treatments-xml/data/A5/C4/4E/A5C44E67EC755DD8A53C6E2F4D41083B.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1026.61554" ID-GBIF-Dataset="0a2d723c-d292-4f0c-9d00-6cdd176c49a6" ID-PMC="PMC8016817" ID-Pensoft-Pub="1313-2970-1026-45" ID-Pensoft-UUID="2EFE20EFF6705E34BFA20CF497608C4D" ID-PubMed="33850418" ID-ZooBank="A3212DAACEF24A1F93CC1823940C0108" ModsDocID="1313-2970-1026-45" checkinTime="1616697129133" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Sumoked, Bob &amp; Balke, Michael" docDate="2021" docId="A5C44E67EC755DD8A53C6E2F4D41083B" docLanguage="en" docName="ZooKeys 1026: 45-67" docOrigin="ZooKeys 1026" docPubDate="2021-03-25" docSource="http://dx.doi.org/10.3897/zookeys.1026.61554" docTitle="Exocelina ekpliktiki Shaverdo, Surbakti &amp; Balke 2021, sp. nov." docType="treatment" docUuid="5869A521-DFF6-45B6-9481-96A9C0D9F3D2" docUuidSource="ZooBank" docVersion="5" id="2EFE20EFF6705E34BFA20CF497608C4D" lastPageNumber="45" masterDocId="2EFE20EFF6705E34BFA20CF497608C4D" masterDocTitle="Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="67" masterPageNumber="45" pageNumber="45" updateTime="1668149974149" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaverdo, Helena</mods:namePart>
<mods:affiliation>Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">shaverdo@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Surbakti, Suriani</mods:namePart>
<mods:affiliation>Department of Biology, Universitas Cendrawasih, Waena, Papua, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sumoked, Bob</mods:namePart>
<mods:affiliation>Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1026</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>45</mods:start>
<mods:end>67</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1026.61554</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1026.61554</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1026-45</mods:identifier>
<mods:identifier type="ZooBank">A3212DAACEF24A1F93CC1823940C0108</mods:identifier>
<mods:identifier type="Pensoft-UUID">2EFE20EFF6705E34BFA20CF497608C4D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="180680994" LSID="urn:lsid:zoobank.org:act:5869A521-DFF6-45B6-9481-96A9C0D9F3D2" httpUri="http://treatment.plazi.org/id/A5C44E67EC755DD8A53C6E2F4D41083B" lastPageNumber="45" pageId="0" pageNumber="45">
<subSubSection pageId="0" pageNumber="45" type="nomenclature">
<paragraph pageId="0" pageNumber="45">
<taxonomicName LSID="http://zoobank.org/5869A521-DFF6-45B6-9481-96A9C0D9F3D2" authority="Shaverdo, Surbakti &amp; Balke" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ekpliktiki" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekpliktiki" status="sp. nov.">Exocelina ekpliktiki Shaverdo, Surbakti &amp; Balke</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 11, 12" captionStartId="F5" captionText="Figures 11, 12. Exocelina ekpliktiki sp. nov., holotype 11 habitus and colouration 12 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures11-12" httpUri="https://binary.pensoft.net/fig/524449" pageId="0" pageNumber="45">Figures 11</figureCitation>
<figureCitation captionStart="Figures 11, 12" captionStartId="F5" captionText="Figures 11, 12. Exocelina ekpliktiki sp. nov., holotype 11 habitus and colouration 12 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures11-12" httpUri="https://binary.pensoft.net/fig/524449" pageId="0" pageNumber="45">, 12</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="reference_group">
<paragraph pageId="0" pageNumber="45">
<taxonomicName authority="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke, 2021" authorityName="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina oceai" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="oceai" status="sp. nov.">Exocelina nr. oceai</taxonomicName>
#6504: 
<bibRefCitation DOI="https://doi.org/10.1186/s12862-021-01764-2" author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" refId="B10" refString="Toussaint, EFA, White, LT, Shaverdo, H, Lam, A, Surbakti, S, Panjaitan, R, Sumoked, B, von Rintelen, T, Sagata, K, Balke, M, 2021. [in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology. https://doi.org/10.1186/s12862-021-01764-2" title="[in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology." url="https://doi.org/10.1186/s12862-021-01764-2" year="2021">Toussaint et al. 2021</bibRefCitation>
