493 lines
71 KiB
XML
493 lines
71 KiB
XML
<document ID-DOI="10.11646/zootaxa.5256.5.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7758872" ID-ZooBank="E5B9708E-D626-4923-825A-A9FD59AE4024" approvalRequired="3" approvalRequired_for_textStreams="2" approvalRequired_for_treatments="1" checkinTime="1679468565563" checkinUser="plazi" docAuthor="Matalin, Andrey V. & Wiesner, Jürgen" docDate="2023" docId="050B87D78D452468E1B818ACFBF37514" docLanguage="en" docName="zootaxa.5256.5.1.pdf" docOrigin="Zootaxa 5256 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Therates fruhstorferi subsp. australis Matalin et Wiesner 2023, ssp. n." docType="treatment" docVersion="2" lastPageNumber="407" masterDocId="F932FFAF8D40246EE12F1976FFC07601" masterDocTitle="Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)" masterLastPageNumber="433" masterPageNumber="401" pageNumber="406" updateTime="1679469078842" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Matalin, Andrey V.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-7790-8709</mods:nameIdentifier>
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<mods:affiliation>Moscow State Pedagogical University, Department of Zoology & Ecology, Kibalchicha str. 6, build. 3, Moscow 129164, Russia. andrei-matalin @ yandex. ru; https: // orcid. org / 0000 - 0002 - 7790 - 8709 & Pirogov National Research Medical University, Department of Biology, Ostrovitianova str. 1, Moscow 117997, Russia.</mods:affiliation>
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<mods:nameIdentifier type="email">andrei-matalin@yandex.ru</mods:nameIdentifier>
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</mods:name>
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||
<mods:name type="personal">
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||
<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wiesner, Jürgen</mods:namePart>
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<mods:affiliation>Dresdener Ring 11, D- 38444 Wolfsburg, Germany.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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||
<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5256</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>401</mods:start>
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<mods:end>433</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.5.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5256.5.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7758872</mods:identifier>
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<mods:identifier type="ZooBank">E5B9708E-D626-4923-825A-A9FD59AE4024</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:050B87D78D452468E1B818ACFBF37514" httpUri="http://treatment.plazi.org/id/050B87D78D452468E1B818ACFBF37514" lastPageId="6" lastPageNumber="407" pageId="5" pageNumber="406">
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<subSubSection box="[151,855,474,501]" pageId="5" pageNumber="406" type="nomenclature">
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<paragraph blockId="5.[151,855,474,537]" box="[151,855,474,501]" pageId="5" pageNumber="406">
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<heading bold="true" box="[151,855,474,501]" fontSize="11" level="1" pageId="5" pageNumber="406" reason="1">
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<emphasis bold="true" box="[151,855,474,501]" pageId="5" pageNumber="406">
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<taxonomicName authority="Matalin et Wiesner" authorityName="Matalin et Wiesner" authorityYear="2023" box="[151,765,474,501]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" status="ssp. n." subSpecies="australis">
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<emphasis bold="true" box="[151,518,474,501]" italics="true" pageId="5" pageNumber="406">Therates fruhstorferi australis</emphasis>
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Matalin et Wiesner
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</taxonomicName>
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,
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<taxonomicNameLabel box="[777,855,475,501]" pageId="5" pageNumber="406" rank="subSpecies">ssp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,620,510,537]" pageId="5" pageNumber="406" type="description">
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<paragraph blockId="5.[151,855,474,537]" box="[151,620,510,537]" pageId="5" pageNumber="406">
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<figureCitation box="[151,249,511,537]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="5" pageNumber="406">Figs 7, 8</figureCitation>
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,
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<figureCitation box="[262,334,511,537]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="5" pageNumber="406">33, 34</figureCitation>
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,
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<figureCitation box="[346,418,510,537]" captionStart="FIGURES 50–57" captionStartId="15.[152,270,1909,1933]" captionTargetBox="[320,1268,195,1860]" captionTargetId="figure-17@15.[303,1285,181,1885]" captionTargetPageId="15" captionText="FIGURES 50–57. Therates fruhstorferi ssp., labrum: 50–55—T. fruhstorferi fruhstorferi (50, 51—Vietnam, Montes Mauson; 52—Vietnam, Na Hang; 53—Vietnam, Tam Dao; 54, 55—Laos, Ban Nape); 56, 57—T. fruhstorferi australis ssp. n. (56—Vietnam, Kon Ka Kinh N.P.; 57—Vietnam, Pak Khe River); 50, 52, 54, 56—males; 51, 53, 55, 57—females; 50, 51—paralectotypes; 56—holotype; 57—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758882" httpUri="https://zenodo.org/record/7758882/files/figure.png" pageId="5" pageNumber="406">56, 57</figureCitation>
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,
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<figureCitation box="[430,502,511,537]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="5" pageNumber="406">88, 89</figureCitation>
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,
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<figureCitation box="[514,616,511,537]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="5" pageNumber="406">126–130</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="406" type="materials_examined">
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<paragraph blockId="5.[151,1437,582,2013]" pageId="5" pageNumber="406">
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<emphasis bold="true" box="[151,328,582,609]" pageId="5" pageNumber="406">Type material.</emphasis>
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<materialsCitation collectingDate="2016-05-09" collectingDateMax="2016-05-22" collectingDateMin="2016-05-09" collectionCode="NEE, ZIN" collectorName="D. Fedorenko" country="Vietnam" elevation="890" latitude="14.203055" location="Kon Ka Kinh Natn. Park" longLatPrecision="21" longitude="108.315" municipality="Pleiku" pageId="5" pageNumber="406" specimenCount="1" stateProvince="Gia Lai" typeStatus="holotype">
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<typeStatus box="[338,456,583,609]" pageId="5" pageNumber="406">HOLOTYPE</typeStatus>
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||
, Ô—
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||
<collectingCountry box="[521,620,584,611]" name="Vietnam" pageId="5" pageNumber="406">Vietnam</collectingCountry>
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||
,
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||
<collectingRegion box="[633,792,583,609]" country="Vietnam" name="Gia Lai" pageId="5" pageNumber="406">Gia Lai Prov.</collectingRegion>
|
||
, ~
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||
<quantity box="[820,895,583,609]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="406" unit="km" value="40.0">40 km</quantity>
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||
<collectionCode box="[904,958,583,609]" pageId="5" pageNumber="406">NEE</collectionCode>
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of
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<collectingMunicipality box="[1002,1076,583,609]" pageId="5" pageNumber="406">Pleiku</collectingMunicipality>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:050B87D78D452468E1B818ACFBF37514:887D601A8D45246BE56D1B31FAA67460" box="[1090,1382,583,609]" country="Vietnam" latitude="14.203055" longLatPrecision="21" longitude="108.315" municipality="Pleiku" name="Kon Ka Kinh Natn. Park" pageId="5" pageNumber="406" stateProvince="Gia Lai">Kon Ka Kinh Natn. Park</location>
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, h =
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||
<quantity box="[151,228,619,645]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="5" pageNumber="406" unit="m" value="890.0">
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||
<elevation box="[151,228,619,645]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="5" pageNumber="406" unit="m" value="890.0">890 m</elevation>
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</quantity>
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,
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<geoCoordinate box="[244,381,619,645]" degrees="14" direction="north" minutes="12" orientation="latitude" pageId="5" pageNumber="406" precision="15" seconds="11" value="14.203055">
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14
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<superScript attach="none" box="[272,280,619,633]" fontSize="6" pageId="5" pageNumber="406">o</superScript>
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12’11’’N
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</geoCoordinate>
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<geoCoordinate box="[392,543,618,645]" degrees="108" direction="east" minutes="18" orientation="longitude" pageId="5" pageNumber="406" precision="15" seconds="54" value="108.315">
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108
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<superScript attach="none" box="[434,442,619,633]" fontSize="6" pageId="5" pageNumber="406">o</superScript>
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18’54’’E
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</geoCoordinate>
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||
,
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||
<date box="[559,704,619,645]" pageId="5" pageNumber="406" value="2016-05-09" valueMax="2016-05-22" valueMin="2016-05-09">
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<collectingDate box="[559,704,619,645]" pageId="5" pageNumber="406" value="2016-05-09" valueMax="2016-05-22" valueMin="2016-05-09">9–22.V.2016</collectingDate>
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||
</date>
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||
, leg.
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||
<collectorName box="[771,930,619,645]" pageId="5" pageNumber="406">D. Fedorenko</collectorName>
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(
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<collectionCode box="[948,1000,619,645]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="406" type="Museum">ZIN</collectionCode>
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)
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||
</materialsCitation>
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||
,
|
||
<materialsCitation collectionCode="MSPU" country="Vietnam" pageId="5" pageNumber="406" specimenCount="1" specimenCount-female="1" stateProvince="Gia Lai" typeStatus="paratype">
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<typeStatus box="[1023,1145,619,645]" pageId="5" pageNumber="406">PARATYPES</typeStatus>
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:
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<specimenCount box="[1159,1172,619,645]" pageId="5" pageNumber="406" type="female">♀</specimenCount>
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||
—same labeled as the
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<typeStatus box="[151,249,655,681]" pageId="5" pageNumber="406">holotype</typeStatus>
|
||
(
|
||
<collectionCode box="[266,349,655,681]" pageId="5" pageNumber="406">MSPU</collectionCode>
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||
)
|
||
</materialsCitation>
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||
;
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<materialsCitation collectingDate="2016-05-24" collectingDateMax="2016-06-02" collectingDateMin="2016-05-24" collectionCode="V, VI, SIEE" collectorName="D. Fedorenko" country="Vietnam" elevation="1020" location="Kon Chu Rang Nat. Reserve" longitude="108.53333" pageId="5" pageNumber="406" specimenCount="1" specimenCount-female="1" stateProvince="Gia Lai" typeStatus="paratype">
|
||
<specimenCount box="[370,393,655,681]" pageId="5" pageNumber="406" type="female">1♀</specimenCount>
|
||
—
|
||
<collectingCountry box="[424,531,655,681]" name="Vietnam" pageId="5" pageNumber="406">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion box="[543,700,655,681]" country="Vietnam" name="Gia Lai" pageId="5" pageNumber="406">Gia Lai Prov.</collectingRegion>
|
||
, ~
|
||
<locationDeviation box="[727,962,654,681]" location="An Khe" pageId="5" pageNumber="406">
|
||
<quantity box="[727,801,654,681]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="406" unit="km" value="50.0">50 km</quantity>
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||
N of An Khe
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||
</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:050B87D78D452468E1B818ACFBF37514:887D601A8D45246BE2E01BF9FAD974A8" box="[975,1305,655,681]" country="Vietnam" longitude="108.53333" name="Kon Chu Rang Nat. Reserve" pageId="5" pageNumber="406" stateProvince="Gia Lai">Kon Chu Rang Nat. Reserve</location>
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||
, h =
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.02" metricValueMax="1.04" metricValueMin="1.0" pageId="5" pageNumber="406" unit="m" value="1020.0" valueMax="1040.0" valueMin="1000.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.02" metricValueMax="1.04" metricValueMin="1.0" pageId="5" pageNumber="406" unit="m" value="1020.0" valueMax="1040.0" valueMin="1000.0">1000– 1040 m</elevation>
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||
</quantity>
|
||
, 14
|
||
<superScript attach="none" box="[279,287,691,705]" fontSize="6" pageId="5" pageNumber="406">o</superScript>
|
||
30–31’N
|
||
<geoCoordinate box="[394,501,691,717]" degrees="108" direction="east" minutes="32" orientation="longitude" pageId="5" pageNumber="406" precision="925" value="108.53333">
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||
108
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||
<superScript attach="none" box="[436,444,691,705]" fontSize="6" pageId="5" pageNumber="406">o</superScript>
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||
32’E
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||
</geoCoordinate>
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||
,
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||
<date box="[513,698,691,717]" pageId="5" pageNumber="406" value="2016-05-24" valueMax="2016-06-02" valueMin="2016-05-24">
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<collectingDate box="[513,698,691,717]" pageId="5" pageNumber="406" value="2016-05-24" valueMax="2016-06-02" valueMin="2016-05-24">
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24.
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<collectionCode box="[546,568,691,717]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="406" type="Museum">V</collectionCode>
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–2.
|
||
<collectionCode box="[601,633,691,717]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="406">VI</collectionCode>
|
||
.2016
|
||
</collectingDate>
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||
</date>
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||
, leg.
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<collectorName box="[759,913,691,717]" pageId="5" pageNumber="406">D. Fedorenko</collectorName>
|
||
(
|
||
<collectionCode box="[928,993,691,717]" pageId="5" pageNumber="406">SIEE</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectingDate="2015-06-08" collectingDateMax="2015-06-23" collectingDateMin="2015-06-08" collectionCode="VI, SIEE, MSPU" collectorName="Ngok Boc I Mt. & D. Fedorenko" country="Vietnam" elevation="1150" latitude="14.733334" location="Kon Plong Distr." longLatPrecision="1287" longitude="108.3" pageId="5" pageNumber="406" specimenCount="1" stateProvince="Gia Lai" typeStatus="paratype">
|
||
1Ô—
|
||
<collectingCountry box="[1072,1174,692,719]" name="Vietnam" pageId="5" pageNumber="406">Vietnam</collectingCountry>
|
||
,
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||
<collectingRegion box="[1185,1273,691,717]" country="Vietnam" name="Gia Lai" pageId="5" pageNumber="406">Gia Lai</collectingRegion>
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, 4.2022 (AA); 2ÔÔ—
|
||
<collectingCountry box="[234,333,728,755]" name="Vietnam" pageId="5" pageNumber="406">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion box="[346,525,727,753]" country="Vietnam" name="Kon Tum" pageId="5" pageNumber="406">Kon Tum Prov.</collectingRegion>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:050B87D78D452468E1B818ACFBF37514:887D601A8D45246BE3351BA1FD1F74F0" box="[538,735,727,753]" country="Vietnam" latitude="14.733334" longLatPrecision="1287" longitude="108.3" name="Kon Plong Distr." pageId="5" pageNumber="406" stateProvince="Gia Lai">Kon Plong Distr.</location>
|
||
, env.
|
||
<collectorName box="[802,988,727,753]" pageId="5" pageNumber="406">Ngok Boc I Mt.</collectorName>
|
||
,
|
||
<geoCoordinate box="[1002,1095,727,753]" degrees="14" direction="north" minutes="44" orientation="latitude" pageId="5" pageNumber="406" precision="925" value="14.733334">
|
||
14
|
||
<superScript attach="none" box="[1030,1038,727,741]" fontSize="6" pageId="5" pageNumber="406">o</superScript>
|
||
44’N
|
||
</geoCoordinate>
|
||
<geoCoordinate box="[1105,1211,727,753]" degrees="108" direction="east" minutes="18" orientation="longitude" pageId="5" pageNumber="406" precision="925" value="108.3">
|
||
108
|
||
<superScript attach="none" box="[1147,1155,727,741]" fontSize="6" pageId="5" pageNumber="406">o</superScript>
|
||
18’E
|
||
</geoCoordinate>
|
||
, h =
|
||
<quantity box="[1273,1431,727,753]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="5" pageNumber="406" unit="m" value="1150.0" valueMax="1200.0" valueMin="1100.0">
|
||
<elevation box="[1273,1431,727,753]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="5" pageNumber="406" unit="m" value="1150.0" valueMax="1200.0" valueMin="1100.0">1100–1200 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date box="[151,309,762,789]" pageId="5" pageNumber="406" value="2015-06-08" valueMax="2015-06-23" valueMin="2015-06-08">
|
||
<collectingDate box="[151,309,762,789]" pageId="5" pageNumber="406" value="2015-06-08" valueMax="2015-06-23" valueMin="2015-06-08">
|
||
8–23.
|
||
<collectionCode box="[212,244,762,789]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="406">VI</collectionCode>
|
||
.2015
|
||
</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[376,533,763,789]" pageId="5" pageNumber="406">D. Fedorenko</collectorName>
|
||
(
|
||
<collectionCode box="[551,614,763,789]" pageId="5" pageNumber="406">SIEE</collectionCode>
|
||
,
|
||
<collectionCode box="[629,710,763,789]" pageId="5" pageNumber="406">MSPU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectingDate="2016-06-04" collectingDateMax="2016-06-12" collectingDateMin="2016-06-04" collectionCode="VI, ZIN, SIEE, MSPU, JW" collectorName="D. Fedorenko" country="Vietnam" latitude="14.722221" location="Pak Khe River" longLatPrecision="21" longitude="108.31612" municipality="Kon Plong Distr." pageId="5" pageNumber="406" specimenCount="14" specimenCount-female="14" stateProvince="Kon Tum" typeStatus="paratype">
|
||
<specimenCount box="[733,775,763,789]" pageId="5" pageNumber="406" type="female">7♀♀</specimenCount>
|
||
—
|
||
<collectingCountry box="[807,914,763,789]" name="Vietnam" pageId="5" pageNumber="406">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion box="[929,1110,763,789]" country="Vietnam" name="Kon Tum" pageId="5" pageNumber="406">Kon Tum Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality box="[1124,1323,763,789]" pageId="5" pageNumber="406">Kon Plong Distr.</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:050B87D78D452468E1B818ACFBF37514:887D601A8D45246BE4161B8DFF177538" country="Vietnam" latitude="14.722221" longLatPrecision="21" longitude="108.31612" municipality="Kon Plong Distr." name="Pak Khe River" pageId="5" pageNumber="406" stateProvince="Kon Tum">Pak Khe River</location>
|
||
,
|
||
<geoCoordinate box="[229,367,799,825]" degrees="14" direction="north" minutes="43" orientation="latitude" pageId="5" pageNumber="406" precision="15" seconds="20" value="14.722221">
|
||
14
|
||
<superScript attach="none" box="[257,265,799,813]" fontSize="6" pageId="5" pageNumber="406">o</superScript>
|
||
43’20’’N
|
||
</geoCoordinate>
|
||
<geoCoordinate box="[376,535,798,825]" degrees="108" direction="east" minutes="18" orientation="longitude" pageId="5" pageNumber="406" precision="15" seconds="58" value="108.31612">
|
||
108
|
||
<superScript attach="none" box="[418,426,799,813]" fontSize="6" pageId="5" pageNumber="406">o</superScript>
|
||
18’58’’ E
|
||
</geoCoordinate>
|
||
, at light,
|
||
<date box="[647,805,799,825]" pageId="5" pageNumber="406" value="2016-06-04" valueMax="2016-06-12" valueMin="2016-06-04">
|
||
<collectingDate box="[647,805,799,825]" pageId="5" pageNumber="406" value="2016-06-04" valueMax="2016-06-12" valueMin="2016-06-04">
|
||
4–12.
|
||
<collectionCode box="[708,740,799,825]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="406">VI</collectionCode>
|
||
.2016
|
||
</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[870,1026,799,825]" pageId="5" pageNumber="406">D. Fedorenko</collectorName>
|
||
(
|
||
<specimenCount box="[1044,1090,799,825]" pageId="5" pageNumber="406" type="female">2♀♀</specimenCount>
|
||
—
|
||
<collectionCode box="[1123,1178,799,825]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="406" type="Museum">ZIN</collectionCode>
|
||
;
|
||
<specimenCount box="[1193,1238,799,825]" pageId="5" pageNumber="406" type="female">2♀♀</specimenCount>
|
||
—
|
||
<collectionCode box="[1271,1338,799,825]" pageId="5" pageNumber="406">SIEE</collectionCode>
|
||
,
|
||
<specimenCount box="[1352,1401,799,825]" pageId="5" pageNumber="406" type="female">2♀♀</specimenCount>
|
||
—
|
||
<collectionCode box="[151,230,835,861]" pageId="5" pageNumber="406">MSPU</collectionCode>
|
||
,
|
||
<specimenCount box="[242,266,835,861]" pageId="5" pageNumber="406" type="female">1♀</specimenCount>
|
||
—
|
||
<collectionCode box="[300,345,835,861]" pageId="5" pageNumber="406">JW</collectionCode>
|
||
); 8ÔÔ
|
||
</materialsCitation>
|
||
<materialsCitation collectingDate="2019-04-23" collectingDateMax="2019-05-11" collectingDateMin="2019-04-23" collectionCode="V, ZIN, SIEE, MSPU, JW" collectorName="D. Fedorenko" country="Vietnam" elevation="1050" latitude="15.5633335" location="Song Thanh Natn. Park" longLatPrecision="21" longitude="107.38944" municipality="Nam Gian distr." pageId="5" pageNumber="406" specimenCount="4" specimenCount-female="4" stateProvince="Quang Nam" typeStatus="paratype">
|
||
<specimenCount box="[422,465,836,863]" pageId="5" pageNumber="406" type="female">2♀♀</specimenCount>
|
||
—
|
||
<collectingCountry box="[497,604,836,863]" name="Vietnam" pageId="5" pageNumber="406">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion box="[615,820,835,861]" country="Vietnam" name="Quang Nam" pageId="5" pageNumber="406">Quang Nam Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality box="[831,1012,835,861]" pageId="5" pageNumber="406">Nam Gian distr.</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:050B87D78D452468E1B818ACFBF37514:887D601A8D45246BE2D01A35FAC9755C" box="[1023,1289,835,861]" country="Vietnam" latitude="15.5633335" longLatPrecision="21" longitude="107.38944" municipality="Nam Gian distr." name="Song Thanh Natn. Park" pageId="5" pageNumber="406" stateProvince="Quang Nam">Song Thanh Natn. Park</location>
|
||
, h =
|
||
<quantity box="[1344,1431,834,861]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="406" unit="m" value="1050.0">
|
||
<elevation box="[1344,1431,834,861]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="406" unit="m" value="1050.0">1050 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate box="[151,290,870,897]" degrees="15" direction="north" minutes="33" orientation="latitude" pageId="5" pageNumber="406" precision="15" seconds="48" value="15.5633335">
|
||
15
|
||
<superScript attach="none" box="[179,187,871,885]" fontSize="6" pageId="5" pageNumber="406">o</superScript>
|
||
33’48’’N
|
||
</geoCoordinate>
|
||
<geoCoordinate box="[301,452,871,897]" degrees="107" direction="east" minutes="23" orientation="longitude" pageId="5" pageNumber="406" precision="15" seconds="22" value="107.38944">
|
||
107
|
||
<superScript attach="none" box="[343,351,871,885]" fontSize="6" pageId="5" pageNumber="406">o</superScript>
|
||
23’22’’E
|
||
</geoCoordinate>
|
||
,
|
||
<date box="[468,662,871,897]" pageId="5" pageNumber="406" value="2019-04-23" valueMax="2019-05-11" valueMin="2019-04-23">
|
||
<collectingDate box="[468,662,871,897]" pageId="5" pageNumber="406" value="2019-04-23" valueMax="2019-05-11" valueMin="2019-04-23">
|
||
23.IV–11.
|
||
<collectionCode box="[576,598,871,897]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="406" type="Museum">V</collectionCode>
|
||
.2019
|
||
</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[731,889,871,897]" pageId="5" pageNumber="406">D. Fedorenko</collectorName>
|
||
(1Ô
|
||
<specimenCount box="[941,966,872,899]" pageId="5" pageNumber="406" type="female">1♀</specimenCount>
|
||
—
|
||
<collectionCode box="[1000,1057,872,899]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="406" type="Museum">ZIN</collectionCode>
|
||
; 3ÔÔ—
|
||
<collectionCode box="[1157,1219,872,899]" pageId="5" pageNumber="406">SIEE</collectionCode>
|
||
, 3ÔÔ
|
||
<specimenCount box="[1291,1314,872,899]" pageId="5" pageNumber="406" type="female">1♀</specimenCount>
|
||
—
|
||
<collectionCode box="[1346,1432,872,899]" pageId="5" pageNumber="406">MSPU</collectionCode>
|
||
, 1Ô—
|
||
<collectionCode box="[214,256,908,935]" pageId="5" pageNumber="406">JW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation box="[276,1169,907,933]" country="Vietnam" elevation="1300" location="Axan mt." municipality="Tay Giang" pageId="5" pageNumber="406" specimenCount="1" specimenCount-female="1" stateProvince="Quang Nam" typeStatus="paratype">
|
||
<specimenCount box="[276,298,907,933]" pageId="5" pageNumber="406" type="female">1♀</specimenCount>
|
||
—
|
||
<collectingCountry box="[329,436,907,933]" name="Vietnam" pageId="5" pageNumber="406">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion box="[447,652,907,933]" country="Vietnam" name="Quang Nam" pageId="5" pageNumber="406">Quang Nam Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality box="[662,782,907,933]" pageId="5" pageNumber="406">Tay Giang</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:050B87D78D452468E1B818ACFBF37514:887D601A8D45246BE2381AFDFC4275A4" box="[791,898,907,933]" country="Vietnam" municipality="Tay Giang" name="Axan mt." pageId="5" pageNumber="406" stateProvince="Quang Nam">Axan mt.</location>
|
||
,
|
||
<quantity box="[910,997,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="406" unit="m" value="1300.0">
|
||
<elevation box="[910,997,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="406" unit="m" value="1300.0">1300 m</elevation>
|
||
</quantity>
|
||
, 06.2017 (AA)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="406" type="diagnosis">
|
||
<paragraph blockId="5.[151,1437,582,2013]" pageId="5" pageNumber="406">
|
||
<emphasis bold="true" box="[199,314,942,968]" pageId="5" pageNumber="406">Diagnosis</emphasis>
|
||
. A new subspecies is distinguished from the nominotypical one by darker colouration, reduced white elytral pattern and narrower elytral apex (
|
||
<figureCitation box="[624,726,979,1005]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="5" pageNumber="406">Figs 7, 8</figureCitation>
|
||
,
|
||
<figureCitation box="[740,811,979,1005]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="5" pageNumber="406">88, 89</figureCitation>
|
||
<emphasis box="[820,850,979,1004]" italics="true" pageId="5" pageNumber="406">vs.</emphasis>
|
||
<figureCitation box="[858,988,978,1005]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="5" pageNumber="406">Figs 1, 3–5</figureCitation>
|
||
,
|
||
<figureCitation box="[1001,1075,978,1005]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="5" pageNumber="406">81–85</figureCitation>
|
||
), by wider pronotum (
|
||
<figureCitation captionStart="FIGURES 179–183" captionStartId="26.[152,270,1538,1562]" captionTargetBox="[151,1435,181,1513]" captionTargetId="figure-28@26.[151,1435,181,1513]" captionTargetPageId="26" captionText="FIGURES 179–183. Morphometric features of Therates spp: 179—relation of the proportions HTbL/HTaL and PW/PL (red quadrates—T. fruhstorferi fruhstorferi; blue circles—T. fruhstorferi australis ssp. n.; open symbols—males; solid symbols— females); 182—relation of the proportions EL/EW and HTbL/HTaL (green rhombus—T. pseudovitalisi sp. n.; blue circles—T. fruhstorferi australis ssp. n.; open symbols—males; solid symbols—females); 180, 183—relation of the proportions HTbL/ HTaL; 181—relation of the proportions PW/PL (small quadrates—medians; rectangles—25–75 percentiles; whiskers—min– max; significantly different samples are marked with different lower letters, according to Mann–Whitney U-test, p <0.05 (180—U = 429.0, z = 3.74, p = 0.0002; 183—U = 40.0, z = 3.79, p = 0.0002; 181—U = 470.0, z = -3.339, p = 0.0007)." figureDoi="http://doi.org/10.5281/zenodo.7758914" httpUri="https://zenodo.org/record/7758914/files/figure.png" pageId="5" pageNumber="406">Figs 179, 181</figureCitation>
|
||
), by the longer hind tarsi—mean HTbL/HTaL =
|
||
<quantity box="[750,830,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="2.9972" pageId="5" pageNumber="406" unit="in" value="1.18">1.18 in</quantity>
|
||
<emphasis box="[837,987,1015,1041]" italics="true" pageId="5" pageNumber="406">T. f. australis</emphasis>
|
||
<emphasis bold="true" box="[995,1070,1015,1041]" pageId="5" pageNumber="406">ssp. n.</emphasis>
|
||
<emphasis box="[1078,1109,1015,1040]" italics="true" pageId="5" pageNumber="406">vs.</emphasis>
|
||
mean HTbL/HTaL =
|
||
<quantity box="[1357,1436,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="3.0987999999999998" pageId="5" pageNumber="406" unit="in" value="1.22">1.22 in</quantity>
|
||
<emphasis box="[151,334,1051,1077]" italics="true" pageId="5" pageNumber="406">
|
||
T. f.
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1902" box="[205,334,1051,1077]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="fruhstorferi">fruhstorferi</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[352,514,1051,1077]" captionStart="FIGURES 179–183" captionStartId="26.[152,270,1538,1562]" captionTargetBox="[151,1435,181,1513]" captionTargetId="figure-28@26.[151,1435,181,1513]" captionTargetPageId="26" captionText="FIGURES 179–183. Morphometric features of Therates spp: 179—relation of the proportions HTbL/HTaL and PW/PL (red quadrates—T. fruhstorferi fruhstorferi; blue circles—T. fruhstorferi australis ssp. n.; open symbols—males; solid symbols— females); 182—relation of the proportions EL/EW and HTbL/HTaL (green rhombus—T. pseudovitalisi sp. n.; blue circles—T. fruhstorferi australis ssp. n.; open symbols—males; solid symbols—females); 180, 183—relation of the proportions HTbL/ HTaL; 181—relation of the proportions PW/PL (small quadrates—medians; rectangles—25–75 percentiles; whiskers—min– max; significantly different samples are marked with different lower letters, according to Mann–Whitney U-test, p <0.05 (180—U = 429.0, z = 3.74, p = 0.0002; 183—U = 40.0, z = 3.79, p = 0.0002; 181—U = 470.0, z = -3.339, p = 0.0007)." figureDoi="http://doi.org/10.5281/zenodo.7758914" httpUri="https://zenodo.org/record/7758914/files/figure.png" pageId="5" pageNumber="406">Figs 179, 180</figureCitation>
|
||
), and by the shorter aedeagus—mean EL/AL =
|
||
<quantity box="[1069,1149,1050,1077]" metricMagnitude="-2" metricUnit="m" metricValue="6.5786" pageId="5" pageNumber="406" unit="in" value="2.59">2.59 in</quantity>
|
||
<emphasis box="[1158,1311,1051,1077]" italics="true" pageId="5" pageNumber="406">T. f. australis</emphasis>
|
||
<emphasis bold="true" box="[1320,1397,1051,1077]" pageId="5" pageNumber="406">ssp. n.</emphasis>
|
||
<emphasis box="[1406,1436,1051,1076]" italics="true" pageId="5" pageNumber="406">vs.</emphasis>
|
||
mean EL/AL =
|
||
<quantity box="[327,405,1087,1113]" metricMagnitude="-2" metricUnit="m" metricValue="6.2738" pageId="5" pageNumber="406" unit="in" value="2.47">2.47 in</quantity>
|
||
<emphasis box="[412,590,1087,1113]" italics="true" pageId="5" pageNumber="406">
|
||
T. f.
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1902" box="[461,590,1087,1113]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="fruhstorferi">fruhstorferi</taxonomicName>
|
||
</emphasis>
|
||
; from dark-coloured specimens of
|
||
<emphasis box="[984,1162,1087,1113]" italics="true" pageId="5" pageNumber="406">
|
||
T. f.
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1902" box="[1033,1162,1087,1113]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="fruhstorferi">fruhstorferi</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[1177,1250,1087,1113]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="5" pageNumber="406">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation box="[1262,1333,1087,1113]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="5" pageNumber="406">86, 87</figureCitation>
|
||
) the new subspecies is readily distinguished by the shape of aedeagus (
|
||
<figureCitation box="[852,1010,1122,1149]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="5" pageNumber="406">Figs 115–125</figureCitation>
|
||
,
|
||
<figureCitation box="[1022,1120,1122,1149]" captionStart="FIGURES 136–151" captionStartId="23.[152,270,1334,1358]" captionTargetBox="[175,1413,195,1300]" captionTargetId="figure-17@23.[151,1435,181,1309]" captionTargetPageId="23" captionText="FIGURES 136–151. Therates fruhstorferi fruhstorferi, variability of aedeagus, left lateral view: 136–143—Vietnam, Phu Tho Province, ~90 km W of Hanoi, Xuan Son Nat. Park, h = 400–700 m, 21o07’01’’N 104o56’39’’E, 6–15.VI.2014; 144–151— Vietnam, Vinh Phu Province, Tam Dao, different years (144, 145—3–11.VI.1985; 146—27.V–2.VI.1986; 147—2–11.VI.1989; 148—6–10.V.1990; 149—12.VI.1994; 150, 151—9–19.V.1996). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.7758904" httpUri="https://zenodo.org/record/7758904/files/figure.png" pageId="5" pageNumber="406">136–151</figureCitation>
|
||
<emphasis box="[1129,1159,1123,1148]" italics="true" pageId="5" pageNumber="406">vs.</emphasis>
|
||
<figureCitation box="[1167,1326,1123,1149]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="5" pageNumber="406">Figs 126–130</figureCitation>
|
||
). From
|
||
<taxonomicName authority="Matalin & Wiesner, 2023" authorityName="Matalin & Wiesner" authorityYear="2023" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="pseudovitalisi" status="sp. n.">
|
||
<emphasis italics="true" pageId="5" pageNumber="406">T. pseudovitalisi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[314,377,1159,1185]" pageId="5" pageNumber="406">
|
||
<taxonomicNameLabel box="[314,377,1159,1185]" pageId="5" pageNumber="406" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the new subspecies is differentiated both by the shape and the size of aedeagus (
|
||
<figureCitation box="[1281,1437,1159,1185]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="5" pageNumber="406">Figs 126–130</figureCitation>
|
||
<emphasis box="[151,182,1195,1220]" italics="true" pageId="5" pageNumber="406">vs.</emphasis>
|
||
<figureCitation box="[187,343,1194,1221]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="5" pageNumber="406">Figs 132–135</figureCitation>
|
||
), by the narrower elytral apex (
|
||
<figureCitation box="[685,782,1195,1221]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="5" pageNumber="406">Figs 7, 8</figureCitation>
|
||
,
|
||
<figureCitation box="[794,863,1195,1221]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="5" pageNumber="406">88, 89</figureCitation>
|
||
<emphasis box="[869,899,1195,1220]" italics="true" pageId="5" pageNumber="406">vs.</emphasis>
|
||
<figureCitation box="[905,1016,1195,1221]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="5" pageNumber="406">Figs 9–11</figureCitation>
|
||
,
|
||
<figureCitation box="[1026,1101,1194,1221]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="5" pageNumber="406">90–95</figureCitation>
|
||
), and by the longer hind tarsus (
|
||
<figureCitation box="[159,318,1231,1257]" captionStart="FIGURES 179–183" captionStartId="26.[152,270,1538,1562]" captionTargetBox="[151,1435,181,1513]" captionTargetId="figure-28@26.[151,1435,181,1513]" captionTargetPageId="26" captionText="FIGURES 179–183. Morphometric features of Therates spp: 179—relation of the proportions HTbL/HTaL and PW/PL (red quadrates—T. fruhstorferi fruhstorferi; blue circles—T. fruhstorferi australis ssp. n.; open symbols—males; solid symbols— females); 182—relation of the proportions EL/EW and HTbL/HTaL (green rhombus—T. pseudovitalisi sp. n.; blue circles—T. fruhstorferi australis ssp. n.; open symbols—males; solid symbols—females); 180, 183—relation of the proportions HTbL/ HTaL; 181—relation of the proportions PW/PL (small quadrates—medians; rectangles—25–75 percentiles; whiskers—min– max; significantly different samples are marked with different lower letters, according to Mann–Whitney U-test, p <0.05 (180—U = 429.0, z = 3.74, p = 0.0002; 183—U = 40.0, z = 3.79, p = 0.0002; 181—U = 470.0, z = -3.339, p = 0.0007)." figureDoi="http://doi.org/10.5281/zenodo.7758914" httpUri="https://zenodo.org/record/7758914/files/figure.png" pageId="5" pageNumber="406">Figs 182, 183</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="407" pageId="5" pageNumber="406" type="description">
|
||
<paragraph blockId="5.[151,1437,582,2013]" pageId="5" pageNumber="406">
|
||
<emphasis bold="true" box="[199,345,1266,1292]" pageId="5" pageNumber="406">Description.</emphasis>
|
||
TL = 10.0–12.0 mm in males (mean =
|
||
<quantity box="[783,899,1266,1293]" metricMagnitude="-2" metricUnit="m" metricValue="1.095" pageId="5" pageNumber="406" unit="mm" value="10.95">10.95 mm</quantity>
|
||
, n = 12), 10.0–
|
||
<quantity box="[1068,1168,1267,1293]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="5" pageNumber="406" unit="mm" value="12.4">12.4 mm</quantity>
|
||
(mean =
|
||
<quantity box="[1273,1388,1267,1293]" metricMagnitude="-2" metricUnit="m" metricValue="1.142" pageId="5" pageNumber="406" unit="mm" value="11.42">11.42 mm</quantity>
|
||
, n = 12) in females.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,582,2013]" pageId="5" pageNumber="406">
|
||
Head shining black-blue with purple-violet reflection, especially bright on vertex, occiput and on smooth, finely or indistinctly striated in posterior third orbital plates; frons convex, semi-circular (see above), slightly downward anteriorly (see at the side), smooth or slightly rugose, frontal sulci deep and slightly convergent in anterior half or 2/3, shallow and distinctly divergent in posterior half or third. Mandibles in males brown underside, yellow or yellow-brown topside with light-brown teeth and apical molar, in females dark-brown except pale base topside. Labial palpomeres 2 and 3 yellow, palpomere 4 dark brown; maxillary palpomeres 1 and 2 yellow, yellow-brown or brown, palpomeres 3 and 4 dark brown or brown-black. Antennae do not extend posteriorly to the shoulders, scape in males yellow or yellow-brown on anterior side and dark-brown on posterior side, in females entirely black or brown-black rarely dark brown on anterior side; antennomeres 3–6 dark brown with narrow light brown apices, antennomeres 9–11 flattened; in males antennomeres 9 and 10 with slightly protruding downward anterior lower margins, antennomere 11 slightly dilated not wider than 10 one, in females antennomeres 9 and 10 slightly dilated on anterior side (
|
||
<figureCitation box="[343,476,1735,1761]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="5" pageNumber="406">Figs 33, 34</figureCitation>
|
||
). Labrum indistinctly transverse, LW/LL = 1.0–1.15 (mean = 1.07, n = 24), in males black-brown with medium-sized or small yellow-brown central spot bordered from central apical teeth (
|
||
<figureCitation box="[1311,1394,1770,1797]" captionStart="FIGURES 50–57" captionStartId="15.[152,270,1909,1933]" captionTargetBox="[320,1268,195,1860]" captionTargetId="figure-17@15.[303,1285,181,1885]" captionTargetPageId="15" captionText="FIGURES 50–57. Therates fruhstorferi ssp., labrum: 50–55—T. fruhstorferi fruhstorferi (50, 51—Vietnam, Montes Mauson; 52—Vietnam, Na Hang; 53—Vietnam, Tam Dao; 54, 55—Laos, Ban Nape); 56, 57—T. fruhstorferi australis ssp. n. (56—Vietnam, Kon Ka Kinh N.P.; 57—Vietnam, Pak Khe River); 50, 52, 54, 56—males; 51, 53, 55, 57—females; 50, 51—paralectotypes; 56—holotype; 57—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758882" httpUri="https://zenodo.org/record/7758882/files/figure.png" pageId="5" pageNumber="406">Fig. 56</figureCitation>
|
||
), in females entirely black or brown-black (
|
||
<figureCitation box="[593,676,1806,1833]" captionStart="FIGURES 50–57" captionStartId="15.[152,270,1909,1933]" captionTargetBox="[320,1268,195,1860]" captionTargetId="figure-17@15.[303,1285,181,1885]" captionTargetPageId="15" captionText="FIGURES 50–57. Therates fruhstorferi ssp., labrum: 50–55—T. fruhstorferi fruhstorferi (50, 51—Vietnam, Montes Mauson; 52—Vietnam, Na Hang; 53—Vietnam, Tam Dao; 54, 55—Laos, Ban Nape); 56, 57—T. fruhstorferi australis ssp. n. (56—Vietnam, Kon Ka Kinh N.P.; 57—Vietnam, Pak Khe River); 50, 52, 54, 56—males; 51, 53, 55, 57—females; 50, 51—paralectotypes; 56—holotype; 57—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758882" httpUri="https://zenodo.org/record/7758882/files/figure.png" pageId="5" pageNumber="406">Fig. 57</figureCitation>
|
||
), rarely with small brown central spot.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,582,2013]" pageId="5" pageNumber="406">
|
||
Pronotum wider than long, PW/PL = 1.0–1.13 (mean = 1.06, n = 24) (
|
||
<figureCitation box="[989,1148,1843,1869]" captionStart="FIGURES 179–183" captionStartId="26.[152,270,1538,1562]" captionTargetBox="[151,1435,181,1513]" captionTargetId="figure-28@26.[151,1435,181,1513]" captionTargetPageId="26" captionText="FIGURES 179–183. Morphometric features of Therates spp: 179—relation of the proportions HTbL/HTaL and PW/PL (red quadrates—T. fruhstorferi fruhstorferi; blue circles—T. fruhstorferi australis ssp. n.; open symbols—males; solid symbols— females); 182—relation of the proportions EL/EW and HTbL/HTaL (green rhombus—T. pseudovitalisi sp. n.; blue circles—T. fruhstorferi australis ssp. n.; open symbols—males; solid symbols—females); 180, 183—relation of the proportions HTbL/ HTaL; 181—relation of the proportions PW/PL (small quadrates—medians; rectangles—25–75 percentiles; whiskers—min– max; significantly different samples are marked with different lower letters, according to Mann–Whitney U-test, p <0.05 (180—U = 429.0, z = 3.74, p = 0.0002; 183—U = 40.0, z = 3.79, p = 0.0002; 181—U = 470.0, z = -3.339, p = 0.0007)." figureDoi="http://doi.org/10.5281/zenodo.7758914" httpUri="https://zenodo.org/record/7758914/files/figure.png" pageId="5" pageNumber="406">Figs 179, 181</figureCitation>
|
||
), shining blue with violet reflection, apical lobe with slightly narrower sides or virtually equal wide throughout; thorax shining blue with light violet reflection.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,582,2013]" lastBlockId="6.[151,1436,151,789]" lastPageId="6" lastPageNumber="407" pageId="5" pageNumber="406">
|
||
Fore and middle femora in males pale-yellow with brown violet-tinged posterior side, in females middle femora with more expanded black colour; hind femora bicolored, pale-yellow in basal half or third in males or in basal quarter in females, and black with violet tinge in other portion; fore tibiae light brown on outer sides and brown on inner sides with dark apices, middle tibiae light brown or yellow-brown with brown apices, in some specimens fore- and middle tibiae dark-brown with slightly lighter basal third, hind tibiae dark brown with black blue tinged apices, rarely all tibiae dark brown or brown-black; all tarsi black with light blue tinge; HTbL/HTaL = 1.09–1.29 (mean = 1.18, n = 23) (
|
||
<figureCitation box="[310,581,295,321]" captionStart="FIGURES 179–183" captionStartId="26.[152,270,1538,1562]" captionTargetBox="[151,1435,181,1513]" captionTargetId="figure-28@26.[151,1435,181,1513]" captionTargetPageId="26" captionText="FIGURES 179–183. Morphometric features of Therates spp: 179—relation of the proportions HTbL/HTaL and PW/PL (red quadrates—T. fruhstorferi fruhstorferi; blue circles—T. fruhstorferi australis ssp. n.; open symbols—males; solid symbols— females); 182—relation of the proportions EL/EW and HTbL/HTaL (green rhombus—T. pseudovitalisi sp. n.; blue circles—T. fruhstorferi australis ssp. n.; open symbols—males; solid symbols—females); 180, 183—relation of the proportions HTbL/ HTaL; 181—relation of the proportions PW/PL (small quadrates—medians; rectangles—25–75 percentiles; whiskers—min– max; significantly different samples are marked with different lower letters, according to Mann–Whitney U-test, p <0.05 (180—U = 429.0, z = 3.74, p = 0.0002; 183—U = 40.0, z = 3.79, p = 0.0002; 181—U = 470.0, z = -3.339, p = 0.0007)." figureDoi="http://doi.org/10.5281/zenodo.7758914" httpUri="https://zenodo.org/record/7758914/files/figure.png" pageId="6" pageNumber="407">Figs 179, 180, 182, 183</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1436,151,789]" pageId="6" pageNumber="407">
|
||
Elytra subparallel, indistinctly divergent toward apex, EL/EW = 1.85–2.06 (mean = 1.97, n = 24); longer in females—EL/PL = 3.41–3.83 (mean = 3.61, n = 12)
|
||
<emphasis box="[740,770,367,392]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
|
||
3.3–3.72 (mean = 3.45, n = 12) in males; shining blue with purple-violet reflection and narrow bright blue or golden-blue reflection along suture; punctuation deep and regular between basal humps and central dot, shallower behind it, entirely absent in apical quarter; scutellum black with greenish-blue tinge; apical margin gradually rounded. Elytral pattern presented by pale-yellow basal portion of humeral lunule and by small basal dot, in some specimens both of them very small or basal dot barely visible or absent, by elongate, slightly oblique, wide-uniformed central dot, semilunar and slightly curved towards to suture in some females as well as by very narrow dark brown apical edge (
|
||
<figureCitation box="[882,982,583,609]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 7, 8</figureCitation>
|
||
,
|
||
<figureCitation box="[995,1069,583,609]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">88, 89</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1436,151,789]" pageId="6" pageNumber="407">
|
||
Aedeagus with moderately short, straight or indistinctly curved, blunt apex without lateral carinae (
|
||
<figureCitation captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="6" pageNumber="407">Figs 126– 130</figureCitation>
|
||
); AL =
|
||
<quantity box="[282,417,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="6" pageNumber="407" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
|
||
(mean = 2.64, n = 12), EL/AL =
|
||
<quantity box="[789,952,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="2.5549999999999997" metricValueMax="2.73" metricValueMin="2.38" pageId="6" pageNumber="407" unit="mm" value="2.5549999999999997" valueMax="2.73" valueMin="2.38">2.38–2.73 mm</quantity>
|
||
(mean = 2.59, n = 12).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="407" type="etymology">
|
||
<paragraph blockId="6.[151,1436,151,789]" pageId="6" pageNumber="407">
|
||
<emphasis bold="true" box="[199,334,691,717]" pageId="6" pageNumber="407">Etymology.</emphasis>
|
||
The name of this new subspecies is derived from the Latin, “
|
||
<emphasis box="[1013,1111,691,717]" italics="true" pageId="6" pageNumber="407">australis</emphasis>
|
||
”, meaning ‘southern’, which indicate the southern border of distribution of
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1902" box="[668,825,727,753]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="fruhstorferi">
|
||
<emphasis box="[668,825,727,753]" italics="true" pageId="6" pageNumber="407">T. fruhstorferi</emphasis>
|
||
</taxonomicName>
|
||
at present time.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1075,762,789]" pageId="6" pageNumber="407" type="distribution">
|
||
<paragraph blockId="6.[151,1436,151,789]" box="[199,1075,762,789]" pageId="6" pageNumber="407">
|
||
<emphasis bold="true" box="[199,353,762,788]" pageId="6" pageNumber="407">Distribution.</emphasis>
|
||
<collectingCountry box="[359,466,763,789]" name="Vietnam" pageId="6" pageNumber="407">VIETNAM</collectingCountry>
|
||
:
|
||
<collectingRegion box="[476,614,763,789]" country="Vietnam" name="Quang Nam" pageId="6" pageNumber="407">Quang Nam</collectingRegion>
|
||
,
|
||
<collectingRegion box="[626,736,763,789]" country="Vietnam" name="Kon Tum" pageId="6" pageNumber="407">Kon Tum</collectingRegion>
|
||
,
|
||
<collectingRegion box="[746,831,763,789]" country="Vietnam" name="Gia Lai" pageId="6" pageNumber="407">Gia Lai</collectingRegion>
|
||
Provinces (
|
||
<figureCitation box="[964,1062,763,789]" captionStart="FIGURES 184" captionStartId="28.[152,270,1579,1603]" captionTargetBox="[151,1435,597,1556]" captionTargetId="figure-226@28.[151,1435,597,1556]" captionTargetPageId="28" captionText="FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png." figureDoi="http://doi.org/10.5281/zenodo.7758916" httpUri="https://zenodo.org/record/7758916/files/figure.png" pageId="6" pageNumber="407">Fig. 184</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |