composed of narrowly conical (males) to broadly lobate (females) denticles, dorsal lateral and lateral supramedian carinae of segments I–IV with terminal denticle enlarged; intercarinal areas essentially smooth, except sparsely granulose on ventrodistal and ventrolateral areas of segment V. Telson vesicle oval and densely setose.
</paragraph>
</subSubSection>
<subSubSection id="C36B6575FB7DFFB4FF51F3DFFB46EC19" lastPageId="20" lastPageNumber="19" pageId="15" pageNumber="14" type="description">
<paragraph id="8BCE36FEFB7DFFAFFF51F3DFFAA2EA87" blockId="15.[192,803,1456,1909]" lastBlockId="15.[830,1442,199,1909]" pageId="15" pageNumber="14">
DESCRIPTION (adult male 
<typeStatus id="54CA885CFB7DFFAFFD86F3DFFD3FEE93" box="[551,649,1456,1479]" pageId="15" pageNumber="14" type="holotype">holotype</typeStatus>
). 
<emphasis id="B905EAECFB7DFFAFFD06F3DFFC94EE93" bold="true" box="[679,802,1456,1479]" pageId="15" pageNumber="14">Coloration</emphasis>
(
<figureCitation id="134A2A7BFB7DFFAFFF66F3A0FEFBEEB2" box="[199,333,1487,1510]" captionStart="Figures 34–37" captionStartId="13.[192,278,1630,1653]" captionTargetBox="[212,1415,153,1571]" captionTargetId="figure-6@13.[192,1439,144,1601]" captionTargetPageId="13" captionText="Figures 34–37: Anomalobuthus krivochatskyi sp. n., adult male holotype (34–35) and adult female paratopotype (36–37): fullbody views, dorsal (34, 36) and ventral (35, 37). Scale bar = 10 mm." pageId="15" pageNumber="14">Figs. 34–35</figureCitation>
) base very vivid yellow, with an orange shade; in general, the base color is paler on pedipalp chelae, legs, and pectines. Chelicerae immaculate, except for blackish finger teeth. Pedipalp femur with a conspicuous blackish spot on dorsodistal apex, which diffusely continues as a broad stripe over almost all internal surface; patella with internal surface blackish; chela immaculate, only with finger denticles blackish. Carapace immaculate, only with a blackish spot under every ocular group. Tergites immaculate. Coxosternal region and genital operculum immaculate. Pectines pale yellowish, immaculate. Sternites immaculate. Legs immaculate; claws with distal half dark brown to blackish. Metasoma conspicuously bicolor: segments I–IV yellow with a diffuse annulated appearance (basal and distal parts of each segment infuscate in the shape of thin blackish rings, which become somewhat larger and more diffuse dorsally and basally), segment V entirely blackish; carinae not infuscate nor underlined with dark pigment, except for darkened terminal denticles on dorsal laterals and lateral supramedians. Telson vesicle blackish, similar to metasomal segment V; aculeus with basal half brownish and distal half blackish.
</paragraph>
<paragraph id="8BCE36FEFB7DFFAFFCCFF7B4FBA6EFC1" blockId="15.[830,1442,199,1909]" pageId="15" pageNumber="14">
<emphasis id="B905EAECFB7DFFAFFCCFF7B4FC5EEAA6" bold="true" box="[878,1000,475,498]" pageId="15" pageNumber="14">Chelicerae</emphasis>
(
<figureCitation id="134A2A7BFB7DFFAFFC5EF7B4FBE3EAA6" box="[1023,1109,475,498]" captionStart="Figures 46–53" captionStartId="17.[192,278,1398,1421]" captionTargetBox="[191,1437,191,1368]" captionTargetId="figure-6@17.[192,1437,192,1367]" captionTargetPageId="17" captionText="Figures 46–53: Anomalobuthus krivochatskyi sp. n., adult male holotype (46–47) and adult female paratopotype (48–53): chelicerae, carapace and tergites I–III, dorsal (46, 48), sternopectinal region and sternites III–V, ventral (47, 49), distal part of right legs I–IV, internal (50–53)." pageId="15" pageNumber="14">Fig. 46</figureCitation>
). Manus smooth and shiny, dorsodistal portion with some weak granules arranged transversally, defining a depressed area; setation very dense ventrally but essentially lacking dorsally, except for many rigid macrosetae on fixed finger and a few around depressed area of manus. Fixed finger of size, shape and curvature ordinary for the family, dorsally with the four standard denticles: single 
<emphasis id="B905EAECFB7DFFAFFAADF4DDFAAFE99D" box="[1292,1305,690,713]" italics="true" pageId="15" pageNumber="14">d</emphasis>
very large, sharp and falcate, single 
<emphasis id="B905EAECFB7DFFAFFBC4F4BFFBCBE9B3" box="[1125,1149,720,743]" italics="true" pageId="15" pageNumber="14">sd</emphasis>
large, sharp and falcate, single 
<emphasis id="B905EAECFB7DFFAFFC29F480FC2DE852" box="[904,923,751,774]" italics="true" pageId="15" pageNumber="14">m</emphasis>
and 
<emphasis id="B905EAECFB7DFFAFFC74F480FC54E852" box="[981,994,751,774]" italics="true" pageId="15" pageNumber="14">b</emphasis>
large, sharp, triangular and fused into a subrectangular bicusp; ventrally with a single denticle, which is medium-sized and placed at a level between dorsal denticles 
<emphasis id="B905EAECFB7DFFAFFC50F524FBB2E836" box="[1009,1028,843,866]" italics="true" pageId="15" pageNumber="14">m</emphasis>
and 
<emphasis id="B905EAECFB7DFFAFFB9CF524FBFCE836" box="[1085,1098,843,866]" italics="true" pageId="15" pageNumber="14">b</emphasis>
. Movable finger of size, shape and curvature ordinary for the family, dorsally with the five standard denticles: single 
<emphasis id="B905EAECFB7DFFAFFB0BF5E7FB75E8CB" box="[1194,1219,904,927]" italics="true" pageId="15" pageNumber="14">ed</emphasis>
very large, sharp, falcate and placed at the same level of 
<emphasis id="B905EAECFB7DFFAFFB57F5C8FABDE8EA" box="[1270,1291,935,958]" italics="true" pageId="15" pageNumber="14">id</emphasis>
, both closely opposable to fixed finger distal denticle, single 
<emphasis id="B905EAECFB7DFFAFFAE7F5A9FAE8E889" box="[1350,1374,966,989]" italics="true" pageId="15" pageNumber="14">sd</emphasis>
large, sharp, triangular, single 
<emphasis id="B905EAECFB7DFFAFFBEAF58BFBE8E8AF" box="[1099,1118,996,1019]" italics="true" pageId="15" pageNumber="14">m</emphasis>
very large, sharp, triangular, two 
<emphasis id="B905EAECFB7DFFAFFCD0F26CFC3CEF4E" box="[881,906,1027,1050]" italics="true" pageId="15" pageNumber="14">ed</emphasis>
small (basalmost smaller), sharp, triangular and adjacent to each other and 
<emphasis id="B905EAECFB7DFFAFFBD8F24DFB3AEF6D" box="[1145,1164,1058,1081]" italics="true" pageId="15" pageNumber="14">m</emphasis>
; ventrally with a single denticle which is medium-sized, small, placed at the level of dorsal denticle 
<emphasis id="B905EAECFB7DFFAFFB98F230FBE7EF22" box="[1081,1105,1119,1142]" italics="true" pageId="15" pageNumber="14">sd</emphasis>
and raises from a conspicuous, translucent lamella.
</paragraph>
<paragraph id="8BCE36FEFB7DFFAFFCCFF2F3FC4CEDAE" blockId="15.[830,1442,199,1909]" pageId="15" pageNumber="14">
<emphasis id="B905EAECFB7DFFAFFCCFF2F3FC6BEFE7" bold="true" box="[878,989,1180,1203]" pageId="15" pageNumber="14">Pedipalps</emphasis>
(
<figureCitation id="134A2A7BFB7DFFAFFC51F2F3FBC5EFE7" box="[1008,1139,1180,1203]" captionStart="Figures 34–37" captionStartId="13.[192,278,1630,1653]" captionTargetBox="[212,1415,153,1571]" captionTargetId="figure-6@13.[192,1439,144,1601]" captionTargetPageId="13" captionText="Figures 34–37: Anomalobuthus krivochatskyi sp. n., adult male holotype (34–35) and adult female paratopotype (36–37): fullbody views, dorsal (34, 36) and ventral (35, 37). Scale bar = 10 mm." pageId="15" pageNumber="14">Figs. 34–35</figureCitation>
). Relatively short but very slender, essentially bare. Femur subtly curved inwards, with carinae weak, granulose to subdenticulate; intercarinal tegument smooth and glossy. Patella somewhat curved inwards, with carinae obsolete to vestigial, smooth; intercarinal tegument smooth and glossy. Chela elongate and very slender; manus conspicuously narrower than patella (ratio 0.78), cylindrical (2.14 times longer than wide, as wide as deep), with carinae obsolete to vestigial, smooth; intercarinal tegument smooth and glossy; fingers very long (movable finger 2.20 times longer than underhand), only subtly curved and with 9– 10 principal rows of denticles (the two basalmost rows are poorly defined), basal lobe/notch combination absent, external accessory denticles absent, internal accessory denticles very large and claw-like (increasing in size distally), numbering seven and six on fixed and movable fingers, respectively, movable finger with one claw-like accessory denticles basal to the very large terminal denticle.
</paragraph>
<paragraph id="8BCE36FEFB7DFFB0FCCFF16DFAB4EC72" blockId="15.[830,1442,199,1909]" lastBlockId="16.[830,1441,1684,1830]" lastPageId="16" lastPageNumber="15" pageId="15" pageNumber="14">
<emphasis id="B905EAECFB7DFFAFFCCFF16DFC6AEC4D" bold="true" box="[878,988,1794,1817]" pageId="15" pageNumber="14">Carapace</emphasis>
(
<figureCitation id="134A2A7BFB7DFFAFFC56F16DFBE0EC4D" box="[1015,1110,1794,1817]" captionStart="Figures 46–53" captionStartId="17.[192,278,1398,1421]" captionTargetBox="[191,1437,191,1368]" captionTargetId="figure-6@17.[192,1437,192,1367]" captionTargetPageId="17" captionText="Figures 46–53: Anomalobuthus krivochatskyi sp. n., adult male holotype (46–47) and adult female paratopotype (48–53): chelicerae, carapace and tergites I–III, dorsal (46, 48), sternopectinal region and sternites III–V, ventral (47, 49), distal part of right legs I–IV, internal (50–53)." pageId="15" pageNumber="14">Fig. 46</figureCitation>
). Very strongly trapezoidal (much narrower anteriorly) and wider than long; anterior margin shallowly convex, with 6–7 pairs of thin macrosetae and some very short microsetae. Carination essentially absent: the only carinae present are the superciliaries, which are weakly granulose to smooth. Furrows: anterior median, median ocular, central median, posterior median and posterior marginal fused, wide and moderately deep; lateral oculars, lateral centrals, central transverse, and posterior laterals long, narrow and shallow. Tegument very finely and densely granulose, with many small to medium-sized granules scattered all over except on both sides of median axis, where the granules are coarser and glossy.