: figs 3-6.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type locality">
<paragraph pageId="0" pageNumber="45">
<typeStatus>Type</typeStatus>
locality.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138329" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" elevation="378" latitude="-3.9056668" location="Iratoi" longLatPrecision="1" longitude="137.20088" municipality="Puncak Regency" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
, 
<collectingMunicipality>Puncak Regency</collectingMunicipality>
, south from 
<location LSID="urn:lsid:plazi:treatment:A5C44E67EC755DD8A53C6E2F4D41083B:E53973782D9801802AA26FD6E7533822" country="Indonesia" latitude="-3.9056668" longLatPrecision="1" longitude="137.20088" municipality="Puncak Regency" name="Iratoi" stateProvince="Papua Barat">Iratoi</location>
, 
<geoCoordinate degrees="03" direction="south" minutes="54" orientation="latitude" precision="1" seconds="20.4" value="-3.9056668">03°54'20.4&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="137" direction="east" minutes="12" orientation="longitude" precision="1" seconds="03.2" value="137.20088">137°12'03.2&quot;E</geoCoordinate>
, 
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.78" unit="m" value="378.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.78" unit="m" value="378.0">378 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type material">
<paragraph pageId="0" pageNumber="45">
<typeStatus>Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3062138302" collectingDate="2015-05-22" collectionCode="MZB" country="Indonesia" elevation="378" latitude="-3.3904028" location="Papua" longLatPrecision="1" longitude="137.3201" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
male &quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, S Iratoi, forest, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.78" unit="m" value="378.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.78" unit="m" value="378.0">378m</elevation>
</quantity>
, 
<date value="2015-05-22">
<collectingDate value="2015-05-22">22.v.2015</collectingDate>
</date>
, 
<geoCoordinate degrees="3,3904028" direction="south" orientation="latitude" precision="1" value="-3.3904028">-3,3904028</geoCoordinate>
<geoCoordinate degrees="137,32009999" direction="east" orientation="longitude" precision="1" value="137.3201">137,32009999</geoCoordinate>
, Pele &amp; Sumoked (Pap037)&quot; (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">
<typeStatus>Paratypes</typeStatus>
</emphasis>
: Puncak Regency
</emphasis>
: 
<materialsCitation ID-GBIF-Occurrence="3062138328" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="MZB, KSP" specimenCount="24" specimenCount-female="8" specimenCount-male="16" typeStatus="paratype">
<specimenCount type="male">16 males</specimenCount>
, 
<specimenCount type="female">8 females</specimenCount>
with the same label as the 
<typeStatus>holotype</typeStatus>
(
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
, 
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3062138304" collectingDate="2015-05-22" collectionCode="KSP" country="Indonesia" elevation="378" location="Papua" specimenCount="3" specimenCount-male="3" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">3 males</specimenCount>
&quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, S Iratoi, forest, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.78" unit="m" value="378.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.78" unit="m" value="378.0">378m</elevation>
</quantity>
, 
<date value="2015-05-22">
<collectingDate value="2015-05-22">22.v.2015</collectingDate>
</date>
, -3,3904028031975 137,320099985226, Pele &amp; Sumoked (Pap037)&quot;, one male an additional label 
<normalizedToken originalValue="“KSP6983”">&quot;KSP6983&quot;</normalizedToken>
[green text] (
