426 lines
54 KiB
XML
426 lines
54 KiB
XML
<document id="9927A2B41235918F34B0EE258DC10CB5" ID-CLB-Dataset="22413" ID-DOI="10.11646/zootaxa.4768.3.2" ID-GBIF-Dataset="43c18a9c-5545-4f3d-b285-956094e43d43" ID-ISSN="1175-5326" ID-Zenodo-Dep="3784036" ID-ZooBank="A005B93A-2070-46F5-A06E-6F6268A207EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588576355177" checkinUser="plazi" docAuthor="Hu, Fang-Shuo" docDate="2020" docId="997687A2FFC4FF94FACC37BC54A1A3D6" docLanguage="en" docName="zootaxa.4768.3.2.pdf" docOrigin="Zootaxa 4768 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobulbus miwai Hu 2020, comb. nov." docType="treatment" docVersion="6" lastPageNumber="343" masterDocId="654FFFDAFFCCFF9DFA5B34445127A409" masterDocTitle="New distributional records of Staphylinina in Taiwan, including a new species of Miobdelus Sharp (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)" masterLastPageNumber="360" masterPageNumber="334" pageNumber="342" updateTime="1698838343837" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="17F896B8DAC66D3B8A661497688533C7">New distributional records of Staphylinina in Taiwan, including a new species of Miobdelus Sharp (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)</mods:title>
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<mods:name id="9A5CEC056E40AE266D45F37BE16E1B80" type="personal">
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<mods:namePart id="56A2B66E8B445B8A807CD25F87C7D848">Hu, Fang-Shuo</mods:namePart>
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<mods:title id="12191E3F3A83ED321878B669E12AD5F6">Zootaxa</mods:title>
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<mods:part id="2683F687C5D59F5C50188BFEC6C9F754">
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<mods:date id="C80953B21D4A47189956F287A11A12B2">2020</mods:date>
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<mods:number id="0C7132AD25B310A171EBE3A649862A17">2020-05-04</mods:number>
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<mods:number id="256F4243D001B8E577B09C5356260FC8">4768</mods:number>
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<treatment id="997687A2FFC4FF94FACC37BC54A1A3D6" ID-DOI="http://doi.org/10.5281/zenodo.3794830" ID-GBIF-Taxon="163885983" ID-Zenodo-Dep="3794830" LSID="urn:lsid:plazi:treatment:997687A2FFC4FF94FACC37BC54A1A3D6" httpUri="http://treatment.plazi.org/id/997687A2FFC4FF94FACC37BC54A1A3D6" lastPageId="9" lastPageNumber="343" pageId="8" pageNumber="342">
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<subSubSection id="59C5653FFFC4FF95FACC37BC5232A01A" box="[151,789,1016,1043]" pageId="8" pageNumber="342" type="nomenclature">
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<paragraph id="116036B4FFC4FF95FACC37BC5232A01A" blockId="8.[151,789,1016,1079]" box="[151,789,1016,1043]" pageId="8" pageNumber="342">
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<heading id="4A2881D8FFC4FF95FACC37BC5232A01A" bold="true" box="[151,789,1016,1043]" fontSize="11" level="1" pageId="8" pageNumber="342" reason="1">
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<emphasis id="23ABEAA6FFC4FF95FACC37BC5232A01A" bold="true" box="[151,789,1016,1043]" pageId="8" pageNumber="342">
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<taxonomicName id="D6DF4D37FFC4FF95FACC37BC53A0A01B" ID-CoL="8Y9QB" authority="(Bernhauer, 1943)" authorityName="Hu" authorityYear="2020" baseAuthorityName="Bernhauer" baseAuthorityYear="1943" box="[151,647,1016,1043]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="miwai" status="comb. nov.">
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<emphasis id="23ABEAA6FFC4FF95FACC37BC50B9A01A" bold="true" box="[151,414,1016,1043]" italics="true" pageId="8" pageNumber="342">Sphaerobulbus miwai</emphasis>
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||
(
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<bibRefCitation id="754E4B45FFC4FF95FBF537BC5358A01B" author="Bernhauer, M." box="[430,639,1016,1042]" pageId="8" pageNumber="342" pagination="169 - 188" refId="ref16487" refString="Bernhauer, M. (1943) Neuheiten der palaearktischen Staphylinidenfauna Zugleich 15 (Beitrag zur japanisch-chinesischen Fauna). Mitteilungen der Munchner Entomologischen Gesellschaft e. V., 13 (1), 169 - 188." type="journal article" year="1943">Bernhauer, 1943</bibRefCitation>
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||
)
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||
</taxonomicName>
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<taxonomicNameLabel id="389857DDFFC4FF95F8D437BD5232A01A" box="[655,789,1017,1043]" pageId="8" pageNumber="342" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="59C5653FFFC4FF95FACC3058508CA03E" box="[151,427,1052,1079]" pageId="8" pageNumber="342" type="description">
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<paragraph id="116036B4FFC4FF95FACC3058508CA03E" blockId="8.[151,789,1016,1079]" box="[151,427,1052,1079]" pageId="8" pageNumber="342">
|
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(
|
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<figureCitation id="89E42A31FFC4FF95FAC430585001A03E" box="[159,294,1052,1079]" captionStart="FIGURES 8–13" captionStartId="11.[151,264,1862,1887]" captionTargetBox="[151,1412,185,1837]" captionTargetId="figure@11.[151,1435,181,1837]" captionTargetPageId="11" captionText="FIGURES 8–13. 8. Miobdelus humilis sp. nov.: male, paratype. 9. Miobdelus taiwanensis taiwanensis: male. 10. Nelmanwasus ornatus: male. 11–12. Sphaerobulbus miwai comb. nov.: male, lectotype of Staphylinus (Xanthocypus) miwai and attached labels; male. 13. Naddia taiwanensis: female. Scale bar: 3 mm" figureDoi="http://doi.org/10.5281/zenodo.3784040" httpUri="https://zenodo.org/record/3784040/files/figure.png" pageId="8" pageNumber="342">Figs. 11–12</figureCitation>
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,
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<figureCitation id="89E42A31FFC4FF95FB693059505CA03E" box="[306,379,1053,1079]" captionStart="FIGURES 28–34" captionStartId="19.[151,264,1945,1970]" captionTargetBox="[202,1386,181,1920]" captionTargetId="figure@19.[202,1386,181,1920]" captionTargetPageId="19" captionText="FIGURES 28–34. Sphaerobulbus miwai comb. nov. 28. Aedeagus in ventral view; 29. Aedeagus in lateral view; 30. Apical portion of underside of paramere with sensory peg setae; 31. Sternite 8, male; 32. Sternite 9, male; 33. Tergite 10, male; 34. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784048" httpUri="https://zenodo.org/record/3784048/files/figure.png" pageId="8" pageNumber="342">28–34</figureCitation>
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,
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<figureCitation id="89E42A31FFC4FF95FBDD30585084A03E" box="[390,419,1052,1079]" captionStart="FIGURES 50–53" captionStartId="23.[151,264,1164,1189]" captionTargetBox="[151,1435,181,1139]" captionTargetId="figure@23.[151,1435,181,1139]" captionTargetPageId="23" captionText="FIGURES 50–53. Live specimens of Staphylinina in Taiwan: 50. Agelosus auricomus (Photo by J.-M. Chen); 51. Miobdelus humilis sp. nov., paratype (Photo by K.-W. Chan); 52. Sphaerobulbus miwai comb. nov.; 53. Thoracostrongylus formosanus." figureDoi="http://doi.org/10.5281/zenodo.3784056" httpUri="https://zenodo.org/record/3784056/files/figure.png" pageId="8" pageNumber="342">52</figureCitation>
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)
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="59C5653FFFC4FF94FACC3021505CA4F2" lastPageId="9" lastPageNumber="343" pageId="8" pageNumber="342" type="materials_examined">
