treatments-xml/data/ED/7D/87/ED7D8797FF813F5DFC7A20D16DBE86DE.xml

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<mods:title id="FFBE3768416F3137232B6D7FA063218E">Karakumosa gen. nov., a new Central Asian genus of fossorial wolf spiders (Araneae: Lycosidae: Lycosinae)</mods:title>
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<mods:namePart id="B60F2EFF81584C0AD87252A3FE4850D1">Logunov, Dmitri V.</mods:namePart>
<mods:affiliation id="8A04AAA006E4CCF185692C4EB8817331">The Manchester Museum, The University of Manchester, Oxford Road, Manchester M 13 9 PL, UK.</mods:affiliation>
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<mods:namePart id="212C2A2F3317B632A0E98A0308492231">Ponomarev, Alexander V.</mods:namePart>
<mods:affiliation id="3CD6239A158271478D56AE6B5EF7A80B">Southern Scientific Centre of the Russian Academy of Sciences, Chekhov street 41, Rostov-on-Don, 344006, Russia. E-mail: ponomarev 1952 @ mail. ru</mods:affiliation>
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<taxonomicName id="A2D44D02FF813F53FC7A20D16DBC84DE" ID-CoL="992FK" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[964,1212,1503,1527]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="295" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF813F53FC7A20D16DBC84DE" bold="true" box="[964,1212,1503,1527]" italics="true" pageId="21" pageNumber="295">Karakumosa shmatkoi</emphasis>
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<paragraph id="656B3681FF813F53FBB2230F6DCF8731" blockId="21.[964,1304,1503,1560]" box="[1036,1231,1537,1560]" pageId="21" pageNumber="295">
<figureCitation id="FDEF2A04FF813F53FBB2230F6D9C8731" box="[1036,1180,1537,1560]" captionStart-0="Figs 103-114" captionStart-1="Figs 115-125" captionStart-2="Figs 126-135" captionStart-3="Figs 136-141" captionStartId-0="23.[151,192,1821,1842]" captionStartId-1="24.[151,192,1685,1706]" captionStartId-2="25.[151,192,1838,1859]" captionStartId-3="26.[151,192,1422,1443]" captionTargetBox-0="[154,1435,260,1777]" captionTargetBox-1="[151,1435,257,1636]" captionTargetBox-2="[151,1435,257,1777]" captionTargetBox-3="[151,1435,257,1362]" captionTargetId-0="figure-9@23.[151,1436,257,1790]" captionTargetId-1="figure-9@24.[151,1436,257,1636]" captionTargetId-2="figure-9@25.[151,1436,257,1777]" captionTargetId-3="figure-218@26.[151,1436,257,1362]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionTargetPageId-3="26" captionText-0="Figs 103-114. Karakumosa shmatkoi sp. nov.; male holotype (105-106), male paratype (103-104, 107-112) and female paratype (113- 114); specimens from Kulaly Island (103-104, 109), Zhuzguntyube stow (105-106, 113-114), Mugodzhary Mts (107), Artezian Vil. (108, 110), Tengiz (111), Tyub-Karagan Peninsula (112). (103, 105, 113) Body, dorsal view. (104, 106, 114) Ditto, ventral view. (107-112) Median tooth of MA, posterior view. Scale bars 1 cm (103-106, 113-114), 0.1 mm (107-112)." captionText-1="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." captionText-2="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." captionText-3="Figs 136-141. Karakumosa shmatkoi sp. nov., female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139)." figureDoi-0="http://doi.org/10.5281/zenodo.5743752" figureDoi-1="http://doi.org/10.5281/zenodo.5743754" figureDoi-2="http://doi.org/10.5281/zenodo.5743756" figureDoi-3="http://doi.org/10.5281/zenodo.5743758" httpUri-0="https://zenodo.org/record/5743752/files/figure.png" httpUri-1="https://zenodo.org/record/5743754/files/figure.png" httpUri-2="https://zenodo.org/record/5743756/files/figure.png" httpUri-3="https://zenodo.org/record/5743758/files/figure.png" pageId="21" pageNumber="295">Figs 103-141</figureCitation>
, 
<figureCitation id="FDEF2A04FF813F53FB19230F6DCF8731" box="[1191,1231,1537,1560]" captionStart="Fig" captionStartId="30.[152,187,999,1020]" captionTargetBox="[151,1435,257,943]" captionTargetId="figure-515@30.[151,1436,257,943]" captionTargetPageId="30" captionText="Fig. 155. Localities of four species in the genus Karakumosa gen. nov." figureDoi="http://doi.org/10.5281/zenodo.5743764" httpUri="https://zenodo.org/record/5743764/files/figure.png" pageId="21" pageNumber="295">155</figureCitation>
</paragraph>
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<treatmentCitationGroup id="45C411AFFF813F53FC8123216C9D8757" pageId="21" pageNumber="295">
<emphasis id="57A0EA93FF813F53FC8123216A82876D" box="[831,898,1583,1604]" italics="true" pageId="21" pageNumber="295">Lycosa</emphasis>
<taxonomicName id="A2D44D02FF813F53FC3723216D8F876D" authority="(Kroneberg, 1875)" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[905,1167,1583,1604]" class="Arachnida" family="Lycosidae" genus="Tarentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="295" phylum="Arthropoda" rank="species" species="alticeps">
<emphasis id="57A0EA93FF813F53FC3723216AD4876D" box="[905,980,1583,1604]" italics="true" pageId="21" pageNumber="295">alticeps</emphasis>
(Kroneberg, 1875)
</taxonomicName>
. – 
<treatmentCitation id="E4751090FF813F53FB1523216C68876D" author="Schmidt P." box="[1195,1384,1583,1604]" page="450" pageId="21" pageNumber="295" year="1895">
<bibRefCitation id="01454B70FF813F53FB1523216C68876D" author="Schmidt P." box="[1195,1384,1583,1604]" pageId="21" pageNumber="295" pagination="439 - 484" refId="ref23590" refString="Schmidt P. 1895. Beitrag zur Kenntnis der Laufspinnen (Araneae Citigradae Thor.) Russlands. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere 8 (4): 439 - 484." type="journal article" year="1895">Schmidt, 1895: 450</bibRefCitation>
</treatmentCitation>
(partim, misidentification). – 
<treatmentCitation id="E4751090FF813F53FB3C23426C248748" author="Dunin P. M." box="[1154,1316,1612,1633]" page="55" pageId="21" pageNumber="295" year="1984">
<bibRefCitation id="01454B70FF813F53FB3C23426C248748" author="Dunin P. M." box="[1154,1316,1612,1633]" pageId="21" pageNumber="295" pagination="45 - 60" refId="ref21954" refString="Dunin P. M. 1984. Fauna and ecology of spiders (Aranei) of Apsheron Peninsula [pp. 45 - 60]. In: Utochkin A. S. et al. (eds), Fauna i ekologiya paukoobraznykh. PGU Press, Perm, 159 pp. [In Russian]" type="book chapter" year="1984">Dunin, 1984: 55</bibRefCitation>
</treatmentCitation>
. – 
<treatmentCitation id="E4751090FF813F53FAFC23426DB78757" author="Minoransky V. A. &amp; Ponomarev A. V." page="85" pageId="21" pageNumber="295" year="1984">
<bibRefCitation id="01454B70FF813F53FAFC23426DB78757" author="Minoransky V. A. &amp; Ponomarev A. V." pageId="21" pageNumber="295" pagination="82 - 92" refId="ref22760" refString="Minoransky V. A., Ponomarev A. V. 1984. Materials on the spider fauna of Kamykia [pp. 82 - 92]. In: Utochkin A. S. et al. (eds), Fauna i ekologiya paukoobraznykh. PGU Press, Perm, 159 pp. [In Russian]" type="book chapter" year="1984">Minoransky &amp; Ponomarev, 1984: 85-86</bibRefCitation>
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(all misidentifications).
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“ 
<emphasis id="57A0EA93FF813F53FCF423896A8D87B5" box="[842,909,1671,1692]" italics="true" pageId="21" pageNumber="295">Lycosa</emphasis>
” sp. 1. – 
<bibRefCitation id="01454B70FF813F53FC5723896C9787B5" author="Ponomarev A. V. &amp; Abdurakhmanov G. M." box="[1001,1431,1671,1692]" pageId="21" pageNumber="295" pagination="76 - 121" refId="ref23336" refString="Ponomarev A. V. &amp; Abdurakhmanov G. M. 2014. Spiders (Aranei) of the shore and islands of the northern part of Caspian Sea. The South of Russia: Ecology and Development 1: 76 - 121. [In Russian]" type="journal article" year="2014">Ponomarev &amp; Abdurakhmanov, 2014: 92-93</bibRefCitation>
.
</paragraph>
<paragraph id="656B3681FF813F53FC8123AA6C7B8790" blockId="21.[831,1437,1583,1721]" box="[831,1403,1700,1721]" pageId="21" pageNumber="295">
“ 
<emphasis id="57A0EA93FF813F53FCF423AA6A8D8790" box="[842,909,1700,1721]" italics="true" pageId="21" pageNumber="295">Lycosa</emphasis>
” sp. 2. – 
<bibRefCitation id="01454B70FF813F53FC5723AA6C788790" author="Ponomarev A. V. &amp; Abdurakhmanov G. M." box="[1001,1400,1700,1721]" pageId="21" pageNumber="295" pagination="76 - 121" refId="ref23336" refString="Ponomarev A. V. &amp; Abdurakhmanov G. M. 2014. Spiders (Aranei) of the shore and islands of the northern part of Caspian Sea. The South of Russia: Ecology and Development 1: 76 - 121. [In Russian]" type="journal article" year="2014">Ponomarev &amp; Abdurakhmanov, 2014: 93</bibRefCitation>
.
</paragraph>
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<typeStatus id="BA6F8823FF813F53FC8123DC6AAB87C0" box="[831,939,1746,1769]" pageId="21" pageNumber="295" type="holotype">Holotype</typeStatus>
:
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(
<figureCitation id="FDEF2A04FF813F53FBE023DC6DF087C0" box="[1118,1264,1746,1769]" captionStart="Figs 103-114" captionStartId="23.[151,192,1821,1842]" captionTargetBox="[154,1435,260,1777]" captionTargetId="figure-9@23.[151,1436,257,1790]" captionTargetPageId="23" captionText="Figs 103-114. Karakumosa shmatkoi sp. nov.; male holotype (105-106), male paratype (103-104, 107-112) and female paratype (113- 114); specimens from Kulaly Island (103-104, 109), Zhuzguntyube stow (105-106, 113-114), Mugodzhary Mts (107), Artezian Vil. (108, 110), Tengiz (111), Tyub-Karagan Peninsula (112). (103, 105, 113) Body, dorsal view. (104, 106, 114) Ditto, ventral view. (107-112) Median tooth of MA, posterior view. Scale bars 1 cm (103-106, 113-114), 0.1 mm (107-112)." figureDoi="http://doi.org/10.5281/zenodo.5743752" httpUri="https://zenodo.org/record/5743752/files/figure.png" pageId="21" pageNumber="295">Figs 105-106</figureCitation>
, 
<figureCitation id="FDEF2A04FF813F53FB4223DC6C9087C0" box="[1276,1424,1746,1769]" captionStart="Figs 126-135" captionStartId="25.[151,192,1838,1859]" captionTargetBox="[151,1435,257,1777]" captionTargetId="figure-9@25.[151,1436,257,1777]" captionTargetPageId="25" captionText="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." figureDoi="http://doi.org/10.5281/zenodo.5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="21" pageNumber="295">128, 132-133</figureCitation>
); 
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, 
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, c. 