</paragraph>
<caption id="DF0E6676FB62FFB0FF61F05FFC80ED0B" pageId="16" pageNumber="15" startId="16.[192,278,1584,1607]" targetBox="[212,1422,194,1547]" targetPageId="16">
<paragraph id="8BCE36FEFB62FFB0FF61F05FFC80ED0B" blockId="16.[192,1440,1584,1631]" pageId="16" pageNumber="15">
<emphasis id="B905EAECFB62FFB0FF61F05FFEDEED13" bold="true" box="[192,360,1584,1607]" pageId="16" pageNumber="15">Figures 42–45:</emphasis>
<taxonomicName id="4C714D7DFB62FFB0FECEF05DFD3AED13" authority="Teruel &amp; Kovařík &amp; Fet, 2018" authorityName="Teruel &amp; Kovařík &amp; Fet" authorityYear="2018" box="[367,652,1586,1607]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="krivochatskyi" status="sp. nov.">
<emphasis id="B905EAECFB62FFB0FECEF05DFD3AED13" box="[367,652,1586,1607]" italics="true" pageId="16" pageNumber="15">Anomalobuthus krivochatskyi</emphasis>
</taxonomicName>
<emphasis id="B905EAECFB62FFB0FD32F05DFD7FED13" bold="true" box="[659,713,1586,1607]" pageId="16" pageNumber="15">
<taxonomicNameLabel id="A2365797FB62FFB0FD32F05DFD7FED13" box="[659,713,1586,1607]" pageId="16" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult female (42–43) and juvenile male paratypes (44–45) from Chardara, Kazakhstan: full-body views, dorsal (42, 44) and ventral (43, 45).
</paragraph>
</caption>
<caption id="DF0E6676FB63FFB1FF61F319FE4DEEEA" pageId="17" pageNumber="16" startId="17.[192,278,1398,1421]" targetBox="[191,1437,191,1368]" targetPageId="17">
<paragraph id="8BCE36FEFB63FFB1FF61F319FE4DEEEA" blockId="17.[192,1441,1398,1470]" pageId="17" pageNumber="16">
<emphasis id="B905EAECFB63FFB1FF61F319FEDAEED9" bold="true" box="[192,364,1398,1421]" pageId="17" pageNumber="16">Figures 46–53:</emphasis>
<taxonomicName id="4C714D7DFB63FFB1FED6F318FD21EED8" authority="Teruel &amp; Kovařík &amp; Fet, 2018" authorityName="Teruel &amp; Kovařík &amp; Fet" authorityYear="2018" box="[375,663,1399,1420]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="krivochatskyi" status="sp. nov.">
<emphasis id="B905EAECFB63FFB1FED6F318FD21EED8" box="[375,663,1399,1420]" italics="true" pageId="17" pageNumber="16">Anomalobuthus krivochatskyi</emphasis>
</taxonomicName>
<emphasis id="B905EAECFB63FFB1FD00F318FD6AEED8" bold="true" box="[673,732,1399,1420]" pageId="17" pageNumber="16">
<taxonomicNameLabel id="A2365797FB63FFB1FD00F318FD6AEED8" box="[673,732,1399,1420]" pageId="17" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult male holotype (46–47) and adult female paratopotype (48–53): chelicerae, carapace and tergites I–III, dorsal (46, 48), sternopectinal region and sternites III–V, ventral (47, 49), distal part of right legs I–IV, internal (50–53).
</paragraph>
</caption>
<paragraph id="8BCE36FEFB63FFB1FF51F39CFD80ED13" blockId="17.[192,803,1523,1883]" pageId="17" pageNumber="16">
<emphasis id="B905EAECFB63FFB1FF51F39CFEE5ED5E" bold="true" box="[240,339,1523,1546]" pageId="17" pageNumber="16">Sternum</emphasis>
(
<figureCitation id="134A2A7BFB63FFB1FECAF39CFE77ED5E" box="[363,449,1523,1546]" captionStart="Figures 46–53" captionStartId="17.[192,278,1398,1421]" captionTargetBox="[191,1437,191,1368]" captionTargetId="figure-6@17.[192,1437,192,1367]" captionTargetPageId="17" captionText="Figures 46–53: Anomalobuthus krivochatskyi sp. n., adult male holotype (46–47) and adult female paratopotype (48–53): chelicerae, carapace and tergites I–III, dorsal (46, 48), sternopectinal region and sternites III–V, ventral (47, 49), distal part of right legs I–IV, internal (50–53)." pageId="17" pageNumber="16">Fig. 47</figureCitation>
). Standard for the genus, relatively small and widely triangular in shape, with two pairs of inconspicuous macrosetae.
</paragraph>
<paragraph id="8BCE36FEFB63FFB1FF51F020FE73ED96" blockId="17.[192,803,1523,1883]" pageId="17" pageNumber="16">
<emphasis id="B905EAECFB63FFB1FF51F020FE71ED32" bold="true" box="[240,455,1615,1638]" pageId="17" pageNumber="16">Genital operculum</emphasis>
(
<figureCitation id="134A2A7BFB63FFB1FE77F020FD95ED32" box="[470,547,1615,1638]" captionStart="Figures 46–53" captionStartId="17.[192,278,1398,1421]" captionTargetBox="[191,1437,191,1368]" captionTargetId="figure-6@17.[192,1437,192,1367]" captionTargetPageId="17" captionText="Figures 46–53: Anomalobuthus krivochatskyi sp. n., adult male holotype (46–47) and adult female paratopotype (48–53): chelicerae, carapace and tergites I–III, dorsal (46, 48), sternopectinal region and sternites III–V, ventral (47, 49), distal part of right legs I–IV, internal (50–53)." pageId="17" pageNumber="16">Fig. 47</figureCitation>
). Relatively large, each half roundly subtriangular in shape, with three pairs of inconspicuous macrosetae, plus a few short microsetae. Genital papillae present.
</paragraph>
<paragraph id="8BCE36FEFB63FFB1FF51F0A5FC25ED7D" blockId="17.[192,803,1523,1883]" lastBlockId="17.[830,1441,1523,1883]" pageId="17" pageNumber="16">
<emphasis id="B905EAECFB63FFB1FF51F0A5FEFBEDB5" bold="true" box="[240,333,1738,1761]" pageId="17" pageNumber="16">Pectines</emphasis>
(
<figureCitation id="134A2A7BFB63FFB1FEC2F0A5FE01EDB5" box="[355,439,1738,1761]" captionStart="Figures 46–53" captionStartId="17.[192,278,1398,1421]" captionTargetBox="[191,1437,191,1368]" captionTargetId="figure-6@17.[192,1437,192,1367]" captionTargetPageId="17" captionText="Figures 46–53: Anomalobuthus krivochatskyi sp. n., adult male holotype (46–47) and adult female paratopotype (48–53): chelicerae, carapace and tergites I–III, dorsal (46, 48), sternopectinal region and sternites III–V, ventral (47, 49), distal part of right legs I–IV, internal (50–53)." pageId="17" pageNumber="16">Fig. 47</figureCitation>
). Standard-sized for the genus: very long, extending beyond leg IV coxa-trochanter joint), subrectangular and densely setose. Tooth count 22/22. Basal plate heavily sclerotized, much wider than long, anterior margin with a very deep, narrow anteromedian furrow, posterior margin very shallowly convex.
</paragraph>
<paragraph id="8BCE36FEFB63FFB1FCCFF05FFA24EDD1" blockId="17.[830,1441,1523,1883]" pageId="17" pageNumber="16">
<emphasis id="B905EAECFB63FFB1FCCFF05FFC15ED13" bold="true" box="[878,931,1584,1607]" pageId="17" pageNumber="16">Legs</emphasis>
(
<figureCitation id="134A2A7BFB63FFB1FC16F05FFB8BED13" box="[951,1085,1584,1607]" captionStart="Figures 34–37" captionStartId="13.[192,278,1630,1653]" captionTargetBox="[212,1415,153,1571]" captionTargetId="figure-6@13.[192,1439,144,1601]" captionTargetPageId="13" captionText="Figures 34–37: Anomalobuthus krivochatskyi sp. n., adult male holotype (34–35) and adult female paratopotype (36–37): fullbody views, dorsal (34, 36) and ventral (35, 37). Scale bar = 10 mm." pageId="17" pageNumber="16">Figs. 34–35</figureCitation>
). Very slender, with all carinae weak to obsolete and subgranulose to smooth; intercarinal tegument smooth and glossy. Tibial spur absent.
</paragraph>
<paragraph id="8BCE36FEFB63FFB2FCCFF0E3FD9FEDB8" blockId="17.[830,1441,1523,1883]" lastBlockId="18.[192,803,1503,1894]" lastPageId="18" lastPageNumber="17" pageId="17" pageNumber="16">
<emphasis id="B905EAECFB63FFB1FCCFF0E3FC50EDF7" bold="true" box="[878,998,1676,1699]" pageId="17" pageNumber="16">Mesosoma</emphasis>
(
<figureCitation id="134A2A7BFB63FFB1FC5AF0E3FB36EDF7" box="[1019,1152,1676,1699]" captionStart="Figures 46–53" captionStartId="17.[192,278,1398,1421]" captionTargetBox="[191,1437,191,1368]" captionTargetId="figure-6@17.[192,1437,192,1367]" captionTargetPageId="17" captionText="Figures 46–53: Anomalobuthus krivochatskyi sp. n., adult male holotype (46–47) and adult female paratopotype (48–53): chelicerae, carapace and tergites I–III, dorsal (46, 48), sternopectinal region and sternites III–V, ventral (47, 49), distal part of right legs I–IV, internal (50–53)." pageId="17" pageNumber="16">Figs. 46–47</figureCitation>
). Tergites with the same sculpture as on carapace; I–VI irregularly tricarinate: the median longitudinal carina is moderately strong, short, and formed by irregular medium-sized granules that do not project beyond posterior margin, but the submedian carinae are undefined on I–IV, very irregular on V, and well defined only on VI; tergite VII with five well- defined carinae (median, submedians and laterals), which are long, strong and finely serrate to crenulate. Sternites essentially bare; III–VI glossy and with subtle vestiges of a pair of smooth submedian carinae, spiracles relatively short and slit-like, transversely arranged (not oblique), V with smooth patch absent; VII with two pairs of carinae (submedians and laterals) which are long and finely crenulate to serrate, intercarinal tegument coriaceous to minutely granulose.
