treatments-xml/data/03/92/3C/03923C45FF8CFF873408FD2AFAAC17FF.xml
2024-06-21 12:22:17 +02:00

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<document id="2A16BA63A42C9FF234F7168CE93149A6" ID-CLB-Dataset="7746" ID-DOI="10.5252/geodiversitas2021v43a5" ID-GBIF-Dataset="e42658a2-0e15-4e90-9bc5-0d00cbcbd781" ID-ISSN="1638-9395" ID-Zenodo-Dep="4605963" ID-ZooBank="urn:lsid:zoobank.org:pub:697FC553-E37B-4EF9-97A4-950E4DEE246C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615828395930" checkinUser="felipe" docAuthor="Zouhri, Samir, Gingerich, Philip D., Khalloufi, Bouziane, Bourdon, Estelle, Adnet, Sylvain, Jouve, Stéphane, Elboudali, Najia, Amane, Ayoub, Rage, Jean-Claude &amp; Tabuce, Rodolphe" docDate="2021" docId="03923C45FF8CFF873408FD2AFAAC17FF" docLanguage="en" docName="Geodiversitas.43.5.121-150.pdf" docOrigin="Geodiversitas 43 (5)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Gen. indet." docType="treatment" docVersion="7" lastPageNumber="134" masterDocId="FFAB443DFF81FF883018FFDFFF95166E" masterDocTitle="Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco" masterLastPageNumber="150" masterPageNumber="121" pageNumber="132" updateTime="1698933443279" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="26C2A6BAB97B249809F0D559C0A81E2A">Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco</mods:title>
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<mods:namePart id="BF339AA786B2D72EB11551766D452272">Khalloufi, Bouziane</mods:namePart>
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<mods:namePart id="A9C5ADA8393E9742188FDCD90075A2CE">Bourdon, Estelle</mods:namePart>
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<mods:namePart id="5B7EE2B7344DB8CD11167CBDD754D0C6">Jouve, Stéphane</mods:namePart>
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<mods:namePart id="ED06511170D058E9B4A800BF04826638">Rage, Jean-Claude</mods:namePart>
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<treatment id="03923C45FF8CFF873408FD2AFAAC17FF" ID-DOI="http://doi.org/10.5281/zenodo.4606637" ID-GBIF-Taxon="180569427" ID-Zenodo-Dep="4606637" LSID="urn:lsid:plazi:treatment:03923C45FF8CFF873408FD2AFAAC17FF" httpUri="http://treatment.plazi.org/id/03923C45FF8CFF873408FD2AFAAC17FF" lastPageId="15" lastPageNumber="134" pageId="13" pageNumber="132">
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<paragraph id="8B848D53FF8CFF853408FD2AFB591561" blockId="13.[1040,1228,757,784]" box="[1040,1228,757,784]" pageId="13" pageNumber="132">
<taxonomicName id="4C3BF6D0FF8CFF853408FD2AFB521561" box="[1040,1223,757,784]" genus="Gen. indet." pageId="13" pageNumber="132" rank="genus">Gen. et sp. indet</taxonomicName>
.
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<paragraph id="8B848D53FF8CFF853335FCF0FC5D15F8" blockId="13.[811,1456,814,918]" pageId="13" pageNumber="132">
EXAMINED MATERIAL. —
<materialsCitation id="3B53870EFF8CFF853433FCF1FC5D15F8" ID-GBIF-Occurrence="3053068315" pageId="13" pageNumber="132">
FSAC Bouj-109 (
<figureCitation id="130091D6FF8CFF8534C5FCF1FAB81528" box="[1245,1325,814,838]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="13" pageNumber="132">Fig. 5H</figureCitation>
), costal fragment; 352 (
<figureCitation id="130091D6FF8CFF8533B8FC96FC74150F" box="[928,993,841,865]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="13" pageNumber="132">Fig. 5I</figureCitation>
), fragmentary hyoplastral process; 351 (
<figureCitation id="130091D6FF8CFF85357FFC96FA3D150F" box="[1383,1448,841,865]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="13" pageNumber="132">Fig. 5J</figureCitation>
) fragmentary hypoplastron; 353 (
<figureCitation id="130091D6FF8CFF85347DFCBCFB381515" box="[1125,1197,867,891]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="13" pageNumber="132">Fig. 5K</figureCitation>
) and 354 (
<figureCitation id="130091D6FF8CFF85350CFCBCFAC91515" box="[1300,1372,867,891]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="13" pageNumber="132">Fig. 5L</figureCitation>
), dermal plate fragments.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C321DED8FF8CFF873335FC6AFEFD145E" lastPageId="15" lastPageNumber="134" pageId="13" pageNumber="132" type="description">