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NE of Ganyushkino Vil.
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, 
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stow (
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, 
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</date>
; leg. 
<collectorName id="C8215357FF813F53FB21223D6C218663" box="[1183,1313,1843,1866]" pageId="21" pageNumber="295">F.A. Saraev.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="656B3681FF813F5CFC81226D6DC08795" blockId="21.[831,1437,1891,2077]" lastBlockId="26.[831,1437,1573,1724]" lastPageId="26" lastPageNumber="300" pageId="21" pageNumber="295">
<materialsCitation id="D5BC3CDCFF813F53FC81226D6C1A86B2" ID-GBIF-Occurrence="3886766302" collectingDate="1977-07-01" collectingDateMax="1977-07-10" collectingDateMin="1977-07-01" collectionCode="ZMMU" collectorName="F. A. Saraev" country="Kazakhstan" isTailEnumeration="true" latitude="46.717056" longLatPrecision="1" longitude="49.425583" pageId="21" pageNumber="295" specimenCount="1" specimenCount-female="1" stateProvince="Atyrau Oblysy" typeStatus="paratype">
<emphasis id="57A0EA93FF813F53FC81226D6ABB8653" bold="true" box="[831,955,1891,1914]" pageId="21" pageNumber="295">
<typeStatus id="BA6F8823FF813F53FC81226D6AB58653" box="[831,949,1891,1914]" pageId="21" pageNumber="295" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="03C5AE44FF813F53FC7A226D6D1A8653" box="[964,1050,1891,1914]" collectionName="ZMMU" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="21" pageNumber="295" type="Museum">ZMMU</collectionCode>
; 
<specimenCount id="73D2FD08FF813F53FB96226D6D8B8653" box="[1064,1163,1891,1914]" pageId="21" pageNumber="295" type="female">1 female</specimenCount>
; same locality as for the holotype; 
<date id="116A1041FF813F53FC0E228A6D5386B2" box="[944,1107,1924,1947]" pageId="21" pageNumber="295" value="1977-07-01" valueMax="1977-07-10" valueMin="1977-07-01">
<collectingDate id="012EE9A9FF813F53FC0E228A6D5386B2" box="[944,1107,1924,1947]" pageId="21" pageNumber="295" value="1977-07-01" valueMax="1977-07-10" valueMin="1977-07-01">1.-10.VII.1977</collectingDate>
</date>
; leg. 
<collectorName id="C8215357FF813F53FB28228A6C1A86B2" box="[1174,1306,1924,1947]" pageId="21" pageNumber="295">F.A. Saraev</collectorName>
</materialsCitation>
. – 
<materialsCitation id="D5BC3CDCFF813F53FAFF228A6A828692" ID-GBIF-Occurrence="3886766322" collectionCode="ISEA" country="Kazakhstan" pageId="21" pageNumber="295" specimenCount="2" specimenCount-male="2" stateProvince="Atyrau Oblysy" typeStatus="paratype">
<collectionCode id="03C5AE44FF813F53FAFF228A6C8086B2" box="[1345,1408,1924,1947]" collectionName="Poland, Krakow, Polish Academy of Sciences, Institute of Systematic Zoology" pageId="21" pageNumber="295">ISEA</collectionCode>
; 
<specimenCount id="73D2FD08FF813F53FA31228A6A828692" pageId="21" pageNumber="295" type="male">2 males</specimenCount>
</materialsCitation>
; 
<materialsCitation id="D5BC3CDCFF813F53FC2622AA6D9986D5" ID-GBIF-Occurrence="3886766334" collectingDate="1986-06-08" collectorName="F. A. Saraev." country="Kazakhstan" isTailEnumeration="true" latitude="46.397198" location="Tengiz" longLatPrecision="1" longitude="53.422947" municipality="Embinsk Distr." pageId="21" pageNumber="295" specimenCount="1" stateProvince="Atyrau Province" typeStatus="paratype">
<collectingCountry id="1DC37611FF813F53FC2622AA6D518692" box="[920,1105,1956,1979]" name="Kazakhstan" pageId="21" pageNumber="295">KAZAKHSTAN</collectingCountry>
, 
<collectingRegion id="A710F863FF813F53FBD822AA6C268692" box="[1126,1318,1956,1979]" country="Kazakhstan" name="Atyrau Oblysy" pageId="21" pageNumber="295">Atyrau Province</collectingRegion>
, 
<collectingMunicipality id="850FACFBFF813F53FA8222AA6A7F86F5" pageId="21" pageNumber="295">Embinsk Distr.</collectingMunicipality>
, near 
<location id="600B605AFF813F53FC6B22CB6D1E86F5" LSID="urn:lsid:plazi:treatment:ED7D8797FF813F5DFC7A20D16DBE86DE:600B605AFF813F53FC6B22CB6D1E86F5" box="[981,1054,1989,2012]" country="Kazakhstan" latitude="46.397198" longLatPrecision="1" longitude="53.422947" municipality="Embinsk Distr." name="Tengiz" pageId="21" pageNumber="295" stateProvince="Atyrau Province">Tengiz</location>
(
<geoCoordinate id="00E05046FF813F53FB8722CB6DDA86F5" box="[1081,1242,1989,2012]" degrees="46" direction="north" minutes="23" orientation="latitude" pageId="21" pageNumber="295" precision="1" seconds="49.9" value="46.397198">46°23’49.9’’N</geoCoordinate>
, 
<geoCoordinate id="00E05046FF813F53FB4F22CB6C8F86F5" box="[1265,1423,1989,2012]" degrees="53" direction="east" minutes="25" orientation="longitude" pageId="21" pageNumber="295" precision="1" seconds="22.6" value="53.422947">53°25’22.6’’E</geoCoordinate>
); 
<date id="116A1041FF813F53FC8122EB6AB186D5" box="[831,945,2021,2044]" pageId="21" pageNumber="295" value="1986-06-08">
<collectingDate id="012EE9A9FF813F53FC8122EB6AB186D5" box="[831,945,2021,2044]" pageId="21" pageNumber="295" value="1986-06-08">8.VI.1986</collectingDate>
</date>
; leg. 
<collectorName id="C8215357FF813F53FBB822EB6D9986D5" box="[1030,1177,2021,2044]" pageId="21" pageNumber="295">F.A. Saraev.</collectorName>
</materialsCitation>
– 
<materialsCitation id="D5BC3CDCFF813F5CFB7022EB68F08775" ID-GBIF-Occurrence="3886766313" collectingDate="1987-06-19" collectingDateMax="1987-06-24" collectingDateMin="1987-06-19" collectionCode="ZMMU" collectorName="F. A. Saraev" country="Kazakhstan" isTailEnumeration="true" lastPageId="26" lastPageNumber="300" latitude="46.804806" location="Koschagyl" longLatPrecision="10" longitude="53.730556" municipality="Embinsk Distr." pageId="21" pageNumber="295" specimenCount="1" specimenCount-male="1" stateProvince="Atyrau Province" typeStatus="paratype">
<collectionCode id="03C5AE44FF813F53FB7022EB6C2586D5" box="[1230,1317,2021,2044]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="21" pageNumber="295" type="Museum">ZMMU</collectionCode>
; 
<specimenCount id="73D2FD08FF813F53FA8022EB6C9886D5" box="[1342,1432,2021,2044]" pageId="21" pageNumber="295" type="male">1 male</specimenCount>
; 
<collectingCountry id="1DC37611FF813F53FC812D086AF88934" box="[831,1016,2054,2077]" name="Kazakhstan" pageId="21" pageNumber="295">KAZAKHSTAN</collectingCountry>
, 
<collectingRegion id="A710F863FF813F53FBB02D086DCF8934" box="[1038,1231,2054,2077]" country="Kazakhstan" name="Atyrau Oblysy" pageId="21" pageNumber="295">Atyrau Province</collectingRegion>
, 
<collectingMunicipality id="850FACFBFF813F53FB582D086C988934" box="[1254,1432,2054,2077]" pageId="21" pageNumber="295">Embinsk Distr.</collectingMunicipality>
, near 
<location id="600B605AFF8E3F5CFF67232B684D8715" LSID="urn:lsid:plazi:treatment:ED7D8797FF813F5DFC7A20D16DBE86DE:600B605AFF8E3F5CFF67232B684D8715" box="[217,333,1573,1596]" country="Kazakhstan" latitude="46.804806" longLatPrecision="10" longitude="53.730556" municipality="Embinsk Distr." name="Koschagyl" pageId="26" pageNumber="300" stateProvince="Atyrau Province">Koschagyl</location>
(
<geoCoordinate id="00E05046FF8E3F5CFED6232B6B088715" box="[360,520,1573,1596]" degrees="46" direction="north" minutes="48" orientation="latitude" pageId="26" pageNumber="300" precision="1" seconds="17.3" value="46.804806">46°48’17.3’’N</geoCoordinate>
, 
<geoCoordinate id="00E05046FF8E3F5CFDA1232B6BA98715" box="[543,681,1573,1596]" degrees="53" direction="east" minutes="43" orientation="longitude" pageId="26" pageNumber="300" precision="15" seconds="50" value="53.730556">53°43’50’’E</geoCoordinate>
); 
<date id="116A1041FF8E3F5CFD77232B68168775" pageId="26" pageNumber="300" value="1987-06-19" valueMax="1987-06-24" valueMin="1987-06-19">
<collectingDate id="012EE9A9FF8E3F5CFD77232B68168775" pageId="26" pageNumber="300" value="1987-06-19" valueMax="1987-06-24" valueMin="1987-06-19">19.- 24.VI.1987</collectingDate>
</date>
; leg. 