</paragraph>
<caption id="DF0E6676FB60FFB2FF61F30DFD1BEEFE" pageId="18" pageNumber="17" startId="18.[192,278,1378,1401]" targetBox="[204,1423,190,1328]" targetPageId="18">
<paragraph id="8BCE36FEFB60FFB2FF61F30DFD1BEEFE" blockId="18.[192,1440,1378,1450]" pageId="18" pageNumber="17">
<emphasis id="B905EAECFB60FFB2FF61F30DFEDDEE2D" bold="true" box="[192,363,1378,1401]" pageId="18" pageNumber="17">Figures 54–58:</emphasis>
<taxonomicName id="4C714D7DFB60FFB2FED5F30BFD22EE2D" authority="Teruel &amp; Kovařík &amp; Fet, 2018" authorityName="Teruel &amp; Kovařík &amp; Fet" authorityYear="2018" box="[372,660,1380,1401]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="krivochatskyi" status="sp. nov.">
<emphasis id="B905EAECFB60FFB2FED5F30BFD22EE2D" box="[372,660,1380,1401]" italics="true" pageId="18" pageNumber="17">Anomalobuthus krivochatskyi</emphasis>
</taxonomicName>
<emphasis id="B905EAECFB60FFB2FD3CF30BFD60EE2D" bold="true" box="[669,726,1380,1401]" pageId="18" pageNumber="17">
<taxonomicNameLabel id="A2365797FB60FFB2FD3CF30BFD60EE2D" box="[669,726,1380,1401]" pageId="18" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult female paratopotype: right pedipalp trochanter and femur, dorsal (54), right pedipalp patella, dorsal (55) and external (56), right pedipalp chela, dorsoexternal (57), right pedipalp movable finger, dorsoexternal (58). Black dots depict trichobothria.
</paragraph>
</caption>
<paragraph id="8BCE36FEFB60FFB3FF51F09CFEBFEDA8" blockId="18.[192,803,1503,1894]" lastBlockId="19.[192,802,1642,1911]" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17">
<emphasis id="B905EAECFB60FFB2FF51F09CFED0EC5E" bold="true" box="[240,358,1779,1802]" pageId="18" pageNumber="17">Metasoma</emphasis>
(
<figureCitation id="134A2A7BFB60FFB2FED6F09CFE4EEC5E" box="[375,504,1779,1802]" captionStart="Figures 59–64" captionStartId="19.[192,278,1542,1565]" captionTargetBox="[204,1417,183,1488]" captionTargetId="figure-6@19.[192,1437,168,1512]" captionTargetPageId="19" captionText="Figures 59–64: Anomalobuthus krivochatskyi sp. n., adult male holotype (59–61) and adult female paratopotype (62–64): metasoma and telson, lateral (59, 62), dorsal (60, 63) and ventral (61, 64)." pageId="18" pageNumber="17">Figs. 59–61</figureCitation>
). Somewhat elongated and slightly wider both basally and distally; with 10/10/ 10/8/5 complete to almost complete carinae, almost all formed by conspicuously isolated, sharply serrate to denticulate granulation: dorsal laterals obsolete to vestigial on I–IV (with one terminal denticles enlarged), absent on V; lateral supramedians obsolete to vestigial on I–IV (with 1–2 terminal denticles enlarged), vestigial as rounded ridges on V; lateral inframedians obsolete to vestigial on I–III, absent on IV–V; ventral laterals obsolete to vestigial on I–IV, moderate to strong on V, where become progressively stronger and somewhat flared distally, formed by sharp, subequal denticles; ventral submedians moderate on I–II, weak on III, obsolete on IV, absent on V (indicated by somewhat raised tegument and irregular granulation on basal half); ventral median absent on I–IV, moderate on V. Intercarinal tegument smooth and glossy, with sparse granulation of different sizes ventrally. Dorsal furrow wide and shallow on all segments. Setation sparse, mostly represented by 4–7 dark macrosetae over every carina.
</paragraph>
<caption id="DF0E6676FB61FFB3FF61F069FC33ED61" pageId="19" pageNumber="18" startId="19.[192,278,1542,1565]" targetBox="[204,1417,183,1488]" targetPageId="19">
<paragraph id="8BCE36FEFB61FFB3FF61F069FC33ED61" blockId="19.[192,1441,1542,1589]" pageId="19" pageNumber="18">
<emphasis id="B905EAECFB61FFB3FF61F069FEDAED49" bold="true" box="[192,364,1542,1565]" pageId="19" pageNumber="18">Figures 59–64:</emphasis>
<taxonomicName id="4C714D7DFB61FFB3FED6F068FD21ED48" authority="Teruel &amp; Kovařík &amp; Fet, 2018" authorityName="Teruel &amp; Kovařík &amp; Fet" authorityYear="2018" box="[375,663,1543,1564]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="krivochatskyi" status="sp. nov.">
<emphasis id="B905EAECFB61FFB3FED6F068FD21ED48" box="[375,663,1543,1564]" italics="true" pageId="19" pageNumber="18">Anomalobuthus krivochatskyi</emphasis>
</taxonomicName>
<emphasis id="B905EAECFB61FFB3FD00F068FD6AED48" bold="true" box="[673,732,1543,1564]" pageId="19" pageNumber="18">
<taxonomicNameLabel id="A2365797FB61FFB3FD00F068FD6AED48" box="[673,732,1543,1564]" pageId="19" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult male holotype (59–61) and adult female paratopotype (62–64): metasoma and telson, lateral (59, 62), dorsal (60, 63) and ventral (61, 64).
</paragraph>
</caption>
<paragraph id="8BCE36FEFB61FFB3FF51F16BFB0BEC4F" blockId="19.[192,802,1642,1911]" lastBlockId="19.[830,1441,1642,1819]" pageId="19" pageNumber="18">
<emphasis id="B905EAECFB61FFB3FF51F16BFE8AEC4F" bold="true" box="[240,316,1796,1819]" pageId="19" pageNumber="18">Telson</emphasis>
(
<figureCitation id="134A2A7BFB61FFB3FEEBF16BFE78EC4F" box="[330,462,1796,1819]" captionStart="Figures 59–64" captionStartId="19.[192,278,1542,1565]" captionTargetBox="[204,1417,183,1488]" captionTargetId="figure-6@19.[192,1437,168,1512]" captionTargetPageId="19" captionText="Figures 59–64: Anomalobuthus krivochatskyi sp. n., adult male holotype (59–61) and adult female paratopotype (62–64): metasoma and telson, lateral (59, 62), dorsal (60, 63) and ventral (61, 64)." pageId="19" pageNumber="18">Figs. 59–61</figureCitation>
). Moderately setose, with many setae scattered all over dorsal and lateral surfaces. Vesicle short oval (1.92 times longer than wide, 0.92 times wider than deep), tegument smooth and glossy, with vestiges of coarse granules arranged into three obsolete longitudinal carinae (ventral median and ventral submedians), and some coarse punctations ventrally. Subaculear tubercle absent, but subtly suggested by a vestigial granule. Aculeus conspicuously shorter than vesicle, thick and shallowly curved.
