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<document id="87E51E32A8ED65EE4FE0FBA6017B8BD1" ID-CLB-Dataset="56108" ID-DOI="10.6620/ZS.2019.58-14" ID-GBIF-Dataset="de0647eb-9b73-4bac-9ff7-f7bff7518dc0" ID-PMC="PMC6759935" ID-PubMed="31966315" ID-Zenodo-Dep="8065869" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1687182873363" checkinUser="valdenar" docAuthor="Nguyen, Anh D., Nguyen, Manh-Ha, Nguyen, Thu-Anh T. &amp; Phung, Hong-Luong T." docDate="2019" docId="03B987C4FFE9AE5CFF06FED8FAD4AAF4" docLanguage="en" docName="ZoolStud.58.14.1-31.pdf" docOrigin="Zoological Studies 58 (14)" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Hylomus asper" docType="treatment" docVersion="7" lastPageNumber="4" masterDocId="FF80FFBCFFEAAE5FFFF0FFA5FF8AAB21" masterDocTitle="Review of Dragon Millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) in the Fauna of Vietnam, with Descriptions of Three New Species" masterLastPageNumber="31" masterPageNumber="1" pageNumber="4" updateTime="1716451626079" updateUser="tatiana">
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<mods:title id="932A6F386489C3B93914CD050AAA684B">Review of Dragon Millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) in the Fauna of Vietnam, with Descriptions of Three New Species</mods:title>
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<mods:namePart id="CA819CA2ACE9712CADAD5BE3E767C850">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="FBE54F038B8C288F7B501E1526A5DE5C">Nguyen, Manh-Ha</mods:namePart>
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<mods:namePart id="3C10F89D8672DC886216D06144A482F7">Nguyen, Thu-Anh T.</mods:namePart>
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<mods:namePart id="3AA1424CF60174C7ED5E92361F9C240B">Phung, Hong-Luong T.</mods:namePart>
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<mods:date id="E70752AD2E28419C2003B318CE8C0431">2019</mods:date>
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<treatment id="03B987C4FFE9AE5CFF06FED8FAD4AAF4" ID-DOI="http://doi.org/10.5281/zenodo.8055825" ID-GBIF-Taxon="209694539" ID-Zenodo-Dep="8055825" LSID="urn:lsid:plazi:treatment:03B987C4FFE9AE5CFF06FED8FAD4AAF4" httpUri="http://treatment.plazi.org/id/03B987C4FFE9AE5CFF06FED8FAD4AAF4" lastPageNumber="4" pageId="3" pageNumber="4">
<subSubSection id="C30A6559FFE9AE5CFF06FED8FDF8AAB7" box="[246,626,381,406]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BAF36D2FFE9AE5CFF06FED8FDF8AAB7" blockId="3.[246,626,381,437]" box="[246,626,381,406]" pageId="3" pageNumber="4">
<heading id="D0E781BEFFE9AE5CFF06FED8FDF8AAB7" bold="true" box="[246,626,381,406]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
<taxonomicName id="4C104D51FFE9AE5CFF06FED8FDF8AAB7" ID-CoL="3NGCR" authority="(Attems, 1937)" baseAuthorityName="Attems" baseAuthorityYear="1937" box="[246,626,381,406]" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="B964EAC0FFE9AE5CFF06FED8FDF8AAB7" bold="true" box="[246,626,381,406]" pageId="3" pageNumber="4">
<emphasis id="B964EAC0FFE9AE5CFF06FED8FE3FAAB7" bold="true" box="[246,437,381,406]" italics="true" pageId="3" pageNumber="4">Hylomus asper</emphasis>
(
<bibRefCitation id="EF814B23FFE9AE5CFE34FED8FDE6AAB7" author="Attems C." box="[452,620,381,406]" pageId="3" pageNumber="4" pagination="1 - 300" refId="ref11356" refString="Attems C. 1937. Myriapoda 3. Polydesmoidea I. Fam. Strongylosomidae. Das Tierreich 68: 1 - 300." type="journal article" year="1937">Attems, 1937</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE9AE5CFE89FE38FE7AAA94" box="[377,496,413,437]" pageId="3" pageNumber="4" type="description">
<paragraph id="8BAF36D2FFE9AE5CFE89FE38FE7AAA94" blockId="3.[246,626,381,437]" box="[377,496,413,437]" pageId="3" pageNumber="4">
(