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3062138315" collectingDate="2015-05-22" collectionCode="KSP" country="Indonesia" elevation="553" latitude="-3.3919227" location="Papua" longLatPrecision="1" longitude="137.32353" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">5 males</specimenCount>
, 
<specimenCount type="female">2 females</specimenCount>
&quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, S Iratoi, forest, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" unit="m" value="553.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" unit="m" value="553.0">553m</elevation>
</quantity>
, 
<date value="2015-05-22">
<collectingDate value="2015-05-22">22.v.2015</collectingDate>
</date>
, 
<geoCoordinate degrees="3,3919226937" direction="south" orientation="latitude" precision="1" value="-3.3919227">-3,3919226937</geoCoordinate>
<geoCoordinate degrees="137,3235277" direction="east" orientation="longitude" precision="1" value="137.32353">137,3235277</geoCoordinate>
, Pele &amp; Sumoked (Pap038)&quot; (
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3062138335" collectingDate="2015-05-22" collectionCode="KSP" country="Indonesia" elevation="553" location="Papua" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">1 male</specimenCount>
, 
<specimenCount type="female">1 female</specimenCount>
&quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, S Iratoi, forest, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" unit="m" value="553.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" unit="m" value="553.0">553m</elevation>
</quantity>
, 
<date value="2015-05-22">
<collectingDate value="2015-05-22">22.v.2015</collectingDate>
</date>
, -3,39192269369959 137,323527764528, Pele &amp; Sumoked (Pap038)&quot;, with additional green text labels 
<normalizedToken originalValue="“6989”">&quot;6989&quot;</normalizedToken>
and 
<normalizedToken originalValue="“6988”">&quot;6988&quot;</normalizedToken>
, respectively (
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3062138319" collectingDate="2015-05-23" collectionCode="MZB, KSP" country="Indonesia" elevation="450" location="Papua" specimenCount="28" specimenCount-female="11" specimenCount-male="17" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">17 males</specimenCount>
, 
<specimenCount type="female">11 females</specimenCount>
&quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, S Iratoi, forest, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">450m</elevation>
</quantity>
, 
<date value="2015-05-23">
<collectingDate value="2015-05-23">23.v.2015</collectingDate>
</date>
, near -3,39192 137,323527764528, Pele &amp; Sumoked (Pap039)&quot; (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
, 
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3062138331" collectingDate="2015-05-23" collectionCode="KSP" country="Indonesia" elevation="450" location="Papua" specimenCount="2" specimenCount-male="2" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">2 males</specimenCount>
&quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, S Iratoi, forest, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">450m</elevation>
</quantity>
, 
<date value="2015-05-23">
<collectingDate value="2015-05-23">23.v.2015</collectingDate>
</date>
, near -3,391922694 137,323527764528, Pele &amp; Sumoked (Pap039)&quot;, with additional green text labels 
<normalizedToken originalValue="“6986”">&quot;6986&quot;</normalizedToken>
and 
<normalizedToken originalValue="“6987”">&quot;6987&quot;</normalizedToken>
(
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3062138313" collectingDate="2014-09-03" collectionCode="MZB, KSP" country="Indonesia" elevation="1100" latitude="-3.587955" location="Wano Land" longLatPrecision="1" longitude="137.51149" specimenCount="18" specimenCount-female="5" specimenCount-male="13" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">13 males</specimenCount>
, 
<specimenCount type="female">5 females</specimenCount>
&quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:A5C44E67EC755DD8A53C6E2F4D41083B:2D349BD9CBA792B33C422E3B9B247081" country="Indonesia" latitude="-3.587955" longLatPrecision="1" longitude="137.51149" name="Wano Land" stateProvince="Papua Barat">Wano Land</location>
, red clay creek nr cave, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100m</elevation>
</quantity>
, 
<date value="2014-09-03">
<collectingDate value="2014-09-03">3.ix.2014</collectingDate>
</date>
, nr 
<geoCoordinate degrees="3.587955" direction="south" orientation="latitude" precision="1" value="-3.587955">-3.587955</geoCoordinate>
<geoCoordinate degrees="137.5114945" direction="east" orientation="longitude" precision="1" value="137.51149">137.5114945</geoCoordinate>
, Bennji (Pap024)&quot;, one male an additional label 
<normalizedToken originalValue="“6517”">&quot;6517&quot;</normalizedToken>
[green text] (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
, 
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Puncak Jaya Regency.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3062138303" collectingDate="2015-05-21" collectionCode="MZB, KSP" country="Indonesia" elevation="220" location="Papua" specimenCount="3" specimenCount-male="3" stateProvince="Papua Barat">
<specimenCount type="male">3 males</specimenCount>
&quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, S Iratoi, forest, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">220m</elevation>
</quantity>
, 
<date value="2015-05-21">
<collectingDate value="2015-05-21">21.v.2015</collectingDate>
</date>
, -3,38095162063837 137,311441982164, Pele &amp; Sumoked (Pap036)&quot;, one male an additional label 
<normalizedToken originalValue="“6982”">&quot;6982&quot;</normalizedToken>
[green text] (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
, 
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3062138305" collectingDate="2014-09-06" collectionCode="KSP" country="Indonesia" elevation="164" latitude="-3.2403085" location="Iratoi" longLatPrecision="1" longitude="137.33298" municipality="Rouaffer" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Papua Barat">
<specimenCount type="male">6 males</specimenCount>
, 
<specimenCount type="female">6 females</specimenCount>
&quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, 
<collectingMunicipality>Rouaffer</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:A5C44E67EC755DD8A53C6E2F4D41083B:1619BEFB9F5DD93ADFACCA9FE14BC78A" country="Indonesia" latitude="-3.2403085" longLatPrecision="1" longitude="137.33298" municipality="Rouaffer" name="Iratoi" stateProvince="Papua Barat">Iratoi</location>
, hill in forest, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.64" unit="m" value="164.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.64" unit="m" value="164.0">164m</elevation>
</quantity>
, 
<date value="2014-09-06">
<collectingDate value="2014-09-06">6.ix.2014</collectingDate>
</date>
, 
<geoCoordinate degrees="3,2403086" direction="south" orientation="latitude" precision="1" value="-3.2403085">-3,2403086</geoCoordinate>
<geoCoordinate degrees="137,3329744" direction="east" orientation="longitude" precision="1" value="137.33298">137,3329744</geoCoordinate>
, Pele &amp; Sumoked (Pap028)&quot;, two males with additional green text labels 
<normalizedToken originalValue="“6504”">&quot;6504&quot;</normalizedToken>
and 
<normalizedToken originalValue="“6505”">&quot;6505&quot;</normalizedToken>
(
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Body size and form</emphasis>
:
</emphasis>
Beetle small: TL-H 2.95-3.35 mm, TL 3.35-3.65 mm, MW 1.6-1.8 mm, excluding the locality Pap024, (
<typeStatus>holotype</typeStatus>
: TL-H 3.15 mm, TL 3.55 mm, 
<collectionCode country="Netherlands" name="Museum Wasmann">MW</collectionCode>
1.7 mm), with oblong-oval habitus (Fig. 
<figureCitation captionStart="Figures 11, 12" captionStartId="F5" captionText="Figures 11, 12. Exocelina ekpliktiki sp. nov., holotype 11 habitus and colouration 12 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures11-12" httpUri="https://binary.pensoft.net/fig/524449" pageId="0" pageNumber="45">11</figureCitation>
), some specimens, especially teneral slightly egg-shaped.