<collectorName id="C8215357FF8E3F5CFEDA234B68F08775" box="[356,496,1605,1628]" pageId="26" pageNumber="300">F.A. Saraev</collectorName>
</materialsCitation>
. – 
<materialsCitation id="D5BC3CDCFF8E3F5CFD9C234B6BF187F5" ID-GBIF-Occurrence="3886766328" collectedFrom="hilly sands," collectingDate="2013-06-17" collectingMethod="pitfall traps" collectionCode="ZMMU" collectorName="G. M. Abdurakhmanov" country="Kazakhstan" isTailEnumeration="true" latitude="44.43886" location="Mangyshlak" longLatPrecision="1" longitude="50.255527" municipality="Fort Shevchenko" pageId="26" pageNumber="300" specimenCount="6" specimenCount-male="6" stateProvince="Mangistau Province" typeStatus="paratype">
<collectionCode id="03C5AE44FF8E3F5CFD9C234B6B798775" box="[546,633,1605,1628]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="26" pageNumber="300" type="Museum">ZMMU</collectionCode>
; 
<specimenCount id="73D2FD08FF8E3F5CFD31234B6BEF8775" box="[655,751,1605,1628]" pageId="26" pageNumber="300" type="male">6 males</specimenCount>
; 
<collectingCountry id="1DC37611FF8E3F5CFF29236B684F8755" box="[151,335,1637,1660]" name="Kazakhstan" pageId="26" pageNumber="300">KAZAKHSTAN</collectingCountry>
, 
<collectingRegion id="A710F863FF8E3F5CFEE0236B6B3C8755" box="[350,572,1637,1660]" inferredBySuffix="true" pageId="26" pageNumber="300">Mangistau Province</collectingRegion>
, 
<location id="600B605AFF8E3F5CFDF5236B6BD28755" LSID="urn:lsid:plazi:treatment:ED7D8797FF813F5DFC7A20D16DBE86DE:600B605AFF8E3F5CFDF5236B6BD28755" box="[587,722,1637,1660]" country="Kazakhstan" latitude="44.43886" longLatPrecision="1" longitude="50.255527" municipality="Fort Shevchenko" name="Mangyshlak" pageId="26" pageNumber="300" stateProvince="Mangistau Province">Mangyshlak</location>
, c. 
<locationDeviation id="74D92FA3FF8E3F5CFF29238B68E487B5" box="[151,484,1669,1692]" pageId="26" pageNumber="300">
<quantity id="A22C9B64FF8E3F5CFF29238B69DF87B5" box="[151,223,1669,1692]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="300" unit="km" value="10.0">10 km</quantity>
S of Fort Shevchenko
</locationDeviation>
, 
<location id="600B605AFF8E3F5CFE4D238B69FF8795" LSID="urn:lsid:plazi:treatment:ED7D8797FF813F5DFC7A20D16DBE86DE:600B605AFF8E3F5CFE4D238B69FF8795" country="Kazakhstan" latitude="44.43886" longLatPrecision="1" longitude="50.255527" municipality="Fort Shevchenko" name="shore of Tyub-Karagan Peninsula" pageId="26" pageNumber="300" stateProvince="Mangistau Province">shore of Tyub-Karagan Peninsula</location>
(
<geoCoordinate id="00E05046FF8E3F5CFEAD23AB68B18795" box="[275,433,1701,1724]" degrees="44" direction="north" minutes="26" orientation="latitude" pageId="26" pageNumber="300" precision="1" seconds="19.9" value="44.43886">44°26’19.9’’N</geoCoordinate>
, 
<geoCoordinate id="00E05046FF8E3F5CFE7F23AB6B5C8795" box="[449,604,1701,1724]" degrees="50" direction="east" minutes="15" orientation="longitude" pageId="26" pageNumber="300" precision="1" seconds="19.9" value="50.255527">50°15’19.9’’E</geoCoordinate>
), 
<collectedFrom id="D6C8757CFF8E3F5CFDCB23AB6BF48795" box="[629,756,1701,1724]" pageId="26" pageNumber="300">hilly sands,</collectedFrom>
<collectingMethod id="BC954E96FF8E3F5CFF2923CB681E87F5" box="[151,286,1733,1756]" pageId="26" pageNumber="300">pitfall traps</collectingMethod>
; 
<date id="116A1041FF8E3F5CFE9123CB68AD87F5" box="[303,429,1733,1756]" pageId="26" pageNumber="300" value="2013-06-17">
<collectingDate id="012EE9A9FF8E3F5CFE9123CB68AD87F5" box="[303,429,1733,1756]" pageId="26" pageNumber="300" value="2013-06-17">17.VI.2013</collectingDate>
</date>
; leg. 
<collectorName id="C8215357FF8E3F5CFE4B23CB6BF187F5" box="[501,753,1733,1756]" pageId="26" pageNumber="300">G.M. Abdurakhmanov</collectorName>
</materialsCitation>
. – 
<materialsCitation id="D5BC3CDCFF8E3F5CFF0523EB6B568675" ID-GBIF-Occurrence="3886766349" collectedFrom="fine hilly sands" collectingDate="2013-06-19" collectingMethod="hand collecting" collectionCode="ZMMU" collectorName="G. M. Abdurakhmanov" country="Kazakhstan" isTailEnumeration="true" latitude="44.892056" location="Island Kulaly" longLatPrecision="1" longitude="50.053665" municipality="Mangyshlak" pageId="26" pageNumber="300" specimenCount="3" specimenCount-male="3" stateProvince="Mangistau Province" typeStatus="paratype">
<collectionCode id="03C5AE44FF8E3F5CFF0523EB681387D5" box="[187,275,1765,1788]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="26" pageNumber="300" type="Museum">ZMMU</collectionCode>
; 
<specimenCount id="73D2FD08FF8E3F5CFE9223EB689387D5" box="[300,403,1765,1788]" pageId="26" pageNumber="300" type="male">3 males</specimenCount>
; 
<collectingCountry id="1DC37611FF8E3F5CFE1323EB6B6787D5" box="[429,615,1765,1788]" name="Kazakhstan" pageId="26" pageNumber="300">KAZAKHSTAN</collectingCountry>
, 
<collectingRegion id="A710F863FF8E3F5CFD3F23EB69F98635" inferredBySuffix="true" pageId="26" pageNumber="300">Mangistau Province</collectingRegion>
, 
<collectingMunicipality id="850FACFBFF8E3F5CFEB5220B68938635" box="[267,403,1797,1820]" pageId="26" pageNumber="300">Mangyshlak</collectingMunicipality>
, 
<location id="600B605AFF8E3F5CFE1D220B6B3B8635" LSID="urn:lsid:plazi:treatment:ED7D8797FF813F5DFC7A20D16DBE86DE:600B605AFF8E3F5CFE1D220B6B3B8635" box="[419,571,1797,1820]" country="Kazakhstan" latitude="44.892056" longLatPrecision="1" longitude="50.053665" municipality="Mangyshlak" name="Island Kulaly" pageId="26" pageNumber="300" stateProvince="Mangistau Province">Island Kulaly</location>
(
<geoCoordinate id="00E05046FF8E3F5CFDEE220B6BEF8635" box="[592,751,1797,1820]" degrees="44" direction="north" minutes="53" orientation="latitude" pageId="26" pageNumber="300" precision="1" seconds="31.4" value="44.892056">44°53’31.4’’N</geoCoordinate>
, 
<geoCoordinate id="00E05046FF8E3F5CFF29222B68358615" box="[151,309,1829,1852]" degrees="50" direction="east" minutes="03" orientation="longitude" pageId="26" pageNumber="300" precision="1" seconds="13.2" value="50.053665">50°03’13.2’’E</geoCoordinate>
), 
<collectedFrom id="D6C8757CFF8E3F5CFEE6222B6B1C8615" box="[344,540,1829,1852]" pageId="26" pageNumber="300">fine hilly sands</collectedFrom>
, 
<collectingMethod id="BC954E96FF8E3F5CFD88222B6BF08615" box="[566,752,1829,1852]" pageId="26" pageNumber="300">hand collecting</collectingMethod>
; 
<date id="116A1041FF8E3F5CFF29224B68158675" box="[151,277,1861,1884]" pageId="26" pageNumber="300" value="2013-06-19">
<collectingDate id="012EE9A9FF8E3F5CFF29224B68158675" box="[151,277,1861,1884]" pageId="26" pageNumber="300" value="2013-06-19">19.VI.2013</collectingDate>
</date>
; leg. 
<collectorName id="C8215357FF8E3F5CFEE5224B6B568675" box="[347,598,1861,1884]" pageId="26" pageNumber="300">G.M. Abdurakhmanov</collectorName>
</materialsCitation>
. – 
<materialsCitation id="D5BC3CDCFF8E3F5CFD3E224B68608655" ID-GBIF-Occurrence="3886766346" collectionCode="MHNG" country="Kazakhstan" isTailEnumeration="true" latitude="44.892056" longLatPrecision="1" longitude="50.053665" municipality="Mangyshlak" pageId="26" pageNumber="300" specimenCount="2" specimenCount-male="2" stateProvince="Mangistau Province" typeStatus="paratype">
<collectionCode id="03C5AE44FF8E3F5CFD3E224B6BD58675" box="[640,725,1861,1884]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="26" pageNumber="300" type="Museum">MHNG</collectionCode>
; 
<specimenCount id="73D2FD08FF8E3F5CFD58224B69D98655" pageId="26" pageNumber="300" type="male">2 males</specimenCount>
; same data
</materialsCitation>
. – 
<materialsCitation id="D5BC3CDCFF8E3F5CFE33226B688C86F5" ID-GBIF-Occurrence="3886766306" collectingDate="2013-06-28" collectionCode="PSU" collectorName="A. O. Shkurikhin" country="Kazakhstan" isTailEnumeration="true" latitude="48.95" location="Mugodzhary Mts" longLatPrecision="1106" longitude="58.65" municipality="Embi Vil." pageId="26" pageNumber="300" specimenCount="1" specimenCount-male="1" stateProvince="Aktobe Province" typeStatus="paratype">
<collectionCode id="03C5AE44FF8E3F5CFE33226B68C28655" box="[397,450,1893,1916]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="26" pageNumber="300" type="Museum">PSU</collectionCode>
; 
<specimenCount id="73D2FD08FF8E3F5CFE6D226B6B268655" box="[467,550,1893,1916]" pageId="26" pageNumber="300" type="male">1 male</specimenCount>
; 
<collectingCountry id="1DC37611FF8E3F5CFD86226B6BF08655" box="[568,752,1893,1916]" name="Kazakhstan" pageId="26" pageNumber="300">KAZAKHSTAN</collectingCountry>
, 
<collectingRegion id="A710F863FF8E3F5CFF29228B685F86B5" box="[151,351,1925,1948]" inferredBySuffix="true" pageId="26" pageNumber="300">Aktobe Province</collectingRegion>
, c. 