<figureCitation id="132B2A57FFE9AE5CFE71FE38FE6DAA94" box="[385,487,413,437]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[175,206,1813,1832]" captionStartId-1="3.[137,168,1952,1971]" captionTargetBox-0="[440,1185,515,1783]" captionTargetBox-1="[233,1316,1032,1927]" captionTargetId-0="figure-96@2.[440,1185,486,1804]" captionTargetId-1="figure-311@3.[233,1316,1032,1927]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Distribution of Hylomus species in mainland Vietnam. 1: Hoang Lien National Park (Sa Pa, Lao Cai Province); 2: Nam Xay Commune (Van Ban District, Lao Cai Province); 3: Duc Xuan Commune (Bac Quang District, Ha Giang Province); 4: Xuan Son National Park (Phu Tho Province); 5: Tam Dao (Vinh Phuc Province); 6: Cuc Phuong National Park (Ninh Binh Province); 7: Phong Nha – Ke Bang National Park (Quang Binh Province); 8: Lien Chieu District (Da Nang Province); 9: Ba Na National Park (Da Nang Province); 10: Lo Xo Pass (Kon Tum Province); 11: Ngoc Linh Mt. (Kon Tum Province); 12: Kon Chu Rang Natural Reserve (Gia Lai Province); 13: Buon Ho District (Dak Lak Province); 14: Hon Ba Mt. (Khanh Hoa Province); 15: Phan Rang (Ninh Thuan Province); 16: Cat Tien National Park (Dong Nai Province)." captionText-1="Fig. 2. Hylomus asper (Attems, 1937) from Ngoc Linh Mt. Right gonopod, lateral view (A), mesal view (B)." figureDoi-0="http://doi.org/10.5281/zenodo.8065873" figureDoi-1="http://doi.org/10.5281/zenodo.8065877" httpUri-0="https://zenodo.org/record/8065873/files/figure.png" httpUri-1="https://zenodo.org/record/8065877/files/figure.png" pageId="3" pageNumber="4">Figs. 1−2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE9AE5CFF78FE7DFAD4AAF4" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="8BAF36D2FFE9AE5CFF78FE7DFEF9A927" blockId="3.[136,738,472,625]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="AB0011FCFFE9AE5CFF78FE7DFD68AACA" box="[136,738,472,491]" pageId="3" pageNumber="4">
<treatmentCitation id="0AB110C3FFE9AE5CFF78FE7DFD9AAACA" author="Attems C." box="[136,528,472,491]" page="126" pageId="3" pageNumber="4" year="1937">
<taxonomicName id="4C104D51FFE9AE5CFF78FE7DFD9AAACA" ID-CoL="15459dcc-20ce-4eef-97ba-07c86f11b2f2" authority="Attems, 1937: 125 - 126" authorityName="Attems" authorityPageNumber="125 - 126" authorityYear="1937" box="[136,528,472,491]" class="Diplopoda" family="Paradoxosomatidae" genus="Centrodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="B964EAC0FFE9AE5CFF78FE7DFEB0AACA" box="[136,314,472,491]" italics="true" pageId="3" pageNumber="4">Centrodesmus asper</emphasis>
<bibRefCitation id="EF814B23FFE9AE5CFECFFE7DFD9AAACA" author="Attems C." box="[319,528,472,491]" pageId="3" pageNumber="4" pagination="1 - 300" refId="ref11356" refString="Attems C. 1937. Myriapoda 3. Polydesmoidea I. Fam. Strongylosomidae. Das Tierreich 68: 1 - 300." type="journal article" year="1937">Attems, 1937: 125−126</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 162;
<bibRefCitation id="EF814B23FFE9AE5CFD9BFE7DFD68AACA" author="Attems C." box="[619,738,472,491]" pageId="3" pageNumber="4" pagination="187 - 353" refId="ref11379" refString="Attems C. 1938. Die von Dr. C. Dawydoff in franzosisch-Indochina gesammelten Myriopoden. Memoires du Museum National d'Histoire Naturelle, Paris {N. S.} 6 (2): 187 - 353." type="journal article" year="1938">Attems 1938:</bibRefCitation>
</treatmentCitationGroup>
240−241, figs. 75−78.
</paragraph>
<paragraph id="8BAF36D2FFE9AE5CFF78FDABFDA5A901" blockId="3.[136,738,472,625]" box="[136,559,525,545]" pageId="3" pageNumber="4">
<taxonomicName id="4C104D51FFE9AE5CFF78FDABFE10A901" ID-CoL="8PRPM" authority=": Jeekel 1964: 63" authorityName="Jeekel" authorityPageNumber="63" authorityYear="1964" box="[136,410,525,545]" class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="B964EAC0FFE9AE5CFF78FDABFE8CA901" box="[136,262,526,544]" italics="true" pageId="3" pageNumber="4">Pratinus asper</emphasis>
:
<bibRefCitation id="EF814B23FFE9AE5CFEE2FDA8FE10A901" author="Jeekel CAW" box="[274,410,525,544]" pageId="3" pageNumber="4" pagination="61 - 73" refId="ref11809" refString="Jeekel CAW. 1964. Two new species of Pratinus Attems, with taxonomoc notes on the genus and a redescription of its typespecies (Diplopoda, Polydesmida). Beaufortia 11 (137): 61 - 73." type="journal article" year="1964">Jeekel 1964: 63</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EF814B23FFE9AE5CFE53FDA8FDA1A901" author="Jeekel CAW" box="[419,555,525,544]" pageId="3" pageNumber="4" refId="ref11849" refString="Jeekel CAW. 1968. On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida). Bronder-Offset Rotterdam." type="book" year="1968">Jeekel 1968: 61</bibRefCitation>
.
