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<document id="DBDFB053EBDA9140B8519FF481227CF2" ID-DOI="10.5281/zenodo.5313274" ID-ISSN="0374-1036" ID-Zenodo-Dep="5313274" ID-ZooBank="60855ED1-C243-4DA1-ADCC-8ADAA4752E9FM" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630120511396" checkinUser="marcus" docAuthor="Kolibáč, Jiří" docDate="2014" docId="03CD87BDDF4EF41FFE450F7FAF3C7087" docLanguage="en" docName="ActaEntMusNatPra.2014.54.suppl.197-210.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Afrocyrona volatilis Kolibáč 2014, sp. nov." docType="treatment" docVersion="4" lastPageNumber="205" masterDocId="FFF4FFC5DF4CF417FFA10E24AC4F7214" masterDocTitle="Two new species of Afrocyrona from Socotra Island (Coleoptera: Trogossitidae)" masterLastPageNumber="210" masterPageNumber="197" pageNumber="199" updateTime="1699160074818" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="D06F398CCAA562C6419F7C3D8747249C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C2BA712D8BB04BC96238A71A7FF3E19A">Two new species of Afrocyrona from Socotra Island (Coleoptera: Trogossitidae)</mods:title>
</mods:titleInfo>
<mods:name id="7534FEEA715E67C2E4F1EB52AA8F8ABB" type="personal">
<mods:role id="169A396D97024DC33532E9B6B16F7CDE">
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</mods:role>
<mods:namePart id="C3303840E16FE2F49F69C11A4D919628">Kolibáč, Jiří</mods:namePart>
</mods:name>
<mods:typeOfResource id="6477EB90D22F61C9701E1D674B37809A">text</mods:typeOfResource>
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<mods:title id="7660F75DD2592D809F050C6CD8B1AA6E">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="23D9DE9D7BEF31FE96A22FEB8515AB5E">
<mods:date id="3395D24BAC90D734579B4756A1AD3FAE">2014</mods:date>
<mods:detail id="7DA1E190A7E710E8983C2C4FBB22A5E5" type="pubDate">
<mods:number id="F97E483F67E10F2B5084A6CCFECF5248">2014-12-30</mods:number>
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<mods:number id="79F03D6D2CCD3888BA2D7C29D2CB0F20">54</mods:number>
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<mods:classification id="7BFC67EAA260F4C20C165602D63DE8FC">journal article</mods:classification>
<mods:identifier id="DBE69E4CDF1AACE0F42C773921DAB4FF" type="DOI">10.5281/zenodo.5313274</mods:identifier>
<mods:identifier id="107FBD1E9C7DA1790C1A7AE111A5D3FD" type="ISSN">0374-1036</mods:identifier>
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<treatment id="03CD87BDDF4EF41FFE450F7FAF3C7087" ID-DOI="http://doi.org/10.5281/zenodo.5449995" ID-Zenodo-Dep="5449995" LSID="urn:lsid:plazi:treatment:03CD87BDDF4EF41FFE450F7FAF3C7087" httpUri="http://treatment.plazi.org/id/03CD87BDDF4EF41FFE450F7FAF3C7087" lastPageId="8" lastPageNumber="205" pageId="2" pageNumber="199">
<subSubSection id="C37E6520DF4EF415FE450F7FAF557367" box="[484,794,347,371]" pageId="2" pageNumber="199" type="nomenclature">
<paragraph id="8BDB36ABDF4EF415FE450F7FAF557367" blockId="2.[484,794,347,371]" box="[484,794,347,371]" pageId="2" pageNumber="199">
<heading id="D09381C7DF4EF415FE450F7FAF557367" bold="true" box="[484,794,347,371]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="199" reason="3">
<emphasis id="B910EAB9DF4EF415FE450F7FAF557367" bold="true" box="[484,794,347,371]" pageId="2" pageNumber="199">
<taxonomicName id="4C644D28DF4EF415FE450F7FAEF17367" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[484,702,347,371]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF4EF415FE450F7FAEF17367" bold="true" box="[484,702,347,371]" italics="true" pageId="2" pageNumber="199">Afrocyrona volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF4EF415FD640F78AF557367" box="[709,794,348,371]" pageId="2" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E6520DF4EF415FD800F5EAE927399" box="[545,733,378,397]" pageId="2" pageNumber="199" type="description">
<paragraph id="8BDB36ABDF4EF415FD800F5EAE927399" blockId="2.[151,1127,378,572]" box="[545,733,378,397]" pageId="2" pageNumber="199">
(
<figureCitation id="135F2A2EDF4EF415FD860F5EAE3C7399" box="[551,627,378,397]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="3.[113,144,1514,1533]" captionStartId-1="4.[151,182,1514,1533]" captionStartId-2="6.[151,182,1509,1528]" captionTargetBox-0="[114,1088,225,1490]" captionTargetBox-1="[151,1126,230,1488]" captionTargetBox-2="[164,1118,223,1493]" captionTargetId-0="figure-13@3.[114,1088,219,1493]" captionTargetId-1="figure-11@4.[151,1129,219,1496]" captionTargetId-2="figure-11@6.[152,1126,219,1493]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="Fig. 1. Afrocyrona volatilis sp. nov., female paratype:A – labium (mentum removed); B – right mandible ventrally; C – maxilla; D – labrum; E – protarsus; F – procoxa with trochanter. Scale bars = 0.1 mm. For abbreviations see Material and methods." captionText-1="Fig. 2. Afrocyrona volatilis sp. nov., female paratype:A – cranium ventrally (with one maxilla, mandible and mentum); B – antenna, detail; C – prothorax ventrally; D – meso- and metathorax ventrally; E – right elytron, ventral view. For abbreviations see Material and methods." captionText-2="Fig. 3. Afrocyrona volatilis sp. nov., female paratype: A – wing; B – sternite VIII; C – tergite VIII; D – internal copulatory organs; E – ovipositor, apical part; F – labium. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5313278" figureDoi-1="http://doi.org/10.5281/zenodo.5313280" figureDoi-2="http://doi.org/10.5281/zenodo.5313286" httpUri-0="https://zenodo.org/record/5313278/files/figure.png" httpUri-1="https://zenodo.org/record/5313280/files/figure.png" httpUri-2="https://zenodo.org/record/5313286/files/figure.png" pageId="2" pageNumber="199">
Figs 1–
<geoCoordinate id="EE50506CDF4EF415FDC90F5EAE3C7399" box="[616,627,378,397]" degrees="3, 4" direction="east" orientation="longitude" pageId="2" pageNumber="199" precision="5555" value="3.4">3</geoCoordinate>
</figureCitation>
,
<figureCitation id="135F2A2EDF4EF415FDDD0F5EAEFA7399" box="[636,693,378,397]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. A–D – Afrocyrona pedestris sp. nov., male holotype:A – aedeagus ventrally; B – aedeagus laterally; C – rudimentary wing; D – protarsus. E – A. volatilis sp. nov., paratype; F – A. volatilis sp. nov., holotype; G – A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="2" pageNumber="199">4E, 4F</figureCitation>
,
<figureCitation id="135F2A2EDF4EF415FD1F0F5EAE997399" box="[702,726,378,397]" captionStart="Fig" captionStartId="11.[113,144,1208,1227]" captionTargetBox="[130,1087,224,1178]" captionTargetId="figure-13@11.[113,1091,219,1183]" captionTargetPageId="11" captionText="Fig. 5. A–D – Afrocyrona pedestris sp. nov., male holotype: A – abdominal segment IX with spiculum of sternite IX (‘spicular fork’); B – tegmen ventrally; C – sternite VIII; D – tergite VIII. E–H – A. volatilis sp. nov., female paratype: E – lacinia (incl. basistipes) and galea (incl. basigalea). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5313292" httpUri="https://zenodo.org/record/5313292/files/figure.png" pageId="2" pageNumber="199">5E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37E6520DF4EF415FF360F80AD297028" pageId="2" pageNumber="199" type="materials_examined">
<paragraph id="8BDB36ABDF4EF415FF360F80AD297028" blockId="2.[151,1127,378,572]" pageId="2" pageNumber="199">
<emphasis id="B910EAB9DF4EF415FF360F80AD5273A3" bold="true" box="[151,285,420,439]" pageId="2" pageNumber="199">Type material.</emphasis>
<materialsCitation id="3B0C3CF6DF4EF415FE830F80AE9973F8" collectingDate="2012-06-16" collectingDateMax="2012-06-18" collectingDateMin="2012-06-16" collectionCode="SCAND, V, NMPC" collectorName="J. Bezdek &amp; J. Hajek &amp; Hula &amp; P. Kment &amp; I. Malenovsky &amp; J. Niedobova &amp; L. Purchart" country="Yemen" latitude="12.576667" location="Hagher Mts." longLatPrecision="129" longitude="54.025" pageId="2" pageNumber="199" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54DF8809DF4EF415FE830F80AD3973A1" box="[290,374,420,439]" pageId="2" pageNumber="199" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D62FD22DF4EF415FE230FBAADDD73A3" box="[386,402,414,439]" pageId="2" pageNumber="199" type="female">♀</specimenCount>
, ‘
<collectingCountry id="F373763BDF4EF415FE010F80ADBD73A3" box="[416,498,420,439]" name="Yemen" pageId="2" pageNumber="199">YEMEN</collectingCountry>
, SOCOTRA Island /
<location id="8EBB6070DF4EF415FD300F80AEB073A3" LSID="urn:lsid:plazi:treatment:03CD87BDDF4EF41FFE450F7FAF3C7087:8EBB6070DF4EF415FD300F80AEB073A3" box="[657,767,420,439]" country="Yemen" latitude="12.576667" longLatPrecision="129" longitude="54.025" name="Hagher Mts." pageId="2" pageNumber="199">Hagher Mts.</location>
,
<collectionCode id="ED75AE6EDF4EF415FCA90F80AF7173A1" box="[776,830,420,439]" pageId="2" pageNumber="199">SCAND</collectionCode>
Mt. env. / broad-leaved evergreen woodland /
<date id="FFDA106BDF4EF415FF580F9BAD3673C6" box="[249,377,447,466]" pageId="2" pageNumber="199" value="2012-06-16" valueMax="2012-06-18" valueMin="2012-06-16">
<collectingDate id="EF9EE983DF4EF415FF580F9BAD3673C6" box="[249,377,447,466]" pageId="2" pageNumber="199" value="2012-06-16" valueMax="2012-06-18" valueMin="2012-06-16">16.–18.vi.2012</collectingDate>
</date>
/
<geoCoordinate id="EE50506CDF4EF415FE260F9BAD9073C6" box="[391,479,447,466]" degrees="12" direction="north" minutes="34.6" orientation="latitude" pageId="2" pageNumber="199" precision="92" value="12.576667">12°34.6′N</geoCoordinate>
,
<geoCoordinate id="EE50506CDF4EF415FE460F9BAE7273C6" box="[487,573,447,466]" degrees="54" direction="east" minutes="01.5" orientation="longitude" pageId="2" pageNumber="199" precision="92" value="54.025">54°01.5′E</geoCoordinate>
, 1450 m’; ‘Socotra expedition 2012 /
<collectorName id="2691537DDF4EF415FCD70F9BAF8773C6" box="[886,968,447,466]" pageId="2" pageNumber="199">J. Bezděk</collectorName>
,
<collectorName id="2691537DDF4EF415FC710F9BA85873C6" box="[976,1047,447,466]" pageId="2" pageNumber="199">J. Hájek</collectorName>
,
<collectionCode id="ED75AE6EDF4EF415FBBC0F9BA86373C6" box="[1053,1068,447,466]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="199" type="Museum">V</collectionCode>
.
