treatments-xml/data/03/B9/87/03B987AEFF92FFF70AE1FA27FB06FEFC.xml

<document id="A964CAF09363C4A49FA3AF349A8E54C4" ID-DOI="10.11646/zootaxa.3633.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5261931" ID-ZooBank="60F0956C-A1F5-44CA-9D14-8C04343AD01F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629849986103" checkinUser="felipe" docAuthor="Tedesco, Anazélia M. &amp; Aguiar, Alexandre P." docDate="2013" docId="03B987AEFF92FFF70AE1FA27FB06FEFC" docLanguage="en" docName="zt03633p138.pdf" docOrigin="Zootaxa 3633 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3633.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Toechorychus melgassus Tedesco &amp; Aguiar 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="53" masterDocId="FF80FFD6FFA0FFC30A76FFC1FF88FFE0" masterDocTitle="&lt;strong&gt; Phylogeny and Revision of &lt;em&gt; Toechorychus &lt;/ em&gt; Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species &lt;/ strong&gt;" masterLastPageNumber="138" masterPageNumber="1" pageNumber="51" updateTime="1699120729359" updateUser="plazi">
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<mods:title id="15650F9982354B9558CF4B5FBF547248">&lt;strong&gt; Phylogeny and Revision of &lt;em&gt; Toechorychus &lt;/ em&gt; Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species &lt;/ strong&gt;</mods:title>
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<mods:namePart id="86E0450302C241BD42C1B9CF0D0C120A">Tedesco, Anazélia M.</mods:namePart>
<mods:affiliation id="CB38627666752087855EC6B501B4379B">Universidade Federal do Espírito Santo, Departamento de Ciências Biológicas, Zoologia, Av. Fernando Ferrari 514 - Goiabeiras,</mods:affiliation>
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<mods:namePart id="273531E47E70B07D8C1F12B8E8C18442">Aguiar, Alexandre P.</mods:namePart>
<mods:affiliation id="F50245F21629E5831259BED92D549578">Universidade Federal do Espírito Santo, Departamento de Ciências Biológicas, Zoologia, Av. Fernando Ferrari 514 - Goiabeiras,</mods:affiliation>
<mods:nameIdentifier id="A4D7549C5B7DB731863EFEBEB8501DDD" type="email">aguiar.2@osu.edu</mods:nameIdentifier>
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<taxonomicName id="4C104D3BFF92FFF10AE1FA27FDA5F9E0" authority="Tedesco" authorityName="Tedesco &amp; Aguiar" authorityYear="2013" box="[151,557,1510,1536]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="melgassus" status="sp. nov.">
<emphasis id="B964EAAAFF92FFF10AE1FA27FE49F9E0" bold="true" box="[151,449,1510,1536]" italics="true" pageId="50" pageNumber="51">Toechorychus melgassus</emphasis>
Tedesco
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, 
<taxonomicNameLabel id="A25757D1FF92FFF1084CFA27FD10F9E0" box="[570,664,1510,1536]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BAF36B8FF92FFF10AE1F9C8FDA9F9C1" blockId="50.[151,664,1510,1569]" box="[151,545,1545,1569]" pageId="50" pageNumber="51">
(
<figureCitation id="132B2A3DFF92FFF10AE9F9C8FF7DF9C1" box="[159,245,1545,1569]" captionStart="FIGURES 36–39" captionStartId="111.[151,264,1919,1942]" captionTargetBox="[151,1435,193,1898]" captionTargetId="figure-65@111.[151,1435,193,1898]" captionTargetPageId="111" captionText="FIGURES 36–39. Lateral pictures of Toechorychus. 36, T. marcapatus sp. nov., holotype. 37, T. melgassus sp. nov., holotype. 38, T. paramaribus sp. nov., holotype. 39, T. callangus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261965" httpUri="https://zenodo.org/record/5261965/files/figure.png" pageId="50" pageNumber="51">Figs 37</figureCitation>
, 
<figureCitation id="132B2A3DFF92FFF10B77F9C8FEA8F9C1" box="[257,288,1545,1569]" captionStart="FIGURES 45–54. 45" captionStartId="113.[151,264,1758,1781]" captionTargetBox="[151,1435,240,1737]" captionTargetId="figure-226@113.[151,1435,179,1737]" captionTargetPageId="113" captionText="FIGURES 45–54. 45, T. callangus sp. nov., holotype, ovipositor in dorsal view, showing dorsal valve with posterior V-shaped carina, a new synapomorphy discovered for the genus. 46, T. vinhaticus sp. nov., holotype, pronotum in lateral view, lateral portion of collar dorsally carinated. 47, T. morelus sp. nov., holotype, pronotum in lateral view, to show lateral portion of collar dorsally swollen. 48, T. fluminensis sp. nov., holotype, to show hind margin of metanotum without teeth or carinae. 49, T. jatainus sp. nov., holotype, to show hind margin of metanotum with two lateral carinae. 50, T. amapaeus sp. nov., holotype, head in posterior view, to show occipital carina dorsally absent. 51, T. paramaribus sp. nov., holotype, to show anterior margin of propodeum with two lateral teeth. 52, T. melgassus sp. nov., holotype, head in posterior view, to show occipital carina dorsally conspicuous. 53, T. zulius sp. nov., holotype, to show first tergite with two basolateral teeth. 54, T. nourageus sp. nov., holotype, to show first tergite without basolateral teeth." figureDoi="http://doi.org/10.5281/zenodo.5261973" httpUri="https://zenodo.org/record/5261973/files/figure.png" pageId="50" pageNumber="51">52</figureCitation>