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<paragraph id="116036B4FFC4FF95FACC30215052A25B" blockId="8.[151,1437,1123,2015]" pageId="8" pageNumber="342">
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<emphasis id="23ABEAA6FFC4FF95FACC30215062A076" bold="true" box="[151,325,1124,1151]" pageId="8" pageNumber="342">Type material.</emphasis>
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<typeStatus id="CE648816FFC4FF95FB103021509CA076" box="[331,443,1125,1151]" pageId="8" pageNumber="342" type="lectotype">Lectotype</typeStatus>
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(male, designated here), in
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<collectionCode id="77CEAE71FFC4FF95F8A930215260A076" box="[754,839,1125,1151]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="342" type="Museum">FMNH</collectionCode>
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: Nokosan [
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<collectingCountry id="69C87624FFC4FF95F9993020553DA077" box="[962,1050,1124,1150]" name="Taiwan" pageId="8" pageNumber="342">Taiwan</collectingCountry>
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,
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<collectingRegion id="D31BF856FFC4FF95FE7E302155F5A076" box="[1061,1234,1125,1151]" country="Taiwan" name="Nantou" pageId="8" pageNumber="342">Nantou County</collectingRegion>
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, Nenggao Mt. (
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<emphasis id="23ABEAA6FFC4FF95FFDB302751E8A097" bold="true" pageId="8" pageNumber="342">
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<collectionCode id="77CEAE71FFC4FF95FFDB302754BBA073" box="[1408,1436,1123,1146]" pageId="8" pageNumber="342">DZ</collectionCode>
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<collectionCode id="77CEAE71FFC4FF95FACC30C351E8A097" box="[151,207,1159,1182]" country="United Kingdom" name="Bristol Museum" pageId="8" pageNumber="342">ḂƜ</collectionCode>
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||
</emphasis>
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||
)], II.
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<date id="65611074FFC4FF95FB5E30CD5072A0AA" box="[261,341,1161,1187]" pageId="8" pageNumber="342" value="1919-05">
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||
<collectionCode id="77CEAE71FFC4FF95FB5E30CD503DA0AA" box="[261,282,1161,1187]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="342" type="Museum">V</collectionCode>
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.1919
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</date>
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//
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<taxonomicName id="D6DF4D37FFC4FF95FB3630CC50DBA0AB" authority="Bernh" authorityName="Bernh" authorityYear="2020" box="[365,508,1160,1186]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="miwai" status="sp. nov.">miwai Bernh</taxonomicName>
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<emphasis id="23ABEAA6FFC4FF95F85830CD5367A0AA" bold="true" box="[515,576,1161,1187]" pageId="8" pageNumber="342">
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||
<taxonomicNameLabel id="389857DDFFC4FF95F85830CD5367A0AA" box="[515,576,1161,1187]" pageId="8" pageNumber="342" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
[hand writing]//
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<taxonomicName id="D6DF4D37FFC4FF95F8A030CC52B5A0AB" authority="Bernh." authorityName="Bernh." box="[763,914,1160,1186]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="miwai">miwai Bernh.</taxonomicName>
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Typus, Xanthocypus [hand writing, red label]// Chicago NHMus, M. Bernhauer Collection// FMNHINS 3855894,
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<collectionCode id="77CEAE71FFC4FF95F9DE30E95571A0CE" box="[901,1110,1197,1223]" pageId="8" pageNumber="342">FILED MUSEUM</collectionCode>
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Pinned [QR code label]. The specimen is in poor condition, remounted on the card, its pronotal disc at the right side bears a hole from the pin. The genital segment was dissected, preserved in glycerin, in a small plastic vial pinned with specimen. Almost both antennae were separated from the head and mounted on the same card, left antenna was missing segment 9–11; right antenna was missing 4 segments, but it is difficult to recognize which segments remain. Tibia of right fore leg and right hind leg, tarsus of right fore leg, right middle leg and right hind leg are missing. Left middle leg was separated from the body and mounted on the same card.
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<typeStatus id="CE648816FFC4FF95F8FD31C05219A197" box="[678,830,1412,1438]" pageId="8" pageNumber="342" type="paralectotype">Paralectotype</typeStatus>
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(female), in
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<collectionCode id="77CEAE71FFC4FF95F98B31C15533A196" box="[976,1044,1413,1439]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="8" pageNumber="342">TARI</collectionCode>
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||
: NUMANOHIRA [
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<collectingCountry id="69C87624FFC4FF95FEB531C05461A197" box="[1262,1350,1412,1438]" name="Taiwan" pageId="8" pageNumber="342">Taiwan</collectingCountry>
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,
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<collectingRegion id="D31BF856FFC4FF95FF0A31C051CCA1CA" country="Taiwan" name="Chiayi" pageId="8" pageNumber="342">Chiayi County</collectingRegion>
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, Chaoping (
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<emphasis id="23ABEAA6FFC4FF95FB2331E35097A1B7" bold="true" box="[376,432,1447,1470]" pageId="8" pageNumber="342">
|
||
<collectionCode id="77CEAE71FFC4FF95FB2331E35097A1B7" box="[376,432,1447,1470]" country="Australia" name="Department of Natural Resources, Environment and the Arts" pageId="8" pageNumber="342" type="Herbarium">ṈṪ</collectionCode>
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</emphasis>
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),
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<date id="65611074FFC4FF95FB9131ED5378A1CA" box="[458,607,1449,1475]" pageId="8" pageNumber="342" value="1927-08-24">24-VIII-1927</date>
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//
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<taxonomicName id="D6DF4D37FFC4FF95F82131EC52BCA1CB" authority="Bernh" authorityName="Bernh" authorityYear="2020" box="[634,923,1448,1474]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="miwai" status="sp. nov.">Staphylinus miwai Bernh</taxonomicName>
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||
<emphasis id="23ABEAA6FFC4FF95F9FE31ED52C1A1CA" bold="true" box="[933,998,1449,1475]" pageId="8" pageNumber="342">
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||
<taxonomicNameLabel id="389857DDFFC4FF95F9FE31ED52C1A1CA" box="[933,998,1449,1475]" pageId="8" pageNumber="342" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
[hand writing]. The specimen in very poor condition, remounted on the plastic card, its right elytron bears a large hole from the pin. The genital segment was dissected, mounted on the same plastic card. Nearly both antennae were missing, only first two segments of left antenna remain. Left hind leg, right middle leg right hind leg, tarsus of left fore leg, right fore leg and middle left leg are missing.