</paragraph>
<paragraph id="8BCE36FEFB61FFB4FCCFF12EFDF1EEA8" blockId="19.[830,1440,1857,1911]" lastBlockId="20.[192,802,1295,1532]" lastPageId="20" lastPageNumber="19" pageId="19" pageNumber="18">
FEMALE (paratopotype). Very similar to described male, sexual dimorphism evident by: 
<emphasis id="B905EAECFB61FFB3FB45F10FFB4CEC23" bold="true" box="[1252,1274,1888,1911]" pageId="19" pageNumber="18">1)</emphasis>
size comparatively larger inside each size-class; 
<emphasis id="B905EAECFB66FFB4FDFBF360FDC6EE72" bold="true" box="[602,624,1295,1318]" pageId="20" pageNumber="19">2)</emphasis>
mesosoma and metasoma slightly less slender; 
<emphasis id="B905EAECFB66FFB4FD89F342FD89EE10" bold="true" box="[552,575,1325,1348]" pageId="20" pageNumber="19">3)</emphasis>
genital papillae absent; 
<emphasis id="B905EAECFB66FFB4FF5CF323FEA5EE37" bold="true" box="[253,275,1356,1379]" pageId="20" pageNumber="19">4)</emphasis>
pectines smaller, with consistently lower tooth counts; 
<emphasis id="B905EAECFB66FFB4FEB5F304FE9CEED6" bold="true" box="[276,298,1387,1410]" pageId="20" pageNumber="19">5)</emphasis>
tibial spur small, sharp and sinuose, present at least on both legs IV (sometimes also on one leg III); 
<emphasis id="B905EAECFB66FFB4FCADF3E6FC97EEF4" bold="true" box="[780,801,1417,1440]" pageId="20" pageNumber="19">6)</emphasis>
metasomal segment V with ventral intercarinal tegument less granulose. See 
<figureCitation id="134A2A7BFB66FFB4FE3CF3A8FD97EE8A" box="[413,545,1479,1502]" captionStart="Figures 34–37" captionStartId="13.[192,278,1630,1653]" captionTargetBox="[212,1415,153,1571]" captionTargetId="figure-6@13.[192,1439,144,1601]" captionTargetPageId="13" captionText="Figures 34–37: Anomalobuthus krivochatskyi sp. n., adult male holotype (34–35) and adult female paratopotype (36–37): fullbody views, dorsal (34, 36) and ventral (35, 37). Scale bar = 10 mm." pageId="20" pageNumber="19">Figs. 36–37</figureCitation>
, 
<figureCitation id="134A2A7BFB66FFB4FD91F3A8FDC0EE8A" box="[560,630,1479,1502]" captionStart-0="Figures 38–41" captionStart-1="Figures 42–45" captionStartId-0="14.[192,278,1754,1777]" captionStartId-1="16.[192,278,1584,1607]" captionTargetBox-0="[201,1418,199,1682]" captionTargetBox-1="[212,1422,194,1547]" captionTargetId-0="figure-12@14.[192,1439,192,1697]" captionTargetId-1="figure-12@16.[192,1440,192,1554]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionText-0="Figures 38–41: Anomalobuthus krivochatskyi sp. n., adult male (38–39) and adult female paratypes (40–41) from between Bukhara and Gazli, Uzbekistan: full-body views, dorsal (38, 40) and ventral (39, 41). Scale bar = 10 mm." captionText-1="Figures 42–45: Anomalobuthus krivochatskyi sp. n., adult female (42–43) and juvenile male paratypes (44–45) from Chardara, Kazakhstan: full-body views, dorsal (42, 44) and ventral (43, 45)." pageId="20" pageNumber="19">40–43</figureCitation>
, 
<figureCitation id="134A2A7BFB66FFB4FD25F3A8FD7FEE8A" box="[644,713,1479,1502]" captionStart-0="Figures 46–53" captionStart-1="Figures 54–58" captionStartId-0="17.[192,278,1398,1421]" captionStartId-1="18.[192,278,1378,1401]" captionTargetBox-0="[191,1437,191,1368]" captionTargetBox-1="[204,1423,190,1328]" captionTargetId-0="figure-6@17.[192,1437,192,1367]" captionTargetId-1="figure-12@18.[192,1437,168,1348]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Figures 46–53: Anomalobuthus krivochatskyi sp. n., adult male holotype (46–47) and adult female paratopotype (48–53): chelicerae, carapace and tergites I–III, dorsal (46, 48), sternopectinal region and sternites III–V, ventral (47, 49), distal part of right legs I–IV, internal (50–53)." captionText-1="Figures 54–58: Anomalobuthus krivochatskyi sp. n., adult female paratopotype: right pedipalp trochanter and femur, dorsal (54), right pedipalp patella, dorsal (55) and external (56), right pedipalp chela, dorsoexternal (57), right pedipalp movable finger, dorsoexternal (58). Black dots depict trichobothria." pageId="20" pageNumber="19">48–58</figureCitation>
, 
<figureCitation id="134A2A7BFB66FFB4FD79F3A8FCABEE8A" box="[728,797,1479,1502]" captionStart="Figures 59–64" captionStartId="19.[192,278,1542,1565]" captionTargetBox="[204,1417,183,1488]" captionTargetId="figure-6@19.[192,1437,168,1512]" captionTargetPageId="19" captionText="Figures 59–64: Anomalobuthus krivochatskyi sp. n., adult male holotype (59–61) and adult female paratopotype (62–64): metasoma and telson, lateral (59, 62), dorsal (60, 63) and ventral (61, 64)." pageId="20" pageNumber="19">62–64</figureCitation>
, 
<figureCitation id="134A2A7BFB66FFB4FF61F38AFE3EEEA8" box="[192,392,1509,1532]" captionStart="Figures 134–140" captionStartId="37.[192,278,1748,1771]" captionTargetBox="[191,1440,191,1719]" captionTargetId="figure-6@37.[192,1439,192,1718]" captionTargetPageId="37" captionText="Figures 134–140: Anomalobuthus spp., SEM views showing size variation of leg IV tibial spur (highlighted in red): large in paratopotype female A. krivochatskyi sp. n. (134), moderately large in paratype female A. krivochatskyi sp. n. from Bukhara (135), small in paratopotype female A. krivochatskyi sp. n. (136), absent in male A. rickmersi from Repetek (137), close-up of a medium-sized leg IV tibial spur of a paratopotype female A. krivochatskyi sp. n. from Zarafshan (138), see spinulate and striate sculpture. Anomalobuthus krivochatskyi sp. n., adult female paratopotype: SEM views of right pecten and adjacent parts of leg coxae and sternite III (139), peg sensilla of tooth from same pecten (140)." pageId="20" pageNumber="19">134–136, 138–140</figureCitation>
and 
<tableCitation id="C6F30345FB66FFB4FE1DF38AFDBAEEA8" box="[444,524,1509,1532]" captionStart="Table 2" captionStartId="20.[192,257,1203,1226]" captionText="Table 2: Measurements of four types of Anomalobuthus krivochatskyi sp. n. Abbreviations: length (L), width (W), posterior width (Wp), depth (D)." pageId="20" pageNumber="19">Tabs. 2</tableCitation>
, 
<tableCitation id="C6F30345FB66FFB4FDB9F38AFDF1EEA8" box="[536,583,1509,1532]" captionStart-0="Table 4" captionStart-1="Table 5" captionStartId-0="35.[192,257,1030,1053]" captionStartId-1="38.[192,257,680,703]" captionText-0="Table 4: Comparison between five species of Anomalobuthus based upon selected morphometric ratios. Abbreviations: length (L)," captionText-1="Table 5: Pectinal tooth count variation in all known species of Anomalobuthus. Abbreviations: number of pectines (N), average (Ave.), standard deviation (SD). This includes counts taken from sources with precise and reliable geographic data (Birula, 1905, 1910)." pageId="20" pageNumber="19">4–5.</tableCitation>
</paragraph>
<paragraph id="8BCE36FEFB66FFB4FCBDF68FFAFDEF2A" pageId="20" pageNumber="19">
<table id="F971C45EFB66005FFF6CF6BEFA26EF2A" box="[205,1424,209,1150]" gridcols="6" gridrows="21" pageId="20" pageNumber="19">
<tr id="354134BCFB66005FFF6CF6BEFA26EA52" box="[205,1424,209,262]" gridrow="0" pageId="20" pageNumber="19" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<th id="76905DC0FB66005FFCBBF6BEFC6DEA52" box="[794,987,209,262]" gridcol="3" gridrow="0" pageId="20" pageNumber="19">
<emphasis id="B905EAECFB66FFB4FCBDF68FFC61EBA3" bold="true" box="[796,983,224,247]" pageId="20" pageNumber="19">UZB, Zarafshan</emphasis>
</th>
<th id="76905DC0FB66005FFB6BF6BEFA26EA52" box="[1226,1424,209,262]" gridcol="5" gridrow="0" pageId="20" pageNumber="19">
<emphasis id="B905EAECFB66FFB4FB6BF6BEFAFAEA52" bold="true" pageId="20" pageNumber="19">UZB, Bukhara to Gazli</emphasis>
</th>
</tr>
<tr id="354134BCFB66005FFF6CF765FA26EA09" box="[205,1424,266,349]" gridrow="1" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF765FD92EA09" box="[205,548,266,349]" colspan="2" colspanRight="1" gridcol="0" gridrow="1" pageId="20" pageNumber="19">
<emphasis id="B905EAECFB66FFB4FEB6F765FE56EA75" bold="true" box="[279,480,266,289]" pageId="20" pageNumber="19">Dimensions (mm)</emphasis>
</th>
<td id="76905DC0FB66005FFDE7F765FD4AEA09" box="[582,764,266,349]" gridcol="2" gridrow="1" pageId="20" pageNumber="19">
♂ 
<emphasis id="B905EAECFB66FFB4FDDCF770FD78EA09" bold="true" pageId="20" pageNumber="19">holotype (FKCP)</emphasis>
</td>
<td id="76905DC0FB66005FFCBBF765FC6DEA09" box="[794,987,266,349]" gridcol="3" gridrow="1" pageId="20" pageNumber="19">
♂ 
<emphasis id="B905EAECFB66FFB4FCF4F770FC6DEA09" bold="true" pageId="20" pageNumber="19">paratype (RTO: Sco-0233)</emphasis>
</td>
<td id="76905DC0FB66005FFC53F765FB05EA09" box="[1010,1203,266,349]" gridcol="4" gridrow="1" pageId="20" pageNumber="19">
♀ 
<emphasis id="B905EAECFB66FFB4FB8CF770FB05EA09" bold="true" pageId="20" pageNumber="19">paratype (RTO: Sco-0233)</emphasis>
</td>
<td id="76905DC0FB66005FFB6BF765FA26EA09" box="[1226,1424,266,349]" gridcol="5" gridrow="1" pageId="20" pageNumber="19">
♀ 
<emphasis id="B905EAECFB66FFB4FAA9F770FA38EA09" bold="true" pageId="20" pageNumber="19">paratype (RTO: Sco-0208)</emphasis>
</td>
</tr>
<tr id="354134BCFB66005FFF6CF719FA26EAD9" box="[205,1424,374,397]" gridrow="2" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF719FE14EAD9" box="[205,418,374,397]" gridcol="0" gridrow="2" pageId="20" pageNumber="19">Carapace</th>