</paragraph>
<paragraph id="8BAF36D2FFE9AE5CFF78FD8CFE0BA977" blockId="3.[136,738,472,625]" pageId="3" pageNumber="4">
<taxonomicName id="4C104D51FFE9AE5CFF78FD8CFDDEA91A" ID-CoL="6CNKX" authority=": Golovatch &amp; Enghoff 1994: 60" authorityName="Golovatch &amp; Enghoff" authorityPageNumber="60" authorityYear="1994" box="[136,596,552,572]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="aspera">
<emphasis id="B964EAC0FFE9AE5CFF78FD8CFEBAA91A" box="[136,304,553,571]" italics="true" pageId="3" pageNumber="4">Desmoxytes aspera</emphasis>
:
<bibRefCitation id="EF814B23FFE9AE5CFECEFD8DFDDEA91A" author="Golovatch SI &amp; Enghoff H." box="[318,596,552,571]" pageId="3" pageNumber="4" pagination="45 - 71" refId="ref11598" refString="Golovatch SI, Enghoff H. 1994. Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45 - 71." type="journal article" year="1994">Golovatch &amp; Enghoff 1994: 60</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EF814B23FFE9AE5CFDAFFD8DFD6BA91D" author="Likhitrakarn N &amp; Golovatch SI &amp; Panha S." box="[607,737,552,572]" pageId="3" pageNumber="4" pagination="483 - 504" refId="ref11904" refString="Likhitrakarn N, Golovatch SI, Panha S. 2015. Two new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from Laos (Diplopoda: Polydesmida: Paradoxosomatidae), with redescriptions of all four species of Attems from Vietnam. Zootaxa 3931: 483 - 504. doi: 10.11646 / zootaxa. 3931.4.2." type="journal article" year="2015">Likhitrakarn et</bibRefCitation>
al. 2015: 493, figs. 7−8.
</paragraph>
<paragraph id="8BAF36D2FFE9AE5CFF78FDF8FD13A950" blockId="3.[136,738,472,625]" box="[136,665,605,625]" pageId="3" pageNumber="4">
<taxonomicName id="4C104D51FFE9AE5CFF78FDF8FE2DA951" authority=": Jeekel 1980: 658" authorityName="Jeekel" authorityPageNumber="658" authorityYear="1980" box="[136,423,605,625]" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="B964EAC0FFE9AE5CFF78FDF8FE82A951" box="[136,264,605,624]" italics="true" pageId="3" pageNumber="4">Hylomus asper</emphasis>
:
<bibRefCitation id="EF814B23FFE9AE5CFEE4FDF8FE2DA951" author="Jeekel CAW" box="[276,423,605,625]" pageId="3" pageNumber="4" pagination="651 - 670" refId="ref11873" refString="Jeekel CAW. 1980. The generic allocation of some little-known Paradoxosomatidae from South-ast Asia (Diplopoda, Polydesmida). Revue suisse de zoologie 87: 651 - 670." type="journal article" year="1980">Jeekel 1980: 658</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EF814B23FFE9AE5CFE40FDF8FD1FA950" author="Srisonchai R &amp; Enghoff H &amp; Likhitrakarn N &amp; Panha S." box="[432,661,605,625]" pageId="3" pageNumber="4" pagination="1 - 177" refId="ref12093" refString="Srisonchai R, Enghoff H, Likhitrakarn N, Panha S. 2018 a. A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae). Zookeys 761: 1 - 177. doi: 10.3897 / zookeys. 761.24214." type="journal article" year="2018">Srisonchai et al. 2018a: 10</bibRefCitation>
.