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||
</paragraph>
|
||
<paragraph id="116036B4FFC4FF94FA9C3218505CA4F2" blockId="8.[151,1437,1123,2015]" lastBlockId="9.[151,1437,151,2015]" lastPageId="9" lastPageNumber="343" pageId="8" pageNumber="342">
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||
<emphasis id="23ABEAA6FFC4FF95FA9C3218530AA27E" bold="true" box="[199,557,1628,1655]" pageId="8" pageNumber="342">
|
||
Material examined.
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||
<collectingCountry id="69C87624FFC4FF95FBE73218530AA27E" box="[444,557,1628,1655]" name="Taiwan" pageId="8" pageNumber="342">TAIWAN</collectingCountry>
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||
</emphasis>
|
||
:
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||
<collectingRegion id="D31BF856FFC4FF95F864321853CDA27E" box="[575,746,1628,1655]" country="Taiwan" name="Chiayi" pageId="8" pageNumber="342">Chiayi County</collectingRegion>
|
||
:
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||
<specimenCount id="07D9FD3DFFC4FF95F8AC3219527AA27F" box="[759,861,1628,1655]" pageId="8" pageNumber="342" type="female">1 female</specimenCount>
|
||
, Tefuye Historic Trail (
|
||
<emphasis id="23ABEAA6FFC4FF95FE36321F55DEA27B" bold="true" box="[1133,1273,1627,1650]" pageId="8" pageNumber="342">Ĥởo古ǎ</emphasis>
|
||
), Alishan To.,
|
||
<geoCoordinate id="74EB5073FFC4FF95FACC32C551D2A292" box="[151,245,1665,1691]" degrees="23.4792" direction="north" orientation="latitude" pageId="8" pageNumber="342" precision="5" value="23.4792">23.4792</geoCoordinate>
|
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,
|
||
<geoCoordinate id="74EB5073FFC4FF95FAA432C4504CA292" box="[255,363,1664,1691]" degrees="120.8252" direction="east" orientation="longitude" pageId="8" pageNumber="342" precision="5" value="120.8252">120.8252</geoCoordinate>
|
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,
|
||
<date id="65611074FFC4FF95FB2E32C550DDA292" box="[373,506,1665,1691]" pageId="8" pageNumber="342" value="2019-04-30">30-IV-2019</date>
|
||
, leg. H. C. Liu (near creek) (FSHc);
|
||
<specimenCount id="07D9FD3DFFC4FF95F9D732C552FEA293" box="[908,985,1664,1691]" pageId="8" pageNumber="342" type="male">1 male</specimenCount>
|
||
, Shi Mt. race (
|
||
<emphasis id="23ABEAA6FFC4FF95FE20323B5420A29F" bold="true" box="[1147,1287,1663,1686]" pageId="8" pageNumber="342">
|
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<collectionCode id="77CEAE71FFC4FF95FE20323B55F8A29F" box="[1147,1247,1663,1686]" pageId="8" pageNumber="342">ƂƜNJ</collectionCode>
|
||
ửǎ
|
||
</emphasis>
|
||
), Alishan To,
|
||
<geoCoordinate id="74EB5073FFC4FF95FACC32E151D2A2B6" box="[151,245,1701,1727]" degrees="23.4731" direction="north" orientation="latitude" pageId="8" pageNumber="342" precision="5" value="23.4731">23.4731</geoCoordinate>
|
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,
|
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<geoCoordinate id="74EB5073FFC4FF95FB5A32E0504AA2B6" box="[257,365,1700,1727]" degrees="120.8458" direction="east" orientation="longitude" pageId="8" pageNumber="342" precision="5" value="120.8458">120.8458</geoCoordinate>
|
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, alt.
|
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<quantity id="D6279B51FFC4FF95FBF832E150DDA2B6" box="[419,506,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="2.32" pageId="8" pageNumber="342" unit="m" value="2320.0">2320 m</quantity>
|
||
,
|
||
<date id="65611074FFC4FF95F85C32E153ACA2B6" box="[519,651,1701,1727]" pageId="8" pageNumber="342" value="2020-03-08">08-III-2020</date>
|
||
, leg. Y. H. Ho (under stone) (FSHc).
|
||
<collectingRegion id="D31BF856FFC4FF95FE6E32E155C1A2B6" box="[1077,1254,1701,1727]" country="Taiwan" name="Nantou" pageId="8" pageNumber="342">Nantou County</collectingRegion>
|
||
:
|
||
<specimenCount id="07D9FD3DFFC4FF95FEAA32E15447A2B7" box="[1265,1376,1700,1727]" pageId="8" pageNumber="342" type="female">2 females</specimenCount>
|
||
, Bilu Mt. Trail (
|
||
<emphasis id="23ABEAA6FFC4FF95FB56328350BEA2D7" bold="true" box="[269,409,1735,1758]" pageId="8" pageNumber="342">ąn'Ɯṃǎ</emphasis>
|
||
), Renai To., ca.
|
||
<quantity id="D6279B51FFC4FF95F80E328C5381A2EB" box="[597,678,1736,1762]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="342" unit="m" value="2600.0">2600m</quantity>
|
||
,
|
||
<geoCoordinate id="74EB5073FFC4FF95F8EA328D5228A2EA" box="[689,783,1737,1763]" degrees="24.1901" direction="north" orientation="latitude" pageId="8" pageNumber="342" precision="5" value="24.1901">24.1901</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="74EB5073FFC4FF95F941328D52A1A2EA" box="[794,902,1737,1763]" degrees="121.3138" direction="east" orientation="longitude" pageId="8" pageNumber="342" precision="5" value="121.3138">121.3138</geoCoordinate>
|
||
,
|
||
<date id="65611074FFC4FF95F9CA328D5530A2EA" box="[913,1047,1737,1763]" pageId="8" pageNumber="342" value="2020-03-31">31-III-2020</date>
|
||
; leg. F. S. Hu & Y. J. Chen (under rock) (FSHc);
|
||
<specimenCount id="07D9FD3DFFC4FF95FB6732A950ADA30F" box="[316,394,1772,1799]" pageId="8" pageNumber="342" type="male">1 male</specimenCount>
|
||
, Meifeng (
|
||
<emphasis id="23ABEAA6FFC4FF95F85D32AF5319A30B" bold="true" box="[518,574,1771,1794]" pageId="8" pageNumber="342">đø</emphasis>
|
||
), Renai To., ca.
