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<document id="BACD448143105C053807022FEAD843E5" ID-CLB-Dataset="22413" ID-DOI="10.11646/zootaxa.4768.3.2" ID-GBIF-Dataset="43c18a9c-5545-4f3d-b285-956094e43d43" ID-ISSN="1175-5326" ID-Zenodo-Dep="3784036" ID-ZooBank="A005B93A-2070-46F5-A06E-6F6268A207EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588576355177" checkinUser="plazi" docAuthor="Hu, Fang-Shuo" docDate="2020" docId="997687A2FFCBFF95FACC324C5302A7AF" docLanguage="en" docName="zootaxa.4768.3.2.pdf" docOrigin="Zootaxa 4768 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nelmanwaslus ornatus A. Smetana 2006" docType="treatment" docVersion="6" lastPageNumber="342" masterDocId="654FFFDAFFCCFF9DFA5B34445127A409" masterDocTitle="New distributional records of Staphylinina in Taiwan, including a new species of Miobdelus Sharp (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)" masterLastPageNumber="360" masterPageNumber="334" pageNumber="341" updateTime="1698838343837" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F5EF69DFC0896BBC81D55FA225B44F8C">New distributional records of Staphylinina in Taiwan, including a new species of Miobdelus Sharp (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)</mods:title>
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<mods:date id="4A71B33E25E899D7206BFA0E27D633ED">2020</mods:date>
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<taxonomicName id="D6DF4D37FFCBFF9AFACC324C534CA22B" ID-CoL="465N4" authority="Smetana, 2006" authorityName="A. Smetana" authorityYear="2006" box="[151,619,1544,1571]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="ornatus">
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<emphasis id="23ABEAA6FFCBFF9AFACC324C508AA22A" bold="true" box="[151,429,1544,1571]" italics="true" pageId="7" pageNumber="341">Nelmanwaslus ornatus</emphasis>
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(
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)
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<emphasis id="23ABEAA6FFCBFF9AFACC32315061A286" bold="true" box="[151,326,1652,1679]" pageId="7" pageNumber="341">Type material.</emphasis>
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(male) in
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:
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, W
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, 16.
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.12.
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. MUYUPING S.env. ~
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,
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leg., 2004//
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,
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Nelmanwaslus ornatus A.
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[red label]. The specimen is in good condition, mounted on the card, the terminal segment was dissected and mounted on a plastic card by Euparal. The
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is missing segments 511 of the left antenna.
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:
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:
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[
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],
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IV/29
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/1/2011, W.
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&amp;
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,
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(
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) (
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[
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],
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IV/6
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/11/2004, C. S.
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&amp; W.
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.
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,
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(
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) (
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;
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<specimenCount id="07D9FD3DFFCBFF9AF97A33095255A36F" box="[801,882,1868,1895]" pageId="7" pageNumber="341" type="male">1 male</specimenCount>
,
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,
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logging rd. (
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), alt. ca.
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,
<geoCoordinate id="74EB5073FFCBFF9AFB2B333450C9A382" box="[368,494,1904,1931]" degrees="120.9145" direction="east" orientation="longitude" pageId="7" pageNumber="341" precision="5" value="120.9145">120.9145E</geoCoordinate>
,
<collectingMethod id="C89E4EA3FFCBFF9AFBA333345349A382" box="[504,622,1904,1931]" pageId="7" pageNumber="341">pitfall trap</collectingMethod>
no. 11.,
<date id="65611074FFCBFF9AF8943335525EA382" box="[719,889,1905,1931]" pageId="7" pageNumber="341" value="2019-09-11" valueMax="2019-09-17" valueMin="2019-09-11">
<collectingDate id="7525E99CFFCBFF9AF8943335525EA382" box="[719,889,1905,1931]" pageId="7" pageNumber="341" value="2019-09-11" valueMax="2019-09-17" valueMin="2019-09-11">1117.IX.2019</collectingDate>
</date>
,
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no. TFBC-1070517 (FSHc)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="59C5653FFFCBFF95FA9C33D050C3A4F3" lastPageId="8" lastPageNumber="342" pageId="7" pageNumber="341" type="diagnosis">
<paragraph id="116036B4FFCBFF9AFA9C33D050EFA3FF" blockId="7.[151,1437,1652,2038]" pageId="7" pageNumber="341">
<emphasis id="23ABEAA6FFCBFF9AFA9C33D05065A3A6" bold="true" box="[199,322,1940,1967]" pageId="7" pageNumber="341">Diagnosis.</emphasis>
<taxonomicName id="D6DF4D37FFCBFF9AFB1C33D05361A3A7" authorityName="A. Smetana" authorityYear="2006" box="[327,582,1940,1966]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="23ABEAA6FFCBFF9AFB1C33D05361A3A7" box="[327,582,1940,1966]" italics="true" pageId="7" pageNumber="341">Nelmanwaslus ornatus</emphasis>
</taxonomicName>
is the only species of the genus in
<collectingCountry id="69C87624FFCBFF9AF99833D0553EA3A7" box="[963,1049,1940,1966]" name="Taiwan" pageId="7" pageNumber="341">Taiwan</collectingCountry>
. In
<collectingCountry id="69C87624FFCBFF9AFE6433D055B3A3A7" box="[1087,1172,1940,1966]" name="Taiwan" pageId="7" pageNumber="341">Taiwan</collectingCountry>
,
<taxonomicName id="D6DF4D37FFCBFF9AFEC633D054BAA3A7" authorityName="A. Smetana" authorityYear="2006" box="[1181,1437,1940,1966]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="23ABEAA6FFCBFF9AFEC633D054BAA3A7" box="[1181,1437,1940,1966]" italics="true" pageId="7" pageNumber="341">Nelmanwaslus ornatus</emphasis>
</taxonomicName>
can be distinguished from the similar genus
<taxonomicName id="D6DF4D37FFCBFF9AF8CB33FC53D2A3DB" authorityName="Sharp" authorityYear="1889" box="[656,757,1976,2002]" class="Insecta" family="Staphylinidae" genus="Agelosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFCBFF9AF8CB33FC53D2A3DB" box="[656,757,1976,2002]" italics="true" pageId="7" pageNumber="341">Agelosus</emphasis>
</taxonomicName>
by the banding of grey tomentose pubescence on the elytra and relatively gracile body.
</paragraph>
<paragraph id="116036B4FFC4FF95FA9C34DC50C3A4F3" blockId="8.[151,1437,152,935]" pageId="8" pageNumber="342">
<emphasis id="23ABEAA6FFC4FF95FA9C34DC506CA4BA" bold="true" box="[199,331,152,179]" pageId="8" pageNumber="342">Bionomics.</emphasis>
Most of specimens of
<taxonomicName id="D6DF4D37FFC4FF95F80434DD53FDA4BB" authorityName="A. Smetana" authorityYear="2006" box="[607,730,153,179]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="23ABEAA6FFC4FF95F80434DD5355A4BB" box="[607,626,153,178]" italics="true" pageId="8" pageNumber="342">N</emphasis>
.
<emphasis id="23ABEAA6FFC4FF95F8DF34DD53FDA4BB" box="[644,730,153,178]" italics="true" pageId="8" pageNumber="342">ornatus</emphasis>
</taxonomicName>
were collected by pitfall trap (
<bibRefCitation id="754E4B45FFC4FF95FE1234DD55D6A4BB" author="Smetana, A." box="[1097,1265,152,179]" pageId="8" pageNumber="342" pagination="33 - 43" refId="ref17775" refString="Smetana, A. (2006) Contributions to the knowledge of the &quot; Staphylinus - complex &quot; (Coleoptera, Staphylinidae, Staphylinini) of China. Part 11. The genus Wasmannellus Bernhauer, 1920 and Nelmanwaslus gen. nov. Zootaxa, 1316 (1), 33 - 43. https: // doi. org / 10.11646 / zootaxa. 1316.1.2" type="journal article" year="2006">Smetana 2006</bibRefCitation>
, this study) or Malaise trap. The specimen from Renlun logging road was collected by a carrion baited pitfall trap set in a middle elevation mountainous forest.
</paragraph>
</subSubSection>
<subSubSection id="59C5653FFFC4FF95FA9C354055A4A54A" pageId="8" pageNumber="342" type="distribution">
<paragraph id="116036B4FFC4FF95FA9C354055A4A54A" blockId="8.[151,1437,152,935]" pageId="8" pageNumber="342">
<emphasis id="23ABEAA6FFC4FF95FA9C35405046A516" bold="true" box="[199,353,260,287]" pageId="8" pageNumber="342">Distribution.</emphasis>
<taxonomicName id="D6DF4D37FFC4FF95FB323540534CA517" authorityName="A. Smetana" authorityYear="2006" box="[361,619,260,286]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="23ABEAA6FFC4FF95FB323540534CA517" box="[361,619,260,286]" italics="true" pageId="8" pageNumber="342">Nelmanwaslus ornatus</emphasis>
</taxonomicName>
is known from mainland
<collectingCountry id="69C87624FFC4FF95F9C9354052F2A517" box="[914,981,260,286]" name="China" pageId="8" pageNumber="342">China</collectingCountry>
(
<collectingRegion id="D31BF856FFC4FF95F9BF35405509A517" box="[996,1070,260,286]" country="China" name="Fujian" pageId="8" pageNumber="342">Fujian</collectingRegion>
and
<collectingRegion id="D31BF856FFC4FF95FE3C3540558CA517" box="[1127,1195,260,286]" country="China" name="Hubei" pageId="8" pageNumber="342">Hubei</collectingRegion>
provinces) (
<bibRefCitation id="754E4B45FFC4FF95FF61354151F3A54A" author="Smetana, A." pageId="8" pageNumber="342" pagination="33 - 43" refId="ref17775" refString="Smetana, A. (2006) Contributions to the knowledge of the &quot; Staphylinus - complex &quot; (Coleoptera, Staphylinidae, Staphylinini) of China. Part 11. The genus Wasmannellus Bernhauer, 1920 and Nelmanwaslus gen. nov. Zootaxa, 1316 (1), 33 - 43. https: // doi. org / 10.11646 / zootaxa. 1316.1.2" type="journal article" year="2006">Smetana 2006</bibRefCitation>
). The species is newly recorded from
<collectingCountry id="69C87624FFC4FF95F820356C53EAA54B" box="[635,717,296,322]" name="Taiwan" pageId="8" pageNumber="342">Taiwan</collectingCountry>
(
<collectingRegion id="D31BF856FFC4FF95F887356C52A8A54A" box="[732,911,296,323]" country="Taiwan" name="Hualien" pageId="8" pageNumber="342">Hualien County</collectingRegion>
and
<collectingRegion id="D31BF856FFC4FF95F99D356D555FA54A" box="[966,1144,297,323]" country="Taiwan" name="Nantou" pageId="8" pageNumber="342">Nantou County</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="59C5653FFFC4FF95FA9C35095302A7AF" pageId="8" pageNumber="342" type="discussion">
<paragraph id="116036B4FFC4FF95FA9C35095472A5A7" blockId="8.[151,1437,152,935]" pageId="8" pageNumber="342">
<emphasis id="23ABEAA6FFC4FF95FA9C3509501CA56E" bold="true" box="[199,315,333,359]" pageId="8" pageNumber="342">Remarks.</emphasis>
When describing
<taxonomicName id="D6DF4D37FFC4FF95F85E35095212A56E" authority="(Smetana 2006)" baseAuthorityName="Smetana" baseAuthorityYear="2006" box="[517,821,332,359]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="23ABEAA6FFC4FF95F85E3509533FA56F" box="[517,536,333,358]" italics="true" pageId="8" pageNumber="342">N</emphasis>
.
<emphasis id="23ABEAA6FFC4FF95F87E3509535CA56F" box="[549,635,333,358]" italics="true" pageId="8" pageNumber="342">ornatus</emphasis>
(
<bibRefCitation id="754E4B45FFC4FF95F8D33509520AA56E" author="Smetana, A." box="[648,813,332,359]" pageId="8" pageNumber="342" pagination="33 - 43" refId="ref17775" refString="Smetana, A. (2006) Contributions to the knowledge of the &quot; Staphylinus - complex &quot; (Coleoptera, Staphylinidae, Staphylinini) of China. Part 11. The genus Wasmannellus Bernhauer, 1920 and Nelmanwaslus gen. nov. Zootaxa, 1316 (1), 33 - 43. https: // doi. org / 10.11646 / zootaxa. 1316.1.2" type="journal article" year="2006">Smetana 2006</bibRefCitation>
)
</taxonomicName>
, one characteristic detail of the pronotum was not mentioned. The pronotal disc of the
<typeStatus id="CE648816FFC4FF95FBA13534537BA583" box="[506,604,368,394]" pageId="8" pageNumber="342" type="holotype">holotype</typeStatus>
of
<taxonomicName id="D6DF4D37FFC4FF95F8DB353553D0A583" baseAuthorityName="Smetana" baseAuthorityYear="2006" box="[640,759,369,395]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="23ABEAA6FFC4FF95F8DB353553B4A583" box="[640,659,369,394]" italics="true" pageId="8" pageNumber="342">N</emphasis>
.
<emphasis id="23ABEAA6FFC4FF95F8FA353553D0A583" box="[673,759,369,394]" italics="true" pageId="8" pageNumber="342">ornatus</emphasis>
</taxonomicName>
bears two spots of golden-yellow pubescence, though in one specimen from
<collectingCountry id="69C87624FFC4FF95FB1E35D050B0A5A7" box="[325,407,404,430]" name="Taiwan" pageId="8" pageNumber="342">Taiwan</collectingCountry>
they are sparser than that of the
<typeStatus id="CE648816FFC4FF95F95F35D05241A5A7" box="[772,870,404,430]" pageId="8" pageNumber="342" type="holotype">holotype</typeStatus>
(
<figureCitation id="89E42A31FFC4FF95F92D35D052EFA5A6" box="[886,968,404,431]" captionStart="FIGURES 813" captionStartId="11.[151,264,1862,1887]" captionTargetBox="[151,1412,185,1837]" captionTargetId="figure@11.[151,1435,181,1837]" captionTargetPageId="11" captionText="FIGURES 813. 8. Miobdelus humilis sp. nov.: male, paratype. 9. Miobdelus taiwanensis taiwanensis: male. 10. Nelmanwasus ornatus: male. 1112. Sphaerobulbus miwai comb. nov.: male, lectotype of Staphylinus (Xanthocypus) miwai and attached labels; male. 13. Naddia taiwanensis: female. Scale bar: 3 mm" figureDoi="http://doi.org/10.5281/zenodo.3784040" httpUri="https://zenodo.org/record/3784040/files/figure.png" pageId="8" pageNumber="342">Fig. 10</figureCitation>
), perhaps due to its poor condition.
</paragraph>
<paragraph id="116036B4FFC4FF95FA9C35FC5302A7AF" blockId="8.[151,1437,152,935]" pageId="8" pageNumber="342">
The systematic position of
<taxonomicName id="D6DF4D37FFC4FF95FBA335FC53BCA5DB" authorityName="Smetana" authorityYear="2006" box="[504,667,440,466]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FBA335FC53BCA5DB" box="[504,667,440,466]" italics="true" pageId="8" pageNumber="342">Nelmanwaslus</emphasis>
</taxonomicName>
within
<taxonomicName id="D6DF4D37FFC4FF95F8AA35FC5258A5DB" authorityName="Latreille" authorityYear="1802" box="[753,895,440,466]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="subTribe" subTribe="Staphylinina">Staphylinina</taxonomicName>
is unclear.
<bibRefCitation id="754E4B45FFC4FF95F9A435FD5596A5DA" author="Smetana, A." box="[1023,1201,440,467]" pageId="8" pageNumber="342" pagination="33 - 43" refId="ref17775" refString="Smetana, A. (2006) Contributions to the knowledge of the &quot; Staphylinus - complex &quot; (Coleoptera, Staphylinidae, Staphylinini) of China. Part 11. The genus Wasmannellus Bernhauer, 1920 and Nelmanwaslus gen. nov. Zootaxa, 1316 (1), 33 - 43. https: // doi. org / 10.11646 / zootaxa. 1316.1.2" type="journal article" year="2006">Smetana (2006)</bibRefCitation>
described this genus based on
<taxonomicName id="D6DF4D37FFC4FF95FB5E3599505AA5FF" baseAuthorityName="Smetana" baseAuthorityYear="2006" box="[261,381,477,503]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="23ABEAA6FFC4FF95FB5E3599503FA5FF" box="[261,280,477,502]" italics="true" pageId="8" pageNumber="342">N</emphasis>
.
<emphasis id="23ABEAA6FFC4FF95FB7C3599505AA5FF" box="[295,381,477,502]" italics="true" pageId="8" pageNumber="342">ornatus</emphasis>
</taxonomicName>
and transferred
<taxonomicName id="D6DF4D37FFC4FF95F86235985535A5FE" authority="(Camercon, 1932)" baseAuthorityName="Camercon" baseAuthorityYear="1932" box="[569,1042,476,503]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="stevensi">
<emphasis id="23ABEAA6FFC4FF95F8623598521BA5FF" box="[569,828,476,502]" italics="true" pageId="8" pageNumber="342">Wasmannellus stevensi</emphasis>
(Camercon, 1932)
</taxonomicName>
to this genus. The general habitus of
<taxonomicName id="D6DF4D37FFC4FF95FAED3644507EA613" authorityName="Smetana" authorityYear="2006" box="[182,345,512,538]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FAED3644507EA613" box="[182,345,512,538]" italics="true" pageId="8" pageNumber="342">Nelmanwaslus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="D6DF4D37FFC4FF95FBB63644552AA612" authority="Bernhauer, 1920" authorityName="Bernhauer" authorityYear="1920" box="[493,1037,512,539]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FBB6364453A9A613" box="[493,654,512,538]" italics="true" pageId="8" pageNumber="342">Wasmannellus</emphasis>
Bernhauer, 1920 (
<bibRefCitation id="754E4B45FFC4FF95F90436455522A612" author="Smetana, A." box="[863,1029,512,539]" pageId="8" pageNumber="342" pagination="33 - 43" refId="ref17775" refString="Smetana, A. (2006) Contributions to the knowledge of the &quot; Staphylinus - complex &quot; (Coleoptera, Staphylinidae, Staphylinini) of China. Part 11. The genus Wasmannellus Bernhauer, 1920 and Nelmanwaslus gen. nov. Zootaxa, 1316 (1), 33 - 43. https: // doi. org / 10.11646 / zootaxa. 1316.1.2" type="journal article" year="2006">Smetana 2006</bibRefCitation>
)
</taxonomicName>
. In the same work,
<bibRefCitation id="754E4B45FFC4FF95FEB1364554BBA612" author="Smetana, A." box="[1258,1436,512,539]" pageId="8" pageNumber="342" pagination="33 - 43" refId="ref17775" refString="Smetana, A. (2006) Contributions to the knowledge of the &quot; Staphylinus - complex &quot; (Coleoptera, Staphylinidae, Staphylinini) of China. Part 11. The genus Wasmannellus Bernhauer, 1920 and Nelmanwaslus gen. nov. Zootaxa, 1316 (1), 33 - 43. https: // doi. org / 10.11646 / zootaxa. 1316.1.2" type="journal article" year="2006">Smetana (2006)</bibRefCitation>
compared
<taxonomicName id="D6DF4D37FFC4FF95FB5536605088A637" authorityName="Bernhauer" authorityYear="1920" box="[270,431,548,574]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FB5536605088A637" box="[270,431,548,574]" italics="true" pageId="8" pageNumber="342">Wasmannellus</emphasis>
</taxonomicName>
with
<taxonomicName id="D6DF4D37FFC4FF95FBAB36605217A636" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[496,816,548,575]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FBAB36605350A637" box="[496,631,548,574]" italics="true" pageId="8" pageNumber="342">Platydracus</emphasis>
Thomson, 1858
</taxonomicName>
stating that the genera are similar in habitus and share several characters but also show differences on the mandible. The male characters of
<taxonomicName id="D6DF4D37FFC4FF95FE15360C55C8A66B" authorityName="Bernhauer" authorityYear="1920" box="[1102,1263,584,610]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FE15360C55C8A66B" box="[1102,1263,584,610]" italics="true" pageId="8" pageNumber="342">Wasmannellus</emphasis>
</taxonomicName>
were not available because only one single female was known.
<bibRefCitation id="754E4B45FFC4FF95F89B36295254A68E" author="Smetana, A." box="[704,883,621,647]" pageId="8" pageNumber="342" pagination="53 - 56" refId="ref17841" refString="Smetana, A. (2008) Contributions to the knowledge of the &quot; Staphylinus - complex &quot; (Insecta: Coleoptera, Staphylinidae, Staphylinini) of China. Part 20. The genus Wasmannellus Bernhauer, 1920. Section 2. Zootaxa, 1791 (1), 53 - 56. https: // doi. org / 10.11646 / zootaxa. 1791.1.2" type="journal article" year="2008">Smetana (2008)</bibRefCitation>
described another species of
<taxonomicName id="D6DF4D37FFC4FF95FEE536285478A68F" authorityName="Bernhauer" authorityYear="1920" box="[1214,1375,620,646]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FEE536285478A68F" box="[1214,1375,620,646]" italics="true" pageId="8" pageNumber="342">Wasmannellus</emphasis>
</taxonomicName>
from
<collectingCountry id="69C87624FFC4FF95FACC36D451FDA6A3" box="[151,218,656,682]" name="China" pageId="8" pageNumber="342">China</collectingCountry>
based on five males. As in
<taxonomicName id="D6DF4D37FFC4FF95F84036D45259A6A2" authority="(Smetana 2006)" baseAuthorityName="Smetana" baseAuthorityYear="2006" box="[539,894,656,683]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95F84036D45399A6A3" box="[539,702,656,682]" italics="true" pageId="8" pageNumber="342">Nelmanwaslus</emphasis>
(
<bibRefCitation id="754E4B45FFC4FF95F89436D55251A6A2" author="Smetana, A." box="[719,886,656,683]" pageId="8" pageNumber="342" pagination="33 - 43" refId="ref17775" refString="Smetana, A. (2006) Contributions to the knowledge of the &quot; Staphylinus - complex &quot; (Coleoptera, Staphylinidae, Staphylinini) of China. Part 11. The genus Wasmannellus Bernhauer, 1920 and Nelmanwaslus gen. nov. Zootaxa, 1316 (1), 33 - 43. https: // doi. org / 10.11646 / zootaxa. 1316.1.2" type="journal article" year="2006">Smetana 2006</bibRefCitation>
)
</taxonomicName>
, the median lobe of aedeagus in this species of
<taxonomicName id="D6DF4D37FFC4FF95FACC36F0501FA6C7" authorityName="Bernhauer" authorityYear="1920" box="[151,312,692,718]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FACC36F0501FA6C7" box="[151,312,692,718]" italics="true" pageId="8" pageNumber="342">Wasmannellus</emphasis>
</taxonomicName>
is slightly asymmetrical and the paramere has asymmetrical torsion. The mandibular subdental indentation in both genera is also shared with the genus
<taxonomicName id="D6DF4D37FFC4FF95F95D369C550DA6FA" authority="(Smetana 2018)" baseAuthorityName="Smetana" baseAuthorityYear="2018" box="[774,1066,728,755]" class="Insecta" family="Staphylinidae" genus="Agelosus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95F95D369C524CA6FB" box="[774,875,728,754]" italics="true" pageId="8" pageNumber="342">Agelosus</emphasis>
(
<bibRefCitation id="754E4B45FFC4FF95F920369D5505A6FA" author="Smetana, A." box="[891,1058,729,755]" pageId="8" pageNumber="342" pagination="201 - 244" refId="ref17962" refString="Smetana, A. (2018) Review of the genera Agelosus Sharp, 1889, Apostenolinus Bernhauer, 1934 and Apecholinus Bernhauer, 1933 (Coleoptera: Staphylinidae: Staphylinini: Staphylinina). Zootaxa, 4471 (2), 201 - 244. https: // doi. org / 10.11646 / zootaxa. 4471.2.1" type="journal article" year="2018">Smetana 2018</bibRefCitation>
)
</taxonomicName>
, but this author did not consider
<taxonomicName id="D6DF4D37FFC4FF95FACC36B851DBA71F" baseAuthorityName="Smetana" baseAuthorityYear="2018" box="[151,252,764,790]" class="Insecta" family="Staphylinidae" genus="Agelosus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FACC36B851DBA71F" box="[151,252,764,790]" italics="true" pageId="8" pageNumber="342">Agelosus</emphasis>
</taxonomicName>
to be closely related to
<taxonomicName id="D6DF4D37FFC4FF95F85C36B8538DA71F" baseAuthorityName="Smetana" baseAuthorityYear="2006" box="[519,682,764,790]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95F85C36B8538DA71F" box="[519,682,764,790]" italics="true" pageId="8" pageNumber="342">Nelmanwaslus</emphasis>
</taxonomicName>
and
<taxonomicName id="D6DF4D37FFC4FF95F8BB36B852A6A71F" authorityName="Bernhauer" authorityYear="1920" box="[736,897,764,790]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95F8BB36B852A6A71F" box="[736,897,764,790]" italics="true" pageId="8" pageNumber="342">Wasmannellus</emphasis>
</taxonomicName>
. Based on the shape of mandibles and the shape of aedeagus and detailed examined of specimens of
<taxonomicName id="D6DF4D37FFC4FF95F8A737655250A733" baseAuthorityName="Smetana" baseAuthorityYear="2006" box="[764,887,801,827]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="23ABEAA6FFC4FF95F8A737655228A733" box="[764,783,801,826]" italics="true" pageId="8" pageNumber="342">N</emphasis>
.
<emphasis id="23ABEAA6FFC4FF95F97A37655250A733" box="[801,887,801,826]" italics="true" pageId="8" pageNumber="342">ornatus</emphasis>
</taxonomicName>
, both
<taxonomicName id="D6DF4D37FFC4FF95F99E3764554FA733" baseAuthorityName="Smetana" baseAuthorityYear="2006" box="[965,1128,800,826]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95F99E3764554FA733" box="[965,1128,800,826]" italics="true" pageId="8" pageNumber="342">Nelmanwaslus</emphasis>
</taxonomicName>
and
<taxonomicName id="D6DF4D37FFC4FF95FEFD37645460A733" authorityName="Bernhauer" authorityYear="1920" box="[1190,1351,800,826]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FEFD37645460A733" box="[1190,1351,800,826]" italics="true" pageId="8" pageNumber="342">Wasmannellus</emphasis>
</taxonomicName>
should belong to the
<taxonomicName id="D6DF4D37FFC4FF95FB6C370150ACA756" authorityName="Leach" authorityYear="1819" box="[311,395,837,863]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FB6C370150ACA756" box="[311,395,837,863]" italics="true" pageId="8" pageNumber="342">Ocypus</emphasis>
</taxonomicName>
group
<emphasis id="23ABEAA6FFC4FF95FB843701533AA757" box="[479,541,837,862]" italics="true" pageId="8" pageNumber="342">sensu</emphasis>
<bibRefCitation id="754E4B45FFC4FF95F87C37005271A756" author="Brunke, A. J. &amp; Smetana, A." box="[551,854,836,863]" pageId="8" pageNumber="342" pagination="745 - 758" refId="ref16526" refString="Brunke, A. J. &amp; Smetana, A. (2019) A new genus of Staphylinina and a review of major lineages (Stpahylinidae: Staphylininae: Staphylinini). Systematics and Biodiversity, 17 (8), 745 - 758. https: // doi. org / 10.1080 / 14772000.2019.1691082" type="journal article" year="2019">Brunke &amp; Smetana (2019)</bibRefCitation>
and are unrelated to
<taxonomicName id="D6DF4D37FFC4FF95FE16370055F3A757" authorityName="Thomson" authorityYear="1858" box="[1101,1236,836,862]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFC4FF95FE16370055F3A757" box="[1101,1236,836,862]" italics="true" pageId="8" pageNumber="342">Platydracus</emphasis>
</taxonomicName>
. To better understand the relationship between these two genera and other members of
<taxonomicName id="D6DF4D37FFC4FF95F9EE372C5562A78B" authorityName="Latreille" authorityYear="1802" box="[949,1093,872,898]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="subTribe" subTribe="Staphylinina">Staphylinina</taxonomicName>
, a phylogenetic approach with broader taxon sampling is required.
</paragraph>
</subSubSection>
</treatment>
</document>