<td id="76905DC0FB66005FFE1FF719FD92EAD9" box="[446,548,374,397]" gridcol="1" gridrow="2" pageId="20" pageNumber="19">L / Wp</td>
<td id="76905DC0FB66005FFDE7F719FD4AEAD9" box="[582,764,374,397]" gridcol="2" gridrow="2" pageId="20" pageNumber="19">3.30 / 3.70</td>
<td id="76905DC0FB66005FFCBBF719FC6DEAD9" box="[794,987,374,397]" gridcol="3" gridrow="2" pageId="20" pageNumber="19">3.40 / 3.80</td>
<td id="76905DC0FB66005FFC53F719FB05EAD9" box="[1010,1203,374,397]" gridcol="4" gridrow="2" pageId="20" pageNumber="19">3.80 / 4.60</td>
<td id="76905DC0FB66005FFB6BF719FA26EAD9" box="[1226,1424,374,397]" gridcol="5" gridrow="2" pageId="20" pageNumber="19">3.00 / 3.40</td>
</tr>
<tr id="354134BCFB66005FFF6CF7F0FA26EAE2" box="[205,1424,415,438]" gridrow="3" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF7F0FE14EAE2" box="[205,418,415,438]" gridcol="0" gridrow="3" pageId="20" pageNumber="19">Mesosoma</th>
<td id="76905DC0FB66005FFE1FF7F0FD92EAE2" box="[446,548,415,438]" gridcol="1" gridrow="3" pageId="20" pageNumber="19">L</td>
<td id="76905DC0FB66005FFDE7F7F0FD4AEAE2" box="[582,764,415,438]" gridcol="2" gridrow="3" pageId="20" pageNumber="19">7.00</td>
<td id="76905DC0FB66005FFCBBF7F0FC6DEAE2" box="[794,987,415,438]" gridcol="3" gridrow="3" pageId="20" pageNumber="19">6.80</td>
<td id="76905DC0FB66005FFC53F7F0FB05EAE2" box="[1010,1203,415,438]" gridcol="4" gridrow="3" pageId="20" pageNumber="19">9.50</td>
<td id="76905DC0FB66005FFB6BF7F0FA26EAE2" box="[1226,1424,415,438]" gridcol="5" gridrow="3" pageId="20" pageNumber="19">7.00</td>
</tr>
<tr id="354134BCFB66005FFF6CF7A7FA26EA8B" box="[205,1424,456,479]" gridrow="4" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF7A7FE14EA8B" box="[205,418,456,479]" gridcol="0" gridrow="4" pageId="20" pageNumber="19">Tergite VII</th>
<td id="76905DC0FB66005FFE1FF7A7FD92EA8B" box="[446,548,456,479]" gridcol="1" gridrow="4" pageId="20" pageNumber="19">L / W</td>
<td id="76905DC0FB66005FFDE7F7A7FD4AEA8B" box="[582,764,456,479]" gridcol="2" gridrow="4" pageId="20" pageNumber="19">2.22 / 3.40</td>
<td id="76905DC0FB66005FFCBBF7A7FC6DEA8B" box="[794,987,456,479]" gridcol="3" gridrow="4" pageId="20" pageNumber="19">2.00 / 3.60</td>
<td id="76905DC0FB66005FFC53F7A7FB05EA8B" box="[1010,1203,456,479]" gridcol="4" gridrow="4" pageId="20" pageNumber="19">2.20 / 4.60</td>
<td id="76905DC0FB66005FFB6BF7A7FA26EA8B" box="[1226,1424,456,479]" gridcol="5" gridrow="4" pageId="20" pageNumber="19">1.80 / 3.70</td>
</tr>
<tr id="354134BCFB66005FFF6CF79DFA26E95D" box="[205,1424,498,521]" gridrow="5" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF79DFE14E95D" box="[205,418,498,521]" gridcol="0" gridrow="5" pageId="20" pageNumber="19">Metasoma + Telson</th>
<td id="76905DC0FB66005FFE1FF79DFD92E95D" box="[446,548,498,521]" gridcol="1" gridrow="5" pageId="20" pageNumber="19">L</td>
<td id="76905DC0FB66005FFDE7F79DFD4AE95D" box="[582,764,498,521]" gridcol="2" gridrow="5" pageId="20" pageNumber="19">21.90</td>
<td id="76905DC0FB66005FFCBBF79DFC6DE95D" box="[794,987,498,521]" gridcol="3" gridrow="5" pageId="20" pageNumber="19">21.30</td>
<td id="76905DC0FB66005FFC53F79DFB05E95D" box="[1010,1203,498,521]" gridcol="4" gridrow="5" pageId="20" pageNumber="19">23.30</td>
<td id="76905DC0FB66005FFB6BF79DFA26E95D" box="[1226,1424,498,521]" gridcol="5" gridrow="5" pageId="20" pageNumber="19">18.40</td>
</tr>
<tr id="354134BCFB66005FFF6CF474FA26E966" box="[205,1424,539,562]" gridrow="6" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF474FE14E966" box="[205,418,539,562]" gridcol="0" gridrow="6" pageId="20" pageNumber="19">Segment I</th>
<td id="76905DC0FB66005FFE1FF474FD92E966" box="[446,548,539,562]" gridcol="1" gridrow="6" pageId="20" pageNumber="19">L / W / D</td>
<td id="76905DC0FB66005FFDE7F474FD4AE966" box="[582,764,539,562]" gridcol="2" gridrow="6" pageId="20" pageNumber="19">2.60 / 1.93 / 1.60</td>
<td id="76905DC0FB66005FFCBBF474FC6DE966" box="[794,987,539,562]" gridcol="3" gridrow="6" pageId="20" pageNumber="19">2.40 / 1.90 / 1.62</td>
<td id="76905DC0FB66005FFC53F474FB05E966" box="[1010,1203,539,562]" gridcol="4" gridrow="6" pageId="20" pageNumber="19">2.50 / 2.30 / 1.96</td>
<td id="76905DC0FB66005FFB6BF474FA26E966" box="[1226,1424,539,562]" gridcol="5" gridrow="6" pageId="20" pageNumber="19">2.10 / 1.70 / 1.47</td>
</tr>
<tr id="354134BCFB66005FFF6CF42BFA26E90F" box="[205,1424,580,603]" gridrow="7" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF42BFE14E90F" box="[205,418,580,603]" gridcol="0" gridrow="7" pageId="20" pageNumber="19">Segment II</th>
<td id="76905DC0FB66005FFE1FF42BFD92E90F" box="[446,548,580,603]" gridcol="1" gridrow="7" pageId="20" pageNumber="19">L / W / D</td>
<td id="76905DC0FB66005FFDE7F42BFD4AE90F" box="[582,764,580,603]" gridcol="2" gridrow="7" pageId="20" pageNumber="19">3.13 / 1.78 / 1.55</td>
<td id="76905DC0FB66005FFCBBF42BFC6DE90F" box="[794,987,580,603]" gridcol="3" gridrow="7" pageId="20" pageNumber="19">3.00 / 1.70 / 1.70</td>
<td id="76905DC0FB66005FFC53F42BFB05E90F" box="[1010,1203,580,603]" gridcol="4" gridrow="7" pageId="20" pageNumber="19">3.30 / 2.20 / 2.00</td>
<td id="76905DC0FB66005FFB6BF42BFA26E90F" box="[1226,1424,580,603]" gridcol="5" gridrow="7" pageId="20" pageNumber="19">2.60 / 1.60 / 1.48</td>
</tr>
<tr id="354134BCFB66005FFF6CF401FA26E9D1" box="[205,1424,622,645]" gridrow="8" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF401FE14E9D1" box="[205,418,622,645]" gridcol="0" gridrow="8" pageId="20" pageNumber="19">Segment III</th>
<td id="76905DC0FB66005FFE1FF401FD92E9D1" box="[446,548,622,645]" gridcol="1" gridrow="8" pageId="20" pageNumber="19">L / W / D</td>
<td id="76905DC0FB66005FFDE7F401FD4AE9D1" box="[582,764,622,645]" gridcol="2" gridrow="8" pageId="20" pageNumber="19">3.45 / 1.85 / 1.65</td>
<td id="76905DC0FB66005FFCBBF401FC6DE9D1" box="[794,987,622,645]" gridcol="3" gridrow="8" pageId="20" pageNumber="19">3.30 / 1.80 / 1.77</td>
<td id="76905DC0FB66005FFC53F401FB05E9D1" box="[1010,1203,622,645]" gridcol="4" gridrow="8" pageId="20" pageNumber="19">3.60 / 2.20 / 2.07</td>
<td id="76905DC0FB66005FFB6BF401FA26E9D1" box="[1226,1424,622,645]" gridcol="5" gridrow="8" pageId="20" pageNumber="19">2.90 / 1.70 / 1.50</td>
</tr>
<tr id="354134BCFB66005FFF6CF4F8FA26E9FA" box="[205,1424,663,686]" gridrow="9" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF4F8FE14E9FA" box="[205,418,663,686]" gridcol="0" gridrow="9" pageId="20" pageNumber="19">Segment IV</th>
<td id="76905DC0FB66005FFE1FF4F8FD92E9FA" box="[446,548,663,686]" gridcol="1" gridrow="9" pageId="20" pageNumber="19">L / W / D</td>
<td id="76905DC0FB66005FFDE7F4F8FD4AE9FA" box="[582,764,663,686]" gridcol="2" gridrow="9" pageId="20" pageNumber="19">4.04 / 1.89 / 1.60</td>
<td id="76905DC0FB66005FFCBBF4F8FC6DE9FA" box="[794,987,663,686]" gridcol="3" gridrow="9" pageId="20" pageNumber="19">4.00 / 1.90 / 1.75</td>
<td id="76905DC0FB66005FFC53F4F8FB05E9FA" box="[1010,1203,663,686]" gridcol="4" gridrow="9" pageId="20" pageNumber="19">4.40 / 2.40 / 2.11</td>
<td id="76905DC0FB66005FFB6BF4F8FA26E9FA" box="[1226,1424,663,686]" gridcol="5" gridrow="9" pageId="20" pageNumber="19">3.40 / 1.80 / 1.51</td>
</tr>
<tr id="354134BCFB66005FFF6CF4AFFA26E983" box="[205,1424,704,727]" gridrow="10" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF4AFFE14E983" box="[205,418,704,727]" gridcol="0" gridrow="10" pageId="20" pageNumber="19">Segment V</th>
<td id="76905DC0FB66005FFE1FF4AFFD92E983" box="[446,548,704,727]" gridcol="1" gridrow="10" pageId="20" pageNumber="19">L / W / D</td>
<td id="76905DC0FB66005FFDE7F4AFFD4AE983" box="[582,764,704,727]" gridcol="2" gridrow="10" pageId="20" pageNumber="19">4.48 / 1.92 / 1.42</td>
<td id="76905DC0FB66005FFCBBF4AFFC6DE983" box="[794,987,704,727]" gridcol="3" gridrow="10" pageId="20" pageNumber="19">4.40 / 1.90 / 1.55</td>
<td id="76905DC0FB66005FFC53F4AFFB05E983" box="[1010,1203,704,727]" gridcol="4" gridrow="10" pageId="20" pageNumber="19">4.80 / 2.40 / 1.80</td>
<td id="76905DC0FB66005FFB6BF4AFFA26E983" box="[1226,1424,704,727]" gridcol="5" gridrow="10" pageId="20" pageNumber="19">3.80 / 1.80 / 1.41</td>
</tr>
<tr id="354134BCFB66005FFF6CF485FA26E855" box="[205,1424,746,769]" gridrow="11" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF485FE14E855" box="[205,418,746,769]" gridcol="0" gridrow="11" pageId="20" pageNumber="19">Telson</th>
<td id="76905DC0FB66005FFE1FF485FD92E855" box="[446,548,746,769]" gridcol="1" gridrow="11" pageId="20" pageNumber="19">L</td>
<td id="76905DC0FB66005FFDE7F485FD4AE855" box="[582,764,746,769]" gridcol="2" gridrow="11" pageId="20" pageNumber="19">4.20</td>
<td id="76905DC0FB66005FFCBBF485FC6DE855" box="[794,987,746,769]" gridcol="3" gridrow="11" pageId="20" pageNumber="19">4.20</td>
<td id="76905DC0FB66005FFC53F485FB05E855" box="[1010,1203,746,769]" gridcol="4" gridrow="11" pageId="20" pageNumber="19">4.70</td>
<td id="76905DC0FB66005FFB6BF485FA26E855" box="[1226,1424,746,769]" gridcol="5" gridrow="11" pageId="20" pageNumber="19">3.60</td>
</tr>
<tr id="354134BCFB66005FFF6CF57CFA26E87E" box="[205,1424,787,810]" gridrow="12" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF57CFE14E87E" box="[205,418,787,810]" gridcol="0" gridrow="12" pageId="20" pageNumber="19">Vesicle</th>