, 
<figureCitation id="132B2A3DFF92FFF10B5DF9C8FEC1F9C1" box="[299,329,1545,1569]" captionStart="FIGURES 55–69" captionStartId="114.[151,264,1840,1863]" captionTargetBox="[151,1435,193,1819]" captionTargetId="figure-163@114.[151,1435,193,1819]" captionTargetPageId="114" captionText="FIGURES 55–69. Head, frontal view. 55, T. surinamus sp. nov., holotype. 56, T. callangus sp. nov., holotype. 57, T. coaracius sp. nov., holotype. 58, T. guarapuavus sp. nov., holotype. 59, T. itapuensis sp. nov., holotype. 60, T. benius sp. nov., holotype. 61, T. melgassus sp. nov., holotype. 62, T. stramineus Taschenberg. 63, T. tumazulus sp. nov., holotype. 64, T. ondensis sp. nov., holotype. 65, T. fluminensis sp. nov., holotype. 66, T. amapaeus sp. nov., holotype. 67, T. barticus sp. nov., holotype. 68, T. albimaculatus Taschenberg. 69, T. cassunungae Brauns." figureDoi="http://doi.org/10.5281/zenodo.5261977" httpUri="https://zenodo.org/record/5261977/files/figure.png" pageId="50" pageNumber="51">61</figureCitation>
, 
<figureCitation id="132B2A3DFF92FFF10B23F9C8FEFBF9C1" box="[341,371,1545,1569]" captionStart="FIGURES 97–104. 97–98" captionStartId="118.[151,264,1839,1862]" captionTargetBox="[151,1435,193,1818]" captionTargetId="figure-108@118.[151,1435,193,1818]" captionTargetPageId="118" captionText="FIGURES 97–104. 97–98: Mesoscutum, dorsal view. 97, T. guarapuavus sp. nov., holotype. 98, T. melgassus sp. nov., holotype; 99–104: Mesopleuron, lateral view. 99, T. vilhenus sp. nov., holotype. 100, T. taperinus sp. nov., holotype. 101, T. catarinus sp. nov., holotype. 102, T. coaracius sp. nov., holotype. 103, T. morelus sp. nov., holotype. 104, T. albimaculatus Taschenberg." figureDoi="http://doi.org/10.5281/zenodo.5261987" httpUri="https://zenodo.org/record/5261987/files/figure.png" pageId="50" pageNumber="51">98</figureCitation>
, 
<figureCitation id="132B2A3DFF92FFF10B09F9C8FE22F9C1" box="[383,426,1545,1569]" captionStart="FIGURE 105–118" captionStartId="119.[151,250,1884,1907]" captionTargetBox="[164,1428,179,1877]" captionTargetId="figure-155@119.[164,1428,179,1877]" captionTargetPageId="119" captionText="FIGURE 105–118. T. albimaculatus Taschenberg, fore and hind wings. FIGURES 106–118. Fore wing. 106, T. cassunungae Brauns. 107, T. stramineus Taschenberg. 108, T. ondensis sp. nov., holotype. 109, T. amapaeus sp. nov., holotype. 110, T. zulius sp. nov., holotype. 111, T. amazonensis sp. nov., holotype. 112, T. itapuensis sp. nov., holotype. 113, T. paramaribus sp. nov., holotype. 114, T. guarapuavus sp. nov., holotype. 115, T. melgassus sp. nov., holotype. 116, T. coaracius sp. nov., holotype. 117, T. marcapatus sp. nov., holotype. 118, T. barticus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261991" httpUri="https://zenodo.org/record/5261991/files/figure.png" pageId="50" pageNumber="51">115</figureCitation>
, 
<figureCitation id="132B2A3DFF92FFF10BC0F9C8FE6AF9C1" box="[438,482,1545,1569]" captionStart="FIGURES 127–132" captionStartId="121.[151,264,1671,1694]" captionTargetBox="[151,1435,193,1650]" captionTargetId="figure-84@121.[151,1435,193,1650]" captionTargetPageId="121" captionText="FIGURES 127–132. Propodeum, dorsal view. 127, T. napus sp. nov., holotype. 128, T. paramaribus sp. nov., holotype. 129, T. callangus sp. nov., holotype. 130, T. melgassus sp. nov., holotype. 131, T. marcapatus sp. nov., holotype. 132, T. catarinus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261997" httpUri="https://zenodo.org/record/5261997/files/figure.png" pageId="50" pageNumber="51">130</figureCitation>
, 
<figureCitation id="132B2A3DFF92FFF10B98F9C8FD91F9C1" box="[494,537,1545,1569]" captionStart="FIGURES 175–186" captionStartId="128.[151,264,1514,1537]" captionTargetBox="[151,1435,193,1493]" captionTargetId="figure-150@128.[151,1435,193,1493]" captionTargetPageId="128" captionText="FIGURES 175–186. Hind coxae and metasoma color patterns, dorsal view. 175, T. paramaribus sp. nov., holotype. 176, T. nourageus sp. nov., holotype. 177, T. callangus sp. nov., holotype. 178, T. melgassus sp. nov., holotype. 179, T. guarapuavus sp. nov., holotype. 180, T. marcapatus sp. nov., holotype. 181, T. vinhaticus sp. nov., holotype. 182, T. coaracius sp. nov., holotype. 183, T. amazonensis sp. nov., holotype. 184, T. sinopus sp. nov., holotype. 185, T. surinamus sp. nov., holotype. 186, T. bombuscarus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5262015" httpUri="https://zenodo.org/record/5262015/files/figure.png" pageId="50" pageNumber="51">178</figureCitation>
)
</paragraph>
<paragraph id="8BAF36B8FF92FFF10AE1F990FCCEF989" blockId="50.[151,1436,1617,2037]" box="[151,838,1617,1641]" pageId="50" pageNumber="51">
Description. 
<typeStatus id="54AB881AFF92FFF10B51F990FE07F989" box="[295,399,1617,1641]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
FEMALE. Fore wing length 
<quantity id="4CE89B5DFF92FFF108ADF990FCCAF989" box="[731,834,1617,1641]" metricMagnitude="-3" metricUnit="m" metricValue="6.32" pageId="50" pageNumber="51" unit="mm" value="6.32">6.32 mm</quantity>
.
</paragraph>
<paragraph id="8BAF36B8FF92FFF10AB1F9B4FD17F815" blockId="50.[151,1436,1617,2037]" pageId="50" pageNumber="51">
<emphasis id="B964EAAAFF92FFF10AB1F9B4FE8CF96D" box="[199,260,1653,1677]" italics="true" pageId="50" pageNumber="51">Head</emphasis>
. Mandible 1.39 as long as basal width, densely pilose; ventral margin projected as crest; dorsal tooth distinctly longer than ventral tooth. Clypeus 1.71 as wide as high, subrectangular, minutely strigulate, apically smooth; apex 1.37 as long as base, truncate; apical margin blunt, medially straight. Supraclypeal area entirely densely punctate, densely pilose, medially slightly prominent; between antennal foramens with U-shaped carina; radicle punctate. Antenna with 24 flagellomeres; white band starting at flagellomere 4, reaching flagellomere 12; flagellum slender; subapical flagellomeres slightly flattened. Supra-antennal area without distinct median carina, medially without longitudinal elevation, near antennal sockets not striate; dorsal half medially smooth. Paraocular area smooth. Vertex strigulate between ocelli, without sulcus; gena and vertex behind ocelli smooth, sparsely pilose; gena in lateral view uniformly wide; occipital carina stout, dorsally uniformly arched, ventrally slightly projected as crest, reaching hypostomal carina far from mandible base; hypostomal carina projected as crest; malar space 1.03 as long as basal width of mandible.
</paragraph>
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<taxonomicName id="4C104D3BFF93FFF00AB1FF58FEB7FF50" box="[199,319,153,176]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAAFF93FFF00AB1FF58FEB7FF50" box="[199,319,153,176]" italics="true" pageId="51" pageNumber="52">Mesosoma</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C104D3BFF93FFF00B3BFF59FE4FFF4F" box="[333,455,152,175]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="51" pageNumber="75" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
in dorsal view 2.12 as long as wide; in lateral view, mesosoma middle width 
<quantity id="4CE89B5DFF93FFF00F47FF56FA10FF4F" box="[1329,1432,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="51" pageNumber="52" unit="mm" value="0.19">0.19 mm</quantity>
. Pronotum centrally smooth, latero-ventrally rugose behind collar, margin near mesopleuron ventrally corrugated; pronotal swelling strigulate; collar dorso-laterally carinated, not swollen, anteriorly punctulate; epomia absent. Mesoscutum sparsely pilose, densely punctate, impression of notaulus corrugated, central lobe without longitudinal sulcus or carina; notauli faintly impressed, posteriorly parallel throughout. Scutoscutellar groove smooth, polished; scutellum 1.3 as long as wide, scarcely punctate; scutellar carina incomplete, restricted approximately to anterior 0.6; postscutellum 0.34 as long as wide; hind margin of metanotum with two lateral teeth. Subalar ridge somewhat rounded; mesopleuron dorsally medially markedly strigate, ventrally strigulate; mesopleural groove corrugated; epicnemial carina stout, dorsally curving forwards; dorsal end of epicnemial carina reaching 0.1 of posterior margin of pronotum, behind epicnemial carina without rounded confused-rugulose area; sternaulus medially shallow, interrupted, reaching base of mid coxa, markedly sinuate, moderately impressed, faint from anterior 0.5 to 0.7, not corrugated; scrobe moderately deep, forming pit with sulcus. Mesothoracic venter punctate; median portion of posterior transverse carina of mesothoracic venter V-shaped, regularly shaped, without projections. Transverse furrow at base of propodeum laterally 0.09 as long as anterior portion of propodeum, shallow and narrow, medially wide, uniformly corrugated; propodeum 1.33 as long as wide medially; anterior margin of propodeum without teeth; anterior portion of propodeum strigulate, medially without longitudinal carinae; lateral longitudinal carina of propodeum indistinct; posterior portion of propodeum strigate, progressively coarse toward posterior margin, medially without distinctly stout longitudinal carina; anterior transverse carina of propodeum complete, stout, medially slightly arched forwards; posterior transverse carina absent; propodeal spiracle 1.71 as long as wide; pleural carina of propodeum absent; lower division of metapleuron strigulate, densely pilose, juxtacoxal carina short and faint.
</paragraph>
<paragraph id="8BAF36B8FF93FFF00AB1FC4CFCB6FBD4" blockId="51.[151,1437,151,1976]" pageId="51" pageNumber="52">
<emphasis id="B964EAAAFF93FFF00AB1FC4CFE83FC44" box="[199,267,909,932]" italics="true" pageId="51" pageNumber="52">Wings</emphasis>
. Fore wing: vein 2+3Rs almost straight; ramellus absent; vein 1M+Rs entirely sinuous; vein 1cu-a posteriorly slightly curved, arising far from vein 1M+Rs base, angle with vein M+Cu distinctly obtuse; vein 2Cu 0.48 as long as vein 2cu-a; vein 4Rs 0.99 as long as vein 4M, straight; cell 1+2Rs 0.74 as high as pterostigma, pentagonal; vein 2M distinctly shorter than vein 3M; vein 3M indistinct. Hind wing: vein 1Cu 1.44 as long as vein cu-a; vein 2-1A reaching 0.8 of distance to posterior margin.
</paragraph>
<paragraph id="8BAF36B8FF93FFF00AB1FB80FCC2FBB7" blockId="51.[151,1437,151,1976]" box="[199,842,1087,1112]" pageId="51" pageNumber="52">
<emphasis id="B964EAAAFF93FFF00AB1FB80FF74FBB8" box="[199,252,1089,1112]" italics="true" pageId="51" pageNumber="52">Legs</emphasis>
. Tibia with sparse short bristles; hind coxa elongate.
</paragraph>
<paragraph id="8BAF36B8FF93FFF00AB1FBA4FEBDFAEB" blockId="51.[151,1437,151,1976]" pageId="51" pageNumber="52">
<emphasis id="B964EAAAFF93FFF00AB1FBA4FEB4FB9C" box="[199,316,1125,1148]" italics="true" pageId="51" pageNumber="52">Metasoma</emphasis>
. T1 0.47 as long as hind femur, without basolateral tooth; anteriorly smooth, posteriorly coriarious; spiracle at anterior 0.53, not prominent; postpetiole dorsally, at level of spiracles, faintly but distinctly concave; T2–8 minutely coriarious; centrally almost glabrous, laterally pilose; T2 1.09 as long as wide at apex; apex of T2 2.37 as wide as base. Dorsal valve of ovipositor with nodus indistinct; apex of lower valve with series of teeth; notch present.
</paragraph>
<paragraph id="8BAF36B8FF93FFF00AB1FAD9FB79F9CC" blockId="51.[151,1437,151,1976]" pageId="51" pageNumber="52">
<emphasis id="B964EAAAFF93FFF00AB1FAD9FE8FFAD0" box="[199,263,1304,1328]" italics="true" pageId="51" pageNumber="52">Color</emphasis>
. Head and metasoma mostly white and black, mesosoma orange. Head: whitish; mandible teeth, supraantennal area only behind scape, interocellar area, vertex and gena, except complete large orbital band, and occiput, dark brown, blackish posteriorly; scape ventrally yellow, scape dorsally, pedicel, and flagellum ventrally, brown; flagellum dorsally completely whitish at f4–10, f3 and f11 partially white. 
<taxonomicName id="4C104D3BFF93FFF00E67FA45FB07FA7B" box="[1041,1167,1412,1435]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="51" pageNumber="75" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
: orange (207,091,021); propleuron, subalar ridge, epicnecium, mesothoracic venter, mesepimeron, and apex of juxtacoxal area, whitish (222,176,094). Legs: orange (207,091,021), yellowish towards apex; all t5 dark brown. Metasoma: T1 orange, both basal and apical 0.1 white; T2 basally orange, blackish toward apex, posterior 0.3 white; T2 basally black, posterior 0.4 white; T3–8 basally black, lateral and posterior 0.4 white; S2–8 white, with lateral brown mark.
</paragraph>
<paragraph id="8BAF36B8FF93FFF00AB1F9F6FAF8F994" blockId="51.[151,1437,151,1976]" pageId="51" pageNumber="52">
<emphasis id="B964EAAAFF93FFF00AB1F9F6FEBFF9B0" bold="true" box="[199,311,1591,1616]" pageId="51" pageNumber="52">Variation</emphasis>
. Fore wing of same length on both females. Antenna with 23 flagellomeres. Supra-antennal area medially orange; around ocelli lighter. T2 orange only at anterior 0.1. Scutoscutellar groove faintly corrugated.
</paragraph>
</subSubSection>
<subSubSection id="C30A6533FF93FFF00AB1F9BEFE00F8AC" pageId="51" pageNumber="52" type="discussion">
<paragraph id="8BAF36B8FF93FFF00AB1F9BEFBFBF8C7" blockId="51.[151,1437,151,1976]" pageId="51" pageNumber="52">
<emphasis id="B964EAAAFF93FFF00AB1F9BEFEC0F978" bold="true" box="[199,328,1663,1688]" pageId="51" pageNumber="52">Comments</emphasis>
. This is the only known species with mesosoma and legs almost entirely orange, except by small white areas (
<figureCitation id="132B2A3DFF93FFF00B50F965FEF3F95C" box="[294,379,1700,1724]" captionStart="FIGURES 36–39" captionStartId="111.[151,264,1919,1942]" captionTargetBox="[151,1435,193,1898]" captionTargetId="figure-65@111.[151,1435,193,1898]" captionTargetPageId="111" captionText="FIGURES 36–39. Lateral pictures of Toechorychus. 36, T. marcapatus sp. nov., holotype. 37, T. melgassus sp. nov., holotype. 38, T. paramaribus sp. nov., holotype. 39, T. callangus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261965" httpUri="https://zenodo.org/record/5261965/files/figure.png" pageId="51" pageNumber="52">Fig. 37</figureCitation>
). Very similar to 
<taxonomicName id="4C104D3BFF93FFF00836F964FD57F95C" authority="Tedesco &amp; Aguiar, 2013" authorityName="Tedesco &amp; Aguiar" authorityYear="2013" box="[576,735,1700,1724]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="marcapatus" status="sp. nov.">
<emphasis id="B964EAAAFF93FFF00836F964FDC5F95C" box="[576,589,1701,1724]" italics="true" pageId="51" pageNumber="52">T</emphasis>
. 
<emphasis id="B964EAAAFF93FFF0082AF964FD57F95C" box="[604,735,1701,1724]" italics="true" pageId="51" pageNumber="52">marcapatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757D1FF93FFF00891F965FCC9F95B" box="[743,833,1700,1724]" pageId="51" pageNumber="52" rank="species">
<emphasis id="B964EAAAFF93FFF00891F965FC89F95C" bold="true" box="[743,769,1700,1724]" pageId="51" pageNumber="52">sp</emphasis>
. 
<emphasis id="B964EAAAFF93FFF00967F965FCB3F95C" bold="true" box="[785,827,1700,1724]" pageId="51" pageNumber="52">nov</emphasis>
.
</taxonomicNameLabel>
from which it can be separated by having mesosoma completely orange (
<emphasis id="B964EAAAFF93FFF00B0BF908FE1CF900" box="[381,404,1737,1760]" italics="true" pageId="51" pageNumber="52">vs</emphasis>
. pronotum, mesoscutum and mesopleuron marked with black areas in 
<taxonomicName id="4C104D3BFF93FFF00EB8F908FAF8F900" authority="Tedesco &amp; Aguiar, 2013" authorityName="Tedesco &amp; Aguiar" authorityYear="2013" box="[1230,1392,1736,1760]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="marcapatus" status="sp. nov.">
<emphasis id="B964EAAAFF93FFF00EB8F908FB53F900" box="[1230,1243,1737,1760]" italics="true" pageId="51" pageNumber="52">T</emphasis>
. 
<emphasis id="B964EAAAFF93FFF00E9BF908FAF8F900" box="[1261,1392,1737,1760]" italics="true" pageId="51" pageNumber="52">marcapatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757D1FF93FFF00F0CF909FF4DF8E4" pageId="51" pageNumber="52" rank="species">
<emphasis id="B964EAAAFF93FFF00F0CF909FA1CF900" bold="true" box="[1402,1428,1736,1760]" pageId="51" pageNumber="52">sp</emphasis>
. 
<emphasis id="B964EAAAFF93FFF00AE1F92DFF49F8E4" bold="true" box="[151,193,1772,1796]" pageId="51" pageNumber="52">nov</emphasis>
.
</taxonomicNameLabel>
); lateral portion of collar carinated, not swollen, as in 
<figureCitation id="132B2A3DFF93FFF00941F92DFC00F8E4" box="[823,904,1772,1796]" captionStart="FIGURES 45–54. 45" captionStartId="113.[151,264,1758,1781]" captionTargetBox="[151,1435,240,1737]" captionTargetId="figure-226@113.[151,1435,179,1737]" captionTargetPageId="113" captionText="FIGURES 45–54. 45, T. callangus sp. nov., holotype, ovipositor in dorsal view, showing dorsal valve with posterior V-shaped carina, a new synapomorphy discovered for the genus. 46, T. vinhaticus sp. nov., holotype, pronotum in lateral view, lateral portion of collar dorsally carinated. 47, T. morelus sp. nov., holotype, pronotum in lateral view, to show lateral portion of collar dorsally swollen. 48, T. fluminensis sp. nov., holotype, to show hind margin of metanotum without teeth or carinae. 49, T. jatainus sp. nov., holotype, to show hind margin of metanotum with two lateral carinae. 50, T. amapaeus sp. nov., holotype, head in posterior view, to show occipital carina dorsally absent. 51, T. paramaribus sp. nov., holotype, to show anterior margin of propodeum with two lateral teeth. 52, T. melgassus sp. nov., holotype, head in posterior view, to show occipital carina dorsally conspicuous. 53, T. zulius sp. nov., holotype, to show first tergite with two basolateral teeth. 54, T. nourageus sp. nov., holotype, to show first tergite without basolateral teeth." figureDoi="http://doi.org/10.5281/zenodo.5261973" httpUri="https://zenodo.org/record/5261973/files/figure.png" pageId="51" pageNumber="52">Fig. 46</figureCitation>
(
<emphasis id="B964EAAAFF93FFF009ECF92CFC39F8E4" box="[922,945,1773,1796]" italics="true" pageId="51" pageNumber="52">vs</emphasis>
. dorsally rounded, distinctly swollen, as in 
<figureCitation id="132B2A3DFF93FFF00AE1F8CEFF62F8C7" box="[151,234,1807,1831]" captionStart="FIGURES 45–54. 45" captionStartId="113.[151,264,1758,1781]" captionTargetBox="[151,1435,240,1737]" captionTargetId="figure-226@113.[151,1435,179,1737]" captionTargetPageId="113" captionText="FIGURES 45–54. 45, T. callangus sp. nov., holotype, ovipositor in dorsal view, showing dorsal valve with posterior V-shaped carina, a new synapomorphy discovered for the genus. 46, T. vinhaticus sp. nov., holotype, pronotum in lateral view, lateral portion of collar dorsally carinated. 47, T. morelus sp. nov., holotype, pronotum in lateral view, to show lateral portion of collar dorsally swollen. 48, T. fluminensis sp. nov., holotype, to show hind margin of metanotum without teeth or carinae. 49, T. jatainus sp. nov., holotype, to show hind margin of metanotum with two lateral carinae. 50, T. amapaeus sp. nov., holotype, head in posterior view, to show occipital carina dorsally absent. 51, T. paramaribus sp. nov., holotype, to show anterior margin of propodeum with two lateral teeth. 52, T. melgassus sp. nov., holotype, head in posterior view, to show occipital carina dorsally conspicuous. 53, T. zulius sp. nov., holotype, to show first tergite with two basolateral teeth. 54, T. nourageus sp. nov., holotype, to show first tergite without basolateral teeth." figureDoi="http://doi.org/10.5281/zenodo.5261973" httpUri="https://zenodo.org/record/5261973/files/figure.png" pageId="51" pageNumber="52">Fig. 47</figureCitation>
); pleural carina of propodeum present only posteriorly (
<emphasis id="B964EAAAFF93FFF0092BF8D0FCFCF8C8" box="[861,884,1809,1832]" italics="true" pageId="51" pageNumber="52">vs</emphasis>
. absent, as in 
<figureCitation id="132B2A3DFF93FFF00E62F8CEFBEFF8C7" box="[1044,1127,1807,1831]" captionStart="FIGURES 41–44. 41" captionStartId="112.[151,264,1842,1865]" captionTargetBox="[151,1435,851,1821]" captionTargetId="figure-109@112.[151,1435,851,1821]" captionTargetPageId="112" captionText="FIGURES 41–44. 41, T. nourageus sp. nov., holotype, propodeum in dorsal view, pleural carina complete. 42, T. marcapatus sp. nov., holotype, propodeum in lateral view, pleural carina absent. 43, T. surinamus sp. nov., holotype, mesopleuron in lateral view, sternaulus incomplete. 44, T. kawus sp. nov., holotype, mesopleuron in lateral view, sternaulus complete, reaching base of mid coxa." figureDoi="http://doi.org/10.5281/zenodo.5261971" httpUri="https://zenodo.org/record/5261971/files/figure.png" pageId="51" pageNumber="52">Fig. 42</figureCitation>
).
</paragraph>
<paragraph id="8BAF36B8FF93FFF00AB1F8F5FE00F8AC" blockId="51.[151,1437,151,1976]" box="[199,392,1844,1868]" pageId="51" pageNumber="52">
<emphasis id="B964EAAAFF93FFF00AB1F8F5FE8CF8AC" bold="true" box="[199,260,1844,1868]" pageId="51" pageNumber="52">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30A6533FF93FFF00AB1F896FE64F874" pageId="51" pageNumber="52" type="distribution">
<paragraph id="8BAF36B8FF93FFF00AB1F896FE64F874" blockId="51.[151,1437,151,1976]" pageId="51" pageNumber="52">
<emphasis id="B964EAAAFF93FFF00AB1F896FED1F890" bold="true" box="[199,345,1879,1904]" pageId="51" pageNumber="52">Distribution</emphasis>
. 
<collectingCountry id="F3077628FF93FFF00B11F896FE27F88F" box="[359,431,1879,1903]" name="Brazil" pageId="51" pageNumber="52">Brazil</collectingCountry>
. Known only from the Floresta Nacional de Caxiuanã (
<geoCoordinate id="EE24507FFF93FFF00E6EF896FB08F88F" box="[1048,1152,1879,1903]" degrees="2" direction="south" minutes="4" orientation="latitude" pageId="51" pageNumber="52" precision="15" seconds="54" value="-2.0816667">2°4’54”S</geoCoordinate>
<geoCoordinate id="EE24507FFF93FFF00EF1F896FA83F88F" box="[1159,1291,1879,1903]" degrees="51" direction="west" minutes="51" orientation="longitude" pageId="51" pageNumber="52" precision="15" seconds="5" value="-51.851387">51°51’5”W</geoCoordinate>
), 
<emphasis id="B964EAAAFF93FFF00F56F899FAD0F890" box="[1312,1368,1880,1904]" italics="true" pageId="51" pageNumber="52">Pará</emphasis>
, from two close localities (
<figureCitation id="132B2A3DFF93FFF00B0BF8BDFE68F874" box="[381,480,1916,1940]" captionStart="FIGURES 208–215" captionStartId="131.[151,264,1944,1967]" captionTargetBox="[172,1416,185,1935]" captionTargetId="figure-102@131.[172,1416,181,1935]" captionTargetPageId="131" captionText="FIGURES 208–215. Distribution records. 208, T. amazonensis sp. nov., holotype. 209, T. itapuensis sp. nov., holotype. 210, T. paramaribus sp. nov., holotype. 211, T. guarapuavus sp. nov., holotype. 212, T. melgassus sp. nov., holotype. 213, T. coaracius sp. nov., holotype. 214, T. vinhaticus sp. nov., holotype. 215, T. morelus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5262025" httpUri="https://zenodo.org/record/5262025/files/figure.png" pageId="51" pageNumber="52">Fig. 212</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A6533FF93FFF00AB1F85EFE2BF858" box="[199,419,1951,1976]" pageId="51" pageNumber="52" type="biology_ecology">
<paragraph id="8BAF36B8FF93FFF00AB1F85EFE2BF858" blockId="51.[151,1437,151,1976]" box="[199,419,1951,1976]" pageId="51" pageNumber="52">
<emphasis id="B964EAAAFF93FFF00AB1F85EFE97F858" bold="true" box="[199,287,1951,1976]" pageId="51" pageNumber="52">Biology</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30A6533FF94FFF70AB1FF56FB06FEFC" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="8BAF36B8FF94FFF70AB1FF56FB06FEFC" blockId="52.[151,1436,151,284]" pageId="52" pageNumber="53">
<emphasis id="B964EAAAFF94FFF70AB1FF56FE21FF50" bold="true" box="[199,425,151,176]" pageId="52" pageNumber="53">Material examined</emphasis>
. 
<materialsCitation id="3B783CE5FF94FFF70BC1FF56FF52FF17" accessionNumber="P05020" collectingDate="2003-11-14" collectingDateMax="2003-11-17" collectingDateMin="2003-11-14" collectionCode="UFES" collectorName="A. P. Aguiar &amp; J. Dias &amp; Complete" country="Brazil" county="Trilha do Igarape Curua" location="Para" municipality="Floresta Nacional de Caxiuana" pageId="52" pageNumber="53" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
<specimenCount id="9D16FD31FF94FFF70BC1FF56FDAFFF4F" box="[439,551,151,175]" pageId="52" pageNumber="53" type="female">2 females</specimenCount>
. 
<typeStatus id="54AB881AFF94FFF70847FF56FD11FF4F" box="[561,665,151,175]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
<specimenCount id="9D16FD31FF94FFF708D6FF59FD3DFF4F" box="[672,693,152,175]" pageId="52" pageNumber="53" type="female">♀</specimenCount>
, 
<emphasis id="B964EAAAFF94FFF708B5FF56FCA6FF50" bold="true" box="[707,814,151,176]" pageId="52" pageNumber="53">
<collectingCountry id="F3077628FF94FFF708B5FF56FCA6FF50" box="[707,814,151,176]" name="Brazil" pageId="52" pageNumber="53">BRAZIL</collectingCountry>
</emphasis>
: 
<emphasis id="B964EAAAFF94FFF7094AFF59FCFCFF50" box="[828,884,152,176]" italics="true" pageId="52" pageNumber="53">
<location id="8ECF6063FF94FFF7094AFF59FCFCFF50" LSID="urn:lsid:plazi:treatment:03B987AEFF92FFF70AE1FA27FB06FEFC:8ECF6063FF94FFF7094AFF59FCFCFF50" box="[828,884,152,176]" country="Brazil" county="Trilha do Igarape Curua" municipality="Floresta Nacional de Caxiuana" name="Para" pageId="52" pageNumber="53">Pará</location>
</emphasis>
, 
<location id="8ECF6063FF94FFF709F4FF56FC6FFF4F" LSID="urn:lsid:plazi:treatment:03B987AEFF92FFF70AE1FA27FB06FEFC:8ECF6063FF94FFF709F4FF56FC6FFF4F" box="[898,999,151,175]" country="Brazil" county="Trilha do Igarape Curua" municipality="Floresta Nacional de Caxiuana" name="Melgaco" pageId="52" pageNumber="53">Melgaço</location>
, 
<collectingMunicipality id="6BCBACC2FF94FFF70984FF56FAC5FF4F" box="[1010,1357,151,175]" pageId="52" pageNumber="53">Floresta Nacional de Caxiuanã</collectingMunicipality>
, 
<collectingCounty id="62CE4E34FF94FFF70F2EFF56FED3FF34" pageId="52" pageNumber="53">Trilha do Igarapé Curua</collectingCounty>
, 
<date id="FFAE1078FF94FFF70B10FF7DFD9AFF34" box="[358,530,188,212]" pageId="52" pageNumber="53" value="2003-11-14" valueMax="2003-11-17" valueMin="2003-11-14">
<collectingDate id="EFEAE990FF94FFF70B10FF7DFD9AFF34" box="[358,530,188,212]" pageId="52" pageNumber="53" value="2003-11-14" valueMax="2003-11-17" valueMin="2003-11-14">14–17.XI.2003</collectingDate>
</date>
, YPT, 
<collectorName id="26E5536EFF94FFF7082BFF7DFD51FF34" box="[605,729,188,212]" pageId="52" pageNumber="53">A.P.Aguiar</collectorName>
&amp; 
<collectorName id="26E5536EFF94FFF7088BFF7DFCCAFF34" box="[765,834,188,212]" pageId="52" pageNumber="53">J.Dias</collectorName>
<emphasis id="B964EAAAFF94FFF7093FFF7DFCE3FF34" box="[841,875,188,212]" italics="true" pageId="52" pageNumber="53">leg</emphasis>
., P05020 [field point] (
<collectionCode id="ED01AE7DFF94FFF70E06FF7DFB32FF34" box="[1136,1210,188,212]" country="Brazil" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="52" pageNumber="53">UFES</collectionCode>
). 
<collectorName id="26E5536EFF94FFF70EBBFF7DFAB5FF34" box="[1229,1341,188,212]" pageId="52" pageNumber="53">Complete</collectorName>
, in good shape
</materialsCitation>
. 
<materialsCitation id="3B783CE5FF94FFF70A98FF21FB02FEFC" accessionNumber="P05038" collectingDate="2003-11-15" collectingDateMax="2003-11-18" collectingDateMin="2003-11-15" collectionCode="UFES" collectorName="A. P. Aguiar &amp; J. Dias" country="Brazil" county="Trilha do Igarape Tijucaquara" location="Para" municipality="Floresta Nacional de Caxiuana" pageId="52" pageNumber="53" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54AB881AFF94FFF70A98FF21FEDCFF18" box="[238,340,224,248]" pageId="52" pageNumber="53" type="paratype">Paratype</typeStatus>
: 
<emphasis id="B964EAAAFF94FFF70B1EFF1EFE5BFF18" bold="true" box="[360,467,223,248]" pageId="52" pageNumber="53">
<collectingCountry id="F3077628FF94FFF70B1EFF1EFE5BFF18" box="[360,467,223,248]" name="Brazil" pageId="52" pageNumber="53">BRAZIL</collectingCountry>
</emphasis>
: 
<specimenCount id="9D16FD31FF94FFF70B9DFF1EFD96FF17" box="[491,542,223,247]" pageId="52" pageNumber="53" type="female">1 ♀</specimenCount>
from 
<emphasis id="B964EAAAFF94FFF70803FF21FD25FF18" box="[629,685,224,248]" italics="true" pageId="52" pageNumber="53">
<location id="8ECF6063FF94FFF70803FF21FD25FF18" LSID="urn:lsid:plazi:treatment:03B987AEFF92FFF70AE1FA27FB06FEFC:8ECF6063FF94FFF70803FF21FD25FF18" box="[629,685,224,248]" country="Brazil" county="Trilha do Igarape Tijucaquara" municipality="Floresta Nacional de Caxiuana" name="Para" pageId="52" pageNumber="53">Pará</location>
</emphasis>
, 
<location id="8ECF6063FF94FFF708B2FF1EFCA1FF17" LSID="urn:lsid:plazi:treatment:03B987AEFF92FFF70AE1FA27FB06FEFC:8ECF6063FF94FFF708B2FF1EFCA1FF17" box="[708,809,223,247]" country="Brazil" county="Trilha do Igarape Tijucaquara" municipality="Floresta Nacional de Caxiuana" name="Melgaco" pageId="52" pageNumber="53">Melgaço</location>
, 
<collectingMunicipality id="6BCBACC2FF94FFF7094BFF1EFB3BFF17" box="[829,1203,223,247]" pageId="52" pageNumber="53">Floresta Nacional de Caxiuanã</collectingMunicipality>
, 
<collectingCounty id="62CE4E34FF94FFF70EBEFF1EFE97FEFC" pageId="52" pageNumber="53">Trilha do Igarapé Tijucaquara</collectingCounty>
, 
<date id="FFAE1078FF94FFF70B5CFEC5FE5EFEFC" box="[298,470,260,284]" pageId="52" pageNumber="53" value="2003-11-15" valueMax="2003-11-18" valueMin="2003-11-15">
<collectingDate id="EFEAE990FF94FFF70B5CFEC5FE5EFEFC" box="[298,470,260,284]" pageId="52" pageNumber="53" value="2003-11-15" valueMax="2003-11-18" valueMin="2003-11-15">15–18.XI.2003</collectingDate>
</date>
, YPT, 
<collectorName id="26E5536EFF94FFF70854FEC5FD16FEFC" box="[546,670,260,284]" pageId="52" pageNumber="53">A.P.Aguiar</collectorName>
&amp; 
<collectorName id="26E5536EFF94FFF708B4FEC5FC8FFEFC" box="[706,775,260,284]" pageId="52" pageNumber="53">J.Dias</collectorName>
<emphasis id="B964EAAAFF94FFF70978FEC5FCB8FEFC" box="[782,816,260,284]" italics="true" pageId="52" pageNumber="53">leg</emphasis>
., P05038 [field point] (
<collectionCode id="ED01AE7DFF94FFF70E41FEC5FB0AFEFC" box="[1079,1154,260,284]" country="Brazil" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="52" pageNumber="53">UFES</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>