<collectorName id="2691537DDF4EF415FB920F9BA81373C6" box="[1075,1116,447,466]" pageId="2" pageNumber="199">Hula</collectorName>
/
<collectorName id="2691537DDF4EF415FF360FFDACA973F8" box="[151,230,473,492]" pageId="2" pageNumber="199">P. Kment</collectorName>
,
<collectorName id="2691537DDF4EF415FF4C0FFDAD2873F8" box="[237,359,473,492]" pageId="2" pageNumber="199">I. Malenovský</collectorName>
/
<collectorName id="2691537DDF4EF415FED40FFDADAA73F8" box="[373,485,473,492]" pageId="2" pageNumber="199">J. Niedobová</collectorName>
&amp;
<collectorName id="2691537DDF4EF415FE5C0FFDAE1373F8" box="[509,604,473,492]" pageId="2" pageNumber="199">L. Purchart</collectorName>
leg.’ (
<collectionCode id="ED75AE6EDF4EF415FD2E0FFDAE9F73F8" box="[655,720,473,492]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="199">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B0C3CF6DF4EF415FD7C0FFDACAE7036" collectionCode="WADI, MADAR" country="Yemen" latitude="12.553333" location="Hagher Mts." longLatPrecision="129" longitude="54.006668" pageId="2" pageNumber="199" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54DF8809DF4EF415FD7C0FFDAF6373FF" box="[733,812,473,492]" pageId="2" pageNumber="199" type="paratype">PARATYPE</typeStatus>
:
<specimenCount id="9D62FD22DF4EF415FC970FF7AF0973F8" box="[822,838,467,492]" pageId="2" pageNumber="199" type="female">♀</specimenCount>
, ‘
<collectingCountry id="F373763BDF4EF415FCF30FFDAFEB73F8" box="[850,932,473,492]" name="Yemen" pageId="2" pageNumber="199">YEMEN</collectingCountry>
, SOCOTRA Island, 18.vi. /
<location id="8EBB6070DF4EF415FF020FD0AD5E7013" LSID="urn:lsid:plazi:treatment:03CD87BDDF4EF41FFE450F7FAF3C7087:8EBB6070DF4EF415FF020FD0AD5E7013" box="[163,273,500,519]" country="Yemen" latitude="12.553333" longLatPrecision="129" longitude="54.006668" name="Hagher Mts." pageId="2" pageNumber="199">Hagher Mts.</location>
,
<collectionCode id="ED75AE6EDF4EF415FEB80FD0ADC47011" box="[281,395,500,519]" pageId="2" pageNumber="199">WADI MADAR</collectionCode>
, 2012 / montane shrubland with /
<taxonomicName id="4C644D28DF4EF415FD140FD0AFD77013" box="[693,920,500,519]" class="Magnoliopsida" family="Euphorbiaceae" genus="Cephalocroton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="199" phylum="Tracheophyta" rank="species" species="socotranus">
<emphasis id="B910EAB9DF4EF415FD140FD0AFD77013" box="[693,920,500,519]" italics="true" pageId="2" pageNumber="199">Cephalocroton socotranus</emphasis>
</taxonomicName>
/
<geoCoordinate id="EE50506CDF4EF415FC0B0FD0A84D7013" box="[938,1026,500,519]" degrees="12" direction="north" minutes="33.2" orientation="latitude" pageId="2" pageNumber="199" precision="92" value="12.553333">12°33.2′N</geoCoordinate>
,
<geoCoordinate id="EE50506CDF4EF415FBAD0FD0A82C7013" box="[1036,1123,500,519]" degrees="54" direction="east" minutes="00.4" orientation="longitude" pageId="2" pageNumber="199" precision="92" value="54.006668">54°00.4′E</geoCoordinate>
, 1170 m’
</materialsCitation>
; ‘
<materialsCitation id="3B0C3CF6DF4EF415FF4F0C2BAD2D7028" collectionCode="V, MMBC" collectorName="J. Bezdek &amp; J. Hajek &amp; P. Kment &amp; I. Malenovsky &amp; J. Niedobova &amp; L. Purchart" country="Yemen" location="Socotra" pageId="2" pageNumber="199" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F373763BDF4EF415FF4F0C2BAD7C7036" box="[238,307,527,546]" name="Yemen" pageId="2" pageNumber="199">Socotra</collectingCountry>
expedition 2012 /
<collectorName id="2691537DDF4EF415FE760C2BAE627036" box="[471,557,527,546]" pageId="2" pageNumber="199">J. Bezděk</collectorName>
,
<collectorName id="2691537DDF4EF415FD970C2BAE307036" box="[566,639,527,546]" pageId="2" pageNumber="199">J. Hájek</collectorName>
,
<collectionCode id="ED75AE6EDF4EF415FD290C2BAED87036" box="[648,663,527,546]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="199" type="Museum">V</collectionCode>
. Hula /
<collectorName id="2691537DDF4EF415FD7D0C2BAF627036" box="[732,813,527,546]" pageId="2" pageNumber="199">P. Kment</collectorName>
,
<collectorName id="2691537DDF4EF415FC960C2BAFFC7036" box="[823,947,527,546]" pageId="2" pageNumber="199">I. Malenovský</collectorName>
/
<collectorName id="2691537DDF4EF415FC670C2BA8787036" box="[966,1079,527,546]" pageId="2" pageNumber="199">J. Niedobová</collectorName>
&amp;
<collectorName id="2691537DDF4EF415FBF50C2BAC907028" pageId="2" pageNumber="199">L. Purchart</collectorName>
leg.’ (
<collectionCode id="ED75AE6EDF4EF415FEB40C0DAD137028" box="[277,348,553,572]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="2" pageNumber="199">MMBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E6520DF4EF41FFF360C71AE557367" lastPageId="8" lastPageNumber="205" pageId="2" pageNumber="199" type="description">
<paragraph id="8BDB36ABDF4EF415FF360C71AE517098" blockId="2.[151,1127,597,1580]" pageId="2" pageNumber="199">
<emphasis id="B910EAB9DF4EF415FF360C71AD6D7078" bold="true" box="[151,290,597,620]" pageId="2" pageNumber="199">Description.</emphasis>
<emphasis id="B910EAB9DF4EF415FE870C71ADEA7078" box="[294,421,597,620]" italics="true" pageId="2" pageNumber="199">Body length</emphasis>
(from elytral apex to clypeus):
<typeStatus id="54DF8809DF4EF415FD4D0C71AF007078" box="[748,847,597,620]" pageId="2" pageNumber="199" type="holotype">Holotype</typeStatus>
<quantity id="4C9C9B4EDF4EF415FCF50C71AFE97078" box="[852,934,597,620]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="199" unit="mm" value="2.7">2.7 mm</quantity>
,
<typeStatus id="54DF8809DF4EF415FC0E0C71A8447078" box="[943,1035,597,620]" pageId="2" pageNumber="199" type="paratype">paratype</typeStatus>
<quantity id="4C9C9B4EDF4EF415FBB10C71A82C7078" box="[1040,1123,597,620]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="199" unit="mm" value="3.2">3.2 mm</quantity>
. For other measurements see
<tableCitation id="C6E60310DF4EF415FE690C51AE587098" box="[456,535,629,652]" captionStart="Table 1" captionStartId="7.[113,160,220,239]" captionText="Table 1. Measurements of Afrocyrona volatilis sp. nov. and A. pedestris sp. nov. (HT – holotype, PT – paratype)." pageId="2" pageNumber="199">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BDB36ABDF4EF415FF140CB1AD3B76B8" blockId="2.[151,1127,597,1580]" pageId="2" pageNumber="199">
<emphasis id="B910EAB9DF4EF415FF140CB1AD8B70B8" box="[181,452,661,684]" italics="true" pageId="2" pageNumber="199">Coloration and sculpture</emphasis>
(
<figureCitation id="135F2A2EDF4EF415FE750CB1AE0570B8" box="[468,586,661,684]" captionStart="Fig" captionStartId="4.[151,182,1514,1533]" captionTargetBox="[151,1126,230,1488]" captionTargetId="figure-11@4.[151,1129,219,1496]" captionTargetPageId="4" captionText="Fig. 2. Afrocyrona volatilis sp. nov., female paratype:A – cranium ventrally (with one maxilla, mandible and mentum); B – antenna, detail; C – prothorax ventrally; D – meso- and metathorax ventrally; E – right elytron, ventral view. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313280" httpUri="https://zenodo.org/record/5313280/files/figure.png" pageId="2" pageNumber="199">Figs 2C–E</figureCitation>
,
<figureCitation id="135F2A2EDF4EF415FDF70CB1AEDA70B8" box="[598,661,661,684]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. A–D – Afrocyrona pedestris sp. nov., male holotype:A – aedeagus ventrally; B – aedeagus laterally; C – rudimentary wing; D – protarsus. E – A. volatilis sp. nov., paratype; F – A. volatilis sp. nov., holotype; G – A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="2" pageNumber="199">4E–F</figureCitation>
): Body oval, moderately convex; all dorsal surface dark brown to black but yellow-brown lateral margins of pronotum, margins of elytra, two transverse patches in each elytron, legs, and antennomeres 1–6 (i.e., excepting club). Head finely, sparsely punctate (interspaces larger than diameter of punctures); punctures with thick decumbent whitish setae. Pronotum densely punctate (interspaces less than diameter of punctures), punctures larger than those on head; pronotum covered with thick semi-erect whitish setae; lateral margins finely punctate, with sparser pubescence. Each elytron with light L-shaped patch in humeral third and transverse patch in apical quarter, the latter not touching suture and side margin; elytral punctation regular, composed of about 14 rows, interspaces among rows inconspicuously elevated; pubescence similar to that on pronotum – composed of thick, semi-erect setae which are whitish in area of light patches and dark on brown-black background. Ventral surface sparsely punctate, punctures relatively large and distinctly separated (interspaces larger than diameter of punctures); anterior and posterior margins of cranium, hypomeral area of prothorax and basal margin of metaventrite without punctation. Pubescence of ventral surface scarce, conspicuous only on metaventrite, formed by short, sparse, decumbent pale setae. Legs without sculpture, sparsely pubescent in same way as metaventrite.
</paragraph>
<paragraph id="8BDB36ABDF4EF415FF140A91AFD27798" blockId="2.[151,1127,597,1580]" pageId="2" pageNumber="199">
<emphasis id="B910EAB9DF4EF415FF140A91ACA076D8" box="[181,239,1205,1228]" italics="true" pageId="2" pageNumber="199">Head</emphasis>
(
<figureCitation id="135F2A2EDF4EF415FF5F0A91AD1A76D8" box="[254,341,1205,1228]" captionStart="Fig" captionStartId="4.[151,182,1514,1533]" captionTargetBox="[151,1126,230,1488]" captionTargetId="figure-11@4.[151,1129,219,1496]" captionTargetPageId="4" captionText="Fig. 2. Afrocyrona volatilis sp. nov., female paratype:A – cranium ventrally (with one maxilla, mandible and mentum); B – antenna, detail; C – prothorax ventrally; D – meso- and metathorax ventrally; E – right elytron, ventral view. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313280" httpUri="https://zenodo.org/record/5313280/files/figure.png" pageId="2" pageNumber="199">Fig. 2A</figureCitation>
): almost hypognathous (position in picture artificial), frons distinctly flat; gular sutures widely separated at base, extending to approximately midpoint of cranium, strongly convergent; frontoclypeal suture inconspicuous or weak, not arcuate or emarginate; antennal grooves on ventral side of cranium short, shallow; antennal sockets not visible from above; eyes relatively small (space between them more than nine times as wide as eye diameter), distinctly elevated, not emarginate, elliptic, their greater part situated ventrally; epicranial acumination inconspicuous; cervical sclerites well-developed.
</paragraph>
<paragraph id="8BDB36ABDF4EF412FF140BB1ADE37327" blockId="2.[151,1127,597,1580]" lastBlockId="5.[113,1089,220,1555]" lastPageId="5" lastPageNumber="202" pageId="2" pageNumber="199">
<emphasis id="B910EAB9DF4EF415FF140BB1AD4577B8" box="[181,266,1429,1452]" italics="true" pageId="2" pageNumber="199">Maxilla</emphasis>
(
<figureCitation id="135F2A2EDF4EF415FEBA0BB1AD3877B8" box="[283,375,1429,1452]" captionStart="Fig" captionStartId="3.[113,144,1514,1533]" captionTargetBox="[114,1088,225,1490]" captionTargetId="figure-13@3.[114,1088,219,1493]" captionTargetPageId="3" captionText="Fig. 1. Afrocyrona volatilis sp. nov., female paratype:A – labium (mentum removed); B – right mandible ventrally; C – maxilla; D – labrum; E – protarsus; F – procoxa with trochanter. Scale bars = 0.1 mm. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313278" httpUri="https://zenodo.org/record/5313278/files/figure.png" pageId="2" pageNumber="199">Figs 1C</figureCitation>
,
<figureCitation id="135F2A2EDF4EF415FE240BB1ADE877B8" box="[389,423,1429,1452]" captionStart="Fig" captionStartId="11.[113,144,1208,1227]" captionTargetBox="[130,1087,224,1178]" captionTargetId="figure-13@11.[113,1091,219,1183]" captionTargetPageId="11" captionText="Fig. 5. A–D – Afrocyrona pedestris sp. nov., male holotype: A – abdominal segment IX with spiculum of sternite IX (‘spicular fork’); B – tegmen ventrally; C – sternite VIII; D – tergite VIII. E–H – A. volatilis sp. nov., female paratype: E – lacinia (incl. basistipes) and galea (incl. basigalea). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5313292" httpUri="https://zenodo.org/record/5313292/files/figure.png" pageId="2" pageNumber="199">5E</figureCitation>
): lacinia relatively robust, with seemingly three spines at apex in pattern 2+1 (two apical spines are part of body of lacinia while spine beneath them is modified seta), and with several thin and soft setae; basigalea relatively large, pigmented; galea weakly clavate, without spines and ciliate setae; with numerous thick and rigid elongate setae and thin soft setae among them; mediostipes perfectly free, not fused with lacinia, quadrangular in shape; basistipes probably fused with lacinia; palpifer relatively small, trapezoidal, not denticulate along outer margin; maxillary palps tetramerous, terminal palpomere conical, elongate.
</paragraph>
<caption id="DF1B6623DF4FF414FFD00BCEAD7D7426" ID-DOI="http://doi.org/10.5281/zenodo.5313278" ID-Zenodo-Dep="5313278" httpUri="https://zenodo.org/record/5313278/files/figure.png" pageId="3" pageNumber="200" startId="3.[113,144,1514,1533]" targetBox="[114,1088,225,1490]" targetPageId="3">
<paragraph id="8BDB36ABDF4FF414FFD00BCEAD7D7426" blockId="3.[113,1089,1514,1586]" pageId="3" pageNumber="200">
Fig. 1.
<taxonomicName id="4C644D28DF4FF414FF0C0BCEAD1877E9" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[173,343,1514,1533]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF4FF414FF0C0BCEAD1877E9" box="[173,343,1514,1533]" italics="true" pageId="3" pageNumber="200">Afrocyrona volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF4FF414FEFA0BCEADEF77E9" box="[347,416,1514,1533]" pageId="3" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype:A – labium (mentum removed); B – right mandible ventrally; C – maxilla; D – labrum; E – protarsus; F – procoxa with trochanter. Scale bars = 0.1 mm. For abbreviations see Material and methods.
</paragraph>
</caption>
<caption id="DF1B6623DF48F413FF360BCEAE087426" ID-DOI="http://doi.org/10.5281/zenodo.5313280" ID-Zenodo-Dep="5313280" httpUri="https://zenodo.org/record/5313280/files/figure.png" pageId="4" pageNumber="201" startId="4.[151,182,1514,1533]" targetBox="[151,1126,230,1488]" targetPageId="4">
<paragraph id="8BDB36ABDF48F413FF360BCEAE087426" blockId="4.[151,1127,1514,1586]" pageId="4" pageNumber="201">
Fig. 2.
<taxonomicName id="4C644D28DF48F413FF750BCEAD3277E9" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[212,381,1514,1533]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="201" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF48F413FF750BCEAD3277E9" box="[212,381,1514,1533]" italics="true" pageId="4" pageNumber="201">Afrocyrona volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF48F413FE230BCEAD8877E9" box="[386,455,1514,1533]" pageId="4" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype:A – cranium ventrally (with one maxilla, mandible and mentum); B – antenna, detail; C – prothorax ventrally; D – meso- and metathorax ventrally; E – right elytron, ventral view. For abbreviations see Material and methods.
</paragraph>
</caption>
<paragraph id="8BDB36ABDF49F412FF310F18AEC173A7" blockId="5.[113,1089,220,1555]" pageId="5" pageNumber="202">
<emphasis id="B910EAB9DF49F412FF310F18ACB97347" box="[144,246,316,339]" italics="true" pageId="5" pageNumber="202">Mandible</emphasis>
(
<figureCitation id="135F2A2EDF49F412FEA20F18AD1A7347" box="[259,341,316,339]" captionStart="Fig" captionStartId="3.[113,144,1514,1533]" captionTargetBox="[114,1088,225,1490]" captionTargetId="figure-13@3.[114,1088,219,1493]" captionTargetPageId="3" captionText="Fig. 1. Afrocyrona volatilis sp. nov., female paratype:A – labium (mentum removed); B – right mandible ventrally; C – maxilla; D – labrum; E – protarsus; F – procoxa with trochanter. Scale bars = 0.1 mm. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313278" httpUri="https://zenodo.org/record/5313278/files/figure.png" pageId="5" pageNumber="202">Fig. 1B</figureCitation>
): with two apical teeth situated side by side (in horizontal axis); medial tooth absent; mola absent; prostheca situated along cutting edge, formed by tuft of long, robust setae; transparent appendage (not pubescent) situated beneath cutting edge (cf.
<taxonomicName id="4C644D28DF49F412FC080F58AD7D73A7" authority=", KOLIBAC 2010: 39" authorityName=", KOLIBAC" authorityPageNumber="39" authorityYear="2010" class="Insecta" family="Trogossitidae" genus="Trichocateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B910EAB9DF49F412FC080F58A8757387" box="[937,1082,380,403]" italics="true" pageId="5" pageNumber="202">Trichocateres</emphasis>
,
<bibRefCitation id="EFF54B5ADF49F412FFD00FB8AD7D73A7" author="KOLIBAC J." box="[113,306,412,435]" pageId="5" pageNumber="202" pagination="34 - 42" refId="ref5491" refString="KOLIBAC J. 2010: Trichocateres fasciculifer, a new genus and species of Trogossitidae: Lophocaterini (Coleoptera). Zootaxa 2353: 34 - 42." type="journal article" year="2010">KOLIBÁČ 2010: 39</bibRefCitation>
</taxonomicName>
); basal notch weakly developed.
</paragraph>
<paragraph id="8BDB36ABDF49F412FF310F98AECB7027" blockId="5.[113,1089,220,1555]" pageId="5" pageNumber="202">
<emphasis id="B910EAB9DF49F412FF310F98ACAC73C7" box="[144,227,444,467]" italics="true" pageId="5" pageNumber="202">Labrum</emphasis>
(
<figureCitation id="135F2A2EDF49F412FF4E0F98AD0F73C7" box="[239,320,444,467]" captionStart="Fig" captionStartId="3.[113,144,1514,1533]" captionTargetBox="[114,1088,225,1490]" captionTargetId="figure-13@3.[114,1088,219,1493]" captionTargetPageId="3" captionText="Fig. 1. Afrocyrona volatilis sp. nov., female paratype:A – labium (mentum removed); B – right mandible ventrally; C – maxilla; D – labrum; E – protarsus; F – procoxa with trochanter. Scale bars = 0.1 mm. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313278" httpUri="https://zenodo.org/record/5313278/files/figure.png" pageId="5" pageNumber="202">Fig. 1D</figureCitation>
): free (not fused with cranium); oval, emarginate in apical part; epipharynx without distinct sclerites; (lateral) tormal processes not separated from body of labrum and not connected in middle; labral (dorsal) sclerite with several thick setae along lateral margins, epipharynx with transverse row of stout sensillae.
</paragraph>
<paragraph id="8BDB36ABDF49F412FF310C18AE637127" blockId="5.[113,1089,220,1555]" pageId="5" pageNumber="202">
<emphasis id="B910EAB9DF49F412FF310C18ACAE7047" box="[144,225,572,595]" italics="true" pageId="5" pageNumber="202">Labium</emphasis>
(
<figureCitation id="135F2A2EDF49F412FF4E0C18AD087047" box="[239,327,572,595]" captionStart="Fig" captionStartId="3.[113,144,1514,1533]" captionTargetBox="[114,1088,225,1490]" captionTargetId="figure-13@3.[114,1088,219,1493]" captionTargetPageId="3" captionText="Fig. 1. Afrocyrona volatilis sp. nov., female paratype:A – labium (mentum removed); B – right mandible ventrally; C – maxilla; D – labrum; E – protarsus; F – procoxa with trochanter. Scale bars = 0.1 mm. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313278" httpUri="https://zenodo.org/record/5313278/files/figure.png" pageId="5" pageNumber="202">Figs 1A</figureCitation>
,
<figureCitation id="135F2A2EDF49F412FEF20C18AD3C7047" box="[339,371,572,595]" captionStart="Fig" captionStartId="6.[151,182,1509,1528]" captionTargetBox="[164,1118,223,1493]" captionTargetId="figure-11@6.[152,1126,219,1493]" captionTargetPageId="6" captionText="Fig. 3. Afrocyrona volatilis sp. nov., female paratype: A – wing; B – sternite VIII; C – tergite VIII; D – internal copulatory organs; E – ovipositor, apical part; F – labium. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5313286" httpUri="https://zenodo.org/record/5313286/files/figure.png" pageId="5" pageNumber="202">3F</figureCitation>
): submentum without ctenidium; mentum trapezoidal, anterior corners extended, anterior margin emarginate, lateral margins rounded; prementum not perfectly divided into two parts; ligula transparent but rigid, weakly emarginate, without ciliate setae, with long pale setae; ‘mental apodeme’ (sclerites beneath mentum or between mentum and prementum) composed of two trapezoidal sclerites; hypopharyngeal sclerite(s) (frequent cleroid structure on hypopharyngeal membrane between mentum and submentum) absent; hypopharynx without longitudinal strip(s) of sensillae; labial palps trimerous, terminal palpomere shorter than labial one, less conical.
</paragraph>
<paragraph id="8BDB36ABDF49F412FF310D18AD3071C7" blockId="5.[113,1089,220,1555]" pageId="5" pageNumber="202">
<emphasis id="B910EAB9DF49F412FF310D18ACBA7147" box="[144,245,828,851]" italics="true" pageId="5" pageNumber="202">Antennae</emphasis>
(
<figureCitation id="135F2A2EDF49F412FEA40D18AD137147" box="[261,348,828,851]" captionStart="Fig" captionStartId="4.[151,182,1514,1533]" captionTargetBox="[151,1126,230,1488]" captionTargetId="figure-11@4.[151,1129,219,1496]" captionTargetPageId="4" captionText="Fig. 2. Afrocyrona volatilis sp. nov., female paratype:A – cranium ventrally (with one maxilla, mandible and mentum); B – antenna, detail; C – prothorax ventrally; D – meso- and metathorax ventrally; E – right elytron, ventral view. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313280" httpUri="https://zenodo.org/record/5313280/files/figure.png" pageId="5" pageNumber="202">Fig. 2B</figureCitation>
): with nine antennomeres; antennal club trimerous, relatively loose; antennomeres 1–6 symmetrical, 7–9 very weakly asymmetrical, without conspicuous sensorial fields, with sparse pubescence; scape robust, pedicel smaller, shorter than antennomere 3, antennomeres 4–6 also shorter than 3; antennae relatively short, extending backwards to midway along pronotum.
</paragraph>
<paragraph id="8BDB36ABDF49F412FF310DF8AF6E7687" blockId="5.[113,1089,220,1555]" pageId="5" pageNumber="202">
<emphasis id="B910EAB9DF49F412FF310DF8ACB271E7" box="[144,253,988,1011]" italics="true" pageId="5" pageNumber="202">Prothorax</emphasis>
(
<figureCitation id="135F2A2EDF49F412FEAB0DF8AD2F71E7" box="[266,352,988,1011]" captionStart="Fig" captionStartId="4.[151,182,1514,1533]" captionTargetBox="[151,1126,230,1488]" captionTargetId="figure-11@4.[151,1129,219,1496]" captionTargetPageId="4" captionText="Fig. 2. Afrocyrona volatilis sp. nov., female paratype:A – cranium ventrally (with one maxilla, mandible and mentum); B – antenna, detail; C – prothorax ventrally; D – meso- and metathorax ventrally; E – right elytron, ventral view. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313280" httpUri="https://zenodo.org/record/5313280/files/figure.png" pageId="5" pageNumber="202">Figs 2C</figureCitation>
,
<figureCitation id="135F2A2EDF49F412FECA0DF8ADE671E7" box="[363,425,988,1011]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. A–D – Afrocyrona pedestris sp. nov., male holotype:A – aedeagus ventrally; B – aedeagus laterally; C – rudimentary wing; D – protarsus. E – A. volatilis sp. nov., paratype; F – A. volatilis sp. nov., holotype; G – A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="5" pageNumber="202">4E–F</figureCitation>
): pronotum transverse; anterior margin weakly emarginate (arcuate) or nearly straight, anterior corners not projecting, rounded; lateral margins rounded, excavate; lateral edge present, finely denticulate; prosternal process distinctly dilated at apex, with sharp side corners; procoxal cavities externally narrowly open, closed to halfway by postcoxal projections, internally open; trochantin slender and small; inner part of hypomeron without longitudinal line; anterior margin with long pubescence.
</paragraph>
<paragraph id="8BDB36ABDF49F412FF310AB8A80E7707" blockId="5.[113,1089,220,1555]" pageId="5" pageNumber="202">
<emphasis id="B910EAB9DF49F412FF310AB8AD4176A7" box="[144,270,1180,1203]" italics="true" pageId="5" pageNumber="202">Mesothorax</emphasis>
(
<figureCitation id="135F2A2EDF49F412FEBB0AB8AD2276A7" box="[282,365,1180,1203]" captionStart="Fig" captionStartId="4.[151,182,1514,1533]" captionTargetBox="[151,1126,230,1488]" captionTargetId="figure-11@4.[151,1129,219,1496]" captionTargetPageId="4" captionText="Fig. 2. Afrocyrona volatilis sp. nov., female paratype:A – cranium ventrally (with one maxilla, mandible and mentum); B – antenna, detail; C – prothorax ventrally; D – meso- and metathorax ventrally; E – right elytron, ventral view. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313280" httpUri="https://zenodo.org/record/5313280/files/figure.png" pageId="5" pageNumber="202">Fig. 2D</figureCitation>
): prepectus present in anterior part of mesepisternum only, absent above mesoventrite; mesocoxal cavities narrowly separated, externally widely open; mesoventral process shorter than coxal diameter; projection of metaventrite reaches approximately towards basal quarter of mesocoxae; mesotrochantin minute; scutum relatively wide, scutellum oval.
</paragraph>
<paragraph id="8BDB36ABDF49F412FF310B38AF2D7787" blockId="5.[113,1089,220,1555]" pageId="5" pageNumber="202">
<emphasis id="B910EAB9DF49F412FF310B38AD4D7727" box="[144,258,1308,1331]" italics="true" pageId="5" pageNumber="202">Hind wing</emphasis>
(
<figureCitation id="135F2A2EDF49F412FEB30B38AD257727" box="[274,362,1308,1331]" captionStart="Fig" captionStartId="6.[151,182,1509,1528]" captionTargetBox="[164,1118,223,1493]" captionTargetId="figure-11@6.[152,1126,219,1493]" captionTargetPageId="6" captionText="Fig. 3. Afrocyrona volatilis sp. nov., female paratype: A – wing; B – sternite VIII; C – tergite VIII; D – internal copulatory organs; E – ovipositor, apical part; F – labium. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5313286" httpUri="https://zenodo.org/record/5313286/files/figure.png" pageId="5" pageNumber="202">Fig. 3A</figureCitation>
): well-developed; radial cell open, moved downwards; r4 missing; pigmented fleck (below Rc) small, pale, not complete; medial field with all four veins, cross-veins between MP3–MP4 and MP4–AA1+2 missing (wedge cell absent); MP3 short (reduced in size); wing weakly pigmented above remnant of vein Rp2.
</paragraph>
<paragraph id="8BDB36ABDF49F412FF310BB8AF877407" blockId="5.[113,1089,220,1555]" pageId="5" pageNumber="202">
<emphasis id="B910EAB9DF49F412FF310BB8AD4377A7" box="[144,268,1436,1459]" italics="true" pageId="5" pageNumber="202">Metathorax</emphasis>
(
<figureCitation id="135F2A2EDF49F412FEBC0BB8AD3977A7" box="[285,374,1436,1459]" captionStart="Fig" captionStartId="4.[151,182,1514,1533]" captionTargetBox="[151,1126,230,1488]" captionTargetId="figure-11@4.[151,1129,219,1496]" captionTargetPageId="4" captionText="Fig. 2. Afrocyrona volatilis sp. nov., female paratype:A – cranium ventrally (with one maxilla, mandible and mentum); B – antenna, detail; C – prothorax ventrally; D – meso- and metathorax ventrally; E – right elytron, ventral view. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313280" httpUri="https://zenodo.org/record/5313280/files/figure.png" pageId="5" pageNumber="202">Fig. 2D</figureCitation>
): metaventrite flat and wide, distinctly narrowed towards anterior portion, widely bordered along lateral margins; discriminal line (discrimen) inconspicuous; paracoxal sutures well-demarcated, parallel with coxae, narrowly separated from base of metaventrite; metepisternum distinctly triangular; metanotum rectangular, wide.
</paragraph>
<caption id="DF1B6623DF4AF411FF360BC1AF0F7406" ID-DOI="http://doi.org/10.5281/zenodo.5313286" ID-Zenodo-Dep="5313286" httpUri="https://zenodo.org/record/5313286/files/figure.png" pageId="6" pageNumber="203" startId="6.[151,182,1509,1528]" targetBox="[164,1118,223,1493]" targetPageId="6">
<paragraph id="8BDB36ABDF4AF411FF360BC1AF0F7406" blockId="6.[151,1126,1509,1554]" pageId="6" pageNumber="203">
Fig. 3.
<taxonomicName id="4C644D28DF4AF411FF760BC1ADCD77EC" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[215,386,1509,1528]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="203" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF4AF411FF760BC1ADCD77EC" box="[215,386,1509,1528]" italics="true" pageId="6" pageNumber="203">Afrocyrona volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF4AF411FE280BC1AD8077EC" box="[393,463,1509,1528]" pageId="6" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype: A – wing; B – sternite VIII; C – tergite VIII; D – internal copulatory organs; E – ovipositor, apical part; F – labium. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="DF1B6623DF4BF410FFD00EF8A87D72FB" box="[113,1074,220,239]" pageId="7" pageNumber="204" startId="7.[113,160,220,239]">
<paragraph id="8BDB36ABDF4BF410FFD00EF8A87D72FB" blockId="7.[113,1074,220,239]" box="[113,1074,220,239]" pageId="7" pageNumber="204">
Table 1. Measurements of
<taxonomicName id="4C644D28DF4BF410FEF50EF8ADB172FB" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[340,510,220,239]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF4BF410FEF50EF8ADB172FB" box="[340,510,220,239]" italics="true" pageId="7" pageNumber="204">Afrocyrona volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF4BF410FDA20EF8AE0972FB" box="[515,582,220,239]" pageId="7" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C644D28DF4BF410FDCF0EF8AE9C72FB" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[622,723,220,239]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="pedestris" status="sp. nov.">
<emphasis id="B910EAB9DF4BF410FDCF0EF8AE9C72FB" box="[622,723,220,239]" italics="true" pageId="7" pageNumber="204">A. pedestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF4BF410FD790EF8AF5472FB" box="[728,795,220,239]" pageId="7" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
(HT – holotype, PT – paratype).
</paragraph>
</caption>
<paragraph id="8BDB36ABDF4BF410FF600F29AFF770D7" pageId="7" pageNumber="204">
<table id="F964C40BDF4B0BE8FF600F28AFA570D7" box="[193,1002,268,707]" gridcols="4" gridrows="15" pageId="7" pageNumber="204">
<tr id="355434E9DF4B0BE8FF600F28AFA57334" box="[193,1002,268,288]" gridrow="0" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600F28ADB27334" box="[193,509,268,288]" gridcol="0" gridrow="0" pageId="7" pageNumber="204">
<emphasis id="B910EAB9DF4BF410FF600F29ADDA7334" bold="true" box="[193,405,269,288]" pageId="7" pageNumber="204">Measurements (in mm)</emphasis>
</th>
<th id="76855D95DF4B0BE8FDB90F28AED67334" box="[536,665,268,288]" gridcol="1" gridrow="0" pageId="7" pageNumber="204">
<taxonomicName id="4C644D28DF4BF410FDB90F29AED67334" authority="HT" box="[536,665,268,288]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="volatilis">
<emphasis id="B910EAB9DF4BF410FDB90F29AED67334" bold="true" box="[536,665,268,288]" pageId="7" pageNumber="204">
<emphasis id="B910EAB9DF4BF410FDB90F29AE39730B" bold="true" box="[536,630,268,288]" italics="true" pageId="7" pageNumber="204">A. volatilis</emphasis>
HT
</emphasis>
</taxonomicName>
</th>
<th id="76855D95DF4B0BE8FD610F28AF717334" box="[704,830,268,288]" gridcol="2" gridrow="0" pageId="7" pageNumber="204">
<taxonomicName id="4C644D28DF4BF410FD610F29AF717334" authority="PT" box="[704,830,268,288]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="volatilis">
<emphasis id="B910EAB9DF4BF410FD610F29AF717334" bold="true" box="[704,830,268,288]" pageId="7" pageNumber="204">
<emphasis id="B910EAB9DF4BF410FD610F29AF51730B" bold="true" box="[704,798,268,288]" italics="true" pageId="7" pageNumber="204">A. volatilis</emphasis>
PT
</emphasis>
</taxonomicName>
</th>
<th id="76855D95DF4B0BE8FCC10F28AFA57334" box="[864,1002,268,288]" gridcol="3" gridrow="0" pageId="7" pageNumber="204">
<taxonomicName id="4C644D28DF4BF410FCC10F29AFA57334" authority="HT" box="[864,1002,268,288]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="pedestris">
<emphasis id="B910EAB9DF4BF410FCC10F29AFA57334" bold="true" box="[864,1002,268,288]" pageId="7" pageNumber="204">
<emphasis id="B910EAB9DF4BF410FCC10F29AF89730B" bold="true" box="[864,966,268,288]" italics="true" pageId="7" pageNumber="204">A. pedestris</emphasis>
HT
</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="355434E9DF4B0BE8FF600F0FAFA5732A" box="[193,1002,299,318]" gridrow="1" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600F0FADB2732A" box="[193,509,299,318]" gridcol="0" gridrow="1" pageId="7" pageNumber="204">Body without mandibles</th>
<td id="76855D95DF4B0BE8FDB90F0FAED6732A" box="[536,665,299,318]" gridcol="1" gridrow="1" pageId="7" pageNumber="204">2.73</td>
<td id="76855D95DF4B0BE8FD610F0FAF71732A" box="[704,830,299,318]" gridcol="2" gridrow="1" pageId="7" pageNumber="204">3.24</td>
<td id="76855D95DF4B0BE8FCC10F0FAFA5732A" box="[864,1002,299,318]" gridcol="3" gridrow="1" pageId="7" pageNumber="204">2.54</td>
</tr>
<tr id="355434E9DF4B0BE8FF600F6DAFA57348" box="[193,1002,329,348]" gridrow="2" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600F6DADB27348" box="[193,509,329,348]" gridcol="0" gridrow="2" pageId="7" pageNumber="204">Head, length of visible part</th>
<td id="76855D95DF4B0BE8FDB90F6DAED67348" box="[536,665,329,348]" gridcol="1" gridrow="2" pageId="7" pageNumber="204">0.28</td>
<td id="76855D95DF4B0BE8FD610F6DAF717348" box="[704,830,329,348]" gridcol="2" gridrow="2" pageId="7" pageNumber="204">0.34</td>
<td id="76855D95DF4B0BE8FCC10F6DAFA57348" box="[864,1002,329,348]" gridcol="3" gridrow="2" pageId="7" pageNumber="204">0.24</td>
</tr>
<tr id="355434E9DF4B0BE8FF600F43AFA5736E" box="[193,1002,359,378]" gridrow="3" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600F43ADB2736E" box="[193,509,359,378]" gridcol="0" gridrow="3" pageId="7" pageNumber="204">Head, max. width incl. eyes</th>
<td id="76855D95DF4B0BE8FDB90F43AED6736E" box="[536,665,359,378]" gridcol="1" gridrow="3" pageId="7" pageNumber="204">0.64</td>
<td id="76855D95DF4B0BE8FD610F43AF71736E" box="[704,830,359,378]" gridcol="2" gridrow="3" pageId="7" pageNumber="204">0.82</td>
<td id="76855D95DF4B0BE8FCC10F43AFA5736E" box="[864,1002,359,378]" gridcol="3" gridrow="3" pageId="7" pageNumber="204">0.67</td>
</tr>
<tr id="355434E9DF4B0BE8FF600FA1AFA5738C" box="[193,1002,389,408]" gridrow="4" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600FA1ADB2738C" box="[193,509,389,408]" gridcol="0" gridrow="4" pageId="7" pageNumber="204">Head, max. width excl. eyes</th>
<td id="76855D95DF4B0BE8FDB90FA1AED6738C" box="[536,665,389,408]" gridcol="1" gridrow="4" pageId="7" pageNumber="204">0.58</td>
<td id="76855D95DF4B0BE8FD610FA1AF71738C" box="[704,830,389,408]" gridcol="2" gridrow="4" pageId="7" pageNumber="204">0.73</td>
<td id="76855D95DF4B0BE8FCC10FA1AFA5738C" box="[864,1002,389,408]" gridcol="3" gridrow="4" pageId="7" pageNumber="204">0.60</td>
</tr>
<tr id="355434E9DF4B0BE8FF600F87AFA573A2" box="[193,1002,419,438]" gridrow="5" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600F87ADB273A2" box="[193,509,419,438]" gridcol="0" gridrow="5" pageId="7" pageNumber="204">Head, mean width between eyes</th>
<td id="76855D95DF4B0BE8FDB90F87AED673A2" box="[536,665,419,438]" gridcol="1" gridrow="5" pageId="7" pageNumber="204">0.51</td>
<td id="76855D95DF4B0BE8FD610F87AF7173A2" box="[704,830,419,438]" gridcol="2" gridrow="5" pageId="7" pageNumber="204">0.65</td>
<td id="76855D95DF4B0BE8FCC10F87AFA573A2" box="[864,1002,419,438]" gridcol="3" gridrow="5" pageId="7" pageNumber="204">0.54</td>
</tr>
<tr id="355434E9DF4B0BE8FF600FE5AFA573C0" box="[193,1002,449,468]" gridrow="6" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600FE5ADB273C0" box="[193,509,449,468]" gridcol="0" gridrow="6" pageId="7" pageNumber="204">Eye, longitudinal diameter</th>
<td id="76855D95DF4B0BE8FDB90FE5AED673C0" box="[536,665,449,468]" gridcol="1" gridrow="6" pageId="7" pageNumber="204">0.11</td>
<td id="76855D95DF4B0BE8FD610FE5AF7173C0" box="[704,830,449,468]" gridcol="2" gridrow="6" pageId="7" pageNumber="204">0.14</td>
<td id="76855D95DF4B0BE8FCC10FE5AFA573C0" box="[864,1002,449,468]" gridcol="3" gridrow="6" pageId="7" pageNumber="204">0.10</td>
</tr>
<tr id="355434E9DF4B0BE8FF600FFAAFA573E5" box="[193,1002,478,497]" gridrow="7" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600FFAADB273E5" box="[193,509,478,497]" gridcol="0" gridrow="7" pageId="7" pageNumber="204">Antenna, total length</th>
<td id="76855D95DF4B0BE8FDB90FFAAED673E5" box="[536,665,478,497]" gridcol="1" gridrow="7" pageId="7" pageNumber="204">0.67</td>
<td id="76855D95DF4B0BE8FD610FFAAF7173E5" box="[704,830,478,497]" gridcol="2" gridrow="7" pageId="7" pageNumber="204">0.78</td>
<td id="76855D95DF4B0BE8FCC10FFAAFA573E5" box="[864,1002,478,497]" gridcol="3" gridrow="7" pageId="7" pageNumber="204">0.61</td>
</tr>
<tr id="355434E9DF4B0BE8FF600FD8AFA5701B" box="[193,1002,508,527]" gridrow="8" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600FD8ADB2701B" box="[193,509,508,527]" gridcol="0" gridrow="8" pageId="7" pageNumber="204">Antenna, length of scape</th>
<td id="76855D95DF4B0BE8FDB90FD8AED6701B" box="[536,665,508,527]" gridcol="1" gridrow="8" pageId="7" pageNumber="204">0.08</td>
<td id="76855D95DF4B0BE8FD610FD8AF71701B" box="[704,830,508,527]" gridcol="2" gridrow="8" pageId="7" pageNumber="204">0.10</td>
<td id="76855D95DF4B0BE8FCC10FD8AFA5701B" box="[864,1002,508,527]" gridcol="3" gridrow="8" pageId="7" pageNumber="204">0.08</td>
</tr>
<tr id="355434E9DF4B0BE8FF600C3EAFA57039" box="[193,1002,538,557]" gridrow="9" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600C3EADB27039" box="[193,509,538,557]" gridcol="0" gridrow="9" pageId="7" pageNumber="204">Antenna, length pedicel</th>
<td id="76855D95DF4B0BE8FDB90C3EAED67039" box="[536,665,538,557]" gridcol="1" gridrow="9" pageId="7" pageNumber="204">0.07</td>
<td id="76855D95DF4B0BE8FD610C3EAF717039" box="[704,830,538,557]" gridcol="2" gridrow="9" pageId="7" pageNumber="204">0.09</td>
<td id="76855D95DF4B0BE8FCC10C3EAFA57039" box="[864,1002,538,557]" gridcol="3" gridrow="9" pageId="7" pageNumber="204">0.07</td>
</tr>
<tr id="355434E9DF4B0BE8FF600C1CAFA5705F" box="[193,1002,568,587]" gridrow="10" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600C1CADB2705F" box="[193,509,568,587]" gridcol="0" gridrow="10" pageId="7" pageNumber="204">Antenna, length of 3-segmented club</th>
<td id="76855D95DF4B0BE8FDB90C1CAED6705F" box="[536,665,568,587]" gridcol="1" gridrow="10" pageId="7" pageNumber="204">0.26</td>
<td id="76855D95DF4B0BE8FD610C1CAF71705F" box="[704,830,568,587]" gridcol="2" gridrow="10" pageId="7" pageNumber="204">0.29</td>
<td id="76855D95DF4B0BE8FCC10C1CAFA5705F" box="[864,1002,568,587]" gridcol="3" gridrow="10" pageId="7" pageNumber="204">0.24</td>
</tr>
<tr id="355434E9DF4B0BE8FF600C72AFA5707D" box="[193,1002,598,617]" gridrow="11" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600C72ADB2707D" box="[193,509,598,617]" gridcol="0" gridrow="11" pageId="7" pageNumber="204">Pronotum, length</th>
<td id="76855D95DF4B0BE8FDB90C72AED6707D" box="[536,665,598,617]" gridcol="1" gridrow="11" pageId="7" pageNumber="204">0.51</td>
<td id="76855D95DF4B0BE8FD610C72AF71707D" box="[704,830,598,617]" gridcol="2" gridrow="11" pageId="7" pageNumber="204">0.67</td>
<td id="76855D95DF4B0BE8FCC10C72AFA5707D" box="[864,1002,598,617]" gridcol="3" gridrow="11" pageId="7" pageNumber="204">0.52</td>
</tr>
<tr id="355434E9DF4B0BE8FF600C50AFA57093" box="[193,1002,628,647]" gridrow="12" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600C50ADB27093" box="[193,509,628,647]" gridcol="0" gridrow="12" pageId="7" pageNumber="204">Pronotum, max. width</th>
<td id="76855D95DF4B0BE8FDB90C50AED67093" box="[536,665,628,647]" gridcol="1" gridrow="12" pageId="7" pageNumber="204">1.12</td>
<td id="76855D95DF4B0BE8FD610C50AF717093" box="[704,830,628,647]" gridcol="2" gridrow="12" pageId="7" pageNumber="204">1.43</td>
<td id="76855D95DF4B0BE8FCC10C50AFA57093" box="[864,1002,628,647]" gridcol="3" gridrow="12" pageId="7" pageNumber="204">1.13</td>
</tr>
<tr id="355434E9DF4B0BE8FF600CB6AFA570B1" box="[193,1002,658,677]" gridrow="13" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600CB6ADB270B1" box="[193,509,658,677]" gridcol="0" gridrow="13" pageId="7" pageNumber="204">Elytra, max. width</th>
<td id="76855D95DF4B0BE8FDB90CB6AED670B1" box="[536,665,658,677]" gridcol="1" gridrow="13" pageId="7" pageNumber="204">1.37</td>
<td id="76855D95DF4B0BE8FD610CB6AF7170B1" box="[704,830,658,677]" gridcol="2" gridrow="13" pageId="7" pageNumber="204">1.76</td>
<td id="76855D95DF4B0BE8FCC10CB6AFA570B1" box="[864,1002,658,677]" gridcol="3" gridrow="13" pageId="7" pageNumber="204">0.68</td>
</tr>
<tr id="355434E9DF4B0BE8FF600C94AFA570D7" box="[193,1002,688,707]" gridrow="14" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FF600C94ADB270D7" box="[193,509,688,707]" gridcol="0" gridrow="14" pageId="7" pageNumber="204">Elytron, max. length</th>
<td id="76855D95DF4B0BE8FDB90C94AED670D7" box="[536,665,688,707]" gridcol="1" gridrow="14" pageId="7" pageNumber="204">1.94</td>
<td id="76855D95DF4B0BE8FD610C94AF7170D7" box="[704,830,688,707]" gridcol="2" gridrow="14" pageId="7" pageNumber="204">2.18</td>
<td id="76855D95DF4B0BE8FCC10C94AFA570D7" box="[864,1002,688,707]" gridcol="3" gridrow="14" pageId="7" pageNumber="204">1.73</td>
</tr>
</table>
</paragraph>
<caption id="DF1B6623DF4BF410FFD00CC9AE11710F" pageId="7" pageNumber="204" startId="7.[113,160,749,768]">
<paragraph id="8BDB36ABDF4BF410FFD00CC9AE11710F" blockId="7.[113,1089,749,795]" pageId="7" pageNumber="204">
Table 2. Differential diagnosis between
<taxonomicName id="4C644D28DF4BF410FE1C0CC9AE2B7114" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[445,612,749,768]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF4BF410FE1C0CC9AE2B7114" box="[445,612,749,768]" italics="true" pageId="7" pageNumber="204">Afrocyrona volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF4BF410FDC90CC9AEE57114" box="[616,682,749,768]" pageId="7" pageNumber="204" rank="species">sp.nov.</taxonomicNameLabel>
and
<taxonomicName id="4C644D28DF4BF410FD6E0CC9AF7D7114" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[719,818,749,768]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="pedestris" status="sp. nov.">
<emphasis id="B910EAB9DF4BF410FD6E0CC9AF7D7114" box="[719,818,749,768]" italics="true" pageId="7" pageNumber="204">A. pedestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF4BF410FC940CC9AF387114" box="[821,887,749,768]" pageId="7" pageNumber="204" rank="species">sp.nov.</taxonomicNameLabel>
The particular character states are explained in detail in descriptions or illustrated.
</paragraph>
</caption>
<paragraph id="8BDB36ABDF4BF410FFD80D1DAFC5762F" pageId="7" pageNumber="204">
<table id="F964C40BDF4B0BE8FFD80D1CA87E762F" box="[121,1073,824,1083]" gridcols="3" gridrows="9" pageId="7" pageNumber="204">
<tr id="355434E9DF4B0BE8FFD80D1CA87E7158" box="[121,1073,824,844]" gridrow="0" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FFD80D1CAD1A7158" box="[121,341,824,844]" gridcol="0" gridrow="0" pageId="7" pageNumber="204">
<emphasis id="B910EAB9DF4BF410FFD80D1DAC997158" bold="true" box="[121,214,825,844]" pageId="7" pageNumber="204">Character</emphasis>
</th>
<th id="76855D95DF4B0BE8FEDB0D1CAEFA7158" box="[378,693,824,844]" gridcol="1" gridrow="0" pageId="7" pageNumber="204">
<emphasis id="B910EAB9DF4BF410FEDB0D1CAE6E7158" bold="true" box="[378,545,824,844]" pageId="7" pageNumber="204">
<taxonomicName id="4C644D28DF4BF410FEDB0D1CAD97715F" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[378,472,824,843]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF4BF410FEDB0D1CAD97715F" bold="true" box="[378,472,824,843]" italics="true" pageId="7" pageNumber="204">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF4BF410FE7C0D1DAE6E7158" box="[477,545,825,844]" pageId="7" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76855D95DF4B0BE8FD400D1CA87E7158" box="[737,1073,824,844]" gridcol="2" gridrow="0" pageId="7" pageNumber="204">
<emphasis id="B910EAB9DF4BF410FD400D1CAFDF7158" bold="true" box="[737,912,824,844]" pageId="7" pageNumber="204">
<taxonomicName id="4C644D28DF4BF410FD400D1CAF08715F" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[737,839,824,843]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="pedestris" status="sp. nov.">
<emphasis id="B910EAB9DF4BF410FD400D1CAF08715F" bold="true" box="[737,839,824,843]" italics="true" pageId="7" pageNumber="204">A. pedestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF4BF410FCED0D1DAFDF7158" box="[844,912,825,844]" pageId="7" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="355434E9DF4B0BE8FFD80D73A87E717E" box="[121,1073,855,874]" gridrow="1" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FFD80D73AD1A717E" box="[121,341,855,874]" gridcol="0" gridrow="1" pageId="7" pageNumber="204">Frons, punctation</th>
<td id="76855D95DF4B0BE8FEDB0D73AEFA717E" box="[378,693,855,874]" gridcol="1" gridrow="1" pageId="7" pageNumber="204">interspace = 2× diameter of puncture</td>
<td id="76855D95DF4B0BE8FD400D73A87E717E" box="[737,1073,855,874]" gridcol="2" gridrow="1" pageId="7" pageNumber="204">interspace = 3–5× diameter of puncture</td>
</tr>
<tr id="355434E9DF4B0BE8FFD80D51A87E719C" box="[121,1073,885,904]" gridrow="2" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FFD80D51AD1A719C" box="[121,341,885,904]" gridcol="0" gridrow="2" pageId="7" pageNumber="204">Head, prothorax ventrally</th>
<td id="76855D95DF4B0BE8FEDB0D51AEFA719C" box="[378,693,885,904]" gridcol="1" gridrow="2" pageId="7" pageNumber="204">punctures separated</td>
<td id="76855D95DF4B0BE8FD400D51A87E719C" box="[737,1073,885,904]" gridcol="2" gridrow="2" pageId="7" pageNumber="204">punctures conjoined</td>
</tr>
<tr id="355434E9DF4B0BE8FFD80DB7A87E71B2" box="[121,1073,915,934]" gridrow="3" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FFD80DB7AD1A71B2" box="[121,341,915,934]" gridcol="0" gridrow="3" pageId="7" pageNumber="204">Dorsal pubescence</th>
<td id="76855D95DF4B0BE8FEDB0DB7AEFA71B2" box="[378,693,915,934]" gridcol="1" gridrow="3" pageId="7" pageNumber="204">semierect thick setae</td>
<td id="76855D95DF4B0BE8FD400DB7A87E71B2" box="[737,1073,915,934]" gridcol="2" gridrow="3" pageId="7" pageNumber="204">decumbent scale-like setae</td>
</tr>
<tr id="355434E9DF4B0BE8FFD80D95A87E71D0" box="[121,1073,945,964]" gridrow="4" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FFD80D95AD1A71D0" box="[121,341,945,964]" gridcol="0" gridrow="4" pageId="7" pageNumber="204">Pronotum</th>
<td id="76855D95DF4B0BE8FEDB0D95AEFA71D0" box="[378,693,945,964]" gridcol="1" gridrow="4" pageId="7" pageNumber="204">densely punctate</td>
<td id="76855D95DF4B0BE8FD400D95A87E71D0" box="[737,1073,945,964]" gridcol="2" gridrow="4" pageId="7" pageNumber="204">sparsely punctate</td>
</tr>
<tr id="355434E9DF4B0BE8FFD80DEAA87E71F5" box="[121,1073,974,993]" gridrow="5" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FFD80DEAAD1A71F5" box="[121,341,974,993]" gridcol="0" gridrow="5" pageId="7" pageNumber="204">Elytra</th>
<td id="76855D95DF4B0BE8FEDB0DEAAEFA71F5" box="[378,693,974,993]" gridcol="1" gridrow="5" pageId="7" pageNumber="204">with two light brown patches</td>
<td id="76855D95DF4B0BE8FD400DEAA87E71F5" box="[737,1073,974,993]" gridcol="2" gridrow="5" pageId="7" pageNumber="204">without patches</td>
</tr>
<tr id="355434E9DF4B0BE8FFD80DC8A87E71EB" box="[121,1073,1004,1023]" gridrow="6" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FFD80DC8AD1A71EB" box="[121,341,1004,1023]" gridcol="0" gridrow="6" pageId="7" pageNumber="204">Elytra, side margins</th>
<td id="76855D95DF4B0BE8FEDB0DC8AEFA71EB" box="[378,693,1004,1023]" gridcol="1" gridrow="6" pageId="7" pageNumber="204">moderately explanate</td>
<td id="76855D95DF4B0BE8FD400DC8A87E71EB" box="[737,1073,1004,1023]" gridcol="2" gridrow="6" pageId="7" pageNumber="204">narrowly explanate</td>
</tr>
<tr id="355434E9DF4B0BE8FFD80A2EA87E7609" box="[121,1073,1034,1053]" gridrow="7" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FFD80A2EAD1A7609" box="[121,341,1034,1053]" gridcol="0" gridrow="7" pageId="7" pageNumber="204">Wings</th>
<td id="76855D95DF4B0BE8FEDB0A2EAEFA7609" box="[378,693,1034,1053]" gridcol="1" gridrow="7" pageId="7" pageNumber="204">well-developed</td>
<td id="76855D95DF4B0BE8FD400A2EA87E7609" box="[737,1073,1034,1053]" gridcol="2" gridrow="7" pageId="7" pageNumber="204">rudimentary</td>
</tr>
<tr id="355434E9DF4B0BE8FFD80A0CA87E762F" box="[121,1073,1064,1083]" gridrow="8" pageId="7" pageNumber="204">
<th id="76855D95DF4B0BE8FFD80A0CAD1A762F" box="[121,341,1064,1083]" gridcol="0" gridrow="8" pageId="7" pageNumber="204">Meso- and metatibiae</th>
<td id="76855D95DF4B0BE8FEDB0A0CAEFA762F" box="[378,693,1064,1083]" gridcol="1" gridrow="8" pageId="7" pageNumber="204">apical spurs small, indistinct</td>
<td id="76855D95DF4B0BE8FD400A0CA87E762F" box="[737,1073,1064,1083]" gridcol="2" gridrow="8" pageId="7" pageNumber="204">apical spurs distinct</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDB36ABDF4BF410FF310AA1AF6176C8" blockId="7.[113,1089,1157,1564]" pageId="7" pageNumber="204">
<emphasis id="B910EAB9DF4BF410FF310AA1AC9C7688" box="[144,211,1157,1180]" italics="true" pageId="7" pageNumber="204">Elytra</emphasis>
(
<figureCitation id="135F2A2EDF4BF410FF400AA1AD787688" box="[225,311,1157,1180]" captionStart="Fig" captionStartId="4.[151,182,1514,1533]" captionTargetBox="[151,1126,230,1488]" captionTargetId="figure-11@4.[151,1129,219,1496]" captionTargetPageId="4" captionText="Fig. 2. Afrocyrona volatilis sp. nov., female paratype:A – cranium ventrally (with one maxilla, mandible and mentum); B – antenna, detail; C – prothorax ventrally; D – meso- and metathorax ventrally; E – right elytron, ventral view. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313280" httpUri="https://zenodo.org/record/5313280/files/figure.png" pageId="7" pageNumber="204">Figs 2E</figureCitation>
,
<figureCitation id="135F2A2EDF4BF410FEE30AA1ADCE7688" box="[322,385,1157,1180]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. A–D – Afrocyrona pedestris sp. nov., male holotype:A – aedeagus ventrally; B – aedeagus laterally; C – rudimentary wing; D – protarsus. E – A. volatilis sp. nov., paratype; F – A. volatilis sp. nov., holotype; G – A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="7" pageNumber="204">4E–F</figureCitation>
): regularly punctate, with moderately wide epipleura conspicuous along whole length of elytra; interlocking mechanism not developed; elytra without distinct longitudinal carinae; lateral margins of elytra narrowly explanate.
</paragraph>
<paragraph id="8BDB36ABDF4BF410FF310AC1AD0277C8" blockId="7.[113,1089,1157,1564]" pageId="7" pageNumber="204">
<emphasis id="B910EAB9DF4BF410FF310AC1AC8D76E8" box="[144,194,1253,1276]" italics="true" pageId="7" pageNumber="204">Legs</emphasis>
(
<figureCitation id="135F2A2EDF4BF410FF6F0AC1AD7276E8" box="[206,317,1253,1276]" captionStart="Fig" captionStartId="3.[113,144,1514,1533]" captionTargetBox="[114,1088,225,1490]" captionTargetId="figure-13@3.[114,1088,219,1493]" captionTargetPageId="3" captionText="Fig. 1. Afrocyrona volatilis sp. nov., female paratype:A – labium (mentum removed); B – right mandible ventrally; C – maxilla; D – labrum; E – protarsus; F – procoxa with trochanter. Scale bars = 0.1 mm. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313278" httpUri="https://zenodo.org/record/5313278/files/figure.png" pageId="7" pageNumber="204">Figs 1E–F</figureCitation>
,
<figureCitation id="135F2A2EDF4BF410FEE70AC1ADC476E8" box="[326,395,1253,1276]" captionStart="Fig" captionStartId="4.[151,182,1514,1533]" captionTargetBox="[151,1126,230,1488]" captionTargetId="figure-11@4.[151,1129,219,1496]" captionTargetPageId="4" captionText="Fig. 2. Afrocyrona volatilis sp. nov., female paratype:A – cranium ventrally (with one maxilla, mandible and mentum); B – antenna, detail; C – prothorax ventrally; D – meso- and metathorax ventrally; E – right elytron, ventral view. For abbreviations see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5313280" httpUri="https://zenodo.org/record/5313280/files/figure.png" pageId="7" pageNumber="204">2C–D</figureCitation>
): procoxae not projecting, transverse; mesocoxae oval, metacoxae extended to lateral margin of metathorax; trochanters relatively small, triangular; femora weakly clavate; tibiae with row of small spines along outer margin; tibial apical spur pattern 1(2)-2-2 (2
<superScript id="7C119BE3DF4BF410FF460B67ACB87745" attach="left" box="[231,247,1347,1361]" fontSize="6" pageId="7" pageNumber="204">nd</superScript>
spur of protibiae reduced in size); large protibial spur distinctly hooked; apices of meso- and metatibiae with row of ca. 3 spines; tarsomere
<quantity id="4C9C9B4EDF4BF410FD440B41AF427768" box="[741,781,1381,1404]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="204" unit="in" value="1.0">1 in</quantity>
all pairs of legs conspicuous, approximately as long as tarsomere 2; tarsomere 5 as long as, or slightly longer than, 1–4 combined; tarsal lobes absent; claws without denticles; empodium large, projecting, bisetose; tarsal formula 5-5-5.
</paragraph>
<paragraph id="8BDB36ABDF4BF41FFF310BC1AE557367" blockId="7.[113,1089,1157,1564]" lastBlockId="8.[151,1127,220,659]" lastPageId="8" lastPageNumber="205" pageId="7" pageNumber="204">
<emphasis id="B910EAB9DF4BF410FF310BC1ACBA77E8" box="[144,245,1509,1532]" italics="true" pageId="7" pageNumber="204">Abdomen</emphasis>
(
<figureCitation id="135F2A2EDF4BF410FEA00BC1AD3877E8" box="[257,375,1509,1532]" captionStart="Fig" captionStartId="6.[151,182,1509,1528]" captionTargetBox="[164,1118,223,1493]" captionTargetId="figure-11@6.[152,1126,219,1493]" captionTargetPageId="6" captionText="Fig. 3. Afrocyrona volatilis sp. nov., female paratype: A – wing; B – sternite VIII; C – tergite VIII; D – internal copulatory organs; E – ovipositor, apical part; F – labium. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5313286" httpUri="https://zenodo.org/record/5313286/files/figure.png" pageId="7" pageNumber="204">Figs 3B–E</figureCitation>
): six ventrites distinctly visible; female sternite VIII weakly emarginate at apical margin, with setae along apical margin and in centre; female tergite VIII straight or rounded at apex, with long spiculum; ovipositor relatively short, reaching posteriorly beyond base of sternite VI; coxitae with row of setae along concave apex; coxital styli relatively large, truncate at apex; vagina and bursa copulatrix connected into single membranous sac; spermatheca not observed, only (spermathecal) gland extending from apex of membranous sac.
<emphasis id="B910EAB9DF44F41FFF360F78ADDD7367" bold="true" box="[151,402,348,371]" pageId="8" pageNumber="205">Differential diagnosis.</emphasis>
See
<tableCitation id="C6E60310DF44F41FFE650F78AE597367" box="[452,534,348,371]" captionStart="Table 2" captionStartId="7.[113,160,749,768]" captionText="Table 2. Differential diagnosis between Afrocyrona volatilis sp.nov.and A. pedestris sp.nov.The particular character states are explained in detail in descriptions or illustrated." pageId="8" pageNumber="205">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E6520DF44F41FFF360F58A8287387" box="[151,1127,380,403]" pageId="8" pageNumber="205" type="etymology">
<paragraph id="8BDB36ABDF44F41FFF360F58A8287387" blockId="8.[151,1127,220,659]" box="[151,1127,380,403]" pageId="8" pageNumber="205">
<emphasis id="B910EAB9DF44F41FFF360F58AD577387" bold="true" box="[151,280,380,403]" pageId="8" pageNumber="205">Etymology.</emphasis>
The species name is Latin adjective
<taxonomicName id="4C644D28DF44F41FFD340F58AEA47387" authorityName="Kolibáč" authorityYear="2014" box="[661,747,380,403]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="volatilis">
<emphasis id="B910EAB9DF44F41FFD340F58AEA47387" box="[661,747,380,403]" italics="true" pageId="8" pageNumber="205">volatilis</emphasis>
</taxonomicName>
(-
<emphasis id="B910EAB9DF44F41FFCA30F58AF5B7387" box="[770,788,380,403]" italics="true" pageId="8" pageNumber="205">is</emphasis>
, -
<emphasis id="B910EAB9DF44F41FFC890F58AF7B7387" box="[808,820,380,403]" italics="true" pageId="8" pageNumber="205">e</emphasis>
) = flying, able to fly, winged.
</paragraph>
</subSubSection>
<subSubSection id="C37E6520DF44F41FFF360FB8AF3C7087" pageId="8" pageNumber="205" type="biology_ecology">
<paragraph id="8BDB36ABDF44F41FFF360FB8ACA873E7" blockId="8.[151,1127,220,659]" pageId="8" pageNumber="205">
<emphasis id="B910EAB9DF44F41FFF360FB8ACBC73A7" bold="true" box="[151,243,412,435]" pageId="8" pageNumber="205">Biology.</emphasis>
Both specimens were knocked down from shrub branches in mountainous forest (see the section on biology of the next species). See
<bibRefCitation id="EFF54B5ADF44F41FFD190F98AF1173C7" author="BATELKA J." box="[696,862,444,467]" pageId="8" pageNumber="205" pagination="1 - 557" refId="ref5318" refString="BATELKA J. 2012: Socotra Archipelago - a lifeboat in the sea of changes: advancement in Socotran insect biodiversity survey. Pp. 1 - 26. In: HAJEK J. &amp; BEZDEK J. (eds): Insect biodiversity of the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52 (supplementum 2): i - vi + 1 - 557." type="journal article" year="2012">BATELKA (2012)</bibRefCitation>
for details on the species habitat.
</paragraph>
<paragraph id="8BDB36ABDF44F41FFF140FD8AF3C7087" blockId="8.[151,1127,220,659]" pageId="8" pageNumber="205">
Gut contents of the dissected
<typeStatus id="54DF8809DF44F41FFE5F0FD8AE157007" box="[510,602,508,531]" pageId="8" pageNumber="205" type="paratype">paratype</typeStatus>
consisted of abundant remnants of detritus and perhaps mould spores. No fragments of insect cuticle were observed. Unlike advanced lophocaterines (
<taxonomicName id="4C644D28DF44F41FFEE40C18ADE37047" authorityName="Reitter" authorityYear="1876" box="[325,428,572,595]" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B910EAB9DF44F41FFEE40C18ADE37047" box="[325,428,572,595]" italics="true" pageId="8" pageNumber="205">Ancyrona</emphasis>
</taxonomicName>
), the species is probably not exclusively predaceous but maybe fungivorous/herbivorous and partially predaceous. Such way of life is also supposed in some other lophocaterines, including South African species of
<taxonomicName id="4C644D28DF44F41FFD540C58AF237087" authorityName="Kolibac" authorityYear="2007" box="[757,876,636,659]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B910EAB9DF44F41FFD540C58AF237087" box="[757,876,636,659]" italics="true" pageId="8" pageNumber="205">Afrocyrona</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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