<locationDeviation id="74D92FA3FF8E3F5CFE20228B6BEC86B5" box="[414,748,1925,1948]" pageId="26" pageNumber="300">
<quantity id="A22C9B64FF8E3F5CFE20228B68EF86B5" box="[414,495,1925,1948]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="300" unit="km" value="25.0">25 km</quantity>
ENE of Embi Vil
</locationDeviation>
., 
<location id="600B605AFF8E3F5CFF2922AB68578695" LSID="urn:lsid:plazi:treatment:ED7D8797FF813F5DFC7A20D16DBE86DE:600B605AFF8E3F5CFF2922AB68578695" box="[151,343,1957,1980]" country="Kazakhstan" latitude="48.95" longLatPrecision="1106" longitude="58.65" municipality="Embi Vil." name="Mugodzhary Mts" pageId="26" pageNumber="300" stateProvince="Aktobe Province">Mugodzhary Mts</location>
(c. 
<geoCoordinate id="00E05046FF8E3F5CFE3522AB68EA8695" box="[395,490,1957,1980]" degrees="48" direction="north" minutes="57" orientation="latitude" pageId="26" pageNumber="300" precision="925" value="48.95">48°57’N</geoCoordinate>
, 
<geoCoordinate id="00E05046FF8E3F5CFE4522AB6B588695" box="[507,600,1957,1980]" degrees="58" direction="east" minutes="39" orientation="longitude" pageId="26" pageNumber="300" precision="925" value="58.65">58°39’E</geoCoordinate>
); 
<date id="116A1041FF8E3F5CFDCC22AB6BEF8695" box="[626,751,1957,1980]" pageId="26" pageNumber="300" value="2013-06-28">
<collectingDate id="012EE9A9FF8E3F5CFDCC22AB6BEF8695" box="[626,751,1957,1980]" pageId="26" pageNumber="300" value="2013-06-28">28.VI.2013</collectingDate>
</date>
; leg. 
<collectorName id="C8215357FF8E3F5CFF7322CB688C86F5" box="[205,396,1989,2012]" pageId="26" pageNumber="300">A.O. Shkurikhin</collectorName>
</materialsCitation>
. – 
<materialsCitation id="D5BC3CDCFF8E3F5CFE0522CB6ABB8775" ID-GBIF-Occurrence="3886766350" collectedFrom="hilly sands with sparse vegetation" collectingDate="1975-06-12" collectionCode="MMUE" collectorName="A. V. Ponomarev" country="Russia" latitude="46.376083" location="Yashkul Distr." longLatPrecision="1" longitude="46.021805" municipality="Yashkul Vil." pageId="26" pageNumber="300" specimenCount="1" specimenCount-male="1" stateProvince="Kalmykia" typeStatus="paratype">
<collectionCode id="03C5AE44FF8E3F5CFE0522CB6B1286F5" box="[443,530,1989,2012]" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="26" pageNumber="300">MMUE</collectionCode>
; 
<specimenCount id="73D2FD08FF8E3F5CFD9822CB6B7A86F5" box="[550,634,1989,2012]" pageId="26" pageNumber="300" type="male">1 male</specimenCount>
; 
<collectingCountry id="1DC37611FF8E3F5CFD3322CB6BEF86F5" box="[653,751,1989,2012]" name="Russia" pageId="26" pageNumber="300">RUSSIA</collectingCountry>
, 
<collectingRegion id="A710F863FF8E3F5CFF2922EB680386D5" box="[151,259,2021,2044]" country="Russia" name="Kalmykiya" pageId="26" pageNumber="300">Kalmykia</collectingRegion>
, 
<location id="600B605AFF8E3F5CFEB022EB68AB86D5" LSID="urn:lsid:plazi:treatment:ED7D8797FF813F5DFC7A20D16DBE86DE:600B605AFF8E3F5CFEB022EB68AB86D5" box="[270,427,2021,2044]" country="Russia" latitude="46.376083" longLatPrecision="1" longitude="46.021805" municipality="Yashkul Vil." name="Yashkul Distr." pageId="26" pageNumber="300" stateProvince="Kalmykia">Yashkul Distr.</location>
, c. 
<locationDeviation id="74D92FA3FF8E3F5CFE6F22EB6BEC86D5" box="[465,748,2021,2044]" pageId="26" pageNumber="300">
<quantity id="A22C9B64FF8E3F5CFE6F22EB6B1686D5" box="[465,534,2021,2044]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="26" pageNumber="300" unit="km" value="55.0">55 km</quantity>
NE of Yashkul Vil
</locationDeviation>
., near 
<location id="600B605AFF8E3F5CFF682D0B683C8935" LSID="urn:lsid:plazi:treatment:ED7D8797FF813F5DFC7A20D16DBE86DE:600B605AFF8E3F5CFF682D0B683C8935" box="[214,316,2053,2076]" country="Russia" latitude="46.376083" longLatPrecision="1" longitude="46.021805" municipality="Yashkul Vil." name="Utta Vil." pageId="26" pageNumber="300" stateProvince="Kalmykia">Utta Vil.</location>
(
<geoCoordinate id="00E05046FF8E3F5CFEEA2D0B68F48935" box="[340,500,2053,2076]" degrees="46" direction="north" minutes="22" orientation="latitude" pageId="26" pageNumber="300" precision="1" seconds="33.9" value="46.376083">46°22’33.9’’N</geoCoordinate>
, 
<geoCoordinate id="00E05046FF8E3F5CFDB62D0B6BA58935" box="[520,677,2053,2076]" degrees="46" direction="east" minutes="01" orientation="longitude" pageId="26" pageNumber="300" precision="1" seconds="18.5" value="46.021805">46°01’18.5’’E</geoCoordinate>
), 
<collectedFrom id="D6C8757CFF8E3F5CFD7C2D0B6D918715" box="[706,1169,1573,2076]" pageId="26" pageNumber="300">hilly sands with sparse vegetation</collectedFrom>
; 
<date id="116A1041FF8E3F5CFB1A232B6C228715" box="[1188,1314,1573,1596]" pageId="26" pageNumber="300" value="1975-06-12">
<collectingDate id="012EE9A9FF8E3F5CFB1A232B6C228715" box="[1188,1314,1573,1596]" pageId="26" pageNumber="300" value="1975-06-12">12.VI.1975</collectingDate>
</date>
; leg. 
<collectorName id="C8215357FF8E3F5CFAD5232B6ABB8775" pageId="26" pageNumber="300">A.V. Ponomarev</collectorName>
</materialsCitation>
. – 
<materialsCitation id="D5BC3CDCFF8E3F5CFC5B234B6DBC8795" ID-GBIF-Occurrence="3886766316" collectingDate="1976-07-01" collectingDateMax="1976-07-04" collectingDateMin="1976-07-01" collectionCode="ZMMU" collectorName="E. A. Khachikov" country="Russia" isTailEnumeration="true" latitude="44.958195" location="Chernozemel'sk Distr." longLatPrecision="1" longitude="46.631527" municipality="Komsomol'sky Vil." pageId="26" pageNumber="300" specimenCount="2" specimenCount-male="2" stateProvince="Kalmykia" typeStatus="paratype">
<collectionCode id="03C5AE44FF8E3F5CFC5B234B6D3C8775" box="[997,1084,1605,1628]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="26" pageNumber="300" type="Museum">ZMMU</collectionCode>
; 
<specimenCount id="73D2FD08FF8E3F5CFBF2234B6DA88775" box="[1100,1192,1605,1628]" pageId="26" pageNumber="300" type="male">2 males</specimenCount>
; 
<collectingCountry id="1DC37611FF8E3F5CFB07234B6C1B8775" box="[1209,1307,1605,1628]" name="Russia" pageId="26" pageNumber="300">RUSSIA</collectingCountry>
, 
<collectingRegion id="A710F863FF8E3F5CFA95234B6C988775" box="[1323,1432,1605,1628]" country="Russia" name="Kalmykiya" pageId="26" pageNumber="300">Kalmykia</collectingRegion>
, 
<location id="600B605AFF8E3F5CFC81236B6D358755" LSID="urn:lsid:plazi:treatment:ED7D8797FF813F5DFC7A20D16DBE86DE:600B605AFF8E3F5CFC81236B6D358755" box="[831,1077,1637,1660]" country="Russia" latitude="44.958195" longLatPrecision="1" longitude="46.631527" municipality="Komsomol'sky Vil." name="Chernozemel'sk Distr." pageId="26" pageNumber="300" stateProvince="Kalmykia">Chernozemel’sk Distr.</location>
, c. 
<locationDeviation id="74D92FA3FF8E3F5CFBE1236B6A6587B5" pageId="26" pageNumber="300">
<quantity id="A22C9B64FF8E3F5CFBE1236B6DA68755" box="[1119,1190,1637,1660]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="26" pageNumber="300" unit="km" value="65.0">65 km</quantity>
SE of Komsomol’sky Vil
</locationDeviation>
., near 
<location id="600B605AFF8E3F5CFC12238B6D3987B5" LSID="urn:lsid:plazi:treatment:ED7D8797FF813F5DFC7A20D16DBE86DE:600B605AFF8E3F5CFC12238B6D3987B5" box="[940,1081,1669,1692]" country="Russia" latitude="44.958195" longLatPrecision="1" longitude="46.631527" municipality="Komsomol'sky Vil." name="Artezian Vil." pageId="26" pageNumber="300" stateProvince="Kalmykia">Artezian Vil.</location>
(
<geoCoordinate id="00E05046FF8E3F5CFBF7238B6DE787B5" box="[1097,1255,1669,1692]" degrees="44" direction="north" minutes="57" orientation="latitude" pageId="26" pageNumber="300" precision="1" seconds="29.5" value="44.958195">44°57’29.5’’N</geoCoordinate>
, 
<geoCoordinate id="00E05046FF8E3F5CFB4A238B6C9087B5" box="[1268,1424,1669,1692]" degrees="46" direction="east" minutes="37" orientation="longitude" pageId="26" pageNumber="300" precision="1" seconds="53.5" value="46.631527">46°37’53.5’’E</geoCoordinate>
); 
<date id="116A1041FF8E3F5CFC8123AB6AD58795" box="[831,981,1701,1724]" pageId="26" pageNumber="300" value="1976-07-01" valueMax="1976-07-04" valueMin="1976-07-01">
<collectingDate id="012EE9A9FF8E3F5CFC8123AB6AD58795" box="[831,981,1701,1724]" pageId="26" pageNumber="300" value="1976-07-01" valueMax="1976-07-04" valueMin="1976-07-01">1.-4.VII.1976</collectingDate>
</date>
; leg. 
<collectorName id="C8215357FF8E3F5CFBB323AB6DBC8795" box="[1037,1212,1701,1724]" pageId="26" pageNumber="300">E.A. Khachikov</collectorName>
</materialsCitation>
.
</paragraph>
<caption id="31AB6609FF823F50FF29224A6A1786E4" ID-DOI="http://doi.org/10.5281/zenodo.5743750" ID-Zenodo-Dep="5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="22" pageNumber="296" startId="22.[151,192,1860,1881]" targetBox="[151,1435,257,1809]" targetPageId="22">
<paragraph id="656B3681FF823F50FF29224A6A1786E4" blockId="22.[151,1437,1860,1997]" pageId="22" pageNumber="296">
Figs 93-102. 