<paragraph id="8B848D53FF8CFF853335FC6AFC2815A3" blockId="13.[811,1457,949,2028]" box="[813,957,949,975]" pageId="13" pageNumber="132">DESCRIPTION</paragraph>
<paragraph id="8B848D53FF8CFF853335FC0AFB041340" blockId="13.[811,1457,949,2028]" pageId="13" pageNumber="132">
FSAC Bouj-109 (
<figureCitation id="130091D6FF8CFF8533F4FC0BFBD41581" box="[1004,1089,980,1007]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="13" pageNumber="132">Fig. 5H</figureCitation>
) is a fragment of costal of a largesized turtle, covered by three scute parts. The thoracic rib is visible, included in the dermal bone, lentoid in cross-section and roundly protruding along the ventral costal face. The direction of the rib and scutes indicate a medial fragment of costal, not far from the neural, covered by two successive vertebrals (medially) and the corresponding pleural scute (laterally). The ornamentation of the plate is formed by irregular dichotomic sulci on a rough and granulous surface, which are features found in the basic ornamentation of
<taxonomicName id="4C3BF6D0FF8CFF853536FB2CFB191340" authority="(Lapparent de Broin et al. 2014)" baseAuthorityName="Lapparent de Broin" baseAuthorityYear="2014" class="Reptilia" family="Cheloniidae" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="132" phylum="Chordata" rank="family">
Cheloniidae (
<bibRefCitation id="EFAAF0A2FF8CFF85332EFACBFB141340" author="LAPPARENT DE BROIN F. DE &amp; MURELAGA X. &amp; FARRES F. &amp; ALTIMIRAS J." box="[822,1153,1299,1326]" pageId="13" pageNumber="132" pagination="111 - 132" refId="ref22556" refString="LAPPARENT DE BROIN F. DE, MURELAGA X., FARRES F. &amp; ALTIMIRAS J. 2014. - An exceptional cheloniid turtle, Osonachelus decorata nov. gen., nov. sp., from the Eocene (Bartonian) of Catalonia (Spain). Geobios 47: 111 - 132. https: // doi. org / 10.1016 / j. geobios. 2014.02.002" type="journal article" year="2014">
Lapparent de Broin
<emphasis id="B94F5141FF8CFF853417FACBFBD61343" box="[1039,1091,1299,1325]" italics="true" pageId="13" pageNumber="132">et al.</emphasis>
2014
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B848D53FF8CFF87335BFAECFD46171F" blockId="13.[811,1457,949,2028]" lastBlockId="15.[130,778,215,560]" lastPageId="15" lastPageNumber="134" pageId="13" pageNumber="132">
FSAC Bouj-352 (
<figureCitation id="130091D6FF8CFF85341AFAECFBD21320" box="[1026,1095,1331,1358]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="13" pageNumber="132">Fig 5I</figureCitation>
), a fragmentary lateral process of right hyoplastron, and Bouj-351 (
<figureCitation id="130091D6FF8CFF853496FA8CFB411300" box="[1166,1236,1363,1390]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="13" pageNumber="132">Fig. 5J</figureCitation>
) a subcomplete right hypoplastron, probably belong to the same individual. Both fragments show a granulous surface with protruding elongated polygones and dichotomic sulci, representing ornamentation features known in cheloniids. Digitations of hyoplastral and hypoplastral lateral processes are broken close to their base. However, they are much enveloped in the dermal callosity and seem to have shortly overtaken the callosity border. Between both lateral processes lies an important lateral fontanelle that was originally rectangular or square. The medial part of the hypoplastron, along with its counterpart, indicates a very short and narrow central fontanelle with a triangular posteromedial border, and posteriorly both hypoplastra were close and lacked digitations on their medial border. The main body of the hypoplastron is posteriorly broken anterior to the contact with the xiphiplastron, close to the area of the abdominofemoral sulcus. Anterior to the inguinal notch on the lateral hypoplastral process, an inguinal scute sulcus (i.e. the posterior inframarginal of the complete series that is present in cheloniids) joins the area of the femoroabdominal sulcus medially. The lateral hyo-hypoplastral processes are narrower than the main medial body of each hypoplastron, and consequently the lateral fontanelles are narrower than the main hypoplastral bodies and they are narrower than high. The base of each lateral, hyoplastral and hypoplastral process is also shorter than the length of the lateral fontanelles.
