treatments-xml/data/03/CB/3A/03CB3A0C5A1F055BFF3C0965FDFFCD2E.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.6266920" ID-GBIF-Dataset="3ded904d-e34f-4cd1-9903-79b8392a7f66" ID-GBIF-Taxon="114924054" ID-Zenodo-Dep="6266920" checkinTime="1436211515535" checkinUser="donat" docAuthor="Urs Thalmann &amp; Thomas Geissmann" docDate="2005" docId="03CB3A0C5A1F055BFF3C0965FDFFCD2E" docLanguage="en" docName="2005avahi_cleesei.pdf" docOrigin="AmerIcan Journal of PrImatology 67" docTitle="Avahi cleesei New, New SpecIes" docType="treatment" docVersion="5" lastPageId="4" lastPageNumber="375" masterDocId="FFF242745A1D055FFF9A0D61FFDFC83A" masterDocTitle="New Species of Woolly Lemur Avahi (Primates: Lemuriformes) in Bemaraha (Central Western Madagascar)" masterLastPageNumber="376" masterPageNumber="371" pageId="2" pageNumber="373" updateTime="1645745716752" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New Species of Woolly Lemur Avahi (Primates: Lemuriformes) in Bemaraha (Central Western Madagascar)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Urs Thalmann</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Thomas Geissmann</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>AmerIcan Journal of PrImatology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2005</mods:date>
<mods:detail type="volume">
<mods:number>67</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>371</mods:start>
<mods:end>376</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1002/ajp.20191</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6266920" ID-GBIF-Taxon="114924054" ID-Zenodo-Dep="6266920" LSID="urn:lsid:plazi:treatment:03CB3A0C5A1F055BFF3C0965FDFFCD2E" httpUri="http://treatment.plazi.org/id/03CB3A0C5A1F055BFF3C0965FDFFCD2E" lastPageId="4" lastPageNumber="375" pageId="2" pageNumber="373">
<subSubSection box="[166,524,1028,1052]" pageId="2" pageNumber="373" type="nomenclature">
<paragraph box="[166,524,1028,1052]" pageId="2" pageNumber="373">
<heading bold="true" box="[166,524,1028,1052]" fontSize="10" level="3" pageId="2" pageNumber="373" reason="0">
<taxonomicName authority="New" authorityName="New" box="[166,414,1028,1052]" class="Mammalia" family="Indriidae" genus="Avahi" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="2" pageNumber="373" phylum="Chordata" rank="species" species="cleesei">
<emphasis bold="true" box="[166,347,1028,1052]" italics="true" pageId="2" pageNumber="373">
<taxonomicName box="[166,244,1028,1052]" pageId="2" pageNumber="373">Avahi</taxonomicName>
cleesei:
</emphasis>
<taxonomicNameLabel box="[356,524,1028,1052]" pageId="2" pageNumber="373" rank="species">New SpecIes</taxonomicNameLabel>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="373" type="description">
<paragraph box="[218,804,1067,1092]" pageId="2" pageNumber="373">
<heading box="[218,804,1067,1092]" fontSize="10" level="4" pageId="2" pageNumber="373" reason="1">Bemaraha form [Thalmann &amp; Geissmann, 2000].</heading>
</paragraph>
<paragraph box="[166,370,1140,1164]" pageId="2" pageNumber="373">
<heading bold="true" box="[166,370,1140,1164]" fontSize="10" level="3" pageId="2" pageNumber="373" reason="0">
<emphasis bold="true" box="[166,370,1140,1164]" italics="true" pageId="2" pageNumber="373">
<typeStatus box="[166,272,1140,1164]" pageId="2" pageNumber="373">Holotype</typeStatus>
(
<figureCitation box="[292,360,1140,1164]" captionStart="Fig. 2" captionStartId="3.[166,201,1175,1195]" captionTargetBox="[175,1115,188,1155]" captionTargetId="figure@3.[173,1117,185,1159]" captionTargetPageId="3" captionText="Fig. 2. Type photographs (AIMZ 13854. b-d) of an adult male Avahi cleesei after it was captured on 3 October 1991 (a and b) and after it was released on 4 October 1991 (c). [Color figure can be viewed in the online issue, which is available at www. interscience. wiley. com.]" pageId="2" pageNumber="373">Fig. 2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph box="[218,1123,1168,1193]" pageId="2" pageNumber="373">
<materialsCitation ID-GBIF-Occurrence="1227780005" box="[218,1119,1168,1193]" collectingDate="1991-10-03" country="Afghanistan" location="Bemaraha" pageId="2" pageNumber="373" specimenCount="1">
<taxonomicName box="[218,287,1169,1193]" pageId="2" pageNumber="373">
<emphasis bold="true" box="[218,287,1169,1193]" italics="true" pageId="2" pageNumber="373">Avahi</emphasis>
</taxonomicName>
male (body weight =
<quantity box="[553,616,1168,1193]" metricMagnitude="-1" metricUnit="kg" metricValue="8.3" pageId="2" pageNumber="373" unit="g" value="830.0">830 g</quantity>
) captured in
<location LSID="urn:lsid:plazi:treatment:03CB3A0C5A1F055BFF3C0965FDFFCD2E:8EBDDDC15A1F055DFC9C09F1FCA0CC93" box="[774,895,1168,1193]" country="Afghanistan" name="Bemaraha" pageId="2" pageNumber="373">Bemaraha</location>
on
<collectingDate box="[938,1119,1168,1193]" pageId="2" pageNumber="373" value="1991-10-03">
<date box="[938,1119,1168,1193]" pageId="2" pageNumber="373" value="1991-10-03">3 October 1991</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph box="[166,270,1241,1264]" pageId="2" pageNumber="373">
<heading bold="true" box="[166,270,1241,1264]" fontSize="10" level="3" pageId="2" pageNumber="373" reason="0">
<emphasis bold="true" box="[166,270,1241,1264]" italics="true" pageId="2" pageNumber="373">Material</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="373" type="materials_examined">
<paragraph pageId="2" pageNumber="373">
<materialsCitation ID-GBIF-Occurrence="1227780004" collectionCode="AIMZ" country="Madagascar" pageId="2" pageNumber="373" specimenCode="AIMZ 13854" specimenCount="1">
Hair samples (
<specimenCode box="[388,541,1269,1294]" pageId="2" pageNumber="373">AIMZ 13854</specimenCode>
.a), accompanying photographs (
<specimenCode box="[914,1064,1269,1294]" pageId="2" pageNumber="373">AIMZ 13854</specimenCode>
.b-n), and videotape (
<specimenCode box="[351,504,1298,1323]" pageId="2" pageNumber="373">AIMZ 13854</specimenCode>
.o) and audiotape recordings (
<specimenCode box="[861,1015,1298,1323]" pageId="2" pageNumber="373">AIMZ 13854</specimenCode>
.p) of the
<specimenCount box="[166,276,1328,1352]" pageId="2" pageNumber="373" type="generic">specimen</specimenCount>
are stored in the Anthropological Institute and Museum of the University of Zurich (
<collectionCode box="[425,499,1357,1381]" collectionName="AIMZ" pageId="2" pageNumber="373">AIMZ</collectionCode>
), Zurich, Switzerland
</materialsCitation>
.
</paragraph>
<paragraph box="[166,321,1429,1453]" pageId="2" pageNumber="373">
<heading bold="true" box="[166,321,1429,1453]" fontSize="10" level="3" pageId="2" pageNumber="373" reason="0">
<emphasis bold="true" box="[166,321,1429,1453]" italics="true" pageId="2" pageNumber="373">
<typeStatus box="[166,224,1430,1453]" pageId="2" pageNumber="373">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<materialsCitation ID-GBIF-Occurrence="1227780001" country="Madagascar" location="Madagascar" pageId="2" pageNumber="373" specimenCount="1">
<paragraph pageId="2" pageNumber="373">
<geoCoordinate box="[218,307,1455,1482]" pageId="2" pageNumber="373">18°59̸S</geoCoordinate>
,
<geoCoordinate box="[331,423,1455,1482]" pageId="2" pageNumber="373">44°45̸E</geoCoordinate>
, approximately
<locationDeviation box="[633,704,1458,1482]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="373" unit="km" value="3.0">
<quantity box="[633,704,1458,1482]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="373" unit="km" value="3.0">3 km</quantity>
</locationDeviation>
east-northeast of the village Ambalarano, western
<collectingCountry box="[429,570,1487,1511]" name="Madagascar" pageId="2" pageNumber="373" value="Madagascar">Madagascar</collectingCountry>
(
<figureCitation box="[587,654,1487,1511]" captionStart="Fig. 1" captionStartId="1.[166,201,1119,1139]" captionTargetBox="[173,1116,187,1095]" captionTargetId="figure@1.[173,1117,185,1103]" captionTargetPageId="1" captionText="Fig. 1. Distribution map of western taxa of woolly lemurs." pageId="2" pageNumber="373">Fig. 1</figureCitation>
).
</paragraph>
<paragraph box="[1101,1123,1458,1482]" pageId="2" pageNumber="373">of</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection pageId="2" pageNumber="373" type="etymology">
<paragraph box="[166,293,1560,1583]" pageId="2" pageNumber="373">
<emphasis bold="true" box="[166,293,1560,1583]" italics="true" pageId="2" pageNumber="373">
<heading bold="true" box="[166,293,1560,1583]" fontSize="10" level="3" pageId="2" pageNumber="373" reason="0">Etymology</heading>
</emphasis>
</paragraph>
<paragraph pageId="2" pageNumber="373">
Named in honor of British actor and comedian John Cleese for his promotion of conservation issues in movies such as
<emphasis bold="true" box="[639,830,1618,1642]" italics="true" pageId="2" pageNumber="373">Fierce Creatures</emphasis>
[Chepisi &amp; Young, 1997] and documentaries such as
<emphasis bold="true" box="[498,1123,1647,1671]" italics="true" pageId="2" pageNumber="373">Born to be Wild: Operation Lemur With John Cleese</emphasis>
[Kershaw &amp; Cleese, 1999].
</paragraph>
</subSubSection>
<caption pageId="3" pageNumber="374" targetBox="[175,1115,188,1155]" targetPageId="3">
<paragraph pageId="3" pageNumber="374">
Fig. 2. Type photographs (AIMZ 13854.b-d) of an adult male
<taxonomicName box="[759,883,1176,1195]" class="Mammalia" family="Indriidae" genus="Avahi" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="cleesei">
<emphasis bold="true" box="[759,883,1176,1195]" italics="true" pageId="3" pageNumber="374">
<taxonomicName box="[759,816,1176,1195]" pageId="3" pageNumber="374">Avahi</taxonomicName>
cleesei
</emphasis>
</taxonomicName>
after it was captured on 3 October 1991 (
<emphasis bold="true" box="[324,336,1198,1217]" pageId="3" pageNumber="374">a</emphasis>
and
<emphasis bold="true" box="[383,396,1198,1217]" pageId="3" pageNumber="374">b</emphasis>
) and after it was released on 4 October 1991 (
<emphasis bold="true" box="[834,845,1198,1217]" pageId="3" pageNumber="374">c</emphasis>
). [Color figure can be viewed in the online issue, which is available at www.interscience.wiley.com.]
</paragraph>
</caption>
<subSubSection pageId="3" pageNumber="374" type="diagnosis">
<paragraph box="[166,287,1302,1326]" pageId="3" pageNumber="374">
<heading bold="true" box="[166,287,1302,1326]" fontSize="10" level="3" pageId="3" pageNumber="374" reason="0">
<emphasis bold="true" box="[166,287,1302,1326]" italics="true" pageId="3" pageNumber="374">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph pageId="3" pageNumber="374">
Distinguished from
<emphasis bold="true" box="[462,638,1331,1355]" italics="true" pageId="3" pageNumber="374">
<taxonomicName box="[462,638,1331,1355]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="1340" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="occidentalis">A. occidentalis</taxonomicName>
</emphasis>
by its lack of a white facial mask and broad dark eye-rings, and from both
<taxonomicName box="[642,820,1360,1384]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="1359" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="occidentalis">
<emphasis bold="true" box="[642,820,1360,1384]" italics="true" pageId="3" pageNumber="374">A. occidentalis</emphasis>
</taxonomicName>
and
<taxonomicName box="[892,1028,1360,1384]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="1347" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="unicolor">
<emphasis bold="true" box="[892,1028,1360,1384]" italics="true" pageId="3" pageNumber="374">A. unicolor</emphasis>
</taxonomicName>
by the presence of a dark chevron pattern on the forehead.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374" type="description">
<paragraph box="[166,402,1472,1496]" pageId="3" pageNumber="374">
<heading bold="true" box="[166,402,1472,1496]" fontSize="10" level="3" pageId="3" pageNumber="374" reason="0">
<emphasis bold="true" box="[166,402,1472,1496]" italics="true" pageId="3" pageNumber="374">
Description (
<figureCitation box="[324,392,1472,1496]" captionStart="Fig. 2" captionStartId="3.[166,201,1175,1195]" captionTargetBox="[175,1115,188,1155]" captionTargetId="figure@3.[173,1117,185,1159]" captionTargetPageId="3" captionText="Fig. 2. Type photographs (AIMZ 13854. b-d) of an adult male Avahi cleesei after it was captured on 3 October 1991 (a and b) and after it was released on 4 October 1991 (c). [Color figure can be viewed in the online issue, which is available at www. interscience. wiley. com.]" pageId="3" pageNumber="374">Fig. 2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374">
The face of
<taxonomicName box="[351,456,1501,1525]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="300" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="cleesei">
<emphasis bold="true" box="[351,456,1501,1525]" italics="true" pageId="3" pageNumber="374">A. cleesei</emphasis>
</taxonomicName>
is only slightly paler than the upper forehead and crown. The facial area above the nose extends upward toward the forehead (
<figureCitation box="[969,1037,1530,1554]" captionStart="Fig. 2" captionStartId="3.[166,201,1175,1195]" captionTargetBox="[175,1115,188,1155]" captionTargetId="figure@3.[173,1117,185,1159]" captionTargetPageId="3" captionText="Fig. 2. Type photographs (AIMZ 13854. b-d) of an adult male Avahi cleesei after it was captured on 3 October 1991 (a and b) and after it was released on 4 October 1991 (c). [Color figure can be viewed in the online issue, which is available at www. interscience. wiley. com.]" pageId="3" pageNumber="374">Fig. 2</figureCitation>
c). This upward extension contrasts with the virtually opposite pattern created by the triangle of forehead pelage that invades the facial area in other western
<emphasis bold="true" box="[1053,1123,1589,1612]" italics="true" pageId="3" pageNumber="374">
<taxonomicName box="[1053,1123,1589,1612]" pageId="3" pageNumber="374">Avahi</taxonomicName>
</emphasis>
(e.g.,
<taxonomicName box="[229,399,1618,1642]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="1450" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="occidentalis">
<emphasis bold="true" box="[229,399,1618,1642]" italics="true" pageId="3" pageNumber="374">A. occidentalis</emphasis>
</taxonomicName>
and
<taxonomicName box="[455,584,1618,1642]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="1438" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="unicolor">
<emphasis bold="true" box="[455,584,1618,1642]" italics="true" pageId="3" pageNumber="374">A. unicolor</emphasis>
</taxonomicName>
). The forehead fur immediately bordering the facial area is blackish and forms a dark chevron pattern above the facial area. The eyes are maroon and have black, hairless eyelids. The snout is also black and hairless. The fur surrounding the corners of mouth is whitish. The fur on the head and body has a brown-gray coloration and a woolly (slightly curled) flecked appearance. The tail is beige or brown-gray, and is slightly reddish only on the dorsal side of the root. The inner dorsal surface of the lower limbs is white. The fur of the chest, belly, and inner surface of the upper limbs is relatively thin, downy, and very light gray.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="375" type="distribution">
<paragraph box="[166,314,400,424]" pageId="4" pageNumber="375">
<emphasis bold="true" box="[166,314,400,424]" italics="true" pageId="4" pageNumber="375">
<heading bold="true" box="[166,314,400,424]" fontSize="10" level="3" pageId="4" pageNumber="375" reason="0">Distribution</heading>
</emphasis>
</paragraph>
<paragraph pageId="4" pageNumber="375">This species has been found only in the Tsingy de Bemaraha region north of the Manambolo River. The northern limit is unclear. There is no evidence from surveys and interviews that it is distributed in the wider surroundings of Bemaraha [Ausilio &amp; Raveloanrinoro, 1998; Sterling, 1998], the Namoroka Strict Nature Reserve (between the Sambao and Mahavavy rivers [Hawkins et al., 1998; Thalmann et al., 1999], or between the Mahavavy and Betsiboka rivers [Curtis, 1997; Thalmann &amp; Geissmann, 2000].</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="375" type="etymology">
<paragraph box="[166,387,672,696]" pageId="4" pageNumber="375">
<emphasis bold="true" box="[166,387,672,696]" italics="true" pageId="4" pageNumber="375">
<heading bold="true" box="[166,387,672,696]" fontSize="10" level="3" pageId="4" pageNumber="375" reason="0">Vernacular names</heading>
</emphasis>
</paragraph>
<paragraph pageId="4" pageNumber="375">
Locally, these animals are called Dadintsifaky, which translates to grandparent of the sifaky (
<emphasis bold="true" box="[429,567,731,754]" italics="true" pageId="4" pageNumber="375">
<taxonomicName box="[429,567,731,754]" class="Mammalia" family="Indriidae" genus="Propithecus" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="4" pageNumber="375" phylum="Chordata" rank="genus">Propithecus</taxonomicName>
</emphasis>
). As a vernacular name in English, we propose Cleeses woolly lemur or Bemaraha woolly lemur.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="375" type="discussion">
<paragraph box="[166,402,828,851]" pageId="4" pageNumber="375">
<heading allCaps="true" bold="true" box="[166,402,828,851]" fontSize="10" level="1" pageId="4" pageNumber="375" reason="0">
<emphasis bold="true" box="[166,402,828,851]" pageId="4" pageNumber="375">CONSERVATION</emphasis>
</heading>
</paragraph>
<paragraph pageId="4" pageNumber="375">
<taxonomicName box="[218,367,867,891]" class="Mammalia" family="Indriidae" genus="Avahi" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="4" pageNumber="375" phylum="Chordata" rank="species" species="cleesei">
<emphasis bold="true" box="[218,367,867,891]" italics="true" pageId="4" pageNumber="375">
<taxonomicName box="[218,287,867,891]" pageId="4" pageNumber="375">Avahi</taxonomicName>
cleesei
</emphasis>
</taxonomicName>
clearly falls into the category of Endangered (
<collectionCode box="[904,945,867,891]" pageId="4" pageNumber="375">EN</collectionCode>
) as established by the IUCN Red List [IUCN, 2001], according to the following criteria (B1ac(i, ii, iii, v)): the extent of occurrence is estimated to be less than
<quantity box="[889,999,925,950]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="375" unit="km" value="5000.0">5,000 km</quantity>
2 (
<collectionCode box="[1028,1050,925,949]" pageId="4" pageNumber="375">B</collectionCode>
1), the species is known in just one location (
<collectionCode box="[607,628,954,979]" pageId="4" pageNumber="375">B</collectionCode>
1a), and the known population is declining (
<collectionCode box="[176,197,984,1008]" pageId="4" pageNumber="375">B</collectionCode>
1c) in the extent of occurrence (i); area of occupancy (ii); area, extent, and/or quality of habitat (iii); and number of mature individuals (v).
</paragraph>
<paragraph pageId="4" pageNumber="375">
However, more detailed analyses and surveys may reveal that the taxon should be moved to the Critically Endangered (
<collectionCode box="[743,782,1071,1095]" pageId="4" pageNumber="375">CR</collectionCode>
) category. For example, the disturbed forest close to the village of Ankinajao, which supported a substantial number of individuals in 1994 [Thalmann &amp; Geissmann, 2000], had been cut completely by 2003 (Thalmann, unpublished data). The subhumid forest at the base of the escarpment of the Tsingy de Bemaraha is under continuous pressure from bush fires that reduce it in many places every yearin some places to only a few meters in width. Such subhumid forests are the only habitat in which
<taxonomicName class="Mammalia" family="Indriidae" genus="Avahi" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="4" pageNumber="375" phylum="Chordata" rank="species" species="cleesei">
<emphasis bold="true" italics="true" pageId="4" pageNumber="375">
<taxonomicName box="[1053,1123,1247,1271]" pageId="4" pageNumber="375">Avahi</taxonomicName>
cleesei
</emphasis>
</taxonomicName>
is known to occur so far.
</paragraph>
</subSubSection>
</treatment>
</document>