</paragraph>
<paragraph id="8BAF36D2FFE9AE5CFF35FD3CFDEBA9D0" blockId="3.[136,737,665,977]" pageId="3" pageNumber="4">
<emphasis id="B964EAC0FFE9AE5CFF35FD3CFE1FA990" box="[197,405,665,689]" italics="true" pageId="3" pageNumber="4">Material examined</emphasis>
:
<specimenCount id="9D16FD5BFFE9AE5CFE56FD3CFE78A990" box="[422,498,665,689]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
(IEBR-68)
<collectingRegion id="49D4F830FFE9AE5CFD88FD3CFF61A9F0" country="Vietnam" name="Kon Tum" pageId="3" pageNumber="4">Kon Tum Province</collectingRegion>
, Ngoc Linh Mt., primary forest,
<quantity id="4CE89B37FFE9AE5CFDABFD1CFD32A9F1" box="[603,696,697,721]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="4" unit="m" value="1800.0">1,800 m</quantity>
,
<date id="FFAE1012FFE9AE5CFD36FD1CFEF8A9D0" pageId="3" pageNumber="4" value="2006-03-21" valueMax="2006-04-09" valueMin="2006-03-21">21 March – 9 April 2006</date>
, coll. Anh D. Nguyen.
</paragraph>
<paragraph id="8BAF36D2FFE9AE5CFF35FD5FFC09AA77" blockId="3.[136,737,665,977]" lastBlockId="3.[813,1413,253,469]" pageId="3" pageNumber="4">
<emphasis id="B964EAC0FFE9AE5CFF35FD5FFEB8A830" box="[197,306,762,785]" italics="true" pageId="3" pageNumber="4">Diagnosis</emphasis>
: The species can be recognised by the combination of the following characters: paraterga antler-shaped; metaterga smooth, without traces of setae; the fifth sternum with a prominent, rounded, caudal protuberance between coxae 4; male femora without modifications; epiproct unmodified, without conspicuous setiferous knobs near tip; gonopod femorite long, slightly expanded distad; postfemoral region shortened; solenophore and solenomere extremely short (
<figureCitation id="132B2A57FFE9AE5CFCC5FE98FCFCAA77" box="[821,886,317,342]" captionStart="Fig" captionStartId="3.[137,168,1952,1971]" captionTargetBox="[233,1316,1032,1927]" captionTargetId="figure-311@3.[233,1316,1032,1927]" captionTargetPageId="3" captionText="Fig. 2. Hylomus asper (Attems, 1937) from Ngoc Linh Mt. Right gonopod, lateral view (A), mesal view (B)." figureDoi="http://doi.org/10.5281/zenodo.8065877" httpUri="https://zenodo.org/record/8065877/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BAF36D2FFE9AE5CFC99FEFBFB1DAA97" blockId="3.[813,1413,253,469]" pageId="3" pageNumber="4">
<emphasis id="B964EAC0FFE9AE5CFC99FEFBFBB9AA54" box="[873,1075,349,373]" italics="true" pageId="3" pageNumber="4">Previous records</emphasis>
:
<collectingRegion id="49D4F830FFE9AE5CFBB7FEFBFAAFAA54" box="[1095,1317,349,374]" country="Vietnam" name="Da Nang" pageId="3" pageNumber="4">Da Nang Province</collectingRegion>
(Ba Na National Park; Hai Van pass);
<collectingRegion id="49D4F830FFE9AE5CFB60FED8FA0FAAB4" box="[1168,1413,381,405]" country="Vietnam" name="Ninh Thuan" pageId="3" pageNumber="4">Ninh Thuan Province</collectingRegion>
(Phan Rang) (
<bibRefCitation id="EF814B23FFE9AE5CFC30FE3BFBC1AA97" author="Attems C." box="[960,1099,414,438]" pageId="3" pageNumber="4" pagination="1 - 300" refId="ref11356" refString="Attems C. 1937. Myriapoda 3. Polydesmoidea I. Fam. Strongylosomidae. Das Tierreich 68: 1 - 300." type="journal article" year="1937">Attems 1937</bibRefCitation>
<bibRefCitation id="EF814B23FFE9AE5CFBA2FE3BFB01AA97" author="Attems C." box="[1106,1163,414,438]" pageId="3" pageNumber="4" pagination="187 - 353" refId="ref11379" refString="Attems C. 1938. Die von Dr. C. Dawydoff in franzosisch-Indochina gesammelten Myriopoden. Memoires du Museum National d'Histoire Naturelle, Paris {N. S.} 6 (2): 187 - 353." type="journal article" year="1938">1938</bibRefCitation>
).
</paragraph>
<paragraph id="8BAF36D2FFE9AE5CFC99FE18FAD4AAF4" blockId="3.[813,1413,253,469]" box="[873,1374,445,469]" pageId="3" pageNumber="4">
<emphasis id="B964EAC0FFE9AE5CFC99FE18FC61AAF4" box="[873,1003,445,469]" italics="true" pageId="3" pageNumber="4">Distribution</emphasis>
: Southern part of central
<collectingCountry id="F3077642FFE9AE5CFB0CFE18FAD0AAF4" box="[1276,1370,445,469]" name="Vietnam" pageId="3" pageNumber="4">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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