<td id="76905DC0FB66005FFE1FF57CFD92E87E" box="[446,548,787,810]" gridcol="1" gridrow="12" pageId="20" pageNumber="19">L / W / D</td>
<td id="76905DC0FB66005FFDE7F57CFD4AE87E" box="[582,764,787,810]" gridcol="2" gridrow="12" pageId="20" pageNumber="19">2.75 / 1.28 / 1.26</td>
<td id="76905DC0FB66005FFCBBF57CFC6DE87E" box="[794,987,787,810]" gridcol="3" gridrow="12" pageId="20" pageNumber="19">2.30 / 1.20 / 1.30</td>
<td id="76905DC0FB66005FFC53F57CFB05E87E" box="[1010,1203,787,810]" gridcol="4" gridrow="12" pageId="20" pageNumber="19">2.70 / 1.50 / 1.40</td>
<td id="76905DC0FB66005FFB6BF57CFA26E87E" box="[1226,1424,787,810]" gridcol="5" gridrow="12" pageId="20" pageNumber="19">2.10 / 1.20 / 1.10</td>
</tr>
<tr id="354134BCFB66005FFF6CF553FA26E807" box="[205,1424,828,851]" gridrow="13" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF553FE14E807" box="[205,418,828,851]" gridcol="0" gridrow="13" pageId="20" pageNumber="19">Aculeus</th>
<td id="76905DC0FB66005FFE1FF553FD92E807" box="[446,548,828,851]" gridcol="1" gridrow="13" pageId="20" pageNumber="19">L</td>
<td id="76905DC0FB66005FFDE7F553FD4AE807" box="[582,764,828,851]" gridcol="2" gridrow="13" pageId="20" pageNumber="19">1.45</td>
<td id="76905DC0FB66005FFCBBF553FC6DE807" box="[794,987,828,851]" gridcol="3" gridrow="13" pageId="20" pageNumber="19">1.90</td>
<td id="76905DC0FB66005FFC53F553FB05E807" box="[1010,1203,828,851]" gridcol="4" gridrow="13" pageId="20" pageNumber="19">2.00</td>
<td id="76905DC0FB66005FFB6BF553FA26E807" box="[1226,1424,828,851]" gridcol="5" gridrow="13" pageId="20" pageNumber="19">1.50</td>
</tr>
<tr id="354134BCFB66005FFF6CF509FA26E829" box="[205,1424,870,893]" gridrow="14" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF509FE14E829" box="[205,418,870,893]" gridcol="0" gridrow="14" pageId="20" pageNumber="19">Pedipalp</th>
<td id="76905DC0FB66005FFE1FF509FD92E829" box="[446,548,870,893]" gridcol="1" gridrow="14" pageId="20" pageNumber="19">L</td>
<td id="76905DC0FB66005FFDE7F509FD4AE829" box="[582,764,870,893]" gridcol="2" gridrow="14" pageId="20" pageNumber="19">10.51</td>
<td id="76905DC0FB66005FFCBBF509FC6DE829" box="[794,987,870,893]" gridcol="3" gridrow="14" pageId="20" pageNumber="19">11.00</td>
<td id="76905DC0FB66005FFC53F509FB05E829" box="[1010,1203,870,893]" gridcol="4" gridrow="14" pageId="20" pageNumber="19">11.40</td>
<td id="76905DC0FB66005FFB6BF509FA26E829" box="[1226,1424,870,893]" gridcol="5" gridrow="14" pageId="20" pageNumber="19">9.50</td>
</tr>
<tr id="354134BCFB66005FFF6CF5E0FA26E8F2" box="[205,1424,911,934]" gridrow="15" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF5E0FE14E8F2" box="[205,418,911,934]" gridcol="0" gridrow="15" pageId="20" pageNumber="19">Femur</th>
<td id="76905DC0FB66005FFE1FF5E0FD92E8F2" box="[446,548,911,934]" gridcol="1" gridrow="15" pageId="20" pageNumber="19">L / W</td>
<td id="76905DC0FB66005FFDE7F5E0FD4AE8F2" box="[582,764,911,934]" gridcol="2" gridrow="15" pageId="20" pageNumber="19">2.78 / 0.68</td>
<td id="76905DC0FB66005FFCBBF5E0FC6DE8F2" box="[794,987,911,934]" gridcol="3" gridrow="15" pageId="20" pageNumber="19">2.80 / 0.70</td>
<td id="76905DC0FB66005FFC53F5E0FB05E8F2" box="[1010,1203,911,934]" gridcol="4" gridrow="15" pageId="20" pageNumber="19">3.10 / 0.90</td>
<td id="76905DC0FB66005FFB6BF5E0FA26E8F2" box="[1226,1424,911,934]" gridcol="5" gridrow="15" pageId="20" pageNumber="19">2.50 / 0.70</td>
</tr>
<tr id="354134BCFB66005FFF6CF5D7FA26E89B" box="[205,1424,952,975]" gridrow="16" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF5D7FE14E89B" box="[205,418,952,975]" gridcol="0" gridrow="16" pageId="20" pageNumber="19">Patella</th>
<td id="76905DC0FB66005FFE1FF5D7FD92E89B" box="[446,548,952,975]" gridcol="1" gridrow="16" pageId="20" pageNumber="19">L / W</td>
<td id="76905DC0FB66005FFDE7F5D7FD4AE89B" box="[582,764,952,975]" gridcol="2" gridrow="16" pageId="20" pageNumber="19">3.33 / 0.93</td>
<td id="76905DC0FB66005FFCBBF5D7FC6DE89B" box="[794,987,952,975]" gridcol="3" gridrow="16" pageId="20" pageNumber="19">3.40 / 0.90</td>
<td id="76905DC0FB66005FFC53F5D7FB05E89B" box="[1010,1203,952,975]" gridcol="4" gridrow="16" pageId="20" pageNumber="19">3.60 / 1.10</td>
<td id="76905DC0FB66005FFB6BF5D7FA26E89B" box="[1226,1424,952,975]" gridcol="5" gridrow="16" pageId="20" pageNumber="19">2.90 / 0.90</td>
</tr>
<tr id="354134BCFB66005FFF6CF58DFA26E8AD" box="[205,1424,994,1017]" gridrow="17" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF58DFE14E8AD" box="[205,418,994,1017]" gridcol="0" gridrow="17" pageId="20" pageNumber="19">Chela</th>
<td id="76905DC0FB66005FFE1FF58DFD92E8AD" box="[446,548,994,1017]" gridcol="1" gridrow="17" pageId="20" pageNumber="19">L</td>
<td id="76905DC0FB66005FFDE7F58DFD4AE8AD" box="[582,764,994,1017]" gridcol="2" gridrow="17" pageId="20" pageNumber="19">4.40</td>
<td id="76905DC0FB66005FFCBBF58DFC6DE8AD" box="[794,987,994,1017]" gridcol="3" gridrow="17" pageId="20" pageNumber="19">4.80</td>
<td id="76905DC0FB66005FFC53F58DFB05E8AD" box="[1010,1203,994,1017]" gridcol="4" gridrow="17" pageId="20" pageNumber="19">4.70</td>
<td id="76905DC0FB66005FFB6BF58DFA26E8AD" box="[1226,1424,994,1017]" gridcol="5" gridrow="17" pageId="20" pageNumber="19">4.10</td>
</tr>
<tr id="354134BCFB66005FFF6CF264FA26EF76" box="[205,1424,1035,1058]" gridrow="18" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF264FE14EF76" box="[205,418,1035,1058]" gridcol="0" gridrow="18" pageId="20" pageNumber="19">Manus</th>
<td id="76905DC0FB66005FFE1FF264FD92EF76" box="[446,548,1035,1058]" gridcol="1" gridrow="18" pageId="20" pageNumber="19">L / W / D</td>
<td id="76905DC0FB66005FFDE7F264FD4AEF76" box="[582,764,1035,1058]" gridcol="2" gridrow="18" pageId="20" pageNumber="19">1.22 / 0.71 / 0.75</td>
<td id="76905DC0FB66005FFCBBF264FC6DEF76" box="[794,987,1035,1058]" gridcol="3" gridrow="18" pageId="20" pageNumber="19">1.50 / 0.70 / 0.70</td>
<td id="76905DC0FB66005FFC53F264FB05EF76" box="[1010,1203,1035,1058]" gridcol="4" gridrow="18" pageId="20" pageNumber="19">1.40 / 0.80 / 0.90</td>
<td id="76905DC0FB66005FFB6BF264FA26EF76" box="[1226,1424,1035,1058]" gridcol="5" gridrow="18" pageId="20" pageNumber="19">1.20 / 0.70 / 0.70</td>
</tr>
<tr id="354134BCFB66005FFF6CF25BFA26EF1F" box="[205,1424,1076,1099]" gridrow="19" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF25BFE14EF1F" box="[205,418,1076,1099]" gridcol="0" gridrow="19" pageId="20" pageNumber="19">Movable finger</th>
<td id="76905DC0FB66005FFE1FF25BFD92EF1F" box="[446,548,1076,1099]" gridcol="1" gridrow="19" pageId="20" pageNumber="19">L</td>
<td id="76905DC0FB66005FFDE7F25BFD4AEF1F" box="[582,764,1076,1099]" gridcol="2" gridrow="19" pageId="20" pageNumber="19">3.18</td>
<td id="76905DC0FB66005FFCBBF25BFC6DEF1F" box="[794,987,1076,1099]" gridcol="3" gridrow="19" pageId="20" pageNumber="19">3.30</td>
<td id="76905DC0FB66005FFC53F25BFB05EF1F" box="[1010,1203,1076,1099]" gridcol="4" gridrow="19" pageId="20" pageNumber="19">3.30</td>
<td id="76905DC0FB66005FFB6BF25BFA26EF1F" box="[1226,1424,1076,1099]" gridcol="5" gridrow="19" pageId="20" pageNumber="19">2.90</td>
</tr>
<tr id="354134BCFB66005FFF6CF208FA26EF2A" box="[205,1424,1127,1150]" gridrow="20" pageId="20" pageNumber="19">
<th id="76905DC0FB66005FFF6CF208FE14EF2A" box="[205,418,1127,1150]" gridcol="0" gridrow="20" pageId="20" pageNumber="19">Total</th>
<td id="76905DC0FB66005FFE1FF208FD92EF2A" box="[446,548,1127,1150]" gridcol="1" gridrow="20" pageId="20" pageNumber="19">L</td>
<td id="76905DC0FB66005FFDE7F208FD4AEF2A" box="[582,764,1127,1150]" gridcol="2" gridrow="20" pageId="20" pageNumber="19">32.20</td>
<td id="76905DC0FB66005FFCBBF208FC6DEF2A" box="[794,987,1127,1150]" gridcol="3" gridrow="20" pageId="20" pageNumber="19">31.50</td>
<td id="76905DC0FB66005FFC53F208FB05EF2A" box="[1010,1203,1127,1150]" gridcol="4" gridrow="20" pageId="20" pageNumber="19">36.60</td>
<td id="76905DC0FB66005FFB6BF208FA26EF2A" box="[1226,1424,1127,1150]" gridcol="5" gridrow="20" pageId="20" pageNumber="19">28.40</td>
</tr>
</table>
</paragraph>
<caption id="DF0E6676FB66FFB4FF61F2DCFE29EFB7" pageId="20" pageNumber="19" startId="20.[192,257,1203,1226]">
<paragraph id="8BCE36FEFB66FFB4FF61F2DCFE29EFB7" blockId="20.[192,1440,1203,1251]" pageId="20" pageNumber="19">
<emphasis id="B905EAECFB66FFB4FF61F2DCFE97EF9E" bold="true" box="[192,289,1203,1226]" pageId="20" pageNumber="19">Table 2:</emphasis>
Measurements of four types of 
<taxonomicName id="4C714D7DFB66FFB4FDC2F2DAFC35EF9E" authority="Teruel &amp; Kovařík &amp; Fet, 2018" authorityName="Teruel &amp; Kovařík &amp; Fet" authorityYear="2018" box="[611,899,1205,1226]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="krivochatskyi" status="sp. nov.">
<emphasis id="B905EAECFB66FFB4FDC2F2DAFC35EF9E" box="[611,899,1205,1226]" italics="true" pageId="20" pageNumber="19">Anomalobuthus krivochatskyi</emphasis>
</taxonomicName>
<emphasis id="B905EAECFB66FFB4FC2DF2DAFC72EF9E" bold="true" box="[908,964,1205,1226]" pageId="20" pageNumber="19">
<taxonomicNameLabel id="A2365797FB66FFB4FC2DF2DAFC72EF9E" box="[908,964,1205,1226]" pageId="20" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Abbreviations: length (L), width (W), posterior width (Wp), depth (D).