|
||
<quantity id="D6279B51FFC4FF95F8A732A9526BA30E" box="[764,844,1773,1799]" metricMagnitude="3" metricUnit="m" metricValue="2.113" pageId="8" pageNumber="342" unit="m" value="2113.0">2113m</quantity>
|
||
,
|
||
<geoCoordinate id="74EB5073FFC4FF95F90332A95291A30E" box="[856,950,1773,1799]" degrees="24.0973" direction="north" orientation="latitude" pageId="8" pageNumber="342" precision="5" value="24.0973">24.0973</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="74EB5073FFC4FF95F99932A85508A30F" box="[962,1071,1772,1798]" degrees="121.1764" direction="east" orientation="longitude" pageId="8" pageNumber="342" precision="5" value="121.1764">121.1764</geoCoordinate>
|
||
;
|
||
<date id="65611074FFC4FF95FE6132A955E6A30E" box="[1082,1217,1773,1799]" pageId="8" pageNumber="342" value="2020-03-31">31-III-2020</date>
|
||
; leg. F. S. Hu & Y. J. Chen (sifting leaf litters) (FSHc);
|
||
<collectingRegion id="D31BF856FFC4FF95F862335453F8A323" box="[569,735,1808,1834]" country="Taiwan" name="Taichung" pageId="8" pageNumber="342">Taichung City</collectingRegion>
|
||
:
|
||
<specimenCount id="07D9FD3DFFC4FF95F8B03355521CA323" box="[747,827,1808,1835]" pageId="8" pageNumber="342" type="male">1 male</specimenCount>
|
||
, Hseuhshan, above Shan-Liu-Gieu Hut [369
|
||
<emphasis id="23ABEAA6FFC4FF95FF63334B54ABA32F" bold="true" box="[1336,1420,1807,1830]" pageId="8" pageNumber="342">Ɯƛ±</emphasis>
|
||
],
|
||
<quantity id="D6279B51FFC4FF95FACC337051CFA346" box="[151,232,1844,1871]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="8" pageNumber="342" unit="m" value="3350.0">3350m</quantity>
|
||
, 10.
|
||
<collectionCode id="77CEAE71FFC4FF95FB4F3371500CA346" box="[276,299,1845,1871]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="342" type="Museum">V</collectionCode>
|
||
.91, A. Smetana [
|
||
<collectionCode id="77CEAE71FFC4FF95FBBE337050D0A346" box="[485,503,1844,1871]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="342">T</collectionCode>
|
||
75]//
|
||
<taxonomicName id="D6DF4D37FFC4FF95F86F33705296A346" authority="Brnh, Smetana" authorityName="Brnh, Smetana" box="[564,945,1844,1871]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="miwai">Staphylinus miwai Brnh, Smetana</taxonomicName>
|
||
det. 1991 [
|
||
<collectionCode id="77CEAE71FFC4FF95FE76337155A6A346" box="[1069,1153,1845,1871]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="342">NMNS</collectionCode>
|
||
ENT 1283-12] (
|
||
<collectionCode id="77CEAE71FFC4FF95FF62337154B7A346" box="[1337,1424,1845,1871]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="342">NMNS</collectionCode>
|
||
);
|
||
<specimenCount id="07D9FD3DFFC4FF95FACC331D51DBA37B" box="[151,252,1880,1907]" pageId="8" pageNumber="342" type="female">1 female</specimenCount>
|
||
, same data as above [
|
||
<collectionCode id="77CEAE71FFC4FF95FBA9331D5361A37A" box="[498,582,1881,1907]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="342">NMNS</collectionCode>
|
||
ENT 1283-2] (
|
||
<collectionCode id="77CEAE71FFC4FF95F8AE331D526BA37A" box="[757,844,1881,1907]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="342">NMNS</collectionCode>
|
||
);
|
||
<specimenCount id="07D9FD3DFFC4FF95F93A331D5295A37A" box="[865,946,1881,1907]" pageId="8" pageNumber="342" type="generic">1 spec.</specimenCount>
|
||
, same data as above//
|
||
<taxonomicName id="D6DF4D37FFC4FF95FEEA331D54BAA37B" authority="Brnh." authorityName="Bernh. Det. A. Smetana" authorityYear="2000" box="[1201,1437,1880,1907]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="miwai">Ocypus miwai Brnh.</taxonomicName>
|
||
det. A. Smetana 1998//
|
||
<taxonomicName id="D6DF4D37FFC4FF95FBF13339550FA39E" authority="Bernh. Det. A. Smetana, 2000" authorityName="Bernh. Det. A. Smetana" authorityYear="2000" box="[426,1064,1916,1943]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="miwai" subGenus="Ocypus">
|
||
Ocypus (Ocypus) miwai Bernh. Det. A.
|
||
<bibRefCitation id="754E4B45FFC4FF95F9253339550FA39E" author="Smetana, A. & Davies, A." box="[894,1064,1917,1943]" pageId="8" pageNumber="342" pagination="1 - 88" refId="ref18020" refString="Smetana, A. & Davies, A. (2000) Reclassification of the North temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1 - 88. https: // doi. org / 10.1206 / 0003 - 0082 (2000) 287 <0001: ROTNTT> 2.0. CO; 2" type="journal article" year="2000">Smetana, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="77CEAE71FFC4FF95FE62333955B7A39E" box="[1081,1168,1917,1943]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="342" type="Museum">FMNH</collectionCode>
|
||
);
|
||
<specimenCount id="07D9FD3DFFC4FF95FEFC333955DDA39E" box="[1191,1274,1917,1943]" pageId="8" pageNumber="342" type="generic">1 spec.</specimenCount>
|
||
, same data as above//
|
||
<taxonomicName id="D6DF4D37FFC4FF95FAA933E55269A3B2" authority="Brnh. Det. A. Smetana 2000" authorityName="Bernh. Det. A. Smetana" authorityYear="2000" box="[242,846,1952,1979]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="342" phylum="Arthropoda" rank="species" species="miwai" subGenus="Ocypus">
|
||
Ocypus (Ocypus) miwai Brnh. Det. A.
|
||
<bibRefCitation id="754E4B45FFC4FF95F8F533E55269A3B2" author="Smetana, A. & Davies, A." box="[686,846,1953,1979]" pageId="8" pageNumber="342" pagination="1 - 88" refId="ref18020" refString="Smetana, A. & Davies, A. (2000) Reclassification of the North temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1 - 88. https: // doi. org / 10.1206 / 0003 - 0082 (2000) 287 <0001: ROTNTT> 2.0. CO; 2" type="journal article" year="2000">Smetana 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="77CEAE71FFC4FF95F90533E55292A3B2" box="[862,949,1953,1979]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="342" type="Museum">FMNH</collectionCode>
|
||
);
|
||
<specimenCount id="07D9FD3DFFC4FF95F99233E5553FA3B3" box="[969,1048,1952,1979]" pageId="8" pageNumber="342" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="07D9FD3DFFC4FF95FE7E33E555ADA3B3" box="[1061,1162,1952,1979]" pageId="8" pageNumber="342" type="female">1 female</specimenCount>
|
||
, same data as above except
|
||
<quantity id="D6279B51FFC4FF95FA9633815039A3D6" box="[205,286,1989,2015]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="342" unit="m" value="3200.0">3200m</quantity>
|
||
, 8–10.
|
||
<collectionCode id="77CEAE71FFC4FF95FB3C33815059A3D6" box="[359,382,1989,2015]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="342" type="Museum">V</collectionCode>
|
||
.91, [
|
||
<collectionCode id="77CEAE71FFC4FF95FBEA338050E3A3D6" box="[433,452,1988,2015]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="342">T</collectionCode>
|
||
70] [
|
||
<collectionCode id="77CEAE71FFC4FF95FBAC3381536CA3D6" box="[503,587,1989,2015]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="342">NMNS</collectionCode>
|
||
ENT 1283-3;
|
||
<collectionCode id="77CEAE71FFC4FF95F8B633815219A3D6" box="[749,830,1989,2015]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="342">NMNS</collectionCode>
|
||
ENT 1283-4] (
|
||
<collectionCode id="77CEAE71FFC4FF95F9B133815566A3D6" box="[1002,1089,1989,2015]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="342">NMNS</collectionCode>
|
||
);
|
||
<specimenCount id="07D9FD3DFFC4FF95FE0E3381559EA3D7" box="[1109,1209,1988,2015]" pageId="8" pageNumber="342" type="female">1 female</specimenCount>
|
||
, same data as above except 8.
