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<document id="C90574BBE6A3486B8F18A4804CEA383B" ID-CLB-Dataset="2977" ID-DOI="10.11646/palaeoentomology.4.6.10" ID-GBIF-Dataset="f49cd992-5bff-4957-8e2c-a52adff778bf" ID-ISSN="2624-2834" ID-Zenodo-Dep="5778474" ID-ZooBank="5EEF1F8F-30AE-4705-9B71-AB42294903A3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639468513686" checkinUser="plazi" docAuthor="Li, Yan-Da, Huang, Di-Ying &amp; Cai, Chen-Yang" docDate="2021" docId="038887CDFFB5FFB3FC9A00DAFE29561A" docLanguage="en" docName="palaeoentomology.4.6.10.pdf" docOrigin="Palaeoentomology 4 (6)" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Coslonatus rasnitsyni Li &amp; Huang &amp; Cai 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="618" masterDocId="FFB1FFB5FFB4FFB7FFB0013DFF805507" masterDocTitle="An aberrant colydiine-like tenebrionoid beetle from mid-Cretaceous Burmese amber (Coleoptera: Tenebrionoidea: Zopheridae)" masterLastPageNumber="619" masterPageNumber="614" pageNumber="615" updateTime="1699472351469" updateUser="ExternalLinkService">
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<mods:title id="5EFD7FBFE09B94CACB21BD439913132B">An aberrant colydiine-like tenebrionoid beetle from mid-Cretaceous Burmese amber (Coleoptera: Tenebrionoidea: Zopheridae)</mods:title>
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<mods:namePart id="B3C5A0A2242D99E1EAE753CF6B952DFC">Li, Yan-Da</mods:namePart>
<mods:affiliation id="69EB9B98E5D559F2911FBBA5DBA97CFF">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Centre for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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<mods:namePart id="588F1EB0A7D4C74E27E128CB2E75AC6A">Huang, Di-Ying</mods:namePart>
<mods:affiliation id="2D45F4AAB150C20B16B88E3E491E5523">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Centre for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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<mods:namePart id="40CB03FA6B5389B77FB790C162725886">Cai, Chen-Yang</mods:namePart>
<mods:affiliation id="A1447159B330A717C402FFDCF6990049">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Centre for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK</mods:affiliation>
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<treatment id="038887CDFFB5FFB3FC9A00DAFE29561A" ID-DOI="http://doi.org/10.5281/zenodo.5779796" ID-GBIF-Taxon="191559957" ID-Zenodo-Dep="5779796" LSID="urn:lsid:plazi:treatment:038887CDFFB5FFB3FC9A00DAFE29561A" httpUri="http://treatment.plazi.org/id/038887CDFFB5FFB3FC9A00DAFE29561A" lastPageId="4" lastPageNumber="618" pageId="1" pageNumber="615">
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<paragraph id="8B9E36DBFFB5FFB6FC9A00DAFB755705" blockId="1.[810,1269,487,550]" box="[810,1269,487,514]" pageId="1" pageNumber="615">
<heading id="D0D681B7FFB5FFB6FC9A00DAFB755705" bold="true" box="[810,1269,487,514]" fontSize="10" level="2" pageId="1" pageNumber="615" reason="3">
<emphasis id="B955EAC9FFB5FFB6FC9A00DAFB755705" bold="true" box="[810,1269,487,514]" pageId="1" pageNumber="615">
<taxonomicName id="4C214D58FFB5FFB6FC9A00DAFB145706" authority="Li &amp; Cai" authorityName="Li &amp; Huang &amp; Cai" authorityYear="2021" box="[810,1172,487,514]" class="Insecta" family="Zopheridae" genus="Coslonatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="615" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
<emphasis id="B955EAC9FFB5FFB6FC9A00DAFBA45705" bold="true" box="[810,1060,487,514]" italics="true" pageId="1" pageNumber="615">Coslonatus rasnitsyni</emphasis>
Li &amp; Cai
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<taxonomicNameLabel id="A26657B2FFB5FFB6FB2B00D5FB755705" box="[1179,1269,488,514]" pageId="1" pageNumber="615" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C33B6550FFB5FFB6FC9A0331FC1E5721" box="[810,926,524,550]" pageId="1" pageNumber="615" type="description">