</paragraph>
</caption>
<paragraph id="8BCE36FEFB66FFB4FF51F04DFDD3ED0C" blockId="20.[192,802,1570,1869]" pageId="20" pageNumber="19">
VARIATION. Adult size varied from 
<quantity id="4C899B1BFB66FFB4FD22F04DFC94ED6D" box="[643,802,1570,1593]" metricMagnitude="-2" metricUnit="m" metricValue="3.185" metricValueMax="3.22" metricValueMin="3.15" pageId="20" pageNumber="19" unit="mm" value="31.85" valueMax="32.2" valueMin="31.5">31.5–32.2 mm</quantity>
in males and 
<quantity id="4C899B1BFB66FFB4FEECF02EFE5EED0C" box="[333,488,1601,1624]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.66" metricValueMin="2.84" pageId="20" pageNumber="19" unit="mm" value="32.5" valueMax="36.6" valueMin="28.4">28.4–36.6 mm</quantity>
in females.
</paragraph>
<paragraph id="8BCE36FEFB66FFB4FF51F00FFF69EDA5" blockId="20.[192,802,1570,1869]" pageId="20" pageNumber="19">
Count of teeth per pecten varied as follows (
<tableCitation id="C6F30345FB66FFB4FD6DF00FFCA5ED23" box="[716,787,1632,1655]" captionStart="Table 5" captionStartId="38.[192,257,680,703]" captionText="Table 5: Pectinal tooth count variation in all known species of Anomalobuthus. Abbreviations: number of pectines (N), average (Ave.), standard deviation (SD). This includes counts taken from sources with precise and reliable geographic data (Birula, 1905, 1910)." pageId="20" pageNumber="19">Tab. 5</tableCitation>
): in males 22 (3), 23 (1), 24 (3), 25 (3), and in females 19 (2), 20 (7), 21 (6), 22 (3). No significant differences detected among specimens from the localities examined by us.
</paragraph>
<paragraph id="8BCE36FEFB66FFB4FF51F096FCABEC19" blockId="20.[192,802,1570,1869]" pageId="20" pageNumber="19">
The tibial spur is always absent in males, but in females it is present always on both leg IV and occasionally also in leg III of one side only (
<figureCitation id="134A2A7BFB66FFB4FD9FF159FD6DEC19" box="[574,731,1846,1869]" captionStart="Figures 126–133" captionStartId="36.[192,278,1688,1711]" captionTargetBox="[191,1440,167,1659]" captionTargetId="figure-12@36.[192,1439,168,1658]" captionTargetPageId="36" captionText="Figures 126–133: Anomalobuthus spp., SEM views of right pedipalp of adult male A. rickmersi from Repetek (126–127) and adult male paratopotype A. krivochatskyi sp. n. (128–133): chela (126, 128), close-up of manus (127, 129), three ventrointernal magnified views of mid part of both fingers in consecutive figures (130–132, movable finger on top), ventrointernal close-up of fingertips (133, movable finger on top). See trichobothria highlighted in red." pageId="20" pageNumber="19">Figs. 130–132</figureCitation>
, 
<figureCitation id="134A2A7BFB66FFB4FD44F159FCA7EC19" box="[741,785,1846,1869]" captionStart="Figures 134–140" captionStartId="37.[192,278,1748,1771]" captionTargetBox="[191,1440,191,1719]" captionTargetId="figure-6@37.[192,1439,192,1718]" captionTargetPageId="37" captionText="Figures 134–140: Anomalobuthus spp., SEM views showing size variation of leg IV tibial spur (highlighted in red): large in paratopotype female A. krivochatskyi sp. n. (134), moderately large in paratype female A. krivochatskyi sp. n. from Bukhara (135), small in paratopotype female A. krivochatskyi sp. n. (136), absent in male A. rickmersi from Repetek (137), close-up of a medium-sized leg IV tibial spur of a paratopotype female A. krivochatskyi sp. n. from Zarafshan (138), see spinulate and striate sculpture. Anomalobuthus krivochatskyi sp. n., adult female paratopotype: SEM views of right pecten and adjacent parts of leg coxae and sternite III (139), peg sensilla of tooth from same pecten (140)." pageId="20" pageNumber="19">134</figureCitation>
).
</paragraph>
<paragraph id="8BCE36FEFB66FFB4FCCFF360FAFDEEEB" blockId="20.[830,1440,1295,1471]" pageId="20" pageNumber="19">
Specimens from Chardara and Bukhara to Gazli are identical to those from the 
<typeStatus id="54CA885CFB66FFB4FBC5F342FB24EE10" box="[1124,1170,1325,1348]" pageId="20" pageNumber="19">type</typeStatus>
locality in all diagnostic characters. They only differ by being slightly smaller and darker, with better-marked dark patterns (
<figureCitation id="134A2A7BFB66FFB4FA98F304FCEAEEF4" captionStart-0="Figures 38–41" captionStart-1="Figures 42–45" captionStartId-0="14.[192,278,1754,1777]" captionStartId-1="16.[192,278,1584,1607]" captionTargetBox-0="[201,1418,199,1682]" captionTargetBox-1="[212,1422,194,1547]" captionTargetId-0="figure-12@14.[192,1439,192,1697]" captionTargetId-1="figure-12@16.[192,1440,192,1554]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionText-0="Figures 38–41: Anomalobuthus krivochatskyi sp. n., adult male (38–39) and adult female paratypes (40–41) from between Bukhara and Gazli, Uzbekistan: full-body views, dorsal (38, 40) and ventral (39, 41). Scale bar = 10 mm." captionText-1="Figures 42–45: Anomalobuthus krivochatskyi sp. n., adult female (42–43) and juvenile male paratypes (44–45) from Chardara, Kazakhstan: full-body views, dorsal (42, 44) and ventral (43, 45)." pageId="20" pageNumber="19">Figs. 38– 45</figureCitation>
). These differences are minor and attributable to standard variation between different populations.
