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<mods:title id="A0B0FD8770268D830976498023185BB8">A new species of Scolelepis (Polychaeta: Spionidae) from sandy beaches in China, with a review of Chinese Scolelepis species</mods:title>
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<mods:namePart id="E6B7E01BD4AA4C53C62AB082B284449B">Zhou, Jin</mods:namePart>
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<treatment id="03A43259FF9D2B38CAC6F88AFC413DC3" ID-DOI="http://doi.org/10.5281/zenodo.6219688" ID-GBIF-Taxon="119624208" ID-Zenodo-Dep="6219688" LSID="urn:lsid:plazi:treatment:03A43259FF9D2B38CAC6F88AFC413DC3" httpUri="http://treatment.plazi.org/id/03A43259FF9D2B38CAC6F88AFC413DC3" lastPageId="9" lastPageNumber="46" pageId="8" pageNumber="45">
<subSubSection id="C317D0C4FF9D2B39CAC6F88AFF413E78" pageId="8" pageNumber="45" type="nomenclature">
<paragraph id="8BB2834FFF9D2B39CAC6F88AFD6E3E1F" blockId="8.[151,725,1814,1879]" box="[151,725,1814,1840]" pageId="8" pageNumber="45">
<heading id="D0FA3423FF9D2B39CAC6F88AFD6E3E1F" bold="true" box="[151,725,1814,1840]" fontSize="11" level="1" pageId="8" pageNumber="45" reason="1">
<taxonomicName id="4C0DF8CCFF9D2B39CAC6F88AFD6E3E1F" ID-CoL="4VRND" authority="Imajima, 1992" authorityName="Imajima" authorityYear="1992" box="[151,725,1814,1840]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="45" phylum="Annelida" rank="species" species="variegata" subGenus="Scolelepis">
<emphasis id="B9795F5DFF9D2B39CAC6F88AFD6E3E1F" bold="true" box="[151,725,1814,1840]" pageId="8" pageNumber="45">
<emphasis id="B9795F5DFF9D2B39CAC6F88AFEAB3E1F" bold="true" box="[151,272,1814,1840]" italics="true" pageId="8" pageNumber="45">Scolelepis</emphasis>
(
<emphasis id="B9795F5DFF9D2B39CB71F88AFE233E1F" bold="true" box="[288,408,1814,1840]" italics="true" pageId="8" pageNumber="45">Scolelepis</emphasis>
)
<emphasis id="B9795F5DFF9D2B39CBFBF88AFDA13E1F" bold="true" box="[426,538,1814,1840]" italics="true" pageId="8" pageNumber="45">variegata</emphasis>
<bibRefCitation id="EF9CFEBEFF9D2B39C870F88AFD6E3E1F" author="Imajima" box="[545,725,1814,1840]" pageId="8" pageNumber="48" refString="Imajima, M. (1992) Spionidae (Annelida, Polychaeta) from Japan VIII. The genus Scolelepis. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 18 (1), 1 - 34." type="journal article" year="1992">Imajima, 1992</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB2834FFF9D2B39CAC6F8A1FF413E78" blockId="8.[151,725,1814,1879]" box="[151,250,1853,1879]" pageId="8" pageNumber="45">
<figureCitation id="13369FCAFF9D2B39CAC6F8A1FF413E78" box="[151,250,1853,1879]" captionStart="FIGURE 4" captionStartId="10.[151,256,1872,1896]" captionTargetBox="[193,1381,187,1822]" captionTargetId="figure@10.[183,1404,172,1839]" captionTargetPageId="10" captionText="FIGURE 4. Scolelepis (Scolelepis) variegata Imajima, 1992. A, anterior end, dorsal view; B, parapodia of setiger 1, anterior view; C, parapodia of setiger 2, anterior view; D, parapodia of setiger 31, anterior view; E, notopodial seta from anterior row of setiger 4; F, notopodial seta from posterior row of setiger 4; G, neuropodial seta from anterior row of setiger 4; H, neuropodial seta from posterior row of setiger 4; I, neuropodial hooded hook from setiger 30. Scales: A, 600 μm; B D, 120 μm; E I, 250 μm." httpUri="https://zenodo.org/record/190438/files/figure.png" pageId="8" pageNumber="45">Figure 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C317D0C4FF9D2B39CAC6F816FCDE3E8E" box="[151,869,1930,1953]" pageId="8" pageNumber="45" type="reference_group">