</paragraph>
<caption id="DF44DDDBFF8FFF86309CF92AFE1D1187" ID-DOI="http://doi.org/10.5281/zenodo.4605975" ID-Zenodo-Dep="4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="14" pageNumber="133" startId="14.[132,143,1781,1798]" targetBox="[132,1455,215,1723]" targetPageId="14">
<paragraph id="8B848D53FF8FFF86309CF92AFE1D1187" blockId="14.[132,1457,1781,2025]" pageId="14" pageNumber="133">
FIG. 5. —
<taxonomicName id="4C3BF6D0FF8FFF8630CEF92AFEDC1168" authorityName="Cope" authorityYear="1887" box="[214,329,1781,1798]" class="Actinopterygii" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="133" phylum="Chordata" rank="class">Actinopterygii</taxonomicName>
and
<taxonomicName id="4C3BF6D0FF8FFF86316CF92AFE5A1168" authorityName="Linnaeus" authorityYear="1758" box="[372,463,1781,1798]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="133" phylum="Chordata" rank="order">Testudines</taxonomicName>
from the Sabkha of Gueran:
<emphasis id="B94F5141FF8FFF8632DDF92AFD721168" bold="true" box="[709,743,1781,1798]" pageId="14" pageNumber="133">A -G</emphasis>
,
<taxonomicName id="4C3BF6D0FF8FFF8632EBF92AFCFC1168" authorityName="Cope" authorityYear="1887" box="[755,873,1781,1798]" class="Actinopterygii" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="133" phylum="Chordata" rank="class">Actinopterygii</taxonomicName>
:
<taxonomicName id="4C3BF6D0FF8FFF863368F92AFC631168" authorityName="Leidy" authorityYear="1856" box="[880,1014,1781,1798]" class="Actinopterygii" family="Scombridae" genus="Cylindracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="133" phylum="Chordata" rank="genus">
<emphasis id="B94F5141FF8FFF863368F92AFC631168" box="[880,1014,1781,1798]" italics="true" pageId="14" pageNumber="133">Cylindracanthus</emphasis>
</taxonomicName>
sp., FSAC Bouj-356, rostrum in (
<emphasis id="B94F5141FF8FFF863517F92AFAB01168" bold="true" box="[1295,1317,1781,1798]" pageId="14" pageNumber="133">A1</emphasis>
) lateral and (
<emphasis id="B94F5141FF8FFF86358BF92AFA3E1168" bold="true" box="[1427,1451,1781,1798]" pageId="14" pageNumber="133">A2</emphasis>
) transversal views;
<emphasis id="B94F5141FF8FFF863105F8D3FEA81173" bold="true" box="[285,317,1804,1821]" pageId="14" pageNumber="133">B -E</emphasis>
, Percomorpha indet.:
<emphasis id="B94F5141FF8FFF8631EDF8D3FD971173" bold="true" box="[501,514,1804,1821]" pageId="14" pageNumber="133">B</emphasis>
, FSAC Bouj-369,vertebra in (
<emphasis id="B94F5141FF8FFF8632E1F8D3FC851173" bold="true" box="[761,784,1804,1821]" pageId="14" pageNumber="133">B1</emphasis>
) lateral and (
<emphasis id="B94F5141FF8FFF863362F8D3FC041173" bold="true" box="[890,913,1804,1821]" pageId="14" pageNumber="133">B2</emphasis>
) transversal views;
<emphasis id="B94F5141FF8FFF86342BF8D3FBD41173" bold="true" box="[1075,1089,1804,1821]" pageId="14" pageNumber="133">C</emphasis>
, FSAC Bouj-363, vertebra in (
<emphasis id="B94F5141FF8FFF86352FF8D3FADA1173" bold="true" box="[1335,1359,1804,1821]" pageId="14" pageNumber="133">C1</emphasis>
) lateral and (
<emphasis id="B94F5141FF8FFF863091F8FDFF34115D" bold="true" box="[137,161,1826,1843]" pageId="14" pageNumber="133">C2</emphasis>
) transversal views;
<emphasis id="B94F5141FF8FFF86315EF8FDFEC1115D" bold="true" box="[326,340,1826,1843]" pageId="14" pageNumber="133">D</emphasis>
, FSAC Bouj-370, basioccipital in left lateral view;
<emphasis id="B94F5141FF8FFF8632EDF8FDFC94115D" bold="true" box="[757,769,1826,1843]" pageId="14" pageNumber="133">E</emphasis>
, FSAC Bouj-371, fin spine in anterior view;
<emphasis id="B94F5141FF8FFF863476F8FDFBEC115D" bold="true" box="[1134,1145,1826,1843]" pageId="14" pageNumber="133">F</emphasis>
,
<taxonomicName id="4C3BF6D0FF8FFF86349CF8FDFB7C115D" authorityName="Rafinesque" authorityYear="1815" box="[1156,1257,1826,1843]" class="Actinopterygii" family="Scombridae" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="133" phylum="Chordata" rank="family">Scombridae</taxonomicName>
indet., FSAC Bouj-358, incomplete jawbone in labial view;
<emphasis id="B94F5141FF8FFF8631B0F8E6FE231124" bold="true" box="[424,438,1849,1866]" pageId="14" pageNumber="133">G</emphasis>
,
<taxonomicName id="4C3BF6D0FF8FFF8631D9F8E6FDB11124" authorityName="Cuvier" authorityYear="1817" box="[449,548,1849,1866]" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="132" phylum="Chordata" rank="order">Siluriformes</taxonomicName>
indet., FSAC Bouj-372, right pectoral spine in anterior view;
<emphasis id="B94F5141FF8FFF863438F8E6FBD41124" bold="true" box="[1056,1089,1849,1866]" pageId="14" pageNumber="133">H -P</emphasis>
,
<taxonomicName id="4C3BF6D0FF8FFF863454F8E6FB3F1124" authorityName="Linnaeus" authorityYear="1758" box="[1100,1194,1849,1866]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="133" phylum="Chordata" rank="order">Testudines</taxonomicName>
:
<emphasis id="B94F5141FF8FFF8634AAF8E6FB471124" bold="true" box="[1202,1234,1849,1866]" pageId="14" pageNumber="133">H -L</emphasis>
,
<taxonomicName id="4C3BF6D0FF8FFF8634C5F8E6FAD51124" box="[1245,1344,1849,1866]" class="Reptilia" family="Cheloniidae" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="133" phylum="Chordata" rank="family">Cheloniidae</taxonomicName>
indet., FSAC Bouj-109,fragmentary costal in (
<emphasis id="B94F5141FF8FFF86318AF88FFE3C110F" bold="true" box="[402,425,1872,1889]" pageId="14" pageNumber="133">H1</emphasis>
) lateral transection, (
<emphasis id="B94F5141FF8FFF86324DF88FFDFB110F" bold="true" box="[597,622,1872,1889]" pageId="14" pageNumber="133">H2</emphasis>
) ventral, (
<emphasis id="B94F5141FF8FFF8632A5F88FFD43110F" bold="true" box="[701,726,1872,1889]" pageId="14" pageNumber="133">H3</emphasis>
) dorsal and (
<emphasis id="B94F5141FF8FFF863358F88FFCCD110F" bold="true" box="[832,856,1872,1889]" pageId="14" pageNumber="133">H4</emphasis>
) medial transection views;
<emphasis id="B94F5141FF8FFF86342FF88FFBA8110F" bold="true" box="[1079,1085,1872,1889]" pageId="14" pageNumber="133">I</emphasis>
, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (
<emphasis id="B94F5141FF8FFF86315DF8B9FEC01119" bold="true" box="[325,341,1894,1911]" pageId="14" pageNumber="133">I1</emphasis>
) ventral and (
<emphasis id="B94F5141FF8FFF8631DEF8B9FE431119" bold="true" box="[454,470,1894,1911]" pageId="14" pageNumber="133">I2</emphasis>
) dorsal views;
<emphasis id="B94F5141FF8FFF86324BF8B9FDC81119" bold="true" box="[595,605,1894,1911]" pageId="14" pageNumber="133">J</emphasis>
, FSAC Bouj-351, subcomplete right hypoplastron in (
<emphasis id="B94F5141FF8FFF863405F8B9FBA41119" bold="true" box="[1053,1073,1894,1911]" pageId="14" pageNumber="133">J1</emphasis>
) ventral and (
<emphasis id="B94F5141FF8FFF8634BBF8B9FB2D1119" bold="true" box="[1187,1208,1894,1911]" pageId="14" pageNumber="133">J2</emphasis>
) dorsal views;
<emphasis id="B94F5141FF8FFF86352CF8B9FAD41119" bold="true" box="[1332,1345,1894,1911]" pageId="14" pageNumber="133">K</emphasis>
, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view;
<emphasis id="B94F5141FF8FFF8632EAF8A2FD6811E0" bold="true" box="[754,765,1917,1934]" pageId="14" pageNumber="133">L</emphasis>
, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (
<emphasis id="B94F5141FF8FFF863551F8A2FACB11E0" bold="true" box="[1353,1374,1917,1934]" pageId="14" pageNumber="133">L1</emphasis>
,
<emphasis id="B94F5141FF8FFF863571F8A2FAEA11E0" bold="true" box="[1385,1407,1917,1934]" pageId="14" pageNumber="133">L2</emphasis>
) both faces;
<emphasis id="B94F5141FF8FFF8630A3F84BFF5911CB" bold="true" box="[187,204,1940,1957]" pageId="14" pageNumber="133">M</emphasis>
,
<taxonomicName id="4C3BF6D0FF8FFF8630CEF84BFEC811CB" authorityName="Fitzinger" authorityYear="1843" box="[214,349,1940,1957]" class="Reptilia" family="Dermochelyidae" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="133" phylum="Chordata" rank="family">Dermochelyidae</taxonomicName>
indet., FSAC Bouj-350, lateral pubic process of right pubis in (
<emphasis id="B94F5141FF8FFF86337AF84BFCE811CB" bold="true" box="[866,893,1940,1957]" pageId="14" pageNumber="133">M1</emphasis>
) ventral and (
<emphasis id="B94F5141FF8FFF8633F4F84BFB9D11CB" bold="true" box="[1004,1032,1940,1957]" pageId="14" pageNumber="133">M2</emphasis>
) dorsal views;
<emphasis id="B94F5141FF8FFF86349AF84BFB0511CB" bold="true" box="[1154,1168,1940,1957]" pageId="14" pageNumber="133">N</emphasis>
,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (
<emphasis id="B94F5141FF8FFF8631ECF875FD9911D5" bold="true" box="[500,524,1962,1979]" pageId="14" pageNumber="133">N1</emphasis>
) dorsal and (
<emphasis id="B94F5141FF8FFF863260F875FD0511D5" bold="true" box="[632,656,1962,1979]" pageId="14" pageNumber="133">N2</emphasis>
) ventral views;
<emphasis id="B94F5141FF8FFF863308F875FC8A11D5" bold="true" box="[784,799,1962,1979]" pageId="14" pageNumber="133">O</emphasis>
, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (
<emphasis id="B94F5141FF8FFF86312BF81EFED911BC" bold="true" box="[307,332,1985,2002]" pageId="14" pageNumber="133">O1</emphasis>
) dorsal and (
<emphasis id="B94F5141FF8FFF8631AFF81EFE4511BC" bold="true" box="[439,464,1985,2002]" pageId="14" pageNumber="133">O2</emphasis>
) ventral views;
<emphasis id="B94F5141FF8FFF863249F81EFDC811BC" bold="true" box="[593,605,1985,2002]" pageId="14" pageNumber="133">P</emphasis>
, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm.
</paragraph>
</caption>
<paragraph id="8B848D53FF8EFF873083FEA8FEFD145E" blockId="15.[130,778,215,560]" pageId="15" pageNumber="134">
FSAC Bouj-353 (
<figureCitation id="130091D6FF8EFF87317EFEA8FE2B17FF" box="[358,446,374,401]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="15" pageNumber="134">Fig. 5K</figureCitation>
) and Bouj-354 (
<figureCitation id="130091D6FF8EFF873299FEA8FD4017FF" box="[641,725,374,401]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="15" pageNumber="134">Fig. 5L</figureCitation>
) are fragments of dermal plate which are not located on the shell. Bouj-354 shows a granular surface, not clearly polygonal, and probably corresponds to the same individual as Bouj-352 (
<figureCitation id="130091D6FF8EFF873095FE29FF42147F" box="[141,215,502,529]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="15" pageNumber="134">Fig. 5I</figureCitation>
) and Bouj-351(
<figureCitation id="130091D6FF8EFF873190FE29FE41147F" box="[392,468,502,529]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="15" pageNumber="134">Fig. 5J</figureCitation>
). Bouj-353 shows a comparable ornamentation.
</paragraph>
</subSubSection>
<subSubSection id="C321DED8FF8EFF87309CFD89FAAC17FF" pageId="15" pageNumber="134" type="discussion">
<paragraph id="8B848D53FF8EFF87309CFD89FF7D1400" blockId="15.[130,777,598,2027]" box="[132,232,598,624]" pageId="15" pageNumber="134">REMARKS</paragraph>
<paragraph id="8B848D53FF8EFF87309CFDA9FEAD1521" blockId="15.[130,777,598,2027]" pageId="15" pageNumber="134">
FSAC Bouj-352 (
<figureCitation id="130091D6FF8EFF873150FDA9FE0614FE" box="[328,403,630,657]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="15" pageNumber="134">Fig. 5I</figureCitation>
), Bouj-351 (
<figureCitation id="130091D6FF8EFF87323BFDA9FDFB14FE" box="[547,622,630,657]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="15" pageNumber="134">Fig. 5J</figureCitation>
) and possibly Bouj-354 (
<figureCitation id="130091D6FF8EFF8730E0FD49FED014DE" box="[248,325,661,688]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="15" pageNumber="134">Fig. 5L</figureCitation>
) are parts of the same cheloniid individual, being found lying together in one piece (
<figureCitation id="130091D6FF8EFF873240FD6AFD3914BE" box="[600,684,693,720]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="15" pageNumber="134">Fig. 5P</figureCitation>
), which also contained Bouj-350 (
<figureCitation id="130091D6FF8EFF873183FD0AFE61149E" box="[411,500,725,752]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="15" pageNumber="134">Fig. 5M</figureCitation>
), a dermochelyid remain. Bouj-353 (
<figureCitation id="130091D6FF8EFF8730E6FD2AFEC6157E" box="[254,339,757,784]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="15" pageNumber="134">Fig. 5K</figureCitation>
) was probably collected near this piece and corresponds to the same cheloniid and possibly to the same individual.