<taxonomicName id="A2D44D02FF823F50FEA4224A68DF8670" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[282,479,1860,1881]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="296" phylum="Arthropoda" rank="species" species="repetek" status="sp. nov.">
<emphasis id="57A0EA93FF823F50FEA4224A68DF8670" box="[282,479,1860,1881]" italics="true" pageId="22" pageNumber="296">Karakumosa repetek</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF823F50FE5A224A6B308670" box="[484,560,1860,1881]" pageId="22" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods.
</paragraph>
</caption>
<caption id="31AB6609FF833F51FF292213688C868C" ID-DOI="http://doi.org/10.5281/zenodo.5743752" ID-Zenodo-Dep="5743752" httpUri="https://zenodo.org/record/5743752/files/figure.png" pageId="23" pageNumber="297" startId="23.[151,192,1821,1842]" targetBox="[154,1435,260,1777]" targetPageId="23">
<paragraph id="656B3681FF833F51FF292213688C868C" blockId="23.[151,1436,1821,1957]" pageId="23" pageNumber="297">
Figs 103-114. 
<taxonomicName id="A2D44D02FF833F51FE98221368FE861B" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[294,510,1821,1842]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="297" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF833F51FE98221368FE861B" box="[294,510,1821,1842]" italics="true" pageId="23" pageNumber="297">Karakumosa shmatkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF833F51FDBD22136B51861B" box="[515,593,1821,1842]" pageId="23" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
; male holotype (105-106), male paratype (103-104, 107-112) and female paratype (113- 114); specimens from Kulaly Island (103-104, 109), Zhuzguntyube stow (105-106, 113-114), Mugodzhary Mts (107), Artezian Vil. (108, 110), Tengiz (111), Tyub-Karagan Peninsula (112). (103, 105, 113) Body, dorsal view. (104, 106, 114) Ditto, ventral view. (107-112) Median tooth of MA, posterior view. Scale bars 1 cm (103-106, 113-114), 0.1 mm (107-112).
</paragraph>
</caption>
<caption id="31AB6609FF8C3F5EFF29239B6B978613" ID-DOI="http://doi.org/10.5281/zenodo.5743754" ID-Zenodo-Dep="5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="24" pageNumber="298" startId="24.[151,192,1685,1706]" targetBox="[151,1435,257,1636]" targetPageId="24">
<paragraph id="656B3681FF8C3F5EFF29239B6B978613" blockId="24.[151,1437,1685,1850]" pageId="24" pageNumber="298">
Figs 115-125. 
<taxonomicName id="A2D44D02FF8C3F5EFE97239B6B038783" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[297,515,1685,1706]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="298" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF8C3F5EFE97239B6B038783" box="[297,515,1685,1706]" italics="true" pageId="24" pageNumber="298">Karakumosa shmatkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8C3F5EFDB4239B6B5A8783" box="[522,602,1685,1706]" pageId="24" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125). Abbreviations as explained in Material and methods.
</paragraph>
</caption>
<caption id="31AB6609FF8D3F5FFF292220685C869F" ID-DOI="http://doi.org/10.5281/zenodo.5743756" ID-Zenodo-Dep="5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="25" pageNumber="299" startId="25.[151,192,1838,1859]" targetBox="[151,1435,257,1777]" targetPageId="25">
<paragraph id="656B3681FF8D3F5FFF292220685C869F" blockId="25.[151,1437,1838,1974]" pageId="25" pageNumber="299">
Figs 126-135. 
<taxonomicName id="A2D44D02FF8D3F5FFE9722206B02866A" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[297,514,1838,1859]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="299" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF8D3F5FFE9722206B02866A" box="[297,514,1838,1859]" italics="true" pageId="25" pageNumber="299">Karakumosa shmatkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8D3F5FFDB622206B57866A" box="[520,599,1838,1859]" pageId="25" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134).
</paragraph>
</caption>
<caption id="31AB6609FF8E3F5CFF2920806D3484F4" ID-DOI="http://doi.org/10.5281/zenodo.5743758" ID-Zenodo-Dep="5743758" httpUri="https://zenodo.org/record/5743758/files/figure.png" pageId="26" pageNumber="300" startId="26.[151,192,1422,1443]" targetBox="[151,1435,257,1362]" targetPageId="26">
<paragraph id="656B3681FF8E3F5CFF2920806D3484F4" blockId="26.[151,1435,1422,1501]" pageId="26" pageNumber="300">
Figs 136-141. 
<taxonomicName id="A2D44D02FF8E3F5CFE99208068FF848A" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[295,511,1422,1443]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="300" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF8E3F5CFE99208068FF848A" box="[295,511,1422,1443]" italics="true" pageId="26" pageNumber="300">Karakumosa shmatkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8E3F5CFDBA20806B50848A" box="[516,592,1422,1443]" pageId="26" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
, female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139).
</paragraph>
</caption>
<paragraph id="656B3681FF8E3F5CFC8123DA6D688602" blockId="26.[831,1437,1748,1835]" pageId="26" pageNumber="300">
<emphasis id="57A0EA93FF8E3F5CFC8123DA6AFD87C2" bold="true" box="[831,1021,1748,1771]" pageId="26" pageNumber="300">Other material:</emphasis>
<materialsCitation id="D5BC3CDCFF8E3F5CFBB223DA6D688602" ID-GBIF-Occurrence="3886766338" collectionCode="ZMMU" country="Azerbaijan" location="Peninsula" municipality="Peninsula" pageId="26" pageNumber="300" specimenCount="1" specimenCount-female="1" stateProvince="Absheron">
<collectionCode id="03C5AE44FF8E3F5CFBB223DA6D6387C2" box="[1036,1123,1748,1771]" collectionName="ZMMU" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="26" pageNumber="300" type="Museum">ZMMU</collectionCode>
; 
<specimenCount id="73D2FD08FF8E3F5CFBC823DA6DDC87C2" box="[1142,1244,1748,1771]" pageId="26" pageNumber="300" type="female">1 female</specimenCount>
(
<figureCitation id="FDEF2A04FF8E3F5CFB4D23DA6C9087C2" box="[1267,1424,1748,1771]" captionStart="Figs 136-141" captionStartId="26.[151,192,1422,1443]" captionTargetBox="[151,1435,257,1362]" captionTargetId="figure-218@26.[151,1436,257,1362]" captionTargetPageId="26" captionText="Figs 136-141. Karakumosa shmatkoi sp. nov., female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139)." figureDoi="http://doi.org/10.5281/zenodo.5743758" httpUri="https://zenodo.org/record/5743758/files/figure.png" pageId="26" pageNumber="300">Figs 136-141</figureCitation>
); 
<collectingCountry id="1DC37611FF8E3F5CFC8123FA6AE68622" box="[831,998,1780,1803]" name="Azerbaijan" pageId="26" pageNumber="300">AZERBAIJAN</collectingCountry>
, 
<collectingRegion id="A710F863FF8E3F5CFC4F23FA6D598622" box="[1009,1113,1780,1803]" country="Azerbaijan" name="Abseron" pageId="26" pageNumber="300">Absheron</collectingRegion>
<collectingMunicipality id="850FACFBFF8E3F5CFBDF23FA6DC98622" box="[1121,1225,1780,1803]" pageId="26" pageNumber="300">Peninsula</collectingMunicipality>
(no exact locality); no date and collector given.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF8E3F5CFC81224A6DAD86D2" pageId="26" pageNumber="300" type="etymology">
<paragraph id="656B3681FF8E3F5CFC81224A6DAD86D2" blockId="26.[831,1437,1860,2043]" pageId="26" pageNumber="300">
<emphasis id="57A0EA93FF8E3F5CFC81224A6AC48672" bold="true" box="[831,964,1860,1883]" pageId="26" pageNumber="300">Etymology:</emphasis>
The species is dedicated to our friend and colleague, Mr 
<collectingRegion id="A710F863FF8E3F5CFC5C226A6D438652" box="[994,1091,1892,1915]" country="Russia" name="Vladimir" pageId="26" pageNumber="300">Vladimir</collectingRegion>
Yu. Shmatko (Rostov-on-Don, 
<collectingCountry id="1DC37611FF8E3F5CFC81228A6A8B86B2" box="[831,907,1924,1947]" name="Russia" pageId="26" pageNumber="300">Russia</collectingCountry>
), who has been collaborating with the second author (AVP) in spider studies for many years and who took the majority of digital photos of 
<taxonomicName id="A2D44D02FF8E3F5CFAAC22CA6A8C86D2" class="Arachnida" family="Lycosidae" genus="Karakumosa" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="300" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="57A0EA93FF8E3F5CFAAC22CA6C9C86F2" box="[1298,1436,1988,2011]" italics="true" pageId="26" pageNumber="300">Karakumosa</emphasis>
species
</taxonomicName>
given in the present paper.
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF8F3F5DFF29240C6BDE8210" pageId="27" pageNumber="301" type="diagnosis">
<paragraph id="656B3681FF8F3F5DFF29240C6BDE8210" blockId="27.[151,756,258,825]" pageId="27" pageNumber="301">
<emphasis id="57A0EA93FF8F3F5DFF29240C680F8030" bold="true" box="[151,271,258,281]" pageId="27" pageNumber="301">Diagnosis:</emphasis>
The male of 
<taxonomicName id="A2D44D02FF8F3F5DFE0E240C6B358030" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[432,565,258,281]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF8F3F5DFE0E240C6B358030" box="[432,565,258,281]" italics="true" pageId="27" pageNumber="301">K. shmatkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8F3F5DFDFE240C6B988030" box="[576,664,258,281]" pageId="27" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to those of 
<taxonomicName id="A2D44D02FF8F3F5DFECB242C68F28010" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[373,498,290,313]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="gromovi" status="sp. nov.">
<emphasis id="57A0EA93FF8F3F5DFECB242C68F28010" box="[373,498,290,313]" italics="true" pageId="27" pageNumber="301">K. gromovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8F3F5DFE40242C6B578010" box="[510,599,290,313]" pageId="27" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="FDEF2A04FF8F3F5DFDD7242C69D78070" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="27" pageNumber="301">Figs 59, 60, 64-69</figureCitation>
) and 
<taxonomicName id="A2D44D02FF8F3F5DFE9E244C68B68070" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[288,438,322,345]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="tashkumyr" status="sp. nov.">
<emphasis id="57A0EA93FF8F3F5DFE9E244C68B68070" box="[288,438,322,345]" italics="true" pageId="27" pageNumber="301">K. tashkumyr</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8F3F5DFE7D244C6B1D8070" box="[451,541,322,345]" pageId="27" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="FDEF2A04FF8F3F5DFD8F244C6BCB8070" box="[561,715,322,345]" captionStart="Figs 142-154" captionStartId="28.[151,192,1972,1993]" captionTargetBox="[169,1419,257,1942]" captionTargetId="figure-9@28.[169,1419,257,1943]" captionTargetPageId="28" captionText="Figs 142-154. Karakumosa tashkumyr sp. nov., male paratypes, in ISEA. (142) Body, ventral view. (143) Ditto, dorsal view. (144) Left chelicera, ventral view. (145, 147, 152-154) Median tooth of MA, posterior view. (146) Left male palp, ventral view. (148, 150) Bulbus, ventral view. (151) Ditto, retrolateral view. (149) Embolic division. Scale bars 1 cm (142-143), 0.5 cm (144, 146, 150-151), 0.2 mm (148), 0.1 mm (145, 147, 149, 152-154)." figureDoi="http://doi.org/10.5281/zenodo.5743760" httpUri="https://zenodo.org/record/5743760/files/figure.png" pageId="27" pageNumber="301">Figs 142-154</figureCitation>
); it can be distinguished from the former by the presence of a low serrate flange at the foot of the median tooth (absent in 
<taxonomicName id="A2D44D02FF8F3F5DFEB024AC68898090" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[270,393,418,441]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="gromovi" status="sp. nov.">
<emphasis id="57A0EA93FF8F3F5DFEB024AC68898090" box="[270,393,418,441]" italics="true" pageId="27" pageNumber="301">K. gromovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8F3F5DFE2A24AC68ED8090" box="[404,493,418,441]" pageId="27" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
; 
<figureCitation id="FDEF2A04FF8F3F5DFE4224AC6B5E8090" box="[508,606,418,441]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="27" pageNumber="301">Figs 125</figureCitation>
, 
<figureCitation id="FDEF2A04FF8F3F5DFDD324AC6B958090" box="[621,661,418,441]" captionStart="Figs 126-135" captionStartId="25.[151,192,1838,1859]" captionTargetBox="[151,1435,257,1777]" captionTargetId="figure-9@25.[151,1436,257,1777]" captionTargetPageId="25" captionText="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." figureDoi="http://doi.org/10.5281/zenodo.5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="27" pageNumber="301">134</figureCitation>
cf. 
<figureCitation id="FDEF2A04FF8F3F5DFD7824AC69D880F0" captionStart="Figs 70-75" captionStartId="17.[151,192,1222,1243]" captionTargetBox="[160,1436,257,1181]" captionTargetId="figure-337@17.[151,1439,257,1191]" captionTargetPageId="17" captionText="Figs 70-75. Karakumosa gromovi sp. nov.; male paratypes (74-75) and female paratype (70-73). (70) Body, dorsal view. (71) Ditto, ventral view. (72) Epigyne, ventral view. (73) Vulva, dorsal view. (74-75) Median tooth of MA, posterior view. Scale bars 1 cm (70-71), 0.5 mm (72), 0.25 mm (73), 0.1 mm (74-75)." figureDoi="http://doi.org/10.5281/zenodo.5743738" httpUri="https://zenodo.org/record/5743738/files/figure.png" pageId="27" pageNumber="301">Figs 74-75</figureCitation>
), and from the latter by a narrower inner plate of the median apophysis, which is almost hidden under the outer plate (
<figureCitation id="FDEF2A04FF8F3F5DFEFB270C68D48330" box="[325,468,514,537]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="27" pageNumber="301">Figs 118-119</figureCitation>
cf. 
<figureCitation id="FDEF2A04FF8F3F5DFDBF270C6B998330" box="[513,665,514,537]" captionStart="Figs 142-154" captionStartId="28.[151,192,1972,1993]" captionTargetBox="[169,1419,257,1942]" captionTargetId="figure-9@28.[169,1419,257,1943]" captionTargetPageId="28" captionText="Figs 142-154. Karakumosa tashkumyr sp. nov., male paratypes, in ISEA. (142) Body, ventral view. (143) Ditto, dorsal view. (144) Left chelicera, ventral view. (145, 147, 152-154) Median tooth of MA, posterior view. (146) Left male palp, ventral view. (148, 150) Bulbus, ventral view. (151) Ditto, retrolateral view. (149) Embolic division. Scale bars 1 cm (142-143), 0.5 cm (144, 146, 150-151), 0.2 mm (148), 0.1 mm (145, 147, 149, 152-154)." figureDoi="http://doi.org/10.5281/zenodo.5743760" httpUri="https://zenodo.org/record/5743760/files/figure.png" pageId="27" pageNumber="301">Figs 148, 150</figureCitation>
) and by the shape of the synembolus (
<figureCitation id="FDEF2A04FF8F3F5DFE5E272C6B3C8310" box="[480,572,546,569]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="27" pageNumber="301">Fig. 116</figureCitation>
cf. 
<figureCitation id="FDEF2A04FF8F3F5DFDD4272C6BC98310" box="[618,713,546,569]" captionStart="Figs 142-154" captionStartId="28.[151,192,1972,1993]" captionTargetBox="[169,1419,257,1942]" captionTargetId="figure-9@28.[169,1419,257,1943]" captionTargetPageId="28" captionText="Figs 142-154. Karakumosa tashkumyr sp. nov., male paratypes, in ISEA. (142) Body, ventral view. (143) Ditto, dorsal view. (144) Left chelicera, ventral view. (145, 147, 152-154) Median tooth of MA, posterior view. (146) Left male palp, ventral view. (148, 150) Bulbus, ventral view. (151) Ditto, retrolateral view. (149) Embolic division. Scale bars 1 cm (142-143), 0.5 cm (144, 146, 150-151), 0.2 mm (148), 0.1 mm (145, 147, 149, 152-154)." figureDoi="http://doi.org/10.5281/zenodo.5743760" httpUri="https://zenodo.org/record/5743760/files/figure.png" pageId="27" pageNumber="301">Fig. 149</figureCitation>
). In the sigmoid lateral edges of the epigynal atrium, the female is similar to those of 
<taxonomicName id="A2D44D02FF8F3F5DFE5E276C6B518350" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[480,593,610,633]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="repetek" status="sp. nov.">
<emphasis id="57A0EA93FF8F3F5DFE5E276C6B518350" box="[480,593,610,633]" italics="true" pageId="27" pageNumber="301">K. repetek</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8F3F5DFDE2276C6BB38350" box="[604,691,610,633]" pageId="27" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="FDEF2A04FF8F3F5DFD7A276C69D783B0" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="27" pageNumber="301">Figs 95-96</figureCitation>
) and 
<taxonomicName id="A2D44D02FF8F3F5DFEA0278C688D83B0" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[286,397,642,665]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="zyuzini" status="sp. nov.">
<emphasis id="57A0EA93FF8F3F5DFEA0278C688D83B0" box="[286,397,642,665]" italics="true" pageId="27" pageNumber="301">K. zyuzini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8F3F5DFE27278C68F283B0" box="[409,498,642,665]" pageId="27" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="FDEF2A04FF8F3F5DFDBB278C6BA783B0" box="[517,679,642,665]" captionStart="Figs 173-186" captionStartId="34.[151,192,1877,1898]" captionTargetBox="[151,1435,257,1821]" captionTargetId="figure-9@34.[151,1436,257,1827]" captionTargetPageId="34" captionText="Figs 173-186. Karakumosa zyuzini sp. nov., male paratype (173-176, 178, 180-181, 183-186) and female paratype (177, 179, 182), in MMUE. (173-174, 176) Bulbus, ventral view. (175) Embolic division. (177) Left female chelicera, ventral view. (178) Bulbus, retrolateral view. (179) Epigyne, ventral view. (180-181, 183-186) Median tooth of MA, posterior view. (182) Vulva, dorsal view. Scale bars 0.5 mm (173, 176-179), 0.25 mm (182), 0.2 (174), 0.1 mm (175, 180-181, 183-186)." figureDoi="http://doi.org/10.5281/zenodo.5743772" httpUri="https://zenodo.org/record/5743772/files/figure.png" pageId="27" pageNumber="301">Figs 179, 182</figureCitation>
), from which it differs in the shape of the posterior transverse plate (developed as a low, inverted triangle; 
<figureCitation id="FDEF2A04FF8F3F5DFD2B27CC6BF483F0" box="[661,756,706,729]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="27" pageNumber="301">Fig. 122</figureCitation>
cf. 
<figureCitation id="FDEF2A04FF8F3F5DFF0427EC680783D0" box="[186,263,738,761]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="27" pageNumber="301">Fig. 96</figureCitation>
and 
<figureCitation id="FDEF2A04FF8F3F5DFE8327EC689883D0" box="[317,408,738,761]" captionStart="Figs 173-186" captionStartId="34.[151,192,1877,1898]" captionTargetBox="[151,1435,257,1821]" captionTargetId="figure-9@34.[151,1436,257,1827]" captionTargetPageId="34" captionText="Figs 173-186. Karakumosa zyuzini sp. nov., male paratype (173-176, 178, 180-181, 183-186) and female paratype (177, 179, 182), in MMUE. (173-174, 176) Bulbus, ventral view. (175) Embolic division. (177) Left female chelicera, ventral view. (178) Bulbus, retrolateral view. (179) Epigyne, ventral view. (180-181, 183-186) Median tooth of MA, posterior view. (182) Vulva, dorsal view. Scale bars 0.5 mm (173, 176-179), 0.25 mm (182), 0.2 (174), 0.1 mm (175, 180-181, 183-186)." figureDoi="http://doi.org/10.5281/zenodo.5743772" httpUri="https://zenodo.org/record/5743772/files/figure.png" pageId="27" pageNumber="301">Fig. 179</figureCitation>
) and by the spermathecae being visibly swollen (
<figureCitation id="FDEF2A04FF8F3F5DFEF2260C68A88230" box="[332,424,770,793]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="27" pageNumber="301">Fig. 121</figureCitation>
cf. 