|
||
<collectionCode id="77CEAE71FFC5FF94FAA234DD5036A4BA" box="[249,273,153,179]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="343" type="Museum">V</collectionCode>
|
||
.91, [
|
||
<collectionCode id="77CEAE71FFC5FF94FB1E34DC507FA4BA" box="[325,344,152,179]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="343">T</collectionCode>
|
||
72] [
|
||
<collectionCode id="77CEAE71FFC5FF94FBD034DD50F8A4BA" box="[395,479,153,179]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="9" pageNumber="343">NMNS</collectionCode>
|
||
ENT 1283-13] (
|
||
<collectionCode id="77CEAE71FFC5FF94F8C334DD53C8A4BA" box="[664,751,153,179]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="9" pageNumber="343">NMNS</collectionCode>
|
||
);
|
||
<specimenCount id="07D9FD3DFFC5FF94F95934DD5277A4BA" box="[770,848,153,179]" pageId="9" pageNumber="343" type="male">1 male</specimenCount>
|
||
, Tian Pond (
|
||
<emphasis id="23ABEAA6FFC5FF94F98534D35531A4A7" bold="true" box="[990,1046,151,174]" pageId="9" pageNumber="343">Ŧô</emphasis>
|
||
), Heping Dist.,
|
||
<geoCoordinate id="74EB5073FFC5FF94FE9134DD540FA4BA" box="[1226,1320,153,179]" degrees="24.2800" direction="north" orientation="latitude" pageId="9" pageNumber="343" precision="5" value="24.28">24.2800</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="74EB5073FFC5FF94FF6834DD54BBA4BA" box="[1331,1436,153,179]" degrees="121.0260" direction="east" orientation="longitude" pageId="9" pageNumber="343" precision="5" value="121.026">121.0260</geoCoordinate>
|
||
(alt.
|
||
<quantity id="D6279B51FFC5FF94FA9034F8503AA4DE" box="[203,285,188,215]" metricMagnitude="3" metricUnit="m" metricValue="2.583" pageId="9" pageNumber="343" unit="m" value="2583.0">2583m</quantity>
|
||
),
|
||
<date id="65611074FFC5FF94FB6A34F95090A4DE" box="[305,439,189,215]" pageId="9" pageNumber="343" value="2019-06-08">08-VI-2019</date>
|
||
, leg. F. S. Hu (under stone) (FSHc);
|
||
<specimenCount id="07D9FD3DFFC5FF94F90E34F95283A4DE" box="[853,932,189,215]" pageId="9" pageNumber="343" type="male">1 male</specimenCount>
|
||
, same data as above except
|
||
<date id="65611074FFC5FF94FEBB34F95440A4DE" box="[1248,1383,189,215]" pageId="9" pageNumber="343" value="2018-09-23">23-IX-2018</date>
|
||
, leg. S.
|
||
<collectionCode id="77CEAE71FFC5FF94FAEF34A551E1A4F2" box="[180,198,225,251]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="343">T</collectionCode>
|
||
. Chang (FSHc).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59C5653FFFC5FF94FA9C354155B4A13A" pageId="9" pageNumber="343" type="description">
|
||
<paragraph id="116036B4FFC5FF94FA9C354152ECA54A" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">
|
||
<emphasis id="23ABEAA6FFC5FF94FA9C35415059A516" bold="true" box="[199,382,261,287]" pageId="9" pageNumber="343">Measurements.</emphasis>
|
||
BL: 12.07 (11.93–12.21); HL: 1.98 (1.78–2.21); HW: 2.44 (2.22–2.82); EyL: 0.63 (0.58–0.66); TL: 0.95 (0.82–1.20); PL: 2.55 (2.28–2.76); ElL: 2.21 (1.94–2.59) (mm).
|
||
</paragraph>
|
||
<paragraph id="116036B4FFC5FF94FA9C35085542A612" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">
|
||
<emphasis id="23ABEAA6FFC5FF94FA9C35085059A56E" bold="true" box="[199,382,332,359]" pageId="9" pageNumber="343">Re-description.</emphasis>
|
||
Small species. Habitus (
|
||
<figureCitation id="89E42A31FFC5FF94F8CF3509523EA56E" box="[660,793,333,359]" captionStart="FIGURES 8–13" captionStartId="11.[151,264,1862,1887]" captionTargetBox="[151,1412,185,1837]" captionTargetId="figure@11.[151,1435,181,1837]" captionTargetPageId="11" captionText="FIGURES 8–13. 8. Miobdelus humilis sp. nov.: male, paratype. 9. Miobdelus taiwanensis taiwanensis: male. 10. Nelmanwasus ornatus: male. 11–12. Sphaerobulbus miwai comb. nov.: male, lectotype of Staphylinus (Xanthocypus) miwai and attached labels; male. 13. Naddia taiwanensis: female. Scale bar: 3 mm" figureDoi="http://doi.org/10.5281/zenodo.3784040" httpUri="https://zenodo.org/record/3784040/files/figure.png" pageId="9" pageNumber="343">Figs. 11–12</figureCitation>
|
||
) with dark reddish to piceous-black body; head and neck covered with black to brownish tomentose pubescence, denser on tempora; pronotum, elytra and tergite 3–5 covered with black tomentose pubescence; tergite 6–8 covered with dense yellowish-golden tomentose pubescence, sometimes this pubescence is sparser on tergite 8; femora and tibiae dark red to piceous-black, tarsus brick red; legs covered with paler yellowish-golden tomentose pubescence, denser on fore legs; antennal segments 1–3 black but basal half of segment 2 brick red; segment 4–11 brownish red, gradually become paler.
|
||
</paragraph>
|
||
<paragraph id="116036B4FFC5FF94FA9C36615407A6C6" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">Head obtusely quadrangular, not dilated posteriad, with rounded posterior angles, wider than long (ratio: 1.23); eyes small, tempora markedly longer than length of eyes in dorsal view (ratio: 1.51); dorsal surface of head densely covered with fine punctation. Antennae moderately long, segment 1 (scape) markedly longer than other segments; segment 2 slightly shorter than segment 1 and segment 3 but markedly longer than other segments; segments 4–10 longer than wide, gradually become shorter; last segment longer than wide, asymmetrically emarginate.</paragraph>
|
||
<paragraph id="116036B4FFC5FF94FA9C369D5389A7AE" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">Thorax. Pronotum longer than wide, not dilated in anterior part; lateral margins of pronotum parallel; disc of pronotum with narrow impunctate midline, not obvious anteriorly, with weak expansion in front of posterior margin of pronotum; punctation similar to that of head. Scutellum large, with dense black setae. First four segments of front tarsus markedly dilated, sub-bilobed, covered with dense setae ventrally, sexes not distinctly different. Elytra markedly wider than long; shorter than pronotum (ratio: 0.87); anterior margin of humeral area not dilated; slightly dilated posteriad. Hind wing markedly reduced.</paragraph>
|
||
<paragraph id="116036B4FFC5FF94FA9C37F55409A7E6" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">Abdomen. Tergite 7 (fifth visible) without apical palisade fringe; tergite 2 (anterior to first fully visible tergite) with dense setae; general punctation of tergites moderately fine and dense, spread evenly on each tergite.</paragraph>
|
||
<paragraph id="116036B4FFC5FF94FA9C37BD55A8A0E2" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">
|
||
Male. Sternite 8 (
|
||
<figureCitation id="89E42A31FFC5FF94FBD537BD50C7A01A" box="[398,480,1017,1043]" captionStart="FIGURES 28–34" captionStartId="19.[151,264,1945,1970]" captionTargetBox="[202,1386,181,1920]" captionTargetId="figure@19.[202,1386,181,1920]" captionTargetPageId="19" captionText="FIGURES 28–34. Sphaerobulbus miwai comb. nov. 28. Aedeagus in ventral view; 29. Aedeagus in lateral view; 30. Apical portion of underside of paramere with sensory peg setae; 31. Sternite 8, male; 32. Sternite 9, male; 33. Tergite 10, male; 34. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784048" httpUri="https://zenodo.org/record/3784048/files/figure.png" pageId="9" pageNumber="343">Fig. 31</figureCitation>
|
||
) with obvious, deep and obtuse medioapical emargination; sternite 9 (
|
||
<figureCitation id="89E42A31FFC5FF94FEA537BD5476A01A" box="[1278,1361,1017,1043]" captionStart="FIGURES 28–34" captionStartId="19.[151,264,1945,1970]" captionTargetBox="[202,1386,181,1920]" captionTargetId="figure@19.[202,1386,181,1920]" captionTargetPageId="19" captionText="FIGURES 28–34. Sphaerobulbus miwai comb. nov. 28. Aedeagus in ventral view; 29. Aedeagus in lateral view; 30. Apical portion of underside of paramere with sensory peg setae; 31. Sternite 8, male; 32. Sternite 9, male; 33. Tergite 10, male; 34. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784048" httpUri="https://zenodo.org/record/3784048/files/figure.png" pageId="9" pageNumber="343">Fig. 32</figureCitation>
|
||
) without extended basal portion, apical portion with deep and narrow medioapical emargination. Tergite 10 (
|
||
<figureCitation id="89E42A31FFC5FF94FF1B305954B3A03E" box="[1344,1428,1053,1079]" captionStart="FIGURES 28–34" captionStartId="19.[151,264,1945,1970]" captionTargetBox="[202,1386,181,1920]" captionTargetId="figure@19.[202,1386,181,1920]" captionTargetPageId="19" captionText="FIGURES 28–34. Sphaerobulbus miwai comb. nov. 28. Aedeagus in ventral view; 29. Aedeagus in lateral view; 30. Apical portion of underside of paramere with sensory peg setae; 31. Sternite 8, male; 32. Sternite 9, male; 33. Tergite 10, male; 34. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784048" httpUri="https://zenodo.org/record/3784048/files/figure.png" pageId="9" pageNumber="343">Fig. 33</figureCitation>
|
||
) subtrapezoidal, with a narrow arcuate apex; numerous brownish setae on posterior margin. Aedeagus (
|
||
<figureCitation id="89E42A31FFC5FF94FF55300554B3A052" box="[1294,1428,1089,1115]" captionStart="FIGURES 28–34" captionStartId="19.[151,264,1945,1970]" captionTargetBox="[202,1386,181,1920]" captionTargetId="figure@19.[202,1386,181,1920]" captionTargetPageId="19" captionText="FIGURES 28–34. Sphaerobulbus miwai comb. nov. 28. Aedeagus in ventral view; 29. Aedeagus in lateral view; 30. Apical portion of underside of paramere with sensory peg setae; 31. Sternite 8, male; 32. Sternite 9, male; 33. Tergite 10, male; 34. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784048" httpUri="https://zenodo.org/record/3784048/files/figure.png" pageId="9" pageNumber="343">Figs. 28–29</figureCitation>
|
||
) large, with both median lobe and paramere asymmetrical; median lobe in ventral view sub-parallel along most of its length, at about midlength, gradually narrowing to left side; bulbus markedly spherical. Paramere slender, not extended beyond median lobe, slightly wider before apical portion; apical portion of underside of paramere (
|
||
<figureCitation id="89E42A31FFC5FF94FF1F30E954B3A0CE" box="[1348,1428,1197,1223]" captionStart="FIGURES 28–34" captionStartId="19.[151,264,1945,1970]" captionTargetBox="[202,1386,181,1920]" captionTargetId="figure@19.[202,1386,181,1920]" captionTargetPageId="19" captionText="FIGURES 28–34. Sphaerobulbus miwai comb. nov. 28. Aedeagus in ventral view; 29. Aedeagus in lateral view; 30. Apical portion of underside of paramere with sensory peg setae; 31. Sternite 8, male; 32. Sternite 9, male; 33. Tergite 10, male; 34. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784048" httpUri="https://zenodo.org/record/3784048/files/figure.png" pageId="9" pageNumber="343">Fig. 30</figureCitation>
|
||
) without sensory peg setae but bearing few long setae on the margins of both sides apically.
|
||
</paragraph>
|
||
<paragraph id="116036B4FFC5FF94FA9C30B155B4A13A" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">
|
||
Female genital segment. Tergite 10 (
|
||
<figureCitation id="89E42A31FFC5FF94F80630B15389A106" box="[605,686,1269,1295]" captionStart="FIGURES 28–34" captionStartId="19.[151,264,1945,1970]" captionTargetBox="[202,1386,181,1920]" captionTargetId="figure@19.[202,1386,181,1920]" captionTargetPageId="19" captionText="FIGURES 28–34. Sphaerobulbus miwai comb. nov. 28. Aedeagus in ventral view; 29. Aedeagus in lateral view; 30. Apical portion of underside of paramere with sensory peg setae; 31. Sternite 8, male; 32. Sternite 9, male; 33. Tergite 10, male; 34. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784048" httpUri="https://zenodo.org/record/3784048/files/figure.png" pageId="9" pageNumber="343">Fig. 34</figureCitation>
|
||
) subtrapezoidal, similar to male tergite 10, apical margin of female tergite 10 more sharply projecting than that of male; setae on apical margin denser than male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59C5653FFFC5FF94FA9C3178509EA2EA" pageId="9" pageNumber="343" type="diagnosis">
|
||
<paragraph id="116036B4FFC5FF94FA9C317854A1A196" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">
|
||
<emphasis id="23ABEAA6FFC5FF94FA9C31785019A15E" bold="true" box="[199,318,1340,1367]" pageId="9" pageNumber="343">Diagnosis.</emphasis>
|
||
Due to the relatively small size of
|
||
<taxonomicName id="D6DF4D37FFC5FF94F8F731795232A15E" authorityName="Bernh. Det. A. Smetana" authorityYear="2000" box="[684,789,1341,1367]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="species" species="miwai">
|
||
<emphasis id="23ABEAA6FFC5FF94F8F7317953E0A15E" box="[684,711,1341,1367]" italics="true" pageId="9" pageNumber="343">Sp</emphasis>
|
||
.
|
||
<emphasis id="23ABEAA6FFC5FF94F888317A5232A15E" box="[723,789,1342,1367]" italics="true" pageId="9" pageNumber="343">miwai</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="D6279B51FFC5FF94F978317952B6A15E" box="[803,913,1341,1367]" metricMagnitude="-2" metricUnit="m" metricValue="1.207" pageId="9" pageNumber="343" unit="mm" value="12.07">12.07 mm</quantity>
|
||
on average), it can only be confused with
|
||
<taxonomicName id="D6DF4D37FFC5FF94FF0A3179507AA172" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="species" species="pusio">
|
||
<emphasis id="23ABEAA6FFC5FF94FF0A3179544BA15E" box="[1361,1388,1341,1367]" italics="true" pageId="9" pageNumber="343">Sp</emphasis>
|
||
.
|
||
<emphasis id="23ABEAA6FFC5FF94FF23317A5190A172" italics="true" pageId="9" pageNumber="343">pusio</emphasis>
|
||
Smetana, 2005
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D6DF4D37FFC5FF94FBCA3125538CA172" authority="Smetana, 2016" authorityName="Smetana" authorityYear="2016" box="[401,683,1377,1403]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="species" species="radani">
|
||
<emphasis id="23ABEAA6FFC5FF94FBCA3125508BA172" box="[401,428,1377,1403]" italics="true" pageId="9" pageNumber="343">Sp</emphasis>
|
||
.
|
||
<emphasis id="23ABEAA6FFC5FF94FBE331255326A172" box="[440,513,1377,1403]" italics="true" pageId="9" pageNumber="343">radani</emphasis>
|
||
Smetana, 2016
|
||
</taxonomicName>
|
||
. However, it can be easily distinguished from these two species by the presence of dense yellowish-golden tomentose pubescence on the tergites 6–8 sometimes even covering tergites 6–7.
|
||
</paragraph>
|
||
<paragraph id="116036B4FFC5FF94FA9C31ED544FA202" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">
|
||
<taxonomicName id="D6DF4D37FFC5FF94FA9C31ED5091A1CA" box="[199,438,1449,1475]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="species" species="miwai">
|
||
<emphasis id="23ABEAA6FFC5FF94FA9C31ED5091A1CA" box="[199,438,1449,1475]" italics="true" pageId="9" pageNumber="343">Sphaerobulbus miwai</emphasis>
|
||
</taxonomicName>
|
||
is the only member of this genus in
|
||
<collectingCountry id="69C87624FFC5FF94F90B31ED5283A1CA" box="[848,932,1449,1475]" name="Taiwan" pageId="9" pageNumber="343">Taiwan</collectingCountry>
|
||
. It can be distinguished from almost all other
|
||
<taxonomicName id="D6DF4D37FFC5FF94FACC31895006A1EE" authorityName="Latreille" authorityYear="1802" box="[151,289,1485,1511]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="subTribe" subTribe="Staphylinina">Staphylinina</taxonomicName>
|
||
by the single simple tooth on each mandible. This character is shared with the genus
|
||
<taxonomicName id="D6DF4D37FFC5FF94FE9531895471A1EE" authorityName="Bernhauer" authorityYear="1933" box="[1230,1366,1485,1511]" class="Insecta" family="Staphylinidae" genus="Apecholinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="23ABEAA6FFC5FF94FE9531895471A1EE" box="[1230,1366,1485,1511]" italics="true" pageId="9" pageNumber="343">Apecholinus</emphasis>
|
||
</taxonomicName>
|
||
but in
|
||
<collectingCountry id="69C87624FFC5FF94FACC31B551CDA202" box="[151,234,1521,1547]" name="Taiwan" pageId="9" pageNumber="343">Taiwan</collectingCountry>
|
||
,
|
||
<taxonomicName id="D6DF4D37FFC5FF94FAAE31B55046A202" box="[245,353,1521,1547]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="species" species="miwai">
|
||
<emphasis id="23ABEAA6FFC5FF94FAAE31B55037A202" box="[245,272,1521,1547]" italics="true" pageId="9" pageNumber="343">Sp</emphasis>
|
||
.
|
||
<emphasis id="23ABEAA6FFC5FF94FB4531B65046A202" box="[286,353,1522,1547]" italics="true" pageId="9" pageNumber="343">miwai</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by the elytra markedly shorter than pronotum and the much smaller size.
|
||
</paragraph>
|
||
<paragraph id="116036B4FFC5FF94FA9C3250509EA2EA" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">
|
||
<emphasis id="23ABEAA6FFC5FF94FA9C3250506FA226" bold="true" box="[199,328,1556,1583]" pageId="9" pageNumber="343">Bionomics.</emphasis>
|
||
Based on an import of locality name into Google Earth from collection data (
|
||
<bibRefCitation id="754E4B45FFC5FF94FEE33251544BA226" author="Bernhauer, M." box="[1208,1388,1557,1583]" pageId="9" pageNumber="343" pagination="169 - 188" refId="ref16487" refString="Bernhauer, M. (1943) Neuheiten der palaearktischen Staphylinidenfauna Zugleich 15 (Beitrag zur japanisch-chinesischen Fauna). Mitteilungen der Munchner Entomologischen Gesellschaft e. V., 13 (1), 169 - 188." type="journal article" year="1943">Bernhauer 1943</bibRefCitation>
|
||
and this study), the species occurs from
|
||
<quantity id="D6279B51FFC5FF94F86B327C53E9A25A" box="[560,718,1592,1619]" metricMagnitude="3" metricUnit="m" metricValue="2.7314999999999996" metricValueMax="3.35" metricValueMin="2.113" pageId="9" pageNumber="343" unit="m" value="2731.5" valueMax="3350.0" valueMin="2113.0">2113–3350 m</quantity>
|
||
. Apparently, this species only occurs on high mountains. Some specimens of
|
||
<taxonomicName id="D6DF4D37FFC5FF94FB76321950BFA27E" box="[301,408,1629,1655]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="species" species="miwai">
|
||
<emphasis id="23ABEAA6FFC5FF94FB763219506FA27E" box="[301,328,1629,1655]" italics="true" pageId="9" pageNumber="343">Sp</emphasis>
|
||
.
|
||
<emphasis id="23ABEAA6FFC5FF94FB0E321A50BFA27E" box="[341,408,1630,1655]" italics="true" pageId="9" pageNumber="343">miwai</emphasis>
|
||
</taxonomicName>
|
||
were collected by pitfall traps set in
|
||
<taxonomicName id="D6DF4D37FFC5FF94F97332195242A27E" box="[808,869,1629,1655]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="343" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="23ABEAA6FFC5FF94F97332195242A27E" box="[808,869,1629,1655]" italics="true" pageId="9" pageNumber="343">Abies</emphasis>
|
||
</taxonomicName>
|
||
forest (as in photos 8 and
|
||
<quantity id="D6279B51FFC5FF94FED83219558BA27E" box="[1155,1196,1629,1655]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="9" pageNumber="343" unit="in" value="9.0">9 in</quantity>
|
||
<bibRefCitation id="754E4B45FFC5FF94FEE832195474A27E" author="Smetana, A." box="[1203,1363,1628,1655]" pageId="9" pageNumber="343" pagination="1 - 145" refId="ref17430" refString="Smetana, A. (1995) Revision of the tribes Quediini and Tanygnathinini. Part III. Taiwan. (Coleoptera: Staphylinidae). National Museum of Natural Science, Taichung city, Special Publication, 6, 1 - 145." type="journal article" year="1995">Smetana 1995</bibRefCitation>
|
||
: p.13). The specimens from Tian Pond were collected from under stones on a grassland near the trail. The specimen from Tefuye Historic Trail was collected from under a stone near a creek. The specimen from Meifeng was collected by sifting leaf litters in forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59C5653FFFC5FF94FA9C32A854A1A346" pageId="9" pageNumber="343" type="distribution">
|
||
<paragraph id="116036B4FFC5FF94FA9C32A854A1A346" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">
|
||
<emphasis id="23ABEAA6FFC5FF94FA9C32A8507BA30E" bold="true" box="[199,348,1772,1799]" pageId="9" pageNumber="343">Distribution.</emphasis>
|
||
<taxonomicName id="D6DF4D37FFC5FF94FB3B32A9536DA30E" box="[352,586,1773,1799]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="species" species="miwai">
|
||
<emphasis id="23ABEAA6FFC5FF94FB3B32A9536DA30E" box="[352,586,1773,1799]" italics="true" pageId="9" pageNumber="343">Sphaerobulbus miwai</emphasis>
|
||
</taxonomicName>
|
||
is restricted to high elevation mountainous areas in central and southern
|
||
<collectingCountry id="69C87624FFC5FF94FF1732A954BBA30E" box="[1356,1436,1773,1799]" name="Taiwan" pageId="9" pageNumber="343">Taiwan</collectingCountry>
|
||
including
|
||
<collectingRegion id="D31BF856FFC5FF94FB5E33555081A322" box="[261,422,1809,1835]" country="Taiwan" name="Chiayi" pageId="9" pageNumber="343">Chiayi County</collectingRegion>
|
||
,
|
||
<collectingRegion id="D31BF856FFC5FF94FBE93355537DA322" box="[434,602,1809,1835]" country="Taiwan" name="Nantou" pageId="9" pageNumber="343">Nantou County</collectingRegion>
|
||
and
|
||
<collectingRegion id="D31BF856FFC5FF94F8CA3355520AA322" box="[657,813,1809,1835]" country="Taiwan" name="Taichung" pageId="9" pageNumber="343">Taichung City</collectingRegion>
|
||
. The reduced hindwings suggest that
|
||
<taxonomicName id="D6DF4D37FFC5FF94FE9133555410A322" box="[1226,1335,1809,1835]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="species" species="miwai">
|
||
<emphasis id="23ABEAA6FFC5FF94FE91335555C2A322" box="[1226,1253,1809,1835]" italics="true" pageId="9" pageNumber="343">Sp</emphasis>
|
||
.
|
||
<emphasis id="23ABEAA6FFC5FF94FEAF33565410A322" box="[1268,1335,1810,1835]" italics="true" pageId="9" pageNumber="343">miwai</emphasis>
|
||
</taxonomicName>
|
||
is an endemic species to the island with distribution restricted to the high mountainous areas of central and southern
|
||
<collectingCountry id="69C87624FFC5FF94FF75337154A6A346" box="[1326,1409,1845,1871]" name="Taiwan" pageId="9" pageNumber="343">Taiwan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59C5653FFFC5FF94FA9C331D54A1A3D6" pageId="9" pageNumber="343" type="discussion">
|
||
<paragraph id="116036B4FFC5FF94FA9C331D54A1A3D6" blockId="9.[151,1437,151,2015]" pageId="9" pageNumber="343">
|
||
<emphasis id="23ABEAA6FFC5FF94FA9C331D501CA37A" bold="true" box="[199,315,1881,1907]" pageId="9" pageNumber="343">Remarks.</emphasis>
|
||
<bibRefCitation id="754E4B45FFC5FF94FB18331D5321A37A" author="Bernhauer, M." box="[323,518,1881,1907]" pageId="9" pageNumber="343" pagination="169 - 188" refId="ref16487" refString="Bernhauer, M. (1943) Neuheiten der palaearktischen Staphylinidenfauna Zugleich 15 (Beitrag zur japanisch-chinesischen Fauna). Mitteilungen der Munchner Entomologischen Gesellschaft e. V., 13 (1), 169 - 188." type="journal article" year="1943">Bernhauer (1943)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="D6DF4D37FFC5FF94F825331D52C8A37A" box="[638,1007,1881,1907]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="species" species="miwai" subGenus="Xanthocypus">
|
||
<emphasis id="23ABEAA6FFC5FF94F825331D53D9A37A" box="[638,766,1881,1907]" italics="true" pageId="9" pageNumber="343">Staphylinus</emphasis>
|
||
(
|
||
<emphasis id="23ABEAA6FFC5FF94F955331D52BBA37A" box="[782,924,1881,1907]" italics="true" pageId="9" pageNumber="343">Xanthocypus</emphasis>
|
||
)
|
||
<emphasis id="23ABEAA6FFC5FF94F9F7331E52C8A37A" box="[940,1007,1882,1907]" italics="true" pageId="9" pageNumber="343">miwai</emphasis>
|
||
</taxonomicName>
|
||
based on two specimens from
|
||
<collectingCountry id="69C87624FFC5FF94FF1E331D54BEA37A" box="[1349,1433,1881,1907]" name="Taiwan" pageId="9" pageNumber="343">Taiwan</collectingCountry>
|
||
. Herein, I re-examined the
|
||
<typeStatus id="CE648816FFC5FF94FBE1333950CEA39E" box="[442,489,1917,1943]" pageId="9" pageNumber="343">type</typeStatus>
|
||
series of this species and discovered that its morphology fits the generic definition of the genus
|
||
<taxonomicName id="D6DF4D37FFC5FF94FB5E33E55375A3B2" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[261,594,1953,1979]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="23ABEAA6FFC5FF94FB5E33E5508EA3B2" box="[261,425,1953,1979]" italics="true" pageId="9" pageNumber="343">Sphaerobulbus</emphasis>
|
||
<bibRefCitation id="754E4B45FFC5FF94FBF433E55375A3B2" author="Smetana, A." box="[431,594,1953,1979]" pageId="9" pageNumber="343" pagination="57 - 135" refId="ref17576" refString="Smetana, A. (2003) Contributions to the knowledge of the genera of the " Staphylinus - complex " (Coleoptera: Staphylinidae) of China Part 4. Key to the Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n., Aulacocypus and Apecholinus, and comments on the genus Protocypus. Folia Heyrovskyana, 11 (2), 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
based on the presence of only one simple tooth on the medial margin of each mandible and remarkably spherical bulbus of the aedeagus. Hence, I move this species to the genus
|
||
<taxonomicName id="D6DF4D37FFC5FF94FE8033815459A3D6" authorityName="Smetana" authorityYear="2003" box="[1243,1406,1989,2015]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="343" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="23ABEAA6FFC5FF94FE8033815459A3D6" box="[1243,1406,1989,2015]" italics="true" pageId="9" pageNumber="343">Sphaerobulbus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |