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<document id="BA8C5FD8152A41EF1A1AFD9DF6C0042C" ID-DOI="10.5281/zenodo.5313274" ID-ISSN="0374-1036" ID-Zenodo-Dep="5313274" ID-ZooBank="60855ED1-C243-4DA1-ADCC-8ADAA4752E9FM" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630120511396" checkinUser="marcus" docAuthor="Kolibáč, Jiří" docDate="2014" docId="03CD87BDDF44F41BFE7F0C9FAEC97307" docLanguage="en" docName="ActaEntMusNatPra.2014.54.suppl.197-210.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Afrocyrona pedestris Kolibáč 2014, sp. nov." docType="treatment" docVersion="4" lastPageNumber="208" masterDocId="FFF4FFC5DF4CF417FFA10E24AC4F7214" masterDocTitle="Two new species of Afrocyrona from Socotra Island (Coleoptera: Trogossitidae)" masterLastPageNumber="210" masterPageNumber="197" pageNumber="205" updateTime="1699160074818" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="901152BA0492579A71F0A7E578AD15B6">Two new species of Afrocyrona from Socotra Island (Coleoptera: Trogossitidae)</mods:title>
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<mods:namePart id="E2D9064D4F3E65FB8430622A209F7027">Kolibáč, Jiří</mods:namePart>
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<mods:title id="DAD850A88D1557AB396139C9D4694334">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="E9F91C7C37AC2318FEC309B8AB595114">2014</mods:date>
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<mods:number id="F756C0E29574BB1EF301D98A50908014">2014-12-30</mods:number>
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<treatment id="03CD87BDDF44F41BFE7F0C9FAEC97307" ID-DOI="http://doi.org/10.5281/zenodo.5449997" ID-Zenodo-Dep="5449997" LSID="urn:lsid:plazi:treatment:03CD87BDDF44F41BFE7F0C9FAEC97307" httpUri="http://treatment.plazi.org/id/03CD87BDDF44F41BFE7F0C9FAEC97307" lastPageId="12" lastPageNumber="208" pageId="8" pageNumber="205">
<subSubSection id="C37E6520DF44F41FFE7F0C9FAF5070C7" box="[478,799,699,723]" pageId="8" pageNumber="205" type="nomenclature">
<paragraph id="8BDB36ABDF44F41FFE7F0C9FAF5070C7" blockId="8.[478,799,699,723]" box="[478,799,699,723]" pageId="8" pageNumber="205">
<heading id="D09381C7DF44F41FFE7F0C9FAF5070C7" bold="true" box="[478,799,699,723]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="205" reason="3">
<emphasis id="B910EAB9DF44F41FFE7F0C9FAF5070C7" bold="true" box="[478,799,699,723]" pageId="8" pageNumber="205">
<taxonomicName id="4C644D28DF44F41FFE7F0C9FAE8C70C7" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[478,707,699,723]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="pedestris" status="sp. nov.">
<emphasis id="B910EAB9DF44F41FFE7F0C9FAE8C70C7" bold="true" box="[478,707,699,723]" italics="true" pageId="8" pageNumber="205">Afrocyrona pedestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF44F41FFD6B0C98AF5070C7" box="[714,799,700,723]" pageId="8" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E6520DF44F41FFDB60CFEAEA870F9" box="[535,743,730,749]" pageId="8" pageNumber="205" type="description">
<paragraph id="8BDB36ABDF44F41FFDB60CFEAEA870F9" blockId="8.[151,1127,730,844]" box="[535,743,730,749]" pageId="8" pageNumber="205">
(
<figureCitation id="135F2A2EDF44F41FFDBC0CFEAEEC70F9" box="[541,675,730,749]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. AD Afrocyrona pedestris sp. nov., male holotype:A aedeagus ventrally; B aedeagus laterally; C rudimentary wing; D protarsus. E A. volatilis sp. nov., paratype; F A. volatilis sp. nov., holotype; G A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="8" pageNumber="205">Figs 4AD, 4G</figureCitation>
,
<figureCitation id="135F2A2EDF44F41FFD0D0CFEAEAE70F9" box="[684,737,730,749]" captionStart="Fig" captionStartId="11.[113,144,1208,1227]" captionTargetBox="[130,1087,224,1178]" captionTargetId="figure-13@11.[113,1091,219,1183]" captionTargetPageId="11" captionText="Fig. 5. AD Afrocyrona pedestris sp. nov., male holotype: A abdominal segment IX with spiculum of sternite IX (spicular fork); B tegmen ventrally; C sternite VIII; D tergite VIII. EH A. volatilis sp. nov., female paratype: E lacinia (incl. basistipes) and galea (incl. basigalea). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5313292" httpUri="https://zenodo.org/record/5313292/files/figure.png" pageId="8" pageNumber="205">5AD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37E6520DF44F41FFF360D20AEBB7158" pageId="8" pageNumber="205" type="materials_examined">
<paragraph id="8BDB36ABDF44F41FFF360D20AEBB7158" blockId="8.[151,1127,730,844]" pageId="8" pageNumber="205">
<emphasis id="B910EAB9DF44F41FFF360D20AD527103" bold="true" box="[151,285,772,791]" pageId="8" pageNumber="205">Type material.</emphasis>
<materialsCitation id="3B0C3CF6DF44F41FFE830D20AEBB7158" collectingDate="2012-06-16" collectingDateMax="2012-06-18" collectingDateMin="2012-06-16" collectionCode="SCAND, V, NMPC" collectorName="Socotra &amp; J. Bezdek &amp; J. Hajek &amp; Hula &amp; P. Kment &amp; I. Malenovsky &amp; J. Niedobova &amp; L. Purchart" country="Yemen" latitude="12.576667" location="Island" longLatPrecision="129" longitude="54.025" municipality="Hagher Mts." pageId="8" pageNumber="205" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DF8809DF44F41FFE830D20AD397101" box="[290,374,772,791]" pageId="8" pageNumber="205" type="holotype">HOLOTYPE</typeStatus>
: J,
<collectingCountry id="F373763BDF44F41FFE010D20ADBD7103" box="[416,498,772,791]" name="Yemen" pageId="8" pageNumber="205">YEMEN</collectingCountry>
, SOCOTRA
<location id="8EBB6070DF44F41FFDEA0D20AE307103" LSID="urn:lsid:plazi:treatment:03CD87BDDF44F41BFE7F0C9FAEC97307:8EBB6070DF44F41FFDEA0D20AE307103" box="[587,639,772,791]" country="Yemen" latitude="12.576667" longLatPrecision="129" longitude="54.025" municipality="Hagher Mts." name="Island" pageId="8" pageNumber="205">Island</location>
/
<collectingMunicipality id="6BBFACD1DF44F41FFD300D20AEB07103" box="[657,767,772,791]" pageId="8" pageNumber="205">Hagher Mts.</collectingMunicipality>
,
<collectionCode id="ED75AE6EDF44F41FFCA90D20AF717101" box="[776,830,772,791]" pageId="8" pageNumber="205">SCAND</collectionCode>
Mt. env. / broad-leaved evergreen woodland /
<date id="FFDA106BDF44F41FFF5B0D3BAD357126" box="[250,378,799,818]" pageId="8" pageNumber="205" value="2012-06-16" valueMax="2012-06-18" valueMin="2012-06-16">
<collectingDate id="EF9EE983DF44F41FFF5B0D3BAD357126" box="[250,378,799,818]" pageId="8" pageNumber="205" value="2012-06-16" valueMax="2012-06-18" valueMin="2012-06-16">16.18.vi.2012</collectingDate>
</date>
/
<geoCoordinate id="EE50506CDF44F41FFE280D3BADAD7126" box="[393,482,799,818]" degrees="12" direction="north" minutes="34.6" orientation="latitude" pageId="8" pageNumber="205" precision="92" value="12.576667">12°34.6N</geoCoordinate>
,
<geoCoordinate id="EE50506CDF44F41FFE4B0D3BAE0E7126" box="[490,577,799,818]" degrees="54" direction="east" minutes="01.5" orientation="longitude" pageId="8" pageNumber="205" precision="92" value="54.025">54°01.5E</geoCoordinate>
, 1450 m;
<collectorName id="2691537DDF44F41FFD3D0D3BAEAE7126" box="[668,737,799,818]" pageId="8" pageNumber="205">Socotra</collectorName>
expedition 2012 /
<collectorName id="2691537DDF44F41FFCDC0D3BAF9E7126" box="[893,977,799,818]" pageId="8" pageNumber="205">J. Bezděk</collectorName>
,
<collectorName id="2691537DDF44F41FFC780D3BA86F7126" box="[985,1056,799,818]" pageId="8" pageNumber="205">J. Hájek</collectorName>
,
<collectionCode id="ED75AE6EDF44F41FFB860D3BA8797126" box="[1063,1078,799,818]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="205" type="Museum">V</collectionCode>
.
<collectorName id="2691537DDF44F41FFB9C0D3BA8297126" box="[1085,1126,799,818]" pageId="8" pageNumber="205">Hula</collectorName>
/
<collectorName id="2691537DDF44F41FFF030D1DACBC7158" box="[162,243,825,844]" pageId="8" pageNumber="205">P. Kment</collectorName>
,
<collectorName id="2691537DDF44F41FFF5A0D1DAD397158" box="[251,374,825,844]" pageId="8" pageNumber="205">I. Malenovský</collectorName>
/
<collectorName id="2691537DDF44F41FFE260D1DADB77158" box="[391,504,825,844]" pageId="8" pageNumber="205">J. Niedobová</collectorName>
&amp;
<collectorName id="2691537DDF44F41FFDB20D1DAE3C7158" box="[531,627,825,844]" pageId="8" pageNumber="205">L. Purchart</collectorName>
leg. (
<collectionCode id="ED75AE6EDF44F41FFD080D1DAEA57158" box="[681,746,825,844]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="205">NMPC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37E6520DF44F41CFF360D41ACB377C4" lastPageId="11" lastPageNumber="208" pageId="8" pageNumber="205" type="description">
<paragraph id="8BDB36ABDF44F41FFF360D41AD5A7188" blockId="8.[151,1127,869,1564]" pageId="8" pageNumber="205">
<emphasis id="B910EAB9DF44F41FFF360D41AD6D7168" bold="true" box="[151,290,869,892]" pageId="8" pageNumber="205">Description.</emphasis>
<emphasis id="B910EAB9DF44F41FFE880D41ADE47168" box="[297,427,869,892]" italics="true" pageId="8" pageNumber="205">Body length</emphasis>
(from elytral apex to clypeus):
<quantity id="4C9C9B4EDF44F41FFCA30D41AF177168" box="[770,856,869,892]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="205" unit="mm" value="2.5">2.5 mm</quantity>
. For other measurements see
<tableCitation id="C6E60310DF44F41FFF1E0DA1AD407188" box="[191,271,901,924]" captionStart="Table 1" captionStartId="7.[113,160,220,239]" captionText="Table 1. Measurements of Afrocyrona volatilis sp. nov. and A. pedestris sp. nov. (HT holotype, PT paratype)." pageId="8" pageNumber="205">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BDB36ABDF44F41FFF140D81ADCE7408" blockId="8.[151,1127,869,1564]" pageId="8" pageNumber="205">
<emphasis id="B910EAB9DF44F41FFF140D81AD8D71A8" box="[181,450,933,956]" italics="true" pageId="8" pageNumber="205">Coloration and sculpture</emphasis>
(
<figureCitation id="135F2A2EDF44F41FFE700D81AE6771A8" box="[465,552,933,956]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. AD Afrocyrona pedestris sp. nov., male holotype:A aedeagus ventrally; B aedeagus laterally; C rudimentary wing; D protarsus. E A. volatilis sp. nov., paratype; F A. volatilis sp. nov., holotype; G A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="8" pageNumber="205">Fig. 4G</figureCitation>
): Body oval, conspicuously convex; all dorsal surface black-brown to black but brown lateral margins of pronotum, margins of elytra, mesonotum, legs, and antennae. Setae covering dorsal surface scale-like, much thicker than those in
<taxonomicName id="4C644D28DF44F41FFBEE0DC1ACA27608" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF44F41FFBEE0DC1ACA27608" italics="true" pageId="8" pageNumber="205">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF44F41FFF550A21AD087608" box="[244,327,1029,1052]" pageId="8" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
Head very finely and very sparsely punctate (interspaces much larger than those in
<taxonomicName id="4C644D28DF44F41FFF4C0A01AD117628" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[237,350,1061,1084]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF44F41FFF4C0A01AD117628" box="[237,350,1061,1084]" italics="true" pageId="8" pageNumber="205">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF44F41FFEC30A01ADF97628" box="[354,438,1061,1084]" pageId="8" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
); punctures with thick erect or decumbent whitish setae. Pronotum finely and sparsely punctate (interspaces larger than diameter of punctures), size of punctures nearly same as that on head; centre of pronotum covered with thick, perfectly decumbent whitish setae; lateral margins with denser erect pubescence composed of black and white clavate setae. Elytral punctation regular, composed of about 14 rows of punctures which are clearly visible from ventral side only (covered with pubescence from above), interspaces among rows inconspicuously elevate; pubescence similar to that in centre of pronotum, composed of thick, decumbent setae which are whitish in humeral and apical areas as well as along lateral margins and black on rest of elytra; margins of elytra also with short erect setae. Sculpture of ventral surface of head and prothorax coarser than that in
<taxonomicName id="4C644D28DF44F41FFC6F0B61A80E7748" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[974,1089,1349,1372]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF44F41FFC6F0B61A80E7748" box="[974,1089,1349,1372]" italics="true" pageId="8" pageNumber="205">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF44F41FFBE90B61AC867768" pageId="8" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
: punctures relatively large, not separated, joined to transverse wrinkles; sculpture and pubescence of meso- and metaventrite as well as legs similar to that in
<taxonomicName id="4C644D28DF44F41FFC330BA1A8497788" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[914,1030,1413,1436]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF44F41FFC330BA1A8497788" box="[914,1030,1413,1436]" italics="true" pageId="8" pageNumber="205">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF44F41FFBAC0BA1A82C7788" box="[1037,1123,1413,1436]" pageId="8" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
: punctation sparse, punctures relatively large and distinctly separated (interspaces larger than diameter of punctures); hypomeral area of prothorax and basal margin of metaventrite without punctation; pubescence scarce, conspicuous only in metaventrite, formed by short, sparse, decumbent pale hairs.