</paragraph>
<caption doi="10.3897/zookeys.1026.61554.figures11-12" httpUri="https://binary.pensoft.net/fig/524449" pageId="0" pageNumber="45" start="Figures 11, 12" startId="F5">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figures 11, 12.</emphasis>
<taxonomicName authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ekpliktiki" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekpliktiki">
<emphasis italics="true" pageId="0" pageNumber="45">Exocelina ekpliktiki</emphasis>
</taxonomicName>
sp. nov., 
<typeStatus>holotype</typeStatus>
<emphasis bold="true" pageId="0" pageNumber="45">11</emphasis>
habitus and colouration 
<specimenCount type="male">
<emphasis bold="true" pageId="0" pageNumber="45">12</emphasis>
male
</specimenCount>
structures 
<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
right protarsomeres 4 and 
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
ventral view 
<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
median lobe in ventral view 
<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
median lobe in lateral view 
<emphasis bold="true" pageId="0" pageNumber="45">D</emphasis>
right paramere in external view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Colouration</emphasis>
:
</emphasis>
Dorsally dark brown to piceous, with paler, reddish brown anterior half of head and lateral sides of pronotum (Fig. 
<figureCitation captionStart="Figures 11, 12" captionStartId="F5" captionText="Figures 11, 12. Exocelina ekpliktiki sp. nov., holotype 11 habitus and colouration 12 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures11-12" httpUri="https://binary.pensoft.net/fig/524449" pageId="0" pageNumber="45">11</figureCitation>
). Head dark brown posteriorly and largely (to half) reddish brown anteriorly; pronotum dark brown to piceous, with reddish to reddish brown lateral sides, sometimes also narrowly anteriorly and posteriorly leaving dark brown disc; elytra dark brown to piceous, with reddish brown sutural lines; head appendages yellow, legs yellowish red to reddish brown. Teneral specimens paler, to pale brown, with yellow anterior half of head and pronotal sides.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Surface sculpture</emphasis>
:
</emphasis>
Shiny dorsally, with inconspicuous, almost invisible elytral punctation and weakly impressed microreticulation. Head with relatively sparse punctation (spaces between punctures 1-3 times size of punctures), evidently finer and sparser anteriorly and posteriorly; diameter of larger punctures almost equal to diameter of cells of microreticulation. Pronotum with finer, sparser, and more evenly distributed punctation than on head, often inconspicuous. Elytra with very sparse and fine punctation, almost invisible. Pronotum and elytra with weakly impressed microreticulation, sometimes stronger on pronotal sides; head with microreticulation much stronger. Metaventrite and metacoxa distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine and sparse punctation.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Structures</emphasis>
:
</emphasis>
Pronotum with distinct but narrow lateral bead, in some specimens reduced at posterior angles. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 broadly rounded.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Male</emphasis>
:
</emphasis>
Antenna simple. Pro- and mesotarsomeres 1-3 narrow. Protarsomere 4 narrow, with medium-sized, slightly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 13 and posterior row of four short, pointed setae (Fig. 
<figureCitation captionStart="Figures 11, 12" captionStartId="F5" captionText="Figures 11, 12. Exocelina ekpliktiki sp. nov., holotype 11 habitus and colouration 12 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures11-12" httpUri="https://binary.pensoft.net/fig/524449" pageId="0" pageNumber="45">12A</figureCitation>
). Median lobe with distinctly discontinuous outline; in lateral view, almost straight, with curved downwards, broadly pointed apex; in ventral view, with weak submedian constriction, distal part only slightly narrower than proximal one, and evenly, shallowly concave apex (Fig. 
<figureCitation captionStart="Figures 11, 12" captionStartId="F5" captionText="Figures 11, 12. Exocelina ekpliktiki sp. nov., holotype 11 habitus and colouration 12 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures11-12" httpUri="https://binary.pensoft.net/fig/524449" pageId="0" pageNumber="45">12B, C</figureCitation>
). Paramere with strong notch on dorsal side and subdistal part short and small; subdistal setae long, dense, curved at apex, few lower ones slightly flattened; proximal setae numerous, dense, but weaker than subdistal ones (Fig. 