<paragraph id="8BB2834FFF9D2B39CAC6F816FCDE3E8E" blockId="8.[151,869,1930,1953]" box="[151,869,1930,1953]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB1DA461FF9D2B39CAC6F816FCDE3E8E" box="[151,869,1930,1953]" pageId="8" pageNumber="45">
<taxonomicName id="4C0DF8CCFF9D2B39CAC6F816FD7B3E8E" ID-CoL="4VRND" authority="Imajima, 1992: 20" authorityName="Imajima" authorityPageNumber="20" authorityYear="1992" box="[151,704,1930,1953]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="45" phylum="Annelida" rank="species" species="variegata" subGenus="Scolelepis">
<emphasis id="B9795F5DFF9D2B39CAC6F816FEB83E8E" box="[151,259,1930,1953]" italics="true" pageId="8" pageNumber="45">Scolelepis</emphasis>
(
<emphasis id="B9795F5DFF9D2B39CB43F816FEC53E8E" box="[274,382,1930,1953]" italics="true" pageId="8" pageNumber="45">Scolelepis</emphasis>
)
<emphasis id="B9795F5DFF9D2B39CBDFF816FE4F3E8E" box="[398,500,1930,1953]" italics="true" pageId="8" pageNumber="45">variegata</emphasis>
<treatmentCitation id="0AACA55EFF9D2B39CBAAF816FD7B3E8E" author="Imajima" box="[507,704,1930,1953]" page="20" pageId="8" pageNumber="45" year="1992">
<bibRefCitation id="EF9CFEBEFF9D2B39CBAAF816FD213E8E" author="Imajima" box="[507,666,1930,1953]" pageId="8" pageNumber="48" refString="Imajima, M. (1992) Spionidae (Annelida, Polychaeta) from Japan VIII. The genus Scolelepis. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 18 (1), 1 - 34." type="journal article" year="1992">Imajima, 1992</bibRefCitation>
: 20
</treatmentCitation>
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22, figs. 3ah.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C317D0C4FF9D2B38CAC6F84CFE4D38B5" lastPageId="9" lastPageNumber="46" pageId="8" pageNumber="45" type="materials_examined">
<paragraph id="8BB2834FFF9D2B38CAC6F84CFE4D38B5" blockId="8.[151,1435,2000,2027]" lastBlockId="9.[151,1437,151,1260]" lastPageId="9" lastPageNumber="46" pageId="8" pageNumber="45">
<emphasis id="B9795F5DFF9D2B39CAC6F84CFE303EC5" bold="true" box="[151,395,2000,2026]" pageId="8" pageNumber="45">Material examined.</emphasis>
MBMCAS 228678 (1 spm), 122°40΄ E, 31°15΄ N, Changjiang River Estuary, sandy mud with debris,
<quantity id="4CF52EAAFF9C2B38CB7BFF04FED2399D" box="[298,361,152,178]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="46" unit="m" value="33.0">33 m</quantity>
,
<date id="FFB3A58FFF9C2B38CB24FF04FDB2399D" box="[373,521,152,178]" pageId="9" pageNumber="46" value="1986-06-30">30 Jun 1986</date>
; MBMCAS 228909 (1 spm), 122°45΄ E, 31°45΄ N, Changjiang River Estuary, sandy mud,
<quantity id="4CF52EAAFF9C2B38CB7BFF22FED139F7" box="[298,362,190,216]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="46" unit="m" value="37.0">37 m</quantity>
,
<date id="FFB3A58FFF9C2B38CB29FF22FDAE39F7" box="[376,533,190,216]" pageId="9" pageNumber="46" value="1985-12-13">13 Dec 1985</date>
, coll. Xianqiu Ren; MBMCAS 228912 (1 spm), 121°45΄ E, 31°30΄ N, East
<collectingCountry id="F31AC3DFFF9C2B38CAC6FF79FF6639D0" box="[151,221,229,255]" name="China" pageId="9" pageNumber="46">China</collectingCountry>
Sea, sandy mud,