</paragraph>
<paragraph id="8BDB36ABDF45F41EFF310EF8ADA773A7" blockId="9.[113,1090,220,1587]" pageId="9" pageNumber="206">
<emphasis id="B910EAB9DF45F41EFF310EF8AC8572E7" box="[144,202,220,243]" italics="true" pageId="9" pageNumber="206">Head</emphasis>
(
<figureCitation id="135F2A2EDF45F41EFF790EF8AD6072E7" box="[216,303,220,243]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. AD Afrocyrona pedestris sp. nov., male holotype:A aedeagus ventrally; B aedeagus laterally; C rudimentary wing; D protarsus. E A. volatilis sp. nov., paratype; F A. volatilis sp. nov., holotype; G A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="9" pageNumber="206">Fig. 4G</figureCitation>
): almost hypognathous (position in picture artificial), frons distinctly flat; gular sutures widely separated at base, extending to approximately midpoint of cranium, strongly convergent; frontoclypeal suture inconspicuous; antennal grooves on ventral side of cranium short but relatively deep, more distinct than those in
<taxonomicName id="4C644D28DF45F41EFCA80F18AF327347" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[777,893,316,339]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="206" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF45F41EFCA80F18AF327347" box="[777,893,316,339]" italics="true" pageId="9" pageNumber="206">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF45F41EFC240F18AF947347" box="[901,987,316,339]" pageId="9" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
; antennal sockets not visible from above; eyes relatively small (space between them about twelve times as wide as eye diameter) but coarsely facetted, distinctly elevated, not emarginate, elliptic, their greater part situated ventrally.
</paragraph>
<paragraph id="8BDB36ABDF45F41EFF310F98A80D7007" blockId="9.[113,1090,220,1587]" pageId="9" pageNumber="206">
<emphasis id="B910EAB9DF45F41EFF310F98AD5873C7" box="[144,279,444,467]" italics="true" pageId="9" pageNumber="206">Mouth parts</emphasis>
of unique
<typeStatus id="54DF8809DF45F41EFE370F98ADBB73C7" box="[406,500,444,467]" pageId="9" pageNumber="206" type="holotype">holotype</typeStatus>
not dissected. Maxillary palps tetramerous, terminal palpomere conical, elongate; labial palps with terminal palpomere distinctly shorter than the labial one, cylindrical to weakly securiform; anterior margin of labrum shallowly emarginate.
</paragraph>
<paragraph id="8BDB36ABDF45F41EFF310C38AF7E70A7" blockId="9.[113,1090,220,1587]" pageId="9" pageNumber="206">
<emphasis id="B910EAB9DF45F41EFF310C38ACBA7027" box="[144,245,540,563]" italics="true" pageId="9" pageNumber="206">Antennae</emphasis>
(
<figureCitation id="135F2A2EDF45F41EFEA40C38AD117027" box="[261,350,540,563]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. AD Afrocyrona pedestris sp. nov., male holotype:A aedeagus ventrally; B aedeagus laterally; C rudimentary wing; D protarsus. E A. volatilis sp. nov., paratype; F A. volatilis sp. nov., holotype; G A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="9" pageNumber="206">Fig. 4G</figureCitation>
): with nine antennomeres; antennal club trimerous, relatively loose; antennomeres 16 symmetrical, 79 very weakly asymmetrical, without conspicuous sensorial fields, with sparse pubescence; scape robust, pedicel smaller, slightly shorter than antennomere 3, antennomere 4 approximately as long as antennomere 4, 56 distinctly shorter; antennae relatively short, extending backwards to midway along pronotum.
</paragraph>
<paragraph id="8BDB36ABDF45F41EFF310C98A80D7127" blockId="9.[113,1090,220,1587]" pageId="9" pageNumber="206">
<emphasis id="B910EAB9DF45F41EFF310C98ACB470C7" box="[144,251,700,723]" italics="true" pageId="9" pageNumber="206">Prothorax</emphasis>
(
<figureCitation id="135F2A2EDF45F41EFEAA0C98AD2B70C7" box="[267,356,700,723]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. AD Afrocyrona pedestris sp. nov., male holotype:A aedeagus ventrally; B aedeagus laterally; C rudimentary wing; D protarsus. E A. volatilis sp. nov., paratype; F A. volatilis sp. nov., holotype; G A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="9" pageNumber="206">Fig. 4G</figureCitation>
): pronotum transverse; anterior margin straight, anterior corners not projecting, rounded; lateral margins rounded, excavate; lateral edge present, finely denticulate; prosternal process distinctly dilated at apex, with sharp side corners; procoxal cavities externally narrowly open, closed to halfway by postcoxal projections; anterior margin with long pubescence.
</paragraph>
<paragraph id="8BDB36ABDF45F41EFF310D18AE4071A7" blockId="9.[113,1090,220,1587]" pageId="9" pageNumber="206">
<emphasis id="B910EAB9DF45F41EFF310D18AD407147" box="[144,271,828,851]" italics="true" pageId="9" pageNumber="206">Mesothorax</emphasis>
: prepectus present in anterior part of mesepisternum only, absent above mesoventrite; mesocoxal cavities externally widely open; mesoventral process reaches towards base of mesocoxae (anterior projection of metaventrite minute), mesoventral process with longitudinal groove; mesonotum wide.
</paragraph>
<paragraph id="8BDB36ABDF45F41EFF310D98AF6471C7" blockId="9.[113,1090,220,1587]" box="[144,811,956,979]" pageId="9" pageNumber="206">
<emphasis id="B910EAB9DF45F41EFF310D98AD4F71C7" box="[144,256,956,979]" italics="true" pageId="9" pageNumber="206">Hind wing</emphasis>
(
<figureCitation id="135F2A2EDF45F41EFEAF0D98AD2C71C7" box="[270,355,956,979]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. AD Afrocyrona pedestris sp. nov., male holotype:A aedeagus ventrally; B aedeagus laterally; C rudimentary wing; D protarsus. E A. volatilis sp. nov., paratype; F A. volatilis sp. nov., holotype; G A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="9" pageNumber="206">Fig. 4C</figureCitation>
): rudimentary, vestigial; venation reduced.
</paragraph>
<paragraph id="8BDB36ABDF45F41EFF310DF8AFB57647" blockId="9.[113,1090,220,1587]" pageId="9" pageNumber="206">
<emphasis id="B910EAB9DF45F41EFF310DF8AD4371E7" box="[144,268,988,1011]" italics="true" pageId="9" pageNumber="206">Metathorax</emphasis>
: metaventrite wide and relatively convex, distinctly narrowed towards anterior portion; discriminal line (discrimen) inconspicuous; paracoxal sutures well-demarcated, parallel with coxae, narrowly separated from base of metaventrite; metepisternum distinctly triangular; metanotum rectangular, well-developed (in spite of nonfunctional wings).
</paragraph>
<paragraph id="8BDB36ABDF45F41EFF310A78AD6E76C7" blockId="9.[113,1090,220,1587]" pageId="9" pageNumber="206">
<emphasis id="B910EAB9DF45F41EFF310A78AC9C7667" box="[144,211,1116,1139]" italics="true" pageId="9" pageNumber="206">Elytra</emphasis>
(
<figureCitation id="135F2A2EDF45F41EFF400A78AD737667" box="[225,316,1116,1139]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. AD Afrocyrona pedestris sp. nov., male holotype:A aedeagus ventrally; B aedeagus laterally; C rudimentary wing; D protarsus. E A. volatilis sp. nov., paratype; F A. volatilis sp. nov., holotype; G A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="9" pageNumber="206">Figs 4G</figureCitation>
): regularly punctate, with moderately wide epipleura conspicuous along whole length of elytra; interlocking mechanism not developed; elytra without distinct longitudinal carinae; explanation of lateral margins of elytra much narrower than those in
<taxonomicName id="4C644D28DF45F41EFB8B0AB8AC8876C7" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="206" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF45F41EFB8B0AB8AC8876C7" italics="true" pageId="9" pageNumber="206">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF45F41EFF6F0A98AD6E76C7" box="[206,289,1212,1235]" pageId="9" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDB36ABDF45F41EFF310AF8AD0277E7" blockId="9.[113,1090,220,1587]" pageId="9" pageNumber="206">
<emphasis id="B910EAB9DF45F41EFF310AF8AC8D76E7" box="[144,194,1244,1267]" italics="true" pageId="9" pageNumber="206">Legs</emphasis>
(
<figureCitation id="135F2A2EDF45F41EFF6E0AF8AD6B76E7" box="[207,292,1244,1267]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. AD Afrocyrona pedestris sp. nov., male holotype:A aedeagus ventrally; B aedeagus laterally; C rudimentary wing; D protarsus. E A. volatilis sp. nov., paratype; F A. volatilis sp. nov., holotype; G A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="9" pageNumber="206">Fig. 4D</figureCitation>
): procoxae not projecting, transverse; mesocoxae oval, metacoxae extended to lateral margin of metathorax; trochanters relatively small, triangular; femora weakly clavate; tibiae with row of small spines along outer margin; tibial apical spur pattern 1(2)-2-2 (2nd spur of protibiae reduced in size) but apical spurs in meso- and metatibiae larger and more conspicuous than those in
<taxonomicName id="4C644D28DF45F41EFE2A0B78ADB07767" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[395,511,1372,1395]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="206" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF45F41EFE2A0B78ADB07767" box="[395,511,1372,1395]" italics="true" pageId="9" pageNumber="206">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF45F41EFDA70B78AE147767" box="[518,603,1372,1395]" pageId="9" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
; large protibial spur distinctly hooked; apices of meso- and metatibiae without conspicuous row of spines; tarsomere
<quantity id="4C9C9B4EDF45F41EFCCD0B58AFDA7787" box="[876,917,1404,1427]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="206" unit="in" value="1.0">1 in</quantity>
all pairs of legs conspicuous, approximately as long as tarsomere 2; tarsomere 5 distinctly longer than 14 combined; tarsal lobes absent; claws without denticles; empodium large, projecting, bisetose; tarsal formula 5-5-5.
</paragraph>
<paragraph id="8BDB36ABDF45F41CFF310BD8ACB377C4" blockId="9.[113,1090,220,1587]" lastBlockId="11.[113,1089,1337,1584]" lastPageId="11" lastPageNumber="208" pageId="9" pageNumber="206">
<emphasis id="B910EAB9DF45F41EFF310BD8ACB67407" box="[144,249,1532,1555]" italics="true" pageId="9" pageNumber="206">Abdomen</emphasis>
(
<figureCitation id="135F2A2EDF45F41EFEAA0BD8ADC27407" box="[267,397,1532,1555]" captionStart="Fig" captionStartId="10.[151,182,1514,1533]" captionTargetBox="[157,1117,219,1474]" captionTargetId="figure-11@10.[153,1124,219,1489]" captionTargetPageId="10" captionText="Fig. 4. AD Afrocyrona pedestris sp. nov., male holotype:A aedeagus ventrally; B aedeagus laterally; C rudimentary wing; D protarsus. E A. volatilis sp. nov., paratype; F A. volatilis sp. nov., holotype; G A. pedestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="9" pageNumber="206">Figs 4AB</figureCitation>
,
<figureCitation id="135F2A2EDF45F41EFE3D0BD8ADA97407" box="[412,486,1532,1555]" captionStart="Fig" captionStartId="11.[113,144,1208,1227]" captionTargetBox="[130,1087,224,1178]" captionTargetId="figure-13@11.[113,1091,219,1183]" captionTargetPageId="11" captionText="Fig. 5. AD Afrocyrona pedestris sp. nov., male holotype: A abdominal segment IX with spiculum of sternite IX (spicular fork); B tegmen ventrally; C sternite VIII; D tergite VIII. EH A. volatilis sp. nov., female paratype: E lacinia (incl. basistipes) and galea (incl. basigalea). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5313292" httpUri="https://zenodo.org/record/5313292/files/figure.png" pageId="9" pageNumber="206">5AD</figureCitation>
): male segment IX incomplete (reduced), only with membranous (pubescent) rudiment of sternite IX and well-developed spiculum, tergite IX inconspicuous; tegmen composed of three parts (originally, i.e. in uninverted aedeagus, dorsal parameres separated from phallobase which is split into separate ventral apodeme and lateral struts), inverted (phallobase ventrally open; i.e., parameres ventrally and apodeme dorsally situated); margins of parameres shortly pubescent; phallus slender, slightly longer than tegmen.
</paragraph>
</subSubSection>
<caption id="DF1B6623DF46F41DFF360BCEAD7A7427" ID-DOI="http://doi.org/10.5281/zenodo.5313288" ID-Zenodo-Dep="5313288" httpUri="https://zenodo.org/record/5313288/files/figure.png" pageId="10" pageNumber="207" startId="10.[151,182,1514,1533]" targetBox="[157,1117,219,1474]" targetPageId="10">
<paragraph id="8BDB36ABDF46F41DFF360BCEAD7A7427" blockId="10.[151,1127,1514,1587]" pageId="10" pageNumber="207">
Fig. 4. AD
<taxonomicName id="4C644D28DF46F41DFEB10BCEAD8D77E9" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[272,450,1514,1533]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="207" phylum="Arthropoda" rank="species" species="pedestris" status="sp. nov.">
<emphasis id="B910EAB9DF46F41DFEB10BCEAD8D77E9" box="[272,450,1514,1533]" italics="true" pageId="10" pageNumber="207">Afrocyrona pedestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF46F41DFE660BCEAE4377E9" box="[455,524,1514,1533]" pageId="10" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype:A aedeagus ventrally; B aedeagus laterally; C rudimentary wing; D protarsus. E
<taxonomicName id="4C644D28DF46F41DFE620821AE51740C" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[451,542,1541,1560]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="207" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF46F41DFE620821AE51740C" box="[451,542,1541,1560]" italics="true" pageId="10" pageNumber="207">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF46F41DFD820821AE29740C" box="[547,614,1541,1560]" pageId="10" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
, paratype; F
<taxonomicName id="4C644D28DF46F41DFD410821AF73740C" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[736,828,1541,1560]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="207" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF46F41DFD410821AF73740C" box="[736,828,1541,1560]" italics="true" pageId="10" pageNumber="207">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF46F41DFCE10821AFCB740C" box="[832,900,1541,1560]" pageId="10" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
, holotype; G
<taxonomicName id="4C644D28DF46F41DFBA20821A829740C" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[1027,1126,1541,1560]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="207" phylum="Arthropoda" rank="species" species="pedestris" status="sp. nov.">
<emphasis id="B910EAB9DF46F41DFBA20821A829740C" box="[1027,1126,1541,1560]" italics="true" pageId="10" pageNumber="207">A. pedestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF46F41DFF360804AC937427" box="[151,220,1568,1587]" pageId="10" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
, holotype.
</paragraph>
</caption>
<caption id="DF1B6623DF47F41CFFD00A9CAF187714" ID-DOI="http://doi.org/10.5281/zenodo.5313292" ID-Zenodo-Dep="5313292" httpUri="https://zenodo.org/record/5313292/files/figure.png" pageId="11" pageNumber="208" startId="11.[113,144,1208,1227]" targetBox="[130,1087,224,1178]" targetPageId="11">
<paragraph id="8BDB36ABDF47F41CFFD00A9CAF187714" blockId="11.[113,1089,1208,1280]" pageId="11" pageNumber="208">
Fig. 5. AD
<taxonomicName id="4C644D28DF47F41CFF4E0A9CADEE76DF" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[239,417,1208,1227]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="208" phylum="Arthropoda" rank="species" species="pedestris" status="sp. nov.">
<emphasis id="B910EAB9DF47F41CFF4E0A9CADEE76DF" box="[239,417,1208,1227]" italics="true" pageId="11" pageNumber="208">Afrocyrona pedestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF47F41CFE060A9CADA276DF" box="[423,493,1208,1227]" pageId="11" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype: A abdominal segment IX with spiculum of sternite IX (spicular fork); B tegmen ventrally; C sternite VIII; D tergite VIII. EH
<taxonomicName id="4C644D28DF47F41CFCF50AF6AFFD76F1" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[852,946,1234,1253]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="208" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF47F41CFCF50AF6AFFD76F1" box="[852,946,1234,1253]" italics="true" pageId="11" pageNumber="208">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF47F41CFC190AF6AFB176F1" box="[952,1022,1234,1253]" pageId="11" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype: E lacinia (incl. basistipes) and galea (incl. basigalea). Scale bars = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C37E6520DF47F41CFFD00BFDADBB77E4" box="[113,500,1497,1520]" pageId="11" pageNumber="208" type="diagnosis">
<paragraph id="8BDB36ABDF47F41CFFD00BFDADBB77E4" blockId="11.[113,1089,1337,1584]" box="[113,500,1497,1520]" pageId="11" pageNumber="208">
<emphasis id="B910EAB9DF47F41CFFD00BFDAD2377E4" bold="true" box="[113,364,1497,1520]" pageId="11" pageNumber="208">Differential diagnosis.</emphasis>
See
<tableCitation id="C6E60310DF47F41CFE3E0BFDADBF77E4" box="[415,496,1497,1520]" captionStart="Table 2" captionStartId="7.[113,160,749,768]" captionText="Table 2. Differential diagnosis between Afrocyrona volatilis sp.nov.and A. pedestris sp.nov.The particular character states are explained in detail in descriptions or illustrated." pageId="11" pageNumber="208">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E6520DF47F41CFFD00BDDA80E7404" box="[113,1089,1529,1552]" pageId="11" pageNumber="208" type="etymology">
<paragraph id="8BDB36ABDF47F41CFFD00BDDA80E7404" blockId="11.[113,1089,1337,1584]" box="[113,1089,1529,1552]" pageId="11" pageNumber="208">
<emphasis id="B910EAB9DF47F41CFFD00BDDACBD7404" bold="true" box="[113,242,1529,1552]" pageId="11" pageNumber="208">Etymology.</emphasis>
The species name is Latin adjective
<taxonomicName id="4C644D28DF47F41CFDD60BDDAE987404" authorityName="Kolibáč" authorityYear="2014" box="[631,727,1529,1552]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="208" phylum="Arthropoda" rank="species" species="pedestris">
<emphasis id="B910EAB9DF47F41CFDD60BDDAE987404" box="[631,727,1529,1552]" italics="true" pageId="11" pageNumber="208">pedestris</emphasis>
</taxonomicName>
(-
<emphasis id="B910EAB9DF47F41CFD4E0BDDAF437404" box="[751,780,1529,1552]" italics="true" pageId="11" pageNumber="208">tre</emphasis>
) = on foot, afoot, pedestrian.
</paragraph>
</subSubSection>
<subSubSection id="C37E6520DF47F41BFFD0083DAEC97307" lastPageId="12" lastPageNumber="209" pageId="11" pageNumber="208" type="biology_ecology">
<paragraph id="8BDB36ABDF47F41BFFD0083DAEC97307" blockId="11.[113,1089,1337,1584]" lastBlockId="12.[151,1127,220,275]" lastPageId="12" lastPageNumber="209" pageId="11" pageNumber="208">
<emphasis id="B910EAB9DF47F41CFFD0083DAC827424" bold="true" box="[113,205,1561,1584]" pageId="11" pageNumber="208">Biology.</emphasis>
<materialsCitation id="3B0C3CF6DF47F41BFF72083DADA172E7" collectingDate="2012-06-16" collectingDateMax="2012-06-18" collectingDateMin="2012-06-16" collectionCode="SCAND, V, NMPC" collectorName="Socotra &amp; J. Bezdek &amp; J. Hajek &amp; Hula &amp; P. Kment &amp; I. Malenovsky &amp; J. Niedobova &amp; L. Purchart" country="Yemen" lastPageId="12" lastPageNumber="209" latitude="12.576667" location="Island" longLatPrecision="129" longitude="54.025" municipality="Hagher Mts." pageId="11" pageNumber="208" specimenCount="1">
The single male specimen was obtained by litter sifting in the same locality as the holotype of
<taxonomicName id="4C644D28DF40F41BFEBD0EF8ADDD72E7" authority="Kolibáč, 2014" authorityName="Kolibáč" authorityYear="2014" box="[284,402,220,243]" class="Insecta" family="Trogossitidae" genus="Afrocyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="volatilis" status="sp. nov.">
<emphasis id="B910EAB9DF40F41BFEBD0EF8ADDD72E7" box="[284,402,220,243]" italics="true" pageId="12" pageNumber="209">A. volatilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357C2DF40F41BFE3B0EF8ADA172E7" box="[410,494,220,243]" pageId="12" pageNumber="209" rank="species">sp. nov</taxonomicNameLabel>
</materialsCitation>
.; however, the
<specimenCount id="9D62FD22DF40F41BFD300EF8AF7E72E7" box="[657,817,220,243]" pageId="12" pageNumber="209" type="generic">two specimens</specimenCount>
were not collected together. Wingless species. Gut empty, without content.
</paragraph>
</subSubSection>
</treatment>
</document>