<figureCitation captionStart="Figures 11, 12" captionStartId="F5" captionText="Figures 11, 12. Exocelina ekpliktiki sp. nov., holotype 11 habitus and colouration 12 male structures A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures11-12" httpUri="https://binary.pensoft.net/fig/524449" pageId="0" pageNumber="45">12D</figureCitation>
). Abdominal ventrite 6 with 5-10 lateral striae on each side.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Female</emphasis>
:
</emphasis>
Pro- and mesotarsi not modified. Abdominal ventrite 6 without lateral striae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="variability">
<paragraph pageId="0" pageNumber="45">Variability.</paragraph>
<paragraph pageId="0" pageNumber="45">
Beetles from the locality Pap024 are larger (TL-H 3.25-3.65 mm, TL 3.55-4.0 mm, 
<collectionCode country="Netherlands" name="Museum Wasmann">MW</collectionCode>
1.75-2.0 mm), with distinctly larger and more robust median lobe and paramere, though of the shape and setation of the median lobe and paramere are the same.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="affinities">
<paragraph pageId="0" pageNumber="45">Affinities.</paragraph>
<paragraph pageId="0" pageNumber="45">
The species evidently belongs to the 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. ekari" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekari">
<emphasis italics="true" pageId="0" pageNumber="45">E. ekari</emphasis>
</taxonomicName>
group due to the discontinuous outline of its median lobe. Based on body size and form, colouration, dorsal surface sculpture, shape of anterolateral hook-like seta of the protarsomere 4, and shape and setation of genitalia, the new species is very similar to 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. soppi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="soppi">
<emphasis italics="true" pageId="0" pageNumber="45">E. soppi</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" pagination="1 - 76" refId="B3" refString="Shaverdo, HV, Surbakti, S, Hendrich, LM, Balke, M, 2012. Introduction of the Exocelina ekari -group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76, DOI: https://doi.org/10.3897/zookeys.250.3715" title="Introduction of the Exocelina ekari - group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al., 2012</bibRefCitation>
and, especially, to 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. weylandensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="weylandensis">
<emphasis italics="true" pageId="0" pageNumber="45">E. weylandensis</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" pagination="1 - 76" refId="B3" refString="Shaverdo, HV, Surbakti, S, Hendrich, LM, Balke, M, 2012. Introduction of the Exocelina ekari -group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76, DOI: https://doi.org/10.3897/zookeys.250.3715" title="Introduction of the Exocelina ekari - group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al., 2012</bibRefCitation>
. However, it differs distinctly from them in presence of the pronotal bead and in that, it is similar to 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. oceai" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="oceai">
<emphasis italics="true" pageId="0" pageNumber="45">E. oceai</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" pagination="1 - 76" refId="B3" refString="Shaverdo, HV, Surbakti, S, Hendrich, LM, Balke, M, 2012. Introduction of the Exocelina ekari -group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76, DOI: https://doi.org/10.3897/zookeys.250.3715" title="Introduction of the Exocelina ekari - group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al., 2012</bibRefCitation>
and can be as well as distinguished from the co-occurring species, 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. bewaniensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="bewaniensis">
<emphasis italics="true" pageId="0" pageNumber="45">E. bewaniensis</emphasis>
</taxonomicName>
and 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. oraia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="oraia">
<emphasis italics="true" pageId="0" pageNumber="45">E. oraia</emphasis>
</taxonomicName>
sp. nov.; from 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. oraia" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="oraia">
<emphasis italics="true" pageId="0" pageNumber="45">E. oraia</emphasis>
</taxonomicName>
sp. nov. also by not having modified male antennae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
. This species is known only from the Wano Land (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="habitat">
<paragraph pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="45">The specimens were collected from small forest creeks.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">
The species name 
<taxonomicName authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ekpliktiki" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekpliktiki">
<emphasis italics="true" pageId="0" pageNumber="45">Exocelina ekpliktiki</emphasis>
</taxonomicName>
derives from Greek 
<normalizedToken originalValue="εκπληκτικός">εκπληκτικός</normalizedToken>
(feminine 
<normalizedToken originalValue="εκπληκτική">εκπληκτική</normalizedToken>
) meaning fantastic. The name is an adjective in the nominative singular.
</paragraph>
</subSubSection>
</treatment>
</document>