</paragraph>
<paragraph id="8B848D53FF8EFF873083FC8AFAAC17FF" blockId="15.[130,777,598,2027]" lastBlockId="15.[811,1456,215,401]" pageId="15" pageNumber="134">
The combination of the nearly flat bridge (not an obliquely elevated bridge), surface ornamentation, and fenestration of the plastron matches the cheloniid pattern. Only few middle to late Eocene cheloniids are known from their plastron in the Old World. Among the various cheloniid clades, some cheloniid genera were grouped in the western- European “
<taxonomicName id="4C3BF6D0FF8EFF8730E7FBCBFDB21240" authority="Moody, 1968" authorityName="Moody" authorityYear="1968" box="[255,551,1044,1070]" class="Reptilia" family="Cheloniidae" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="134" phylum="Chordata" rank="subFamily" subFamily="Eochelyinae">
Eochelyinae
<bibRefCitation id="EFAAF0A2FF8EFF873196FBCBFDB21240" author="MOODY R. T. J." box="[398,551,1044,1070]" pageId="15" pageNumber="134" pagination="129 - 140" refId="ref23902" refString="MOODY R. T. J. 1968. - A Turtle, Eochelys crassicostata (Owen), from the London Clay of the Isle of Sheppey. Proceedings of the Geological Association 79: 129 - 140. https: // doi. org / 10.1016 / S 0016 - 7878 (68) 80031 - 8" type="journal article" year="1968">Moody, 1968</bibRefCitation>
</taxonomicName>
”, now considered as a paraphyletic taxon representing an evolutionary grade (
<bibRefCitation id="EFAAF0A2FF8EFF873096FB8AFE7C1200" author="LAPPARENT DE BROIN F. DE &amp; MURELAGA X. &amp; FARRES F. &amp; ALTIMIRAS J." box="[142,489,1108,1135]" pageId="15" pageNumber="134" pagination="111 - 132" refId="ref22556" refString="LAPPARENT DE BROIN F. DE, MURELAGA X., FARRES F. &amp; ALTIMIRAS J. 2014. - An exceptional cheloniid turtle, Osonachelus decorata nov. gen., nov. sp., from the Eocene (Bartonian) of Catalonia (Spain). Geobios 47: 111 - 132. https: // doi. org / 10.1016 / j. geobios. 2014.02.002" type="journal article" year="2014">
Lapparent de Broin
<emphasis id="B94F5141FF8EFF87316BFB8AFE3F1200" box="[371,426,1108,1134]" italics="true" pageId="15" pageNumber="134">et al.</emphasis>
2014
</bibRefCitation>
,
<bibRefCitation id="EFAAF0A2FF8EFF8731E2FB8BFDA61200" author="LAPPARENT DE BROIN F. DE &amp; MURELAGA X. &amp; GARCIA A. &amp; FRANCESC FARRES F. &amp; ALTIMIRAS J." box="[506,563,1108,1134]" pageId="15" pageNumber="134" pagination="237 - 284" refId="ref22627" refString="LAPPARENT DE BROIN F. DE, MURELAGA X., PEREZ- GARCIA A., FRANCESC FARRES F. &amp; ALTIMIRAS J. 2018. - The turtles from the upper Eocene, Osona County (Ebro Basin, Catalonia, Spain): new material and its faunistic and environmental context. Fossil Record 21: 237 - 284. https: // doi. org / 10.5194 / fr- 21 - 237 - 2018" type="journal article" year="2018">2018</bibRefCitation>
). The Gueran species, represented by Bouj-351 and Bouj-352, represents this grade. It differs from the defined species included in the group in the relative proportions of the hyo-hypoplastral main body, in relation to the lateral and central fontanelles. There are several species in the early Eocene of the London Clay basin (Ypresian of the Isle of Sheppey (
<collectingRegion id="49FF43B1FF8EFF873245FACBFD031340" box="[605,662,1300,1326]" country="United Kingdom" name="Kent" pageId="15" pageNumber="134">Kent</collectingRegion>
) and Harwich (
<collectingRegion id="49FF43B1FF8EFF8730DFFAECFE931323" box="[199,262,1331,1357]" country="United Kingdom" name="Essex" pageId="15" pageNumber="134">Essex</collectingRegion>
) (
<collectingCountry id="F32CCDC3FF8EFF873105FAECFEDF1323" box="[285,330,1331,1357]" name="United Kingdom" pageId="15" pageNumber="134">UK</collectingCountry>
), described by
<bibRefCitation id="EFAAF0A2FF8EFF8731E8FAECFD431320" author="OWEN R. &amp; BELL T." box="[496,726,1331,1358]" pageId="15" pageNumber="134" pagination="77 - 79" refId="ref24451" refString="OWEN R. &amp; BELL T. 1849. - Monograph on the Fossil Reptilia of the London Clay. Part I. Chelonia: 1 - 76, Supplement to the Eocene Chelonia. Palaeontographical Society of London, London: 77 - 79. https: // doi. org / 10.5962 / bhl. title. 106755" type="book chapter" year="1849">Owen &amp; Bell (1849)</bibRefCitation>
and Owen (1849-1884). Boujdour species is similar to “
<taxonomicName id="4C3BF6D0FF8EFF8732ABFA8CFEE013E3" authority="Owen, 1842" authorityName="Owen" authorityYear="1842" class="Magnoliopsida" family="Plantaginaceae" genus="Chelone" kingdom="Animalia" order="Lamiales" pageId="15" pageNumber="134" phylum="Tracheophyta" rank="species" species="breviceps">
<emphasis id="B94F5141FF8EFF8732ABFA8CFF7413E3" italics="true" pageId="15" pageNumber="134">Chelone breviceps</emphasis>
Owen, 1842
</taxonomicName>
” (see
<bibRefCitation id="EFAAF0A2FF8EFF8731A0FAACFD1813E3" author="OWEN R. &amp; BELL T." box="[440,653,1395,1421]" pageId="15" pageNumber="134" pagination="77 - 79" refId="ref24451" refString="OWEN R. &amp; BELL T. 1849. - Monograph on the Fossil Reptilia of the London Clay. Part I. Chelonia: 1 - 76, Supplement to the Eocene Chelonia. Palaeontographical Society of London, London: 77 - 79. https: // doi. org / 10.5962 / bhl. title. 106755" type="book chapter" year="1849">Owen &amp; Bell 1849</bibRefCitation>
: pl. 2), i.e. a junior synonym of
<taxonomicName id="4C3BF6D0FF8EFF87316FFA4CFC9113C0" authority="(Konig, 1825)" baseAuthorityName="Konig" baseAuthorityYear="1825" box="[375,772,1427,1454]" class="Reptilia" family="Cheloniidae" genus="Argillochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="134" phylum="Chordata" rank="species" species="antiqua">
<emphasis id="B94F5141FF8EFF87316FFA4CFDCD13C3" box="[375,600,1427,1453]" italics="true" pageId="15" pageNumber="134">Argillochelys antiqua</emphasis>
(König, 1825)
</taxonomicName>
, in the shape of the quadrangular lateral fontanelles, but it differs in the wider proportions of these lateral fontanelles correlated with a narrower hypoplastral main body, and it differs in the robustness of the plates. However, the central fontanelle was narrow and short in both species. By contrast and as
<taxonomicName id="4C3BF6D0FF8EFF8730D5F98CFEAA1002" baseAuthorityName="Konig" baseAuthorityYear="1825" box="[205,319,1619,1645]" class="Reptilia" family="Cheloniidae" genus="Argillochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="134" phylum="Chordata" rank="species" species="antiqua">
<emphasis id="B94F5141FF8EFF8730D5F98CFEAA1002" box="[205,319,1619,1645]" italics="true" pageId="15" pageNumber="134">A. antiqua</emphasis>
</taxonomicName>
, the Gueran species differs from “
<taxonomicName id="4C3BF6D0FF8EFF8732AAF98DFF4210E2" baseAuthorityName="Owen" baseAuthorityYear="1841" class="Magnoliopsida" family="Plantaginaceae" genus="Chelone" kingdom="Animalia" order="Lamiales" pageId="15" pageNumber="134" phylum="Tracheophyta" rank="species" species="convexa">
<emphasis id="B94F5141FF8EFF8732AAF98DFF4210E2" italics="true" pageId="15" pageNumber="134">Chelone convexa</emphasis>
</taxonomicName>
” (undefined taxon, perhaps a junior synonym of
<taxonomicName id="4C3BF6D0FF8EFF87309EF94DFE5610C2" authority="Owen" authorityName="Owen" box="[134,451,1682,1708]" class="Reptilia" family="Cheloniidae" genus="Argillochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="134" phylum="Chordata" rank="species" species="cuneiceps">
<emphasis id="B94F5141FF8EFF87309EF94DFEE110C2" box="[134,372,1682,1708]" italics="true" pageId="15" pageNumber="134">Argillochelys cuneiceps</emphasis>
Owen
</taxonomicName>
,
<emphasis id="B94F5141FF8EFF8731CAF94CFE7D10C2" box="[466,488,1683,1708]" italics="true" pageId="15" pageNumber="134">in</emphasis>
<bibRefCitation id="EFAAF0A2FF8EFF8731E8F94DFD4610C2" author="OWEN R. &amp; BELL T." box="[496,723,1682,1709]" pageId="15" pageNumber="134" pagination="77 - 79" refId="ref24451" refString="OWEN R. &amp; BELL T. 1849. - Monograph on the Fossil Reptilia of the London Clay. Part I. Chelonia: 1 - 76, Supplement to the Eocene Chelonia. Palaeontographical Society of London, London: 77 - 79. https: // doi. org / 10.5962 / bhl. title. 106755" type="book chapter" year="1849">Owen &amp; Bell, 1849</bibRefCitation>
, the shell of which is not defined),
<taxonomicName id="4C3BF6D0FF8EFF8731CFF96DFF2B1082" authority="Dollo, 1903" authorityName="Dollo" authorityYear="1903" class="Reptilia" family="Cheloniidae" genus="Eochelone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="134" phylum="Chordata" rank="species" species="brabantica">
<emphasis id="B94F5141FF8EFF8731CFF96DFD2C10A2" box="[471,697,1714,1740]" italics="true" pageId="15" pageNumber="134">Eochelone brabantica</emphasis>
Dollo, 1903
</taxonomicName>
(Lutetian, middle Eocene of Brabant,
<collectingCountry id="F32CCDC3FF8EFF873268F90DFD5B1083" box="[624,718,1746,1773]" name="Belgium" pageId="15" pageNumber="134">Belgium</collectingCountry>
) and
<taxonomicName id="4C3BF6D0FF8EFF87309EF92DFD2C1162" authority="Lapparent de Broin, Murelaga" authorityName="Lapparent de Broin, Murelaga" box="[134,697,1778,1804]" class="Reptilia" family="Cheloniidae" genus="Eochelone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="134" phylum="Chordata" rank="species" species="voltregana">
<emphasis id="B94F5141FF8EFF87309EF92DFEF71162" box="[134,354,1778,1804]" italics="true" pageId="15" pageNumber="134">Eochelone voltregana</emphasis>
Lapparent de Broin, Murelaga
</taxonomicName>
, Pérez- García, Francesc Farrés &amp; Altimiras, 2018 (Priabonian of Osona county,
<collectingCountry id="F32CCDC3FF8EFF873136F8EEFEF81122" box="[302,365,1841,1868]" name="Spain" pageId="15" pageNumber="134">Spain</collectingCountry>
) in the much narrower central fontanelle, the main body of each hypoplastron being wider, but it is similar in the quadrangular lateral fontanelle general proportions. And in the robustness of the plastral elements, it is similar to
<taxonomicName id="4C3BF6D0FF8EFF873138F86DFE3F11A5" box="[288,426,1969,1995]" class="Reptilia" family="Cheloniidae" genus="Eochelone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="134" phylum="Chordata" rank="species" species="voltregana">
<emphasis id="B94F5141FF8EFF873138F86DFE3F11A5" box="[288,426,1969,1995]" italics="true" pageId="15" pageNumber="134">E. voltregana</emphasis>
</taxonomicName>
. The Gueran species also differs from
<taxonomicName id="4C3BF6D0FF8EFF8730D8F80DFD881185" authority="(Gray, 1831)" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[192,541,2001,2027]" class="Reptilia" family="Cheloniidae" genus="Puppigerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="134" phylum="Chordata" rank="species" species="camperi">
<emphasis id="B94F5141FF8EFF8730D8F80DFE181185" box="[192,397,2002,2027]" italics="true" pageId="15" pageNumber="134">Puppigerus camperi</emphasis>
(Gray, 1831)
</taxonomicName>
(Ypresian of London Clay and Lutetian of Brabant) in the lateral fontanelle size and shape, this structure being much smaller and triangular in
<taxonomicName id="4C3BF6D0FF8EFF873350FEC7FBAB175F" authority="Cope, 1871" authorityName="Cope" authorityYear="1871" box="[840,1086,279,305]" class="Reptilia" family="Cheloniidae" genus="Puppigerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="134" phylum="Chordata" rank="genus">
<emphasis id="B94F5141FF8EFF873350FEC7FC2F175F" box="[840,954,280,305]" italics="true" pageId="15" pageNumber="134">Puppigerus</emphasis>
Cope, 1871
</taxonomicName>
adults with much wider hypoplastra, in a much wider plastron as a whole (due to the more developed shell ossification characteristic of this taxon) (
<bibRefCitation id="EFAAF0A2FF8EFF87332EFEA8FC5317FF" author="MOODY R. T. J." box="[822,966,375,401]" pageId="15" pageNumber="134" pagination="155 - 186" refId="ref23961" refString="MOODY R. T. J. 1974. - The taxonomy and morphology of Puppigerus camperi (Gray), an Eocene sea-turtle from northern Europe. Bulletin of the British Museum (Natural History) Geology 25: 155 - 186. https: // www. biodiversitylibrary. org / page / 2239204" type="journal article" year="1974">Moody 1974</bibRefCitation>
;
<bibRefCitation id="EFAAF0A2FF8EFF8733CDFEA8FABC17FF" author="LAPPARENT DE BROIN F. DE &amp; MURELAGA X. &amp; GARCIA A. &amp; FRANCESC FARRES F. &amp; ALTIMIRAS J." box="[981,1321,375,401]" pageId="15" pageNumber="134" pagination="237 - 284" refId="ref22627" refString="LAPPARENT DE BROIN F. DE, MURELAGA X., PEREZ- GARCIA A., FRANCESC FARRES F. &amp; ALTIMIRAS J. 2018. - The turtles from the upper Eocene, Osona County (Ebro Basin, Catalonia, Spain): new material and its faunistic and environmental context. Fossil Record 21: 237 - 284. https: // doi. org / 10.5194 / fr- 21 - 237 - 2018" type="journal article" year="2018">
Lapparent de Broin
<emphasis id="B94F5141FF8EFF8734ACFEA8FB7C17FF" box="[1204,1257,375,401]" italics="true" pageId="15" pageNumber="134">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>