<figureCitation id="FDEF2A04FF8F3F5DFE6B260C6B238230" box="[469,547,770,793]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="27" pageNumber="301">Fig. 95</figureCitation>
and 
<figureCitation id="FDEF2A04FF8F3F5DFDE4260C6BB98230" box="[602,697,770,793]" captionStart="Figs 173-186" captionStartId="34.[151,192,1877,1898]" captionTargetBox="[151,1435,257,1821]" captionTargetId="figure-9@34.[151,1436,257,1827]" captionTargetPageId="34" captionText="Figs 173-186. Karakumosa zyuzini sp. nov., male paratype (173-176, 178, 180-181, 183-186) and female paratype (177, 179, 182), in MMUE. (173-174, 176) Bulbus, ventral view. (175) Embolic division. (177) Left female chelicera, ventral view. (178) Bulbus, retrolateral view. (179) Epigyne, ventral view. (180-181, 183-186) Median tooth of MA, posterior view. (182) Vulva, dorsal view. Scale bars 0.5 mm (173, 176-179), 0.25 mm (182), 0.2 (174), 0.1 mm (175, 180-181, 183-186)." figureDoi="http://doi.org/10.5281/zenodo.5743772" httpUri="https://zenodo.org/record/5743772/files/figure.png" pageId="27" pageNumber="301">Fig. 182</figureCitation>
). See also comments above under ‘Diagnosis’ of 
<taxonomicName id="A2D44D02FF8F3F5DFDDB262C6BD78210" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[613,727,802,825]" class="Arachnida" family="Lycosidae" genus="Tarentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="alticeps">
<emphasis id="57A0EA93FF8F3F5DFDDB262C6BD78210" box="[613,727,802,825]" italics="true" pageId="27" pageNumber="301">K. alticeps</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF8F3F5DFF29265F6C138430" pageId="27" pageNumber="301" type="description">
<paragraph id="656B3681FF8F3F5DFF29265F6BDB8581" blockId="27.[151,757,849,2024]" pageId="27" pageNumber="301">
<emphasis id="57A0EA93FF8F3F5DFF29265F68248241" bold="true" box="[151,292,849,872]" pageId="27" pageNumber="301">Description:</emphasis>
Male (
<typeStatus id="BA6F8823FF8F3F5DFECF265F68D58241" box="[369,469,849,872]" pageId="27" pageNumber="301" type="holotype">holotype</typeStatus>
). 
<emphasis id="57A0EA93FF8F3F5DFE59265F6B828241" box="[487,642,849,872]" italics="true" pageId="27" pageNumber="301">Measurements</emphasis>
: Carapace 11.00 long, 9.30 wide. Eye sizes and interdistances: AME 0.45, ALE 0.40, PME 0.90, PLE 0.90, AME-AME 0.38, AME-ALE 0.15, PME-PME 1.20, PME-PLE 1.15. Width of anterior eye row 2.25, of second row 2.95, of third row 3.65. Clypeus height 0.35; chelicera length 4.00. Abdomen 10.50 long, 7.20 wide. Length of leg segments: I 10.00 + 5.00 + 9.80 + 10.00 + 4.80 (39.60); II 11.00 + 4.50 + 9.50 + 10.30 + 4.90 (40.20); III 10.00 + 3.80 + 7.60 + 9.80 + 4.20 (35.40); IV 12.00 + 4.10 + 9.70 + 13.50 + 5.40 (44.70). Leg formula: IV, II, I, III.
</paragraph>
<paragraph id="656B3681FF8F3F5DFF2921BF6B8987A1" blockId="27.[151,757,849,2024]" pageId="27" pageNumber="301">
<emphasis id="57A0EA93FF8F3F5DFF2921BF68AE85E1" box="[151,430,1201,1224]" italics="true" pageId="27" pageNumber="301">Colouration in alcohol</emphasis>
(
<figureCitation id="FDEF2A04FF8F3F5DFE7321BF6B6E85E1" box="[461,622,1201,1224]" captionStart="Figs 103-114" captionStartId="23.[151,192,1821,1842]" captionTargetBox="[154,1435,260,1777]" captionTargetId="figure-9@23.[151,1436,257,1790]" captionTargetPageId="23" captionText="Figs 103-114. Karakumosa shmatkoi sp. nov.; male holotype (105-106), male paratype (103-104, 107-112) and female paratype (113- 114); specimens from Kulaly Island (103-104, 109), Zhuzguntyube stow (105-106, 113-114), Mugodzhary Mts (107), Artezian Vil. (108, 110), Tengiz (111), Tyub-Karagan Peninsula (112). (103, 105, 113) Body, dorsal view. (104, 106, 114) Ditto, ventral view. (107-112) Median tooth of MA, posterior view. Scale bars 1 cm (103-106, 113-114), 0.1 mm (107-112)." figureDoi="http://doi.org/10.5281/zenodo.5743752" httpUri="https://zenodo.org/record/5743752/files/figure.png" pageId="27" pageNumber="301">Figs 103-106</figureCitation>
, 
<figureCitation id="FDEF2A04FF8F3F5DFD3521BF6BE385E1" box="[651,739,1201,1224]" captionStart="Figs 126-135" captionStartId="25.[151,192,1838,1859]" captionTargetBox="[151,1435,257,1777]" captionTargetId="figure-9@25.[151,1436,257,1777]" captionTargetPageId="25" captionText="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." figureDoi="http://doi.org/10.5281/zenodo.5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="27" pageNumber="301">126-127</figureCitation>
): Carapace dark brown, densely covered with yellowish white setae, without two wide brownish longitudinal paramedian bands. Sternum yellow-orange-coloured, densely covered with white setae. Maxillae and labium yellowish brown, with yellow tips. Chelicerae dark brown, their frontal and lateral sides densely covered with yellowish white setae. Abdomen: dorsum densely covered with white setae, with an indistinct cardiac mark outlined by brown lines; sides and venter yellow, densely covered with white setae. Book-lung covers yellow, densely covered with white setae. Spinnerets white. All legs yellow brownish, densely covered with white setae; metatarsi and tarsi of all legs ventrally darker (brown). Palp yellow, densely covered with white setae.
</paragraph>
<paragraph id="656B3681FF8F3F5DFF29239F681786C1" blockId="27.[151,757,849,2024]" pageId="27" pageNumber="301">
<emphasis id="57A0EA93FF8F3F5DFF29239F68348781" box="[151,308,1681,1704]" italics="true" pageId="27" pageNumber="301">Palp structure</emphasis>
(
<figureCitation id="FDEF2A04FF8F3F5DFEF6239F68DE8781" box="[328,478,1681,1704]" captionStart="Figs 103-114" captionStartId="23.[151,192,1821,1842]" captionTargetBox="[154,1435,260,1777]" captionTargetId="figure-9@23.[151,1436,257,1790]" captionTargetPageId="23" captionText="Figs 103-114. Karakumosa shmatkoi sp. nov.; male holotype (105-106), male paratype (103-104, 107-112) and female paratype (113- 114); specimens from Kulaly Island (103-104, 109), Zhuzguntyube stow (105-106, 113-114), Mugodzhary Mts (107), Artezian Vil. (108, 110), Tengiz (111), Tyub-Karagan Peninsula (112). (103, 105, 113) Body, dorsal view. (104, 106, 114) Ditto, ventral view. (107-112) Median tooth of MA, posterior view. Scale bars 1 cm (103-106, 113-114), 0.1 mm (107-112)." figureDoi="http://doi.org/10.5281/zenodo.5743752" httpUri="https://zenodo.org/record/5743752/files/figure.png" pageId="27" pageNumber="301">Figs 107-112</figureCitation>
, 
<figureCitation id="FDEF2A04FF8F3F5DFE53239F6BB38781" box="[493,691,1681,1704]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="27" pageNumber="301">115-120, 123-125</figureCitation>
, 
<figureCitation id="FDEF2A04FF8F3F5DFD7D239F69C387E1" captionStart="Figs 126-135" captionStartId="25.[151,192,1838,1859]" captionTargetBox="[151,1435,257,1777]" captionTargetId="figure-9@25.[151,1436,257,1777]" captionTargetPageId="25" captionText="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." figureDoi="http://doi.org/10.5281/zenodo.5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="27" pageNumber="301">128- 135</figureCitation>
): Acutely pointed synembolic lamellae visibly convergent; median tooth developed as a large median claw with additional small teeth on its prolateral side and a prominent serrate ventral flange; proximal extension wide and hook-shaped; distance between proximal extension and median tooth wide, equal to two proximal extension widths; inner plate large, transverse-ovoid, its retrolateral shoulder extended and bent ventrad; conductor triangular, obtuse at its tip (pointed in other specimens).
</paragraph>
<paragraph id="656B3681FF8F3F5DFC81240C6C2B8370" blockId="27.[831,1437,258,1305]" pageId="27" pageNumber="301">
Female (
<typeStatus id="BA6F8823FF8F3F5DFC15240C6D0F8030" box="[939,1039,258,281]" pageId="27" pageNumber="301" type="paratype">paratype</typeStatus>
). 
<emphasis id="57A0EA93FF8F3F5DFB90240C6DCB8030" box="[1070,1227,258,281]" italics="true" pageId="27" pageNumber="301">Measurements</emphasis>
: Carapace 11.50 long, 8.60 wide. Eye sizes and interdistances: AME 0.45, ALE 0.35, PME 1.15, PLE 1.00, AME-AME 0.30, AME-ALE 0.18, PME-PME 1.20, PME-PLE 1.45. Width of anterior eye row 2.00, of second row 3.15, of third row 3.90. Clypeus height 0.50; chelicera length 4.85. Abdomen 14.00 long, 11.00 wide. Length of leg segments: I 9.80 + 4.10 + 7.00 + 6.20 + 3.50 (30.60); II 8.70 + 3.50 + 6.70 + 6.30 + 3.60 (28.80); III 8.00 + 3.60 + 5.80 + 6.80 + 3.60 (27.80); IV 10.00 + 4.50 + 7.00 + 9.50 + 3.90 (34.90). Leg formula: IV, I, II, III.
</paragraph>
<paragraph id="656B3681FF8F3F5DFC81276C6D408510" blockId="27.[831,1437,258,1305]" pageId="27" pageNumber="301">
<emphasis id="57A0EA93FF8F3F5DFC81276C6D3D8350" box="[831,1085,610,633]" italics="true" pageId="27" pageNumber="301">Colouration in alcohol</emphasis>
(
<figureCitation id="FDEF2A04FF8F3F5DFBF1276C6DEE8350" box="[1103,1262,610,633]" captionStart="Figs 103-114" captionStartId="23.[151,192,1821,1842]" captionTargetBox="[154,1435,260,1777]" captionTargetId="figure-9@23.[151,1436,257,1790]" captionTargetPageId="23" captionText="Figs 103-114. Karakumosa shmatkoi sp. nov.; male holotype (105-106), male paratype (103-104, 107-112) and female paratype (113- 114); specimens from Kulaly Island (103-104, 109), Zhuzguntyube stow (105-106, 113-114), Mugodzhary Mts (107), Artezian Vil. (108, 110), Tengiz (111), Tyub-Karagan Peninsula (112). (103, 105, 113) Body, dorsal view. (104, 106, 114) Ditto, ventral view. (107-112) Median tooth of MA, posterior view. Scale bars 1 cm (103-106, 113-114), 0.1 mm (107-112)." figureDoi="http://doi.org/10.5281/zenodo.5743752" httpUri="https://zenodo.org/record/5743752/files/figure.png" pageId="27" pageNumber="301">Figs 113, 114</figureCitation>
): Damaged and shabby specimen with abdomen detached from carapace. Carapace brown, densely covered with yellowish white setae and with two longitudinal lateral bands of brownish setae; carapace sides with wide marginal bands of white setae. Sternum brownish, densely covered with white setae. Maxillae and labium brown, with yellowish tips. Chelicerae dark brown, proximal half of frontal side densely covered with yellowish white setae. Abdomen: dorsum densely covered with yellowish white setae and with a large, wide, brownish cardiac mark; sides and venter, including book-lung covers, densely covered with white setae. Spinnerets brown. All legs and palps brownish yellow, densely covered with white setae. Palps with a claw at their tips.
</paragraph>
<paragraph id="656B3681FF8F3F5DFC81214C6C138430" blockId="27.[831,1437,258,1305]" pageId="27" pageNumber="301">
<emphasis id="57A0EA93FF8F3F5DFC81214C6D068570" box="[831,1030,1090,1113]" italics="true" pageId="27" pageNumber="301">Epigyne and vulva</emphasis>
(
<figureCitation id="FDEF2A04FF8F3F5DFBAA214C6DA68570" box="[1044,1190,1090,1113]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="27" pageNumber="301">Figs 121-122</figureCitation>
, 
<figureCitation id="FDEF2A04FF8F3F5DFB0F214C6DDD8570" box="[1201,1245,1090,1113]" captionStart="Figs 136-141" captionStartId="26.[151,192,1422,1443]" captionTargetBox="[151,1435,257,1362]" captionTargetId="figure-218@26.[151,1436,257,1362]" captionTargetPageId="26" captionText="Figs 136-141. Karakumosa shmatkoi sp. nov., female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139)." figureDoi="http://doi.org/10.5281/zenodo.5743758" httpUri="https://zenodo.org/record/5743758/files/figure.png" pageId="27" pageNumber="301">139</figureCitation>
): Epigynal atrium twice as long as wide, with markedly sigmoid lateral edges (almost S-shaped); posterior transverse plate developed as a low, inverted triangle, its central part slightly elevated and pointed backwards; spermathecae straight and visibly swollen in anterior portion, directed antero-mediad, inclined towards each other.
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF8F3F5DFC81203F6D4487C1" pageId="27" pageNumber="301" type="discussion">
<paragraph id="656B3681FF8F3F5DFC81203F6D4487C1" blockId="27.[831,1437,1329,1768]" pageId="27" pageNumber="301">
<emphasis id="57A0EA93FF8F3F5DFC81203F6AC68461" bold="true" box="[831,966,1329,1352]" pageId="27" pageNumber="301">Comments:</emphasis>
The males of 
<taxonomicName id="A2D44D02FF8F3F5DFB29203F6C268461" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1175,1318,1329,1352]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF8F3F5DFB29203F6C268461" box="[1175,1318,1329,1352]" italics="true" pageId="27" pageNumber="301">K. shmatkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8F3F5DFA85203F6C9D8461" box="[1339,1437,1329,1352]" pageId="27" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
examined display a noticeable variation in the shape of the proximal extension of the pedipalp which has either an obtuse (
<figureCitation id="FDEF2A04FF8F3F5DFC04209F6D188481" box="[954,1048,1425,1448]" captionStart="Figs 126-135" captionStartId="25.[151,192,1838,1859]" captionTargetBox="[151,1435,257,1777]" captionTargetId="figure-9@25.[151,1436,257,1777]" captionTargetPageId="25" captionText="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." figureDoi="http://doi.org/10.5281/zenodo.5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="27" pageNumber="301">Fig. 132</figureCitation>
) or pointed (
<figureCitation id="FDEF2A04FF8F3F5DFB15209F6C3D8481" box="[1195,1341,1425,1448]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="27" pageNumber="301">Figs 118-119</figureCitation>
) median shoulder. In the latter case, the proximal extension varies in its width, being visibly narrower (
<figureCitation id="FDEF2A04FF8F3F5DFA8A20DF6C9484C1" box="[1332,1428,1489,1512]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="27" pageNumber="301">Fig. 118</figureCitation>
) or wider (
<figureCitation id="FDEF2A04FF8F3F5DFC0F20FF6D138721" box="[945,1043,1521,1544]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="27" pageNumber="301">Fig. 119</figureCitation>
). It is possible that more than one closely related species are here placed under the name of 
<taxonomicName id="A2D44D02FF8F3F5DFCDA233F6AEB8761" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[868,1003,1585,1608]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF8F3F5DFCDA233F6AEB8761" box="[868,1003,1585,1608]" italics="true" pageId="27" pageNumber="301">K. shmatkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF8F3F5DFC44233F6D548761" box="[1018,1108,1585,1608]" pageId="27" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
Unfortunately, the available material does not allow us to scrutinize the problem further; currently only one variety is known from both sexes. The problem needs special attention in the future when more material of both sexes from a larger number of localities is available.
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF8F3F5DFC81220E6DBE86DE" pageId="27" pageNumber="301" type="distribution">
<paragraph id="656B3681FF8F3F5DFC81220E6DBE86DE" blockId="27.[831,1436,1792,2039]" pageId="27" pageNumber="301">
<emphasis id="57A0EA93FF8F3F5DFC81220E6AD4863E" bold="true" box="[831,980,1792,1815]" pageId="27" pageNumber="301">Distribution:</emphasis>
Known from two localities in 
<collectingRegion id="A710F863FF8F3F5DFA92220E6C98863E" box="[1324,1432,1792,1815]" country="Russia" name="Kalmykiya" pageId="27" pageNumber="301">Kalmykia</collectingRegion>
, 
<collectingCountry id="1DC37611FF8F3F5DFC81222E6A87861E" box="[831,903,1824,1847]" name="Russia" pageId="27" pageNumber="301">Russia</collectingCountry>
(
<bibRefCitation id="01454B70FF8F3F5DFC24222E6C10861E" author="Minoransky V. A. &amp; Ponomarev A. V." box="[922,1296,1824,1847]" pageId="27" pageNumber="301" pagination="82 - 92" refId="ref22760" refString="Minoransky V. A., Ponomarev A. V. 1984. Materials on the spider fauna of Kamykia [pp. 82 - 92]. In: Utochkin A. S. et al. (eds), Fauna i ekologiya paukoobraznykh. PGU Press, Perm, 159 pp. [In Russian]" type="book chapter" year="1984">Minoransky &amp; Ponomarev, 1984</bibRefCitation>
: sub 
<emphasis id="57A0EA93FF8F3F5DFAEF222E6A93867E" italics="true" pageId="27" pageNumber="301">
Lycosa 
<taxonomicName id="A2D44D02FF8F3F5DFC81224E6A93867E" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[831,915,1856,1879]" class="Arachnida" family="Lycosidae" genus="Tarentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="alticeps">alticeps</taxonomicName>
</emphasis>
; present data), and from several localities in western 
<collectingCountry id="1DC37611FF8F3F5DFC1D226E6D21865E" box="[931,1057,1888,1911]" name="Kazakhstan" pageId="27" pageNumber="301">Kazakhstan</collectingCountry>
(
<bibRefCitation id="01454B70FF8F3F5DFB86226E6A7986BE" author="Ponomarev A. V. &amp; Abdurakhmanov G. M." pageId="27" pageNumber="301" pagination="76 - 121" refId="ref23336" refString="Ponomarev A. V. &amp; Abdurakhmanov G. M. 2014. Spiders (Aranei) of the shore and islands of the northern part of Caspian Sea. The South of Russia: Ecology and Development 1: 76 - 121. [In Russian]" type="journal article" year="2014">Ponomarev &amp; Abdurakhmanov 2014</bibRefCitation>
: sub “ 
<emphasis id="57A0EA93FF8F3F5DFC79228E6D1286BE" box="[967,1042,1920,1943]" italics="true" pageId="27" pageNumber="301">Lycosa</emphasis>
” sp. 1 &amp; “ 
<emphasis id="57A0EA93FF8F3F5DFB20228E6DE986BE" box="[1182,1257,1920,1943]" italics="true" pageId="27" pageNumber="301">Lycosa</emphasis>
” sp. 2; present data) and 
<collectingCountry id="1DC37611FF8F3F5DFC0A22AE6D2A869E" box="[948,1066,1952,1975]" name="Azerbaijan" pageId="27" pageNumber="301">Azerbaijan</collectingCountry>
(
<collectingRegion id="A710F863FF8F3F5DFB8322AE6DA7869E" box="[1085,1191,1952,1975]" country="Azerbaijan" name="Abseron" pageId="27" pageNumber="301">Absheron</collectingRegion>
Peninsula, 
<collectingRegion id="A710F863FF8F3F5DFA8E22AE6C69869E" box="[1328,1385,1952,1975]" country="Azerbaijan" name="Baku" pageId="27" pageNumber="301">Baku</collectingRegion>
and Chilov Island) (
<bibRefCitation id="01454B70FF8F3F5DFBBC22CE6DB186FE" author="Schmidt P." box="[1026,1201,1984,2007]" pageId="27" pageNumber="301" pagination="439 - 484" refId="ref23590" refString="Schmidt P. 1895. Beitrag zur Kenntnis der Laufspinnen (Araneae Citigradae Thor.) Russlands. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere 8 (4): 439 - 484." type="journal article" year="1895">Schmidt, 1895</bibRefCitation>
; 
<bibRefCitation id="01454B70FF8F3F5DFB7722CE6C6086FE" author="Dunin P. M." box="[1225,1376,1984,2007]" pageId="27" pageNumber="301" pagination="45 - 60" refId="ref21954" refString="Dunin P. M. 1984. Fauna and ecology of spiders (Aranei) of Apsheron Peninsula [pp. 45 - 60]. In: Utochkin A. S. et al. (eds), Fauna i ekologiya paukoobraznykh. PGU Press, Perm, 159 pp. [In Russian]" type="book chapter" year="1984">Dunin, 1984</bibRefCitation>
: sub 
<emphasis id="57A0EA93FF8F3F5DFC8122EE6AAF86DE" box="[831,943,2016,2039]" italics="true" pageId="27" pageNumber="301">
L. 
<taxonomicName id="A2D44D02FF8F3F5DFCE222EE6AAF86DE" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[860,943,2016,2039]" class="Arachnida" family="Lycosidae" genus="Tarentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="301" phylum="Arthropoda" rank="species" species="alticeps">alticeps</taxonomicName>
</emphasis>
; present data) (
<figureCitation id="FDEF2A04FF8F3F5DFBEA22EE6DB286DE" box="[1108,1202,2016,2039]" captionStart="Fig" captionStartId="30.[152,187,999,1020]" captionTargetBox="[151,1435,257,943]" captionTargetId="figure-515@30.[151,1436,257,943]" captionTargetPageId="30" captionText="Fig. 155. Localities of four species in the genus Karakumosa gen. nov." figureDoi="http://doi.org/10.5281/zenodo.5743764" httpUri="https://zenodo.org/record/5743764/files/figure.png" pageId="27" pageNumber="301">Fig. 155</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>