</paragraph>
<paragraph id="8BCE36FEFB66FFB4FCCFF38AFB4FEDE0" blockId="20.[830,1441,1509,1869]" pageId="20" pageNumber="19">
COMPARISON. Adults of 
<taxonomicName id="4C714D7DFB66FFB4FBDCF38AFA9DEEA8" authority="Teruel &amp; Kovařík &amp; Fet, 2018" authorityName="Teruel &amp; Kovařík &amp; Fet" authorityYear="2018" box="[1149,1323,1509,1532]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="krivochatskyi" status="sp. nov.">
<emphasis id="B905EAECFB66FFB4FBDCF38AFA9DEEA8" box="[1149,1323,1509,1532]" italics="true" pageId="20" pageNumber="19">A. krivochatskyi</emphasis>
</taxonomicName>
<emphasis id="B905EAECFB66FFB4FA95F38AFAC4EEA8" bold="true" box="[1332,1394,1509,1532]" pageId="20" pageNumber="19">
<taxonomicNameLabel id="A2365797FB66FFB4FA95F38AFAC4EEA8" box="[1332,1394,1509,1532]" pageId="20" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be very easily distinguished by the almost absolute smoothness of the metasomal intercarinal spaces, especially in males (
<figureCitation id="134A2A7BFB66FFB4FBB2F02EFB2FED0C" box="[1043,1177,1601,1624]" captionStart="Figures 59–64" captionStartId="19.[192,278,1542,1565]" captionTargetBox="[204,1417,183,1488]" captionTargetId="figure-6@19.[192,1437,168,1512]" captionTargetPageId="19" captionText="Figures 59–64: Anomalobuthus krivochatskyi sp. n., adult male holotype (59–61) and adult female paratopotype (62–64): metasoma and telson, lateral (59, 62), dorsal (60, 63) and ventral (61, 64)." pageId="20" pageNumber="19">Figs. 59–64</figureCitation>
, 
<figureCitation id="134A2A7BFB66FFB4FB0BF02EFB63ED0C" box="[1194,1237,1601,1624]" captionStart="Figures 120–125" captionStartId="34.[192,278,1734,1757]" captionTargetBox="[191,1440,200,1705]" captionTargetId="figure-12@34.[192,1439,201,1704]" captionTargetPageId="34" captionText="Figures 120–125: Anomalobuthus krivochatskyi sp. n., adult male paratopotype: SEM views of chelicerae, dorsal and ventral (120), metasoma V and telson, lateral (121); anterior half of carapace, dorsal (122); close-up of trichobothria (123); left leg IV basitarsus andtelotarsus, ventrointernal (124); left leg IV basitarsus and telotarsus, external (125). Images courtesy Jan Štundl." pageId="20" pageNumber="19">121</figureCitation>
); almost all other congeners have at least the ventral and lateral surfaces of segments IV–V moderately to densely granulose (
<figureCitation id="134A2A7BFB66FFB4FACAF011FC35EDE0" captionStart="Figures 14–26" captionStartId="9.[192,278,1609,1632]" captionTargetBox="[196,1409,215,1573]" captionTargetId="figure-6@9.[192,1439,192,1580]" captionTargetPageId="9" captionText="Figures 14–26: Anomalobuthus rickmersi, adult male holotype: right pedipalp trochanter and femur, dorsal (14) and ventral (15), right pedipalp patella, dorsal (16), external (17) and ventral (18), right pedipalp chela, dorsal (19–20), external (21) and ventrointernal (22), metasoma and telson, lateral (23), dorsal (24) and ventral (25), telson, lateral (26)." pageId="20" pageNumber="19">Figs. 23–25</figureCitation>
, 
<figureCitation id="134A2A7BFB66FFB4FC2FF0F2FB95EDE0" box="[910,1059,1693,1716]" captionStart="Figures 27–33" captionStartId="10.[192,278,1394,1417]" captionTargetBox="[192,1437,192,1364]" captionTargetId="figure-12@10.[192,1437,192,1364]" captionTargetPageId="10" captionText="Figures 27–33: Anomalobuthus rickmersi from Turkmenistan, adult male from Repetek (27–30) and adult female from Chainury (31–33): metasoma and telson, lateral (27, 31), dorsal (28, 32) and ventral (29, 33), telson, lateral (30)." pageId="20" pageNumber="19">27–29, 31–33</figureCitation>
, 
<figureCitation id="134A2A7BFB66FFB4FB8FF0F2FBFDEDE0" box="[1070,1099,1693,1716]" captionStart="Figures 65–74" captionStartId="22.[192,278,1590,1613]" captionTargetBox="[191,1437,215,1561]" captionTargetId="figure-12@22.[192,1437,216,1560]" captionTargetPageId="22" captionText="Figures 65–74: Anomalobuthus lowei sp. n., adult female holotype. Figures 65–66. full-body dorsal views, under white light (65) and under UV light (66). Figures 67–68. Mesosoma dorsal (67) and ventral (68) under UV light. Figure 69. Metasoma and telson under UV light, lateral. Figures 70–74. Pedipalp under UV light: right pedipalp trochanter and femur, dorsal (70), right pedipalp patella, dorsal (71), right pedipalp chela, dorsal (72), external (73) and ventral (74). Scale bars = 10 mm (65–66) and 1 mm (67–68, 70–74). Photographs courtesy Graeme Lowe." pageId="20" pageNumber="19">69</figureCitation>
, 
<figureCitation id="134A2A7BFB66FFB4FBF7F0F2FB2DEDE0" box="[1110,1179,1693,1716]" captionStart="Figures 75–78" captionStartId="23.[192,278,1060,1083]" captionTargetBox="[191,1437,191,1030]" captionTargetId="figure-9@23.[192,1437,595,1029]" captionTargetPageId="23" captionText="Figures 75–78: Anomalobuthus lowei sp. n., adult female holotype. Figures 75–76. Metasoma and telson, dorsal, under white light (75) and under UV light (76). Figures 77–78. Metasoma and telson, ventral, under white light (77) and under UV light (78). Scale bar = 2 mm. Photographs courtesy Graeme Lowe." pageId="20" pageNumber="19">75–78</figureCitation>
, 
<figureCitation id="134A2A7BFB66FFB4FB07F0F2FB58EDE0" box="[1190,1262,1693,1716]" captionStart="Figures 83–89" captionStartId="27.[192,278,1682,1705]" captionTargetBox="[199,1433,180,1618]" captionTargetId="figure-6@27.[192,1439,168,1653]" captionTargetPageId="27" captionText="Figures 83–89: Anomalobuthus pavlovskyi sp. n., adult male (83–86) and adult female paratopotypes (87–89): right pedipalp (83), metasoma and telson, lateral (84, 87), dorsal (85, 88) and ventral (86, 89)." pageId="20" pageNumber="19">84–89</figureCitation>
).
</paragraph>
<paragraph id="8BCE36FEFB66FFB4FCCFF0D3FB46EC19" blockId="20.[830,1441,1509,1869]" pageId="20" pageNumber="19">
The single exception is 
<taxonomicName id="4C714D7DFB66FFB4FBCEF0D3FB67ED87" authorityName="Teruel, Kovarik, Navidpour et Fet" authorityYear="2014" box="[1135,1233,1724,1747]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="talebii">
<emphasis id="B905EAECFB66FFB4FBCEF0D3FB67ED87" box="[1135,1233,1724,1747]" italics="true" pageId="20" pageNumber="19">A. talebii</emphasis>
</taxonomicName>
, which is similarly smooth (
<figureCitation id="134A2A7BFB66FFB4FC3EF0B5FB84EDA5" box="[927,1074,1754,1777]" captionStart="Figures 92–100" captionStartId="31.[192,278,1669,1692]" captionTargetBox="[191,1440,191,1639]" captionTargetId="figure-6@31.[192,1439,192,1638]" captionTargetPageId="31" captionText="Figures 92–100: Anomalobuthus talebii, adult female holotype: chelicerae, carapace and tergites I–III, dorsal (92), sternopectinal region and sternites III–V, ventral (93), distal part of right legs I–IV, internal (94–97), metasoma and telson, lateral (98), dorsal (99) and ventral (100)." pageId="20" pageNumber="19">Figs. 98–100</figureCitation>
), but the only female known has much higher pectinal tooth count (26/26 
<emphasis id="B905EAECFB66FFB4FAA1F096FAABEC44" box="[1280,1309,1785,1808]" italics="true" pageId="20" pageNumber="19">vs.</emphasis>
20–23) and several quite distinct morphometric ratios (see 
<tableCitation id="C6F30345FB66FFB4FAF7F177FA16EC7B" box="[1366,1440,1816,1839]" captionStart="Table 4" captionStartId="35.[192,257,1030,1053]" captionText="Table 4: Comparison between five species of Anomalobuthus based upon selected morphometric ratios. Abbreviations: length (L)," pageId="20" pageNumber="19">Tab. 4</tableCitation>
and Comparison section of that species).
</paragraph>
</subSubSection>
<subSubSection id="C36B6575FB67FFB5FF51F6A8FDD7EA87" pageId="21" pageNumber="20" type="distribution">
<paragraph id="8BCE36FEFB67FFB5FF51F6A8FE61EA4F" blockId="21.[192,802,199,283]" pageId="21" pageNumber="20">
DISTRIBUTION (
<figureCitation id="134A2A7BFB67FFB5FE3CF6A8FE49EB8A" box="[413,511,199,222]" captionStart="Figures 134–140" captionStartId="37.[192,278,1748,1771]" captionTargetBox="[191,1440,191,1719]" captionTargetId="figure-6@37.[192,1439,192,1718]" captionTargetPageId="37" captionText="Figures 134–140: Anomalobuthus spp., SEM views showing size variation of leg IV tibial spur (highlighted in red): large in paratopotype female A. krivochatskyi sp. n. (134), moderately large in paratype female A. krivochatskyi sp. n. from Bukhara (135), small in paratopotype female A. krivochatskyi sp. n. (136), absent in male A. rickmersi from Repetek (137), close-up of a medium-sized leg IV tibial spur of a paratopotype female A. krivochatskyi sp. n. from Zarafshan (138), see spinulate and striate sculpture. Anomalobuthus krivochatskyi sp. n., adult female paratopotype: SEM views of right pecten and adjacent parts of leg coxae and sternite III (139), peg sensilla of tooth from same pecten (140)." pageId="21" pageNumber="20">Fig. 137</figureCitation>
). Sands of the Kizylkum Desert, from central 
<collectingCountry id="F366766EFB67FFB5FE0DF689FD92EBA9" box="[428,548,230,253]" name="Uzbekistan" pageId="21" pageNumber="20">Uzbekistan</collectingCountry>
through adjacent southern edge of 
<collectingCountry id="F366766EFB67FFB5FEF2F76BFE65EA4F" box="[339,467,260,283]" name="Kazakhstan" pageId="21" pageNumber="20">Kazakhstan</collectingCountry>
.
</paragraph>
<paragraph id="8BCE36FEFB67FFB5FF51F72DFDD7EA87" blockId="21.[192,802,322,467]" pageId="21" pageNumber="20">
NOTES. The Bukhara Province (between Bukhara and Gazli) specimens of this new species were included (under 
<taxonomicName id="4C714D7DFB67FFB5FEADF710FE39EAC2" authorityName=": Soleglad &amp; Fet" authorityYear="2003" box="[268,399,383,406]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB67FFB5FEADF710FE39EAC2" box="[268,399,383,406]" italics="true" pageId="21" pageNumber="20">A. rickmersi</emphasis>
</taxonomicName>
) in a DNA phylogeographic study by 
<bibRefCitation id="EFE04B0FFB67FFB5FF61F7F1FE03EAE1" author="GRAHAM" box="[192,437,414,437]" pageId="21" pageNumber="20" pagination="95 - 110" refId="ref24752" refString="GRAHAM, M. R., V. OLAH-HEMMINGS &amp; V. FET. 2012. Phylogeography of co-distributed dune scorpions identifies the Amu Darya River as a longstanding component of Central Asian biogeography (Scorpiones: Buthidae). Zoology in the Middle East, 55: 95 - 110." type="journal article" year="2012">Graham et al. (2012)</bibRefCitation>
, which demonstrated a derived position of this taxon within the genus.
</paragraph>
</subSubSection>
</treatment>
</document>