<quantity id="4CF52EAAFF9C2B38CBFFFF79FDB839D0" box="[430,515,229,255]" metricMagnitude="1" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="46" unit="m" value="10.5">10.5 m</quantity>
,
<date id="FFB3A58FFF9C2B38C841FF79FD1D39D0" box="[528,678,229,255]" pageId="9" pageNumber="46" value="1959-10-21">21 Oct 1959</date>
; MBMCAS 228988 (2 spms), 121°15΄ E, 38°00΄ N, Yellow Sea, sandy mud,
<quantity id="4CF52EAAFF9C2B38CB74FE90FED83809" box="[293,355,268,294]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="46" unit="m" value="44.0">44 m</quantity>
,
<date id="FFB3A58FFF9C2B38CB3FFE90FE443809" box="[366,511,268,294]" pageId="9" pageNumber="46" value="1959-01-28">28 Jan 1959</date>
; MBMCAS 228990 (2 spms), 123°00΄ E, 37°00΄ N, Yellow Sea, mud,
<quantity id="4CF52EAAFF9C2B38CF64FE90FAC93809" box="[1333,1394,268,294]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="46" unit="m" value="30.0">30 m</quantity>
,
<date id="FFB3A58FFF9C2B38CF2EFE90FEB73863" pageId="9" pageNumber="46" value="1959-04-18">18 Apr 1959</date>
; MBMCAS 228996 (1 spm), 123°00΄E, 31°45΄N, Changjiang River Estuary, sandy mud,
<quantity id="4CF52EAAFF9C2B38CF11FEAEFA3B3863" box="[1344,1408,306,332]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="46" unit="m" value="38.0">38 m</quantity>
,
<date id="FFB3A58FFF9C2B38CFDCFEAEFEB4385C" pageId="9" pageNumber="46" value="1986-03-04">4 Mar 1986</date>
, coll. Xianqiu Ren; MBMCAS 228997 (1 spm), 122°40΄ E, 31°15΄ N, Changjiang River Estuary, silty mud,
<quantity id="4CF52EAAFF9C2B38CB42FE1CFEE938B5" box="[275,338,384,410]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="46" unit="m" value="37.0">37 m</quantity>
,
<date id="FFB3A58FFF9C2B38CB0FFE1CFE4938B5" box="[350,498,384,410]" pageId="9" pageNumber="46" value="1985-10-15">15 Oct 1985</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C317D0C4FF9C2B38CA97FE3AFD383D04" pageId="9" pageNumber="46" type="description">
<paragraph id="8BB2834FFF9C2B38CA97FE3AFBC33B21" blockId="9.[151,1437,151,1260]" pageId="9" pageNumber="46">
<emphasis id="B9795F5DFF9C2B38CA97FE3AFEE438EF" bold="true" box="[198,351,422,448]" pageId="9" pageNumber="46">Description.</emphasis>
Prostomium elongated, with pointed anterior margin; posterior with rounded swelling just behind 2 pairs of small eyes. Occipital tentacle absent. Two pairs of eyes arranged in a transverse row. Peristomium distinct from setiger 1, forming well-developed lateral wings (
<figureCitation id="13369FCAFF9C2B38CE5CFE68FBEF3B21" box="[1037,1108,500,526]" captionStart="FIGURE 4" captionStartId="10.[151,256,1872,1896]" captionTargetBox="[193,1381,187,1822]" captionTargetId="figure@10.[183,1404,172,1839]" captionTargetPageId="10" captionText="FIGURE 4. Scolelepis (Scolelepis) variegata Imajima, 1992. A, anterior end, dorsal view; B, parapodia of setiger 1, anterior view; C, parapodia of setiger 2, anterior view; D, parapodia of setiger 31, anterior view; E, notopodial seta from anterior row of setiger 4; F, notopodial seta from posterior row of setiger 4; G, neuropodial seta from anterior row of setiger 4; H, neuropodial seta from posterior row of setiger 4; I, neuropodial hooded hook from setiger 30. Scales: A, 600 μm; B D, 120 μm; E I, 250 μm." httpUri="https://zenodo.org/record/190438/files/figure.png" pageId="9" pageNumber="46">Fig. 4</figureCitation>
A).
</paragraph>
<paragraph id="8BB2834FFF9C2B38CA94FD86FA283A45" blockId="9.[151,1437,151,1260]" pageId="9" pageNumber="46">
Parapodia of setiger 1 well developed, with small, oval notopodial and neuropodial postsetal lamellae; capillary setae present on neuropodia, lacking on notopodia (
<figureCitation id="13369FCAFF9C2B38C9D8FDDDFC6E3B74" box="[905,981,577,603]" captionStart="FIGURE 4" captionStartId="10.[151,256,1872,1896]" captionTargetBox="[193,1381,187,1822]" captionTargetId="figure@10.[183,1404,172,1839]" captionTargetPageId="10" captionText="FIGURE 4. Scolelepis (Scolelepis) variegata Imajima, 1992. A, anterior end, dorsal view; B, parapodia of setiger 1, anterior view; C, parapodia of setiger 2, anterior view; D, parapodia of setiger 31, anterior view; E, notopodial seta from anterior row of setiger 4; F, notopodial seta from posterior row of setiger 4; G, neuropodial seta from anterior row of setiger 4; H, neuropodial seta from posterior row of setiger 4; I, neuropodial hooded hook from setiger 30. Scales: A, 600 μm; B D, 120 μm; E I, 250 μm." httpUri="https://zenodo.org/record/190438/files/figure.png" pageId="9" pageNumber="46">Fig. 4</figureCitation>
B). Branchiae present from setiger 2, continuing posteriorly to end of body. Notopodial postsetal lamellae completely fused with branchiae from setiger 2 to setiger 29, thereafter lamellae separated from branchiae, posterior notopodial lamellae triangular with elongated pointed tips (
<figureCitation id="13369FCAFF9C2B38CBB8FD29FD943BE0" box="[489,559,693,719]" captionStart="FIGURE 4" captionStartId="10.[151,256,1872,1896]" captionTargetBox="[193,1381,187,1822]" captionTargetId="figure@10.[183,1404,172,1839]" captionTargetPageId="10" captionText="FIGURE 4. Scolelepis (Scolelepis) variegata Imajima, 1992. A, anterior end, dorsal view; B, parapodia of setiger 1, anterior view; C, parapodia of setiger 2, anterior view; D, parapodia of setiger 31, anterior view; E, notopodial seta from anterior row of setiger 4; F, notopodial seta from posterior row of setiger 4; G, neuropodial seta from anterior row of setiger 4; H, neuropodial seta from posterior row of setiger 4; I, neuropodial hooded hook from setiger 30. Scales: A, 600 μm; B D, 120 μm; E I, 250 μm." httpUri="https://zenodo.org/record/190438/files/figure.png" pageId="9" pageNumber="46">Fig. 4</figureCitation>
CD). Notopodial postsetal lamellae long, narrow with slight serrations in anterior setigers, gradually reduced to subtriangular lamellae in middle and posterior region. Notopodial presetal lamellae rounded in anterior setigers, becoming reduced in posterior setigers. Neuropodial postsetal lamellae rounded in anterior setigers, becoming low and round in posterior setigers. Neuropodial presetal lamellae thick and round, from posterior setigers; smaller than corresponding neuropodial postsetal lamellae.
</paragraph>
<paragraph id="8BB2834FFF9C2B38CA94FCEAFD9B3D2B" blockId="9.[151,1437,151,1260]" pageId="9" pageNumber="46">
All anterior setae capillaries, arranged in two rows on both rami. Anterior row thick, heavily granulated, posterior row thin capillaries without obvious sheaths (
<figureCitation id="13369FCAFF9C2B38C972FC01FCD13A98" box="[803,874,925,951]" captionStart="FIGURE 4" captionStartId="10.[151,256,1872,1896]" captionTargetBox="[193,1381,187,1822]" captionTargetId="figure@10.[183,1404,172,1839]" captionTargetPageId="10" captionText="FIGURE 4. Scolelepis (Scolelepis) variegata Imajima, 1992. A, anterior end, dorsal view; B, parapodia of setiger 1, anterior view; C, parapodia of setiger 2, anterior view; D, parapodia of setiger 31, anterior view; E, notopodial seta from anterior row of setiger 4; F, notopodial seta from posterior row of setiger 4; G, neuropodial seta from anterior row of setiger 4; H, neuropodial seta from posterior row of setiger 4; I, neuropodial hooded hook from setiger 30. Scales: A, 600 μm; B D, 120 μm; E I, 250 μm." httpUri="https://zenodo.org/record/190438/files/figure.png" pageId="9" pageNumber="46">Fig. 4</figureCitation>
EH). Neuropodial hooded hooks from setigers 2427, bidentate, with distinct secondary tooth (
<figureCitation id="13369FCAFF9C2B38C8A6FC58FCFE3AF1" box="[759,837,964,990]" captionStart="FIGURE 4" captionStartId="10.[151,256,1872,1896]" captionTargetBox="[193,1381,187,1822]" captionTargetId="figure@10.[183,1404,172,1839]" captionTargetPageId="10" captionText="FIGURE 4. Scolelepis (Scolelepis) variegata Imajima, 1992. A, anterior end, dorsal view; B, parapodia of setiger 1, anterior view; C, parapodia of setiger 2, anterior view; D, parapodia of setiger 31, anterior view; E, notopodial seta from anterior row of setiger 4; F, notopodial seta from posterior row of setiger 4; G, neuropodial seta from anterior row of setiger 4; H, neuropodial seta from posterior row of setiger 4; I, neuropodial hooded hook from setiger 30. Scales: A, 600 μm; B D, 120 μm; E I, 250 μm." httpUri="https://zenodo.org/record/190438/files/figure.png" pageId="9" pageNumber="46">
Fig.
<date id="FFB3A58FFF9C2B38C962FC58FCFE3AF1" box="[819,837,964,990]" pageId="9" pageNumber="46">4</date>
</figureCitation>
I),
<quantity id="4CF52EAAFF9C2B38C938FC58FC7C3AF1" box="[873,967,964,990]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="4.064" metricValueMin="1.524" pageId="9" pageNumber="46" unit="in" value="11.0" valueMax="16.0" valueMin="6.0">616 in</quantity>
a series, accompanied by capillaries. Notopodial hooded hooks absent.
</paragraph>
<paragraph id="8BB2834FFF9C2B38CA94FB8DFD383D04" blockId="9.[151,1437,151,1260]" box="[197,643,1041,1067]" pageId="9" pageNumber="46">Pygidium missing in present material.</paragraph>
</subSubSection>
<subSubSection id="C317D0C4FF9C2B38CA97FBABFC713DE9" pageId="9" pageNumber="46" type="discussion">
<paragraph id="8BB2834FFF9C2B38CA97FBABFC713DE9" blockId="9.[151,1437,151,1260]" pageId="9" pageNumber="46">
<emphasis id="B9795F5DFF9C2B38CA97FBABFEFB3D7E" bold="true" box="[198,320,1079,1105]" pageId="9" pageNumber="46">Remarks.</emphasis>
The specimens agree well with previous descriptions and illustrations.
<taxonomicName id="4C0DF8CCFF9C2B38CE2FFBA4FEFC3D57" authority="Imajima, 1992" authorityName="Imajima" authorityYear="1992" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="9" pageNumber="46" phylum="Annelida" rank="species" species="variegata" subGenus="Scolelepis">
<emphasis id="B9795F5DFF9C2B38CE2FFBA4FB4E3D7E" box="[1150,1269,1080,1105]" italics="true" pageId="9" pageNumber="46">Scolelepis</emphasis>
(
<emphasis id="B9795F5DFF9C2B38CF54FBA4FAAF3D7E" box="[1285,1300,1080,1105]" italics="true" pageId="9" pageNumber="46">S</emphasis>
.)
<emphasis id="B9795F5DFF9C2B38CF7DFBA4FA273D7E" box="[1324,1436,1080,1105]" italics="true" pageId="9" pageNumber="46">variegata</emphasis>
<bibRefCitation id="EF9CFEBEFF9C2B38CAC6FBC2FEFC3D57" author="Imajima" box="[151,327,1118,1144]" pageId="9" pageNumber="48" refString="Imajima, M. (1992) Spionidae (Annelida, Polychaeta) from Japan VIII. The genus Scolelepis. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 18 (1), 1 - 34." type="journal article" year="1992">Imajima, 1992</bibRefCitation>
</taxonomicName>
is unique among
<taxonomicName id="4C0DF8CCFF9C2B38C84EFBC3FD223D57" box="[543,665,1119,1144]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="9" pageNumber="46" phylum="Annelida" rank="genus">
<emphasis id="B9795F5DFF9C2B38C84EFBC3FD223D57" box="[543,665,1119,1144]" italics="true" pageId="9" pageNumber="46">Scolelepis</emphasis>
</taxonomicName>
from
<collectingCountry id="F31AC3DFFF9C2B38C8B5FBC2FC973D57" box="[740,812,1118,1144]" name="China" pageId="9" pageNumber="46">China</collectingCountry>
in its relative plump body and lack of notosetae on setiger 1. Additionally, the branchiae and notopodial lamellae in the anterior part of the body are completely fused. The species is newly reported for the Chinese
<taxonomicName id="4C0DF8CCFF9C2B38C952FB30FCC13DEA" box="[771,890,1196,1221]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="9" pageNumber="46" phylum="Annelida" rank="genus">
<emphasis id="B9795F5DFF9C2B38C952FB30FCC13DEA" box="[771,890,1196,1221]" italics="true" pageId="9" pageNumber="46">Scolelepis</emphasis>
</taxonomicName>
fauna.
</paragraph>
</subSubSection>
<subSubSection id="C317D0C4FF9C2B38CA97FB4EFC413DC3" box="[198,1018,1234,1260]" pageId="9" pageNumber="46" type="distribution">
<paragraph id="8BB2834FFF9C2B38CA97FB4EFC413DC3" blockId="9.[151,1437,151,1260]" box="[198,1018,1234,1260]" pageId="9" pageNumber="46">
<emphasis id="B9795F5DFF9C2B38CA97FB4EFEDC3DC3" bold="true" box="[198,359,1234,1260]" pageId="9" pageNumber="46">Distribution.</emphasis>
<collectingCountry id="F31AC3DFFF9C2B38CB3CFB4EFE083DC3" box="[365,435,1234,1260]" name="China" pageId="9" pageNumber="46">China</collectingCountry>
(Yellow Sea, East
<collectingCountry id="F31AC3DFFF9C2B38C8C3FB4EFD633DC3" box="[658,728,1234,1260]" name="China" pageId="9" pageNumber="46">China</collectingCountry>
Sea;
<quantity id="4CF52EAAFF9C2B38C948FB4EFC203DC3" box="[793,923,1234,1260]" metricMagnitude="1" metricUnit="m" metricValue="2.725" metricValueMax="4.4" metricValueMin="1.05" pageId="9" pageNumber="46" unit="m" value="27.25" valueMax="44.0" valueMin="10.5">10.544 m</quantity>
);
<collectingCountry id="F31AC3DFFF9C2B38C9E1FB4EFC4D3DC3" box="[944,1014,1234,1260]" name="Japan" pageId="9" pageNumber="46">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>