<paragraph id="8B9E36DBFFB5FFB6FC9A0331FC1E5721" blockId="1.[810,1269,487,550]" box="[810,926,524,550]" pageId="1" pageNumber="615">
(
<figureCitation id="131A2A5EFFB5FFB6FC820331FC155721" box="[818,917,524,550]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[136,235,2004,2028]" captionStartId-1="2.[136,235,886,910]" captionStartId-2="2.[136,235,2015,2039]" captionStartId-3="3.[136,235,1703,1727]" captionTargetBox-0="[315,1272,1282,1980]" captionTargetBox-1="[322,1266,174,862]" captionTargetBox-2="[323,1265,980,1992]" captionTargetBox-3="[151,1436,189,1680]" captionTargetId-0="figure-371@1.[315,1272,1282,1980]" captionTargetId-1="figure-18@2.[322,1266,174,862]" captionTargetId-2="figure-50@2.[323,1265,980,1992]" captionTargetId-3="figure-81@3.[151,1436,189,1680]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="3" captionText-0="FIGURE 1. General habitus of Coslonatus rasnitsyni gen. et sp. nov., holotype, NIGP177330, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 500 μm." captionText-1="FIGURE 2. General habitus of Coslonatus rasnitsyni gen. et sp. nov., holotype, NIGP177330, under widefield fluorescence. A, Dorsal view. B, Ventral view. Scale bars: 500 μm." captionText-2="FIGURE 3. General habitus of Coslonatus rasnitsyni gen. et sp. nov., holotype, NIGP177330, under confocal microscopy. A, Dorsal view. B, Ventral view. Scale bars: 400 μm." captionText-3="FIGURE 4. Details of Coslonatus rasnitsyni gen. et sp. nov., holotype, NIGP177330, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Elytral base, dorsal view. D, Middle portion of elytra, dorsal view. E, Antennal club, ventral view. F, Protarsus, ventral view. G, Mesotarsus, ventral view. H, Metatarsus, ventral view. Abbreviations: an1,911, antennomeres 1,911; es, elytral suture; ey, compound eye; lbp, labial palp; md, mandible; mst14, mesotarsomeres 14; mstb, mesotibia; mtt14, metatarsomeres 14; mttb, metatibia; mxp, maxillary palp; pt14, protarsomeres14;ptb, protibia; sc, scutellum.Scale bars:100 μm." figureDoi-0="http://doi.org/10.5281/zenodo.5778476" figureDoi-1="http://doi.org/10.5281/zenodo.5778478" figureDoi-2="http://doi.org/10.5281/zenodo.5778480" figureDoi-3="http://doi.org/10.5281/zenodo.5778482" httpUri-0="https://zenodo.org/record/5778476/files/figure.png" httpUri-1="https://zenodo.org/record/5778478/files/figure.png" httpUri-2="https://zenodo.org/record/5778480/files/figure.png" httpUri-3="https://zenodo.org/record/5778482/files/figure.png" pageId="1" pageNumber="615">Figs 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33B6550FFB5FFB6FC9A0368FBF35793" pageId="1" pageNumber="615" type="materials_examined">
<paragraph id="8B9E36DBFFB5FFB6FC9A0368FBF35793" blockId="1.[810,1452,597,1246]" pageId="1" pageNumber="615">
<emphasis id="B955EAC9FFB5FFB6FC9A0368FC195768" bold="true" box="[810,921,597,623]" pageId="1" pageNumber="615">Material.</emphasis>
<materialsCitation id="3B493C86FFB5FFB6FC130368FBF35793" ID-GBIF-Occurrence="3418229301" collectionCode="NIGP" pageId="1" pageNumber="615" specimenCode="NIGP177330" specimenCount="1" typeStatus="holotype">
<typeStatus id="549A8879FFB5FFB6FC130368FB8E5768" box="[931,1038,597,623]" pageId="1" pageNumber="615" type="holotype">Holotype</typeStatus>
,
<specimenCode id="DB879EA0FFB5FFB6FBAC036BFB345777" box="[1052,1204,598,624]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="1" pageNumber="615">NIGP177330</specimenCode>
, crazed and probably wrinkled during fossilisation.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B6550FFB5FFB6FCEF03A2FAD55677" pageId="1" pageNumber="615" type="etymology">
<paragraph id="8B9E36DBFFB5FFB6FCEF03A2FA2457DA" blockId="1.[810,1452,597,1246]" pageId="1" pageNumber="615">
<emphasis id="B955EAC9FFB5FFB6FCEF03A2FC6657BE" bold="true" box="[863,998,671,697]" pageId="1" pageNumber="615">Etymology.</emphasis>
The species is named for Prof.Alexander P. Rasnitsyn, a well-known Russian palaeoentomologist.
</paragraph>
<paragraph id="8B9E36DBFFB5FFB6FCEF03D5FAD55677" blockId="1.[810,1452,597,1246]" pageId="1" pageNumber="615">
<emphasis id="B955EAC9FFB5FFB6FCEF03D5FBED5605" bold="true" box="[863,1133,744,770]" pageId="1" pageNumber="615">Locality and horizon.</emphasis>
Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
<collectingRegion id="49E5F839FFB5FFB6FC9A020CFC50564C" box="[810,976,817,843]" country="Myanmar" name="Kachin" pageId="1" pageNumber="615">Kachin State</collectingRegion>
,
<collectingCountry id="F336764BFFB5FFB6FC5F020FFBE0564B" box="[1007,1120,818,844]" name="Myanmar" pageId="1" pageNumber="615">Myanmar</collectingCountry>
; unnamed horizon, mid- Cretaceous, Upper Albian to Lower Cenomanian.
</paragraph>
</subSubSection>
<subSubSection id="C33B6550FFB5FFB6FCEF0247FB1E5692" box="[863,1182,890,917]" pageId="1" pageNumber="615" type="diagnosis">
<paragraph id="8B9E36DBFFB5FFB6FCEF0247FB1E5692" blockId="1.[810,1452,597,1246]" box="[863,1182,890,917]" pageId="1" pageNumber="615">
<emphasis id="B955EAC9FFB5FFB6FCEF0247FC595693" bold="true" box="[863,985,890,916]" pageId="1" pageNumber="615">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C33B6550FFB5FFB3FCEF02A2FE29561A" lastPageId="4" lastPageNumber="618" pageId="1" pageNumber="615" type="description">
<paragraph id="8B9E36DBFFB5FFB6FCEF02A2FACA56D9" blockId="1.[810,1452,597,1246]" pageId="1" pageNumber="615">
<emphasis id="B955EAC9FFB5FFB6FCEF02A2FC7156BE" bold="true" box="[863,1009,927,953]" pageId="1" pageNumber="615">Description.</emphasis>
Body elongate, parallel-sided,
<quantity id="4CD99B3EFFB5FFB6FAF902A2FA2C56BE" box="[1353,1452,927,953]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="1" pageNumber="615" unit="mm" value="1.93">1.93 mm</quantity>
long,
<quantity id="4CD99B3EFFB5FFB6FCD902FEFC4D56D9" box="[873,973,963,990]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="1" pageNumber="615" unit="mm" value="0.54">0.54 mm</quantity>
wide, covered with distinct setae.
</paragraph>
<paragraph id="8B9E36DBFFB5FFB4FCEF02D5FB5052D2" blockId="1.[810,1452,597,1246]" lastBlockId="3.[810,1451,1943,2077]" lastPageId="3" lastPageNumber="617" pageId="1" pageNumber="615">
Head (
<figureCitation id="131A2A5EFFB5FFB6FC1702D5FB9A5104" box="[935,1050,1000,1027]" captionStart="FIGURE 4" captionStartId="3.[136,235,1703,1727]" captionTargetBox="[151,1436,189,1680]" captionTargetId="figure-81@3.[151,1436,189,1680]" captionTargetPageId="3" captionText="FIGURE 4. Details of Coslonatus rasnitsyni gen. et sp. nov., holotype, NIGP177330, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Elytral base, dorsal view. D, Middle portion of elytra, dorsal view. E, Antennal club, ventral view. F, Protarsus, ventral view. G, Mesotarsus, ventral view. H, Metatarsus, ventral view. Abbreviations: an1,911, antennomeres 1,911; es, elytral suture; ey, compound eye; lbp, labial palp; md, mandible; mst14, mesotarsomeres 14; mstb, mesotibia; mtt14, metatarsomeres 14; mttb, metatibia; mxp, maxillary palp; pt14, protarsomeres14;ptb, protibia; sc, scutellum.Scale bars:100 μm." figureDoi="http://doi.org/10.5281/zenodo.5778482" httpUri="https://zenodo.org/record/5778482/files/figure.png" pageId="1" pageNumber="615">Fig. 4A, B</figureCitation>
) prognathous, without distinct neck; dorsal surface with a pair of longitudinal protuberances. Compound eyes lateral, entire, with short interfacetal setae. Antennal insertions dorsally located (
<figureCitation id="131A2A5EFFB5FFB6FA8C056BFA1F5177" box="[1340,1439,1110,1136]" captionStart="FIGURE 4" captionStartId="3.[136,235,1703,1727]" captionTargetBox="[151,1436,189,1680]" captionTargetId="figure-81@3.[151,1436,189,1680]" captionTargetPageId="3" captionText="FIGURE 4. Details of Coslonatus rasnitsyni gen. et sp. nov., holotype, NIGP177330, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Elytral base, dorsal view. D, Middle portion of elytra, dorsal view. E, Antennal club, ventral view. F, Protarsus, ventral view. G, Mesotarsus, ventral view. H, Metatarsus, ventral view. Abbreviations: an1,911, antennomeres 1,911; es, elytral suture; ey, compound eye; lbp, labial palp; md, mandible; mst14, mesotarsomeres 14; mstb, mesotibia; mtt14, metatarsomeres 14; mttb, metatibia; mxp, maxillary palp; pt14, protarsomeres14;ptb, protibia; sc, scutellum.Scale bars:100 μm." figureDoi="http://doi.org/10.5281/zenodo.5778482" httpUri="https://zenodo.org/record/5778482/files/figure.png" pageId="1" pageNumber="615">Fig. 4A</figureCitation>
), separated by less than twice diameter of antennomere 1. Antennae 11-segmented, long, almost reaching elytral apex when posteriorly directed; antennomere 1 long and wide; antennomere 2 shorter; antennomere 38 elongate; antennomeres 911 short, asymmetrical (serrate in 9 10), forming a club (
<figureCitation id="131A2A5EFFB7FFB4FE3006E2FE5C52FE" box="[384,476,2015,2041]" captionStart="FIGURE 4" captionStartId="3.[136,235,1703,1727]" captionTargetBox="[151,1436,189,1680]" captionTargetId="figure-81@3.[151,1436,189,1680]" captionTargetPageId="3" captionText="FIGURE 4. Details of Coslonatus rasnitsyni gen. et sp. nov., holotype, NIGP177330, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Elytral base, dorsal view. D, Middle portion of elytra, dorsal view. E, Antennal club, ventral view. F, Protarsus, ventral view. G, Mesotarsus, ventral view. H, Metatarsus, ventral view. Abbreviations: an1,911, antennomeres 1,911; es, elytral suture; ey, compound eye; lbp, labial palp; md, mandible; mst14, mesotarsomeres 14; mstb, mesotibia; mtt14, metatarsomeres 14; mttb, metatibia; mxp, maxillary palp; pt14, protarsomeres14;ptb, protibia; sc, scutellum.Scale bars:100 μm." figureDoi="http://doi.org/10.5281/zenodo.5778482" httpUri="https://zenodo.org/record/5778482/files/figure.png" pageId="3" pageNumber="617">Fig. 4E</figureCitation>
). Mandibles apically with two vertically arranged teeth (
<figureCitation id="131A2A5EFFB7FFB4FE54093EFDBF5D1A" box="[484,575,2051,2077]" captionStart="FIGURE 4" captionStartId="3.[136,235,1703,1727]" captionTargetBox="[151,1436,189,1680]" captionTargetId="figure-81@3.[151,1436,189,1680]" captionTargetPageId="3" captionText="FIGURE 4. Details of Coslonatus rasnitsyni gen. et sp. nov., holotype, NIGP177330, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Elytral base, dorsal view. D, Middle portion of elytra, dorsal view. E, Antennal club, ventral view. F, Protarsus, ventral view. G, Mesotarsus, ventral view. H, Metatarsus, ventral view. Abbreviations: an1,911, antennomeres 1,911; es, elytral suture; ey, compound eye; lbp, labial palp; md, mandible; mst14, mesotarsomeres 14; mstb, mesotibia; mtt14, metatarsomeres 14; mttb, metatibia; mxp, maxillary palp; pt14, protarsomeres14;ptb, protibia; sc, scutellum.Scale bars:100 μm." figureDoi="http://doi.org/10.5281/zenodo.5778482" httpUri="https://zenodo.org/record/5778482/files/figure.png" pageId="3" pageNumber="617">Fig. 4B</figureCitation>
). Maxillary palps relatively long; apical palpomere fusiform. Labial palps present; apical palpomere cylindrical.
</paragraph>
<caption id="DF5E6653FFB5FFB6FF3806E9FDB75D17" ID-DOI="http://doi.org/10.5281/zenodo.5778476" ID-Zenodo-Dep="5778476" httpUri="https://zenodo.org/record/5778476/files/figure.png" pageId="1" pageNumber="615" startId="1.[136,235,2004,2028]" targetBox="[315,1272,1282,1980]" targetPageId="1">
<paragraph id="8B9E36DBFFB5FFB6FF3806E9FDB75D17" blockId="1.[136,1452,2004,2064]" pageId="1" pageNumber="615">
<emphasis id="B955EAC9FFB5FFB6FF3806E9FE8652EB" bold="true" box="[136,262,2004,2028]" pageId="1" pageNumber="615">FIGURE 1.</emphasis>
General habitus of
<taxonomicName id="4C214D58FFB5FFB6FE6606E9FD3552EB" authority="Li &amp; Huang &amp; Cai, 2021" authorityName="Li &amp; Huang &amp; Cai" authorityYear="2021" box="[470,693,2004,2028]" class="Insecta" family="Zopheridae" genus="Coslonatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="615" phylum="Arthropoda" rank="species" species="rasnitsyni" status="gen. et sp. nov.">
<emphasis id="B955EAC9FFB5FFB6FE6606E9FD3552EB" box="[470,693,2004,2028]" italics="true" pageId="1" pageNumber="615">Coslonatus rasnitsyni</emphasis>
</taxonomicName>
<emphasis id="B955EAC9FFB5FFB6FD0D06E9FCE152EB" bold="true" box="[701,865,2004,2028]" pageId="1" pageNumber="615">
<taxonomicNameLabel id="A26657B2FFB5FFB6FD0D06E9FCE152EB" box="[701,865,2004,2028]" pageId="1" pageNumber="615" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP177330, under incident light.
<emphasis id="B955EAC9FFB5FFB6FAF706E9FAD952EB" bold="true" box="[1351,1369,2004,2028]" pageId="1" pageNumber="615">A</emphasis>
, Dorsal view.
<emphasis id="B955EAC9FFB5FFB6FF7406C5FF555D17" bold="true" box="[196,213,2040,2064]" pageId="1" pageNumber="615">B</emphasis>
, Ventral view. Scale bars: 500 μm.
</paragraph>
</caption>
<caption id="DF5E6653FFB6FFB5FF38024BFD0156B4" ID-DOI="http://doi.org/10.5281/zenodo.5778478" ID-Zenodo-Dep="5778478" httpUri="https://zenodo.org/record/5778478/files/figure.png" pageId="2" pageNumber="616" startId="2.[136,235,886,910]" targetBox="[322,1266,174,862]" targetPageId="2">
<paragraph id="8B9E36DBFFB6FFB5FF38024BFD0156B4" blockId="2.[136,1451,886,947]" pageId="2" pageNumber="616">
<emphasis id="B955EAC9FFB6FFB5FF38024BFE855689" bold="true" box="[136,261,886,910]" pageId="2" pageNumber="616">FIGURE 2.</emphasis>
General habitus of
<taxonomicName id="4C214D58FFB6FFB5FE60024BFD2D5689" authority="Li &amp; Huang &amp; Cai, 2021" authorityName="Li &amp; Huang &amp; Cai" authorityYear="2021" box="[464,685,886,910]" class="Insecta" family="Zopheridae" genus="Coslonatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="616" phylum="Arthropoda" rank="species" species="rasnitsyni" status="gen. et sp. nov.">
<emphasis id="B955EAC9FFB6FFB5FE60024BFD2D5689" box="[464,685,886,910]" italics="true" pageId="2" pageNumber="616">Coslonatus rasnitsyni</emphasis>
</taxonomicName>
<emphasis id="B955EAC9FFB6FFB5FD05024AFCD45688" bold="true" box="[693,852,887,911]" pageId="2" pageNumber="616">
<taxonomicNameLabel id="A26657B2FFB6FFB5FD05024AFCD45688" box="[693,852,887,911]" pageId="2" pageNumber="616" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP177330, under widefield fluorescence.
<emphasis id="B955EAC9FFB6FFB5FA23024BFA255689" bold="true" box="[1427,1445,886,910]" pageId="2" pageNumber="616">A</emphasis>
, Dorsal view.
<emphasis id="B955EAC9FFB6FFB5FEBE02A6FE9F56B4" bold="true" box="[270,287,923,947]" pageId="2" pageNumber="616">B</emphasis>
, Ventral view. Scale bars: 500 μm.
</paragraph>
</caption>
<caption id="DF5E6653FFB6FFB5FF3806E2FD015D1B" ID-DOI="http://doi.org/10.5281/zenodo.5778480" ID-Zenodo-Dep="5778480" httpUri="https://zenodo.org/record/5778480/files/figure.png" pageId="2" pageNumber="616" startId="2.[136,235,2015,2039]" targetBox="[323,1265,980,1992]" targetPageId="2">
<paragraph id="8B9E36DBFFB6FFB5FF3806E2FD015D1B" blockId="2.[136,1451,2015,2076]" pageId="2" pageNumber="616">
<emphasis id="B955EAC9FFB6FFB5FF3806E2FE8652FF" bold="true" box="[136,262,2015,2040]" pageId="2" pageNumber="616">FIGURE 3.</emphasis>
General habitus of
<taxonomicName id="4C214D58FFB6FFB5FE6606DDFD3552F0" authority="Li &amp; Huang &amp; Cai, 2021" authorityName="Li &amp; Huang &amp; Cai" authorityYear="2021" box="[470,693,2016,2040]" class="Insecta" family="Zopheridae" genus="Coslonatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="616" phylum="Arthropoda" rank="species" species="rasnitsyni" status="gen. et sp. nov.">
<emphasis id="B955EAC9FFB6FFB5FE6606DDFD3552F0" box="[470,693,2016,2040]" italics="true" pageId="2" pageNumber="616">Coslonatus rasnitsyni</emphasis>
</taxonomicName>
<emphasis id="B955EAC9FFB6FFB5FD0D06DDFCE152FF" bold="true" box="[701,865,2016,2040]" pageId="2" pageNumber="616">
<taxonomicNameLabel id="A26657B2FFB6FFB5FD0D06DDFCE152FF" box="[701,865,2016,2040]" pageId="2" pageNumber="616" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP177330, under confocal microscopy.
<emphasis id="B955EAC9FFB6FFB5FA2306E2FA2552F0" bold="true" box="[1427,1445,2015,2039]" pageId="2" pageNumber="616">A</emphasis>
, Dorsal view.
<emphasis id="B955EAC9FFB6FFB5FEBE0939FE9F5D1B" bold="true" box="[270,287,2052,2076]" pageId="2" pageNumber="616">B</emphasis>
, Ventral view. Scale bars: 400 μm.
</paragraph>
</caption>
<caption id="DF5E6653FFB7FFB4FF38079AFA2C5257" ID-DOI="http://doi.org/10.5281/zenodo.5778482" ID-Zenodo-Dep="5778482" httpUri="https://zenodo.org/record/5778482/files/figure.png" pageId="3" pageNumber="617" startId="3.[136,235,1703,1727]" targetBox="[151,1436,189,1680]" targetPageId="3">
<paragraph id="8B9E36DBFFB7FFB4FF38079AFA2C5257" blockId="3.[136,1452,1703,1872]" pageId="3" pageNumber="617">
<emphasis id="B955EAC9FFB7FFB4FF38079AFE8353C7" bold="true" box="[136,259,1703,1728]" pageId="3" pageNumber="617">FIGURE 4.</emphasis>
Details of
<taxonomicName id="4C214D58FFB7FFB4FEC10795FDCD53B8" authority="Li &amp; Huang &amp; Cai, 2021" authorityName="Li &amp; Huang &amp; Cai" authorityYear="2021" box="[369,589,1704,1728]" class="Insecta" family="Zopheridae" genus="Coslonatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="617" phylum="Arthropoda" rank="species" species="rasnitsyni" status="gen. et sp. nov.">
<emphasis id="B955EAC9FFB7FFB4FEC10795FDCD53B8" box="[369,589,1704,1728]" italics="true" pageId="3" pageNumber="617">Coslonatus rasnitsyni</emphasis>
</taxonomicName>
<emphasis id="B955EAC9FFB7FFB4FDE20795FD6D53C7" bold="true" box="[594,749,1704,1728]" pageId="3" pageNumber="617">
<taxonomicNameLabel id="A26657B2FFB7FFB4FDE20795FD6D53C7" box="[594,749,1704,1728]" pageId="3" pageNumber="617" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP177330, under confocal microscopy.
<emphasis id="B955EAC9FFB7FFB4FABE079AFAA053B8" bold="true" box="[1294,1312,1703,1727]" pageId="3" pageNumber="617">A</emphasis>
, Head, dorsal view.
<emphasis id="B955EAC9FFB7FFB4FF7407F1FF5553E3" bold="true" box="[196,213,1740,1764]" pageId="3" pageNumber="617">B</emphasis>
, Mouthparts, ventral view.
<emphasis id="B955EAC9FFB7FFB4FE5B07F6FE7D53E4" bold="true" box="[491,509,1739,1763]" pageId="3" pageNumber="617">C</emphasis>
, Elytral base, dorsal view.
<emphasis id="B955EAC9FFB7FFB4FCBF07F1FCA153E3" bold="true" box="[783,801,1740,1764]" pageId="3" pageNumber="617">D</emphasis>
, Middle portion of elytra, dorsal view.
<emphasis id="B955EAC9FFB7FFB4FB1E07F1FB3F53E3" bold="true" box="[1198,1215,1740,1764]" pageId="3" pageNumber="617">E</emphasis>
, Antennal club, ventral view.
<emphasis id="B955EAC9FFB7FFB4FF7407CDFF53520F" bold="true" box="[196,211,1776,1800]" pageId="3" pageNumber="617">F</emphasis>
, Protarsus, ventral view.
<emphasis id="B955EAC9FFB7FFB4FE6107D2FE655200" bold="true" box="[465,485,1775,1799]" pageId="3" pageNumber="617">G</emphasis>
, Mesotarsus, ventral view.
<emphasis id="B955EAC9FFB7FFB4FD4807CDFC8C520F" bold="true" box="[760,780,1776,1800]" pageId="3" pageNumber="617">H</emphasis>
, Metatarsus, ventral view. Abbreviations: an1,911, antennomeres 1,911; es, elytral suture; ey, compound eye; lbp, labial palp; md, mandible; mst14, mesotarsomeres 14; mstb, mesotibia; mtt14, metatarsomeres 14; mttb, metatibia; mxp, maxillary palp; pt14, protarsomeres 14; ptb, protibia; sc, scutellum. Scale bars: 100 μm.
</paragraph>
</caption>
<paragraph id="8B9E36DBFFB7FFB3FCEF06E2FD0B544E" blockId="3.[810,1451,1943,2077]" lastBlockId="4.[136,778,158,797]" lastPageId="4" lastPageNumber="618" pageId="3" pageNumber="617">Pronotal disc quadrate; lateral margins straight, denticulate; anterior and posterior angles right- angled; surface with a pair of longitudinal elevated platforms. Hypomera without antennal grooves or impressions. Notosternal sutures not evident, probably absent. Prosternal process raised. Procoxae seemingly countersunk, concealed by prosternal process.</paragraph>
<paragraph id="8B9E36DBFFB0FFB3FF0D006FFF64576E" blockId="4.[136,778,158,797]" pageId="4" pageNumber="618">
Scutellar shield(
<figureCitation id="131A2A5EFFB0FFB3FEC4006FFE46546B" box="[372,454,338,364]" captionStart="FIGURE 4" captionStartId="3.[136,235,1703,1727]" captionTargetBox="[151,1436,189,1680]" captionTargetId="figure-81@3.[151,1436,189,1680]" captionTargetPageId="3" captionText="FIGURE 4. Details of Coslonatus rasnitsyni gen. et sp. nov., holotype, NIGP177330, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Elytral base, dorsal view. D, Middle portion of elytra, dorsal view. E, Antennal club, ventral view. F, Protarsus, ventral view. G, Mesotarsus, ventral view. H, Metatarsus, ventral view. Abbreviations: an1,911, antennomeres 1,911; es, elytral suture; ey, compound eye; lbp, labial palp; md, mandible; mst14, mesotarsomeres 14; mstb, mesotibia; mtt14, metatarsomeres 14; mttb, metatibia; mxp, maxillary palp; pt14, protarsomeres14;ptb, protibia; sc, scutellum.Scale bars:100 μm." figureDoi="http://doi.org/10.5281/zenodo.5778482" httpUri="https://zenodo.org/record/5778482/files/figure.png" pageId="4" pageNumber="618">Fig. 4C</figureCitation>
) with anterior margin slightly emarginate; posterior margin rounded. Elytra elongate, parallel-sided; each with five subtle carinae (including sutural carina); scutellar striole present. Mesocoxae longer than wide; separated by approximately width of mesocoxae. Mesocoxal cavities not evident, probably closed laterally. Metacoxae separated by intercoxal process.
</paragraph>
<paragraph id="8B9E36DBFFB0FFB3FF0D034FFD9B57D3" blockId="4.[136,778,158,797]" pageId="4" pageNumber="618">
Tibiae with two spurs. Tarsi 4-4-4 (
<figureCitation id="131A2A5EFFB0FFB3FDCC034FFD7D578B" box="[636,765,626,652]" captionStart="FIGURE 4" captionStartId="3.[136,235,1703,1727]" captionTargetBox="[151,1436,189,1680]" captionTargetId="figure-81@3.[151,1436,189,1680]" captionTargetPageId="3" captionText="FIGURE 4. Details of Coslonatus rasnitsyni gen. et sp. nov., holotype, NIGP177330, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Elytral base, dorsal view. D, Middle portion of elytra, dorsal view. E, Antennal club, ventral view. F, Protarsus, ventral view. G, Mesotarsus, ventral view. H, Metatarsus, ventral view. Abbreviations: an1,911, antennomeres 1,911; es, elytral suture; ey, compound eye; lbp, labial palp; md, mandible; mst14, mesotarsomeres 14; mstb, mesotibia; mtt14, metatarsomeres 14; mttb, metatibia; mxp, maxillary palp; pt14, protarsomeres14;ptb, protibia; sc, scutellum.Scale bars:100 μm." figureDoi="http://doi.org/10.5281/zenodo.5778482" httpUri="https://zenodo.org/record/5778482/files/figure.png" pageId="4" pageNumber="618">Fig. 4FH</figureCitation>
); tarsomeres 13 similar in form, together shorter than tarsomere 4. Pretarsal claws simple.
</paragraph>
<paragraph id="8B9E36DBFFB0FFB3FF0D03E3FE29561A" blockId="4.[136,778,158,797]" pageId="4" pageNumber="618">Abdomen with five ventrites, separated by wide and deeply impressed sutures.</paragraph>
</subSubSection>
</treatment>
</document>