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<document id="49AF030354ABDBA79FAD5225C76AC523" ID-DOI="10.15298/rusentj.28.3.04" ID-ISSN="0132-8069" ID-Zenodo-Dep="13164041" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722579516313" checkinUser="felipe" docAuthor="Balke, M., Garcia, M. &amp; Hendrich, L." docDate="2019" docId="03B0879DFFE8FF8FAD1637AC84850906" docLanguage="en" docName="RussEntomolJ.28.3.258-262.pdf" docOrigin="Russian Entomological Journal 28 (3)" docSource="http://dx.doi.org/10.15298/rusentj.28.3.04" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.2:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="2" docTitle="Rhantus andinus Balke 1998" docType="treatment" docVersion="2" lastPageNumber="262" masterDocId="FF89FFE5FFECFF8BAC543572863D0E17" masterDocTitle="Two new species of diving beetles from the Duida-Marahuaca Tepui in Venezuela (Coleoptera: Dytiscidae: Colymbetinae)" masterLastPageNumber="262" masterPageNumber="258" pageNumber="262" updateTime="1722581003254" updateUser="ExternalLinkService">
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<mods:title id="FA1647723B4BE2CEDC3480B9CB0C45A6">Two new species of diving beetles from the Duida-Marahuaca Tepui in Venezuela (Coleoptera: Dytiscidae: Colymbetinae)</mods:title>
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<mods:namePart id="FE1A853FF0DA4C10FB50FCE31B498C33">Balke, M.</mods:namePart>
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<mods:namePart id="99653637E0B018481AF69AA9B3DA05C0">Garcia, M.</mods:namePart>
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<mods:date id="102F3D8622F9DCAEC7B7599BA7B4C36F">2019</mods:date>
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<taxonomicName id="4C194D08FFE8FF8FAD1637AC84BD0CE1" ID-CoL="4S3LN" authority="Balke, 1998" authorityName="Balke" authorityYear="1998" box="[322,640,734,758]" class="Insecta" family="Dytiscidae" genus="Rhantus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="262" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B96DEA99FFE8FF8FAD1637AC87CA0CE1" box="[322,503,734,758]" italics="true" pageId="4" pageNumber="262">Rhantus andinus</emphasis>
<bibRefCitation id="EF884B7AFFE8FF8FADAA37AC84BD0CE1" author="Balke M." box="[510,640,734,758]" pageId="4" pageNumber="262" pagination="71 - 79" refId="ref3689" refString="Balke M. 1998. Updating the Pacific, Indomalayan and Neotropical Rhantus - fauna Coleoptera: Dytiscidae) // Koleopterologische Rundschau. Vol. 68. P. 71 - 79." type="journal article" year="1998">Balke, 1998</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3036500FFE8FF8FADFB378E84290D03" box="[431,532,764,788]" pageId="4" pageNumber="262" type="description">
<paragraph id="8BA6368BFFE8FF8FADFB378E84290D03" blockId="4.[189,775,734,1484]" box="[431,532,764,788]" pageId="4" pageNumber="262">
<figureCitation id="13222A0EFFE8FF8FADFB378E842D0D03" box="[431,528,764,788]" captionStart="Figs 15" captionStartId="1.[226,263,1969,1988]" captionTargetBox="[194,1391,1457,1935]" captionTargetId="figure-673@1.[189,1399,1448,1948]" captionTargetPageId="1" captionText="Figs 15. Habitus of Rhantus spp.: 1 — R. lattkei sp.n.; 2 — R. marahuaca sp.n.; 3 — R. elegans; 4 — R. andinus (Colombia, Bogota, La Florida); 5 — R. andinus (Venezuela, Merida, Azulita). Рис. 15. Rhantus spp., внешний вид: 1 — R. lattkei sp.n.; 2 — R. marahuaca sp.n.; 3 — R. elegans; 4 — R. andinus (Colombia, Bogota, La Florida); 5 — R. andinus (Venezuela, Merida, Azulita)." figureDoi="http://doi.org/10.5281/zenodo.13164043" httpUri="https://zenodo.org/record/13164043/files/figure.png" pageId="4" pageNumber="262">Figs 45</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FFE8FF8FACB6365384CE0D23" box="[226,755,801,820]" pageId="4" pageNumber="262" type="reference_group">
<paragraph id="8BA6368BFFE8FF8FACB6365384CE0D23" blockId="4.[189,775,734,1484]" box="[226,755,801,820]" pageId="4" pageNumber="262">
<treatmentCitationGroup id="AB0911A5FFE8FF8FACB6365384CE0D23" box="[226,755,801,820]" pageId="4" pageNumber="262">
<treatmentCitation id="0AB8109AFFE8FF8FACB6365384360D23" author="Balke M." box="[226,523,801,820]" page="73" pageId="4" pageNumber="262" year="1998">
<taxonomicName id="4C194D08FFE8FF8FACB6365384360D23" ID-CoL="4S3LN" authority="Balke, 1998: 73" authorityName="Balke" authorityPageNumber="73" authorityYear="1998" box="[226,523,801,820]" class="Insecta" family="Dytiscidae" genus="Rhantus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="262" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B96DEA99FFE8FF8FACB6365387490D23" box="[226,372,801,820]" italics="true" pageId="4" pageNumber="262">Rhantus andinus</emphasis>
<bibRefCitation id="EF884B7AFFE8FF8FAD2F365384360D23" author="Balke M." box="[379,523,801,820]" pageId="4" pageNumber="262" pagination="71 - 79" refId="ref3689" refString="Balke M. 1998. Updating the Pacific, Indomalayan and Neotropical Rhantus - fauna Coleoptera: Dytiscidae) // Koleopterologische Rundschau. Vol. 68. P. 71 - 79." type="journal article" year="1998">Balke, 1998: 73</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AB8109AFFE8FF8FAE41365384D20D23" author="Nilsson A. N. &amp; Hajek J." box="[533,751,801,820]" page="41" pageId="4" pageNumber="262" year="2019">
<bibRefCitation id="EF884B7AFFE8FF8FAE41365384D20D23" author="Nilsson A. N. &amp; Hajek J." box="[533,751,801,820]" pageId="4" pageNumber="262" refId="ref4116" refString="Nilsson A. N., Hajek J. 2019. A world catalogue of the family Dytiscidae (Coleoptera, Adephaga). 307 pp. Internet version 1. I. 2019. Available from: http: // www. waterbeetles. eu." type="book" year="2019">Nilsson, Hajek 2019: 41</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FFE8FF8FACB6364887300BDB" pageId="4" pageNumber="262" type="materials_examined">
<paragraph id="8BA6368BFFE8FF8FACB63648846F0D5A" blockId="4.[189,775,734,1484]" box="[226,594,826,845]" pageId="4" pageNumber="262">
<materialsCitation id="3B713CD6FFE8FF8FACB6364884730D5A" box="[226,590,826,845]" collectionCode="LOCALITY" country="Colombia" location="Bogota" pageId="4" pageNumber="262" specimenCount="1" stateProvince="Bogota" typeStatus="holotype">
<typeStatus id="54A28829FFE8FF8FACB6364887250D5A" box="[226,280,826,845]" pageId="4" pageNumber="262">TYPE</typeStatus>
<collectionCode id="ED08AE4EFFE8FF8FAD4A364887B40D5A" box="[286,393,826,845]" pageId="4" pageNumber="262">LOCALITY</collectionCode>
<collectingCountry id="F30E761BFFE8FF8FADC4364887DA0D5A" box="[400,487,826,845]" name="Colombia" pageId="4" pageNumber="262">Colombia</collectingCountry>
,
<collectingRegion id="49DDF869FFE8FF8FADA4364884130D5A" box="[496,558,826,845]" country="Colombia" name="Bogota" pageId="4" pageNumber="262">Bogotá</collectingRegion>
[?]
</materialsCitation>
.
</paragraph>
<paragraph id="8BA6368BFFE8FF8FACB6362087350DD0" blockId="4.[189,775,734,1484]" pageId="4" pageNumber="262">
MATERIAL.
<materialsCitation id="3B713CD6FFE8FF8FAD093620874C0D81" collectingDate="1979-07-13" collectionCode="MIZA" collectorName="B. Bechyne" country="Venezuela" location="Azulita" municipality="Estado" pageId="4" pageNumber="262" specimenCount="1" specimenCount-male="1" stateProvince="Merida">
<specimenCount id="9D1FFD02FFE8FF8FAD09362087BF0D71" box="[349,386,848,870]" count="1" pageId="4" pageNumber="262" type="male">1 ♂</specimenCount>
(
<figureCitation id="13222A0EFFE8FF8FADDA362087FE0D72" box="[398,451,850,869]" captionStart="Figs 15" captionStartId="1.[226,263,1969,1988]" captionTargetBox="[194,1391,1457,1935]" captionTargetId="figure-673@1.[189,1399,1448,1948]" captionTargetPageId="1" captionText="Figs 15. Habitus of Rhantus spp.: 1 — R. lattkei sp.n.; 2 — R. marahuaca sp.n.; 3 — R. elegans; 4 — R. andinus (Colombia, Bogota, La Florida); 5 — R. andinus (Venezuela, Merida, Azulita). Рис. 15. Rhantus spp., внешний вид: 1 — R. lattkei sp.n.; 2 — R. marahuaca sp.n.; 3 — R. elegans; 4 — R. andinus (Colombia, Bogota, La Florida); 5 — R. andinus (Venezuela, Merida, Azulita)." figureDoi="http://doi.org/10.5281/zenodo.13164043" httpUri="https://zenodo.org/record/13164043/files/figure.png" pageId="4" pageNumber="262">Fig. 5</figureCitation>
),
<collectingCountry id="F30E761BFFE8FF8FAD813620840E0D72" box="[469,563,850,869]" name="Venezuela" pageId="4" pageNumber="262">Venezuela</collectingCountry>
:
<collectingMunicipality id="6BC2ACF1FFE8FF8FAE683620844A0D72" box="[572,631,850,869]" pageId="4" pageNumber="262">Estado</collectingMunicipality>
<collectingRegion id="49DDF869FFE8FF8FAE29362084830D72" box="[637,702,850,869]" country="Venezuela" name="Merida" pageId="4" pageNumber="262">Merida</collectingRegion>
,
<location id="8EC66050FFE8FF8FAE933620853B0D72" LSID="urn:lsid:plazi:treatment:03B0879DFFE8FF8FAD1637AC84850906:8EC66050FFE8FF8FAE933620853B0D72" box="[711,774,850,869]" country="Venezuela" municipality="Estado" name="Azulita" pageId="4" pageNumber="262" stateProvince="Merida">Azulita</location>
(Universidad de los Andes agricultural research station),
<date id="FFA7104BFFE8FF8FAECB3619853E0D69" box="[671,771,875,894]" pageId="4" pageNumber="262" value="1979-07-13">
<collectingDate id="EFE3E9A3FFE8FF8FAECB3619853E0D69" box="[671,771,875,894]" pageId="4" pageNumber="262" value="1979-07-13">13.vii.1979</collectingDate>
</date>
,
<collectorName id="26EC535DFFE8FF8FACE936F6871F0D81" box="[189,290,899,919]" pageId="4" pageNumber="262">B. Bechyne</collectorName>
(
<collectionCode id="ED08AE4EFFE8FF8FAD7B36F187560D81" box="[303,363,899,918]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="4" pageNumber="262" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CD6FFE8FF8FAD2F36F187390DB8" collectingDate="1976-08-18" collectionCode="MIZA" collectorName="El Joque &amp; I. Jaspe" country="Venezuela" location="El Joque" pageId="4" pageNumber="262" specimenCount="1" specimenCount-male="1" stateProvince="Merida">
<specimenCount id="9D1FFD02FFE8FF8FAD2F36F1879C0D80" box="[379,417,897,919]" count="1" pageId="4" pageNumber="262" type="male">1 ♂</specimenCount>
,
<collectingRegion id="49DDF869FFE8FF8FADF936F187D30D81" box="[429,494,899,918]" country="Venezuela" name="Merida" pageId="4" pageNumber="262">Merida</collectingRegion>
,
<collectorName id="26EC535DFFE8FF8FADA336F1847B0D80" box="[503,582,899,919]" pageId="4" pageNumber="262">
<location id="8EC66050FFE8FF8FADA336F1847B0D80" LSID="urn:lsid:plazi:treatment:03B0879DFFE8FF8FAD1637AC84850906:8EC66050FFE8FF8FADA336F1847B0D80" box="[503,582,899,919]" country="Venezuela" name="El Joque" pageId="4" pageNumber="262" stateProvince="Merida">El Joque</location>
</collectorName>
,
<date id="FFA7104BFFE8FF8FAE0436F184870D81" box="[592,698,899,918]" pageId="4" pageNumber="262" value="1976-08-18">
<collectingDate id="EFE3E9A3FFE8FF8FAE0436F184870D81" box="[592,698,899,918]" pageId="4" pageNumber="262" value="1976-08-18">18.viii.1976</collectingDate>
</date>
,
<collectorName id="26EC535DFFE8FF8FAE9036F6853A0D80" box="[708,775,900,919]" pageId="4" pageNumber="262">I. Jaspe</collectorName>
(
<collectionCode id="ED08AE4EFFE8FF8FAC9736EE86C20DB8" box="[195,255,924,943]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="4" pageNumber="262" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CD6FFE8FF8FAD4536EE87390DD0" collectingDate="1968-09-08" collectionCode="MIZA" collectorName="J. &amp; B. Bechyne" country="Venezuela" location="Sanare" municipality="Estado" pageId="4" pageNumber="262" specimenCount="1" specimenCount-male="1" stateProvince="Lara">
<specimenCount id="9D1FFD02FFE8FF8FAD4536EE87040DB8" box="[273,313,921,943]" count="1" pageId="4" pageNumber="262" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="6BC2ACF1FFE8FF8FAD1C36EE87BE0DB8" box="[328,387,924,943]" pageId="4" pageNumber="262">Estado</collectingMunicipality>
<collectingRegion id="49DDF869FFE8FF8FADD836EE878B0DB8" box="[396,438,924,943]" country="Venezuela" name="Lara" pageId="4" pageNumber="262">Lara</collectingRegion>
,
<location id="8EC66050FFE8FF8FAD9736EE843D0DB8" LSID="urn:lsid:plazi:treatment:03B0879DFFE8FF8FAD1637AC84850906:8EC66050FFE8FF8FAD9736EE843D0DB8" box="[451,512,924,943]" country="Venezuela" municipality="Estado" name="Sanare" pageId="4" pageNumber="262" stateProvince="Lara">Sanare</location>
,
<date id="FFA7104BFFE8FF8FAE5836EE84620DB8" box="[524,607,924,943]" pageId="4" pageNumber="262" value="1968-09-08">
<collectingDate id="EFE3E9A3FFE8FF8FAE5836EE84620DB8" box="[524,607,924,943]" pageId="4" pageNumber="262" value="1968-09-08">8.ix.1968</collectingDate>
</date>
,
<collectorName id="26EC535DFFE8FF8FAE3936EE853B0DB8" box="[621,774,924,943]" pageId="4" pageNumber="262">J. &amp; B. Bechyne</collectorName>
(
<collectionCode id="ED08AE4EFFE8FF8FAC9736C686C20DD0" box="[195,255,948,967]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="4" pageNumber="262" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA6368BFFE8FF8FACB636BF87300BDB" blockId="4.[189,775,734,1484]" pageId="4" pageNumber="262">
<collectionCode id="ED08AE4EFFE8FF8FACB636BF87080DF5" box="[226,309,973,994]" pageId="4" pageNumber="262">NOTES</collectionCode>
. This is the first record for
<collectingCountry id="F30E761BFFE8FF8FAE0036BF84FC0DF4" box="[596,705,973,995]" name="Venezuela" pageId="4" pageNumber="262">Venezuela</collectingCountry>
.
<bibRefCitation id="EF884B7AFFE8FF8FAE9836BF86C20DEA" author="Balke M." pageId="4" pageNumber="262" pagination="71 - 79" refId="ref3689" refString="Balke M. 1998. Updating the Pacific, Indomalayan and Neotropical Rhantus - fauna Coleoptera: Dytiscidae) // Koleopterologische Rundschau. Vol. 68. P. 71 - 79." type="journal article" year="1998">Balke [1998]</bibRefCitation>
observed: “
<specimenCount id="9D1FFD02FFE8FF8FADD6369A87C00DE9" box="[386,509,1000,1022]" count="1" pageId="4" pageNumber="262" type="female">One female</specimenCount>
may also belong to this species:
<collectingRegion id="49DDF869FFE8FF8FAD433171875B0A0E" box="[279,358,1027,1049]" country="Venezuela" name="Merida" pageId="4" pageNumber="262">Merida</collectingRegion>
, Venezuela /
<taxonomicName id="4C194D08FFE8FF8FADAE3171846C0A0E" authorityName="Balke &amp; Garcia &amp; Hendrich" authorityYear="2019" box="[506,593,1027,1049]" class="Insecta" family="Dytiscidae" genus="Rhantus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="262" phylum="Arthropoda" rank="genus" status="n.sp.">Rhantus</taxonomicName>
<taxonomicNameLabel id="A25E57E2FFE8FF8FAE0C317184B60A0F" box="[600,651,1027,1048]" pageId="4" pageNumber="262" rank="species">n.sp.</taxonomicNameLabel>
[hand writing M. Régimbart] (
<collectionCode id="ED08AE4EFFE8FF8FADDF316C87E00A24" box="[395,477,1054,1075]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="262">MNHN</collectionCode>
). This specimen differs from the
<typeStatus id="54A28829FFE8FF8FACB6314887300A58" box="[226,269,1082,1103]" pageId="4" pageNumber="262">type</typeStatus>
by: 1) Elytra rather dark due to fusion of most of the black speckles; 2) dorsal surface, especially elytra, matt because the meshes are deeply impressed; 3) base of pronotum distinctly narrower than base of elytra; 4) metasternal wings slightly narrower. This individual may represent a high altitude form of
<taxonomicName id="4C194D08FFE8FF8FAE4731B084420AC1" authorityName="Balke" authorityYear="1998" box="[531,639,1217,1239]" class="Insecta" family="Dytiscidae" genus="Rhantus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="262" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B96DEA99FFE8FF8FAE4731B084420AC1" box="[531,639,1217,1239]" italics="true" pageId="4" pageNumber="262">R. andinus</emphasis>
</taxonomicName>
, or a closely related species.” The
<specimenCount id="9D1FFD02FFE8FF8FADC631AF84390AE4" box="[402,516,1245,1267]" count="3" pageId="4" pageNumber="262" type="male">three males</specimenCount>
are also of slightly darker coloration than specimens from Bogota (
<figureCitation id="13222A0EFFE8FF8FAE31318A84980B1A" box="[613,677,1272,1294]" captionStart="Figs 15" captionStartId="1.[226,263,1969,1988]" captionTargetBox="[194,1391,1457,1935]" captionTargetId="figure-673@1.[189,1399,1448,1948]" captionTargetPageId="1" captionText="Figs 15. Habitus of Rhantus spp.: 1 — R. lattkei sp.n.; 2 — R. marahuaca sp.n.; 3 — R. elegans; 4 — R. andinus (Colombia, Bogota, La Florida); 5 — R. andinus (Venezuela, Merida, Azulita). Рис. 15. Rhantus spp., внешний вид: 1 — R. lattkei sp.n.; 2 — R. marahuaca sp.n.; 3 — R. elegans; 4 — R. andinus (Colombia, Bogota, La Florida); 5 — R. andinus (Venezuela, Merida, Azulita)." figureDoi="http://doi.org/10.5281/zenodo.13164043" httpUri="https://zenodo.org/record/13164043/files/figure.png" pageId="4" pageNumber="262">Fig. 4</figureCitation>
, from La Florida, in
<collectionCode id="ED08AE4EFFE8FF8FAD61306187510B3F" box="[309,364,1299,1320]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="4" pageNumber="262" type="Museum">ZSM</collectionCode>
), with slightly more impressed surface sculpture, habitus slightly stouter, posterior fore claw a bit wider and the median lobe slightly straighter along anteri- or 1/3. Due to the great overall similarity of specimens, we suggest that this case deserves further investigation based on more material from the area in between Bogota and
<collectingRegion id="49DDF869FFE8FF8FACE930C487340BDB" box="[189,265,1462,1484]" country="Venezuela" name="Merida" pageId="4" pageNumber="262">Merida</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FFE8FF8FACB6309E84850906" pageId="4" pageNumber="262" type="description">
<paragraph id="8BA6368BFFE8FF8FACB6309E84850906" blockId="4.[189,775,1516,1810]" pageId="4" pageNumber="262">
<emphasis id="B96DEA99FFE8FF8FACB6309E87970815" bold="true" box="[226,426,1516,1538]" pageId="4" pageNumber="262">Acknowledgements</emphasis>
. Michael Balke was supported by the The Linnean Society of London / Systematics Association Small Grants Scheme and a Marie Curie Postdoctoral Fellowship under contract number HPMF-CT-2001- 01365 of the European Commission. We thank Harald Schillhammer (Vienna) for the habitus fotographs of the tepui species, and Max Barclay, curator of the
<taxonomicName id="4C194D08FFE8FF8FAECC33FD853B08B2" authorityName="Linnaeus" authorityYear="1758" box="[664,774,1679,1701]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" pageNumber="262" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
of the Natural History Museum in London, for kind assistance in the examination of Neotropical
<taxonomicName id="4C194D08FFE8FF8FAE1C33B484A708CC" authorityName="Balke &amp; Garcia &amp; Hendrich" authorityYear="2019" box="[584,666,1734,1755]" class="Insecta" family="Dytiscidae" genus="Rhantus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEA99FFE8FF8FAE1C33B484A708CC" box="[584,666,1734,1755]" italics="true" pageId="4" pageNumber="262">Rhantus</emphasis>
</taxonomicName>
. We thank José Clavijo, John Lattke and Luis Joly for giving us a warm welcome at MIZA during our visit in 2003.
</paragraph>
</subSubSection>
</treatment>
</document>