155 lines
18 KiB
XML
155 lines
18 KiB
XML
<document id="82CF5043F90D3F418EAE533B60788433" ID-CLB-Dataset="31530" ID-DOI="10.5281/zenodo.1050060" ID-GBIF-Dataset="2d28bebf-e3c5-457b-9152-8a6877ded281" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510742923475" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="C063786AFFEEFFBCFF0D13E55DCD9757" docLanguage="en" docName="zt03074p198.pdf.imf" docOrigin="Zootaxa 3074" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena flagella Perkins, 2011, new species" docType="treatment" docVersion="7" lastPageNumber="66" masterDocId="3C5A0012FFAEFFFDFF9A11335F219540" masterDocTitle="New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)" masterLastPageNumber="198" masterPageNumber="1" pageNumber="65" updateTime="1698485333395" updateUser="ExternalLinkService">
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<mods:title id="9F8B33C78FFE0118740E09D79C83747F">New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="ADEB8D8D1F016B8EBAC4DD71F725FBD9">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="9F17441423A268A38FA5540ACC08FA3A">text</mods:typeOfResource>
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<mods:title id="0ED55DE9166D89C30CF30EEAC1A10322">Zootaxa</mods:title>
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<mods:part id="3D04A5557B79F82BFA41D9F16CB0C08F">
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<mods:date id="7C9EC4B786BF1425E012549F359A24A8">2011</mods:date>
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<mods:number id="14C3751E2071DAD3223B6B2D95A95156">2011-10-28</mods:number>
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<mods:classification id="4BDFA36B43590F0FC300C035AF257A87">journal article</mods:classification>
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<mods:identifier id="B88C52B529361B4BEDF5BF79930BD1A9" type="DOI">10.5281/zenodo.1050060</mods:identifier>
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<treatment id="C063786AFFEEFFBCFF0D13E55DCD9757" ID-DOI="http://doi.org/10.5281/zenodo.6184582" ID-GBIF-Taxon="137306938" ID-Zenodo-Dep="6184582" LSID="urn:lsid:plazi:treatment:C063786AFFEEFFBCFF0D13E55DCD9757" httpUri="http://treatment.plazi.org/id/C063786AFFEEFFBCFF0D13E55DCD9757" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
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<subSubSection id="00D09AF7FFEEFFBDFF0D13E55CA39651" pageId="64" pageNumber="65" type="nomenclature">
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<paragraph id="4875C97CFFEEFFBDFF0D13E55D3197B0" blockId="64.[151,898,726,785]" box="[151,528,726,752]" pageId="64" pageNumber="65">
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<heading id="133D7E10FFEEFFBDFF0D13E55D3197B0" bold="true" box="[151,528,726,752]" fontSize="11" level="1" pageId="64" pageNumber="65" reason="1">
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<emphasis id="7ABE156EFFEEFFBDFF0D13E55D3197B0" bold="true" box="[151,528,726,752]" pageId="64" pageNumber="65">
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<taxonomicName id="8FCAB2FFFFEEFFBDFF0D13E55E5097B0" ID-CoL="6MHH2" box="[151,369,726,752]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="flagella" status="sp. nov.">
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<emphasis id="7ABE156EFFEEFFBDFF0D13E55E5097B0" bold="true" box="[151,369,726,752]" italics="true" pageId="64" pageNumber="65">Hydraena flagella</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="618DA815FFEEFFBDFE1A13E55D3197B0" box="[384,528,726,752]" pageId="64" pageNumber="65" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4875C97CFFEEFFBDFF0D13CA5CA39651" blockId="64.[151,898,726,785]" box="[151,898,761,785]" pageId="64" pageNumber="65">
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<figureCitation id="D0F1D5F9FFEEFFBDFF0D13CA5FDE9651" box="[151,255,761,785]" captionStart="FIGURE 138" captionStartId="168.[151,250,1650,1673]" captionTargetBox="[252,1334,193,1626]" captionTargetId="figure@168.[252,1334,193,1629]" captionTargetPageId="168" captionText="FIGURE 138. Hydraena flagella, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050212/files/figure.png" pageId="64" pageNumber="65">Figs. 138</figureCitation>
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(habitus), 140 (aedeagus), 190 (spermatheca), 202 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FFEEFFBDFF0D12715D9991DD" pageId="64" pageNumber="65" type="materials_examined">
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<paragraph id="4875C97CFFEEFFBDFF0D12715C91963D" blockId="64.[151,1437,833,2009]" pageId="64" pageNumber="65">
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<emphasis id="7ABE156EFFEEFFBDFF0D12715E6B961A" bold="true" box="[151,330,833,858]" pageId="64" pageNumber="65">
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<typeStatus id="977177DEFFEEFFBDFF0D12715FF3961A" box="[151,210,834,858]" pageId="64" pageNumber="65">Type</typeStatus>
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Material.
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</emphasis>
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<typeStatus id="977177DEFFEEFFBDFECE12725E9D9619" box="[340,444,833,857]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
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(male):
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<collectingCountry id="30DD89ECFFEEFFBDFDB012715DBC961A" box="[554,669,834,858]" name="Paraguay" pageId="64" pageNumber="65">
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<emphasis id="7ABE156EFFEEFFBDFDB012715DBC961A" bold="true" box="[554,669,834,858]" pageId="64" pageNumber="65">Paraguay</emphasis>
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</collectingCountry>
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: Dep. Guaira, Mbocayaty, Arroyo Gervasio,
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<date id="3C74EFBCFFEEFFBDFB2812725A059619" box="[1202,1316,833,857]" pageId="64" pageNumber="65" value="1995-11-04">4 xi 1995</date>
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, Drechsel. Deposited in the
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<collectionCode id="2EDB51B9FFEEFFBDFECC12555EBE963D" box="[342,415,870,893]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="64" pageNumber="65">NMW</collectionCode>
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.
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<emphasis id="7ABE156EFFEEFFBDFE3212565D08963D" bold="true" box="[424,553,869,893]" pageId="64" pageNumber="65">
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<typeStatus id="977177DEFFEEFFBDFE3212565D05963D" box="[424,548,869,893]" pageId="64" pageNumber="65" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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Same data as
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<typeStatus id="977177DEFFEEFFBDFD4A12565C13963D" box="[720,818,869,893]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
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(2
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<collectionCode id="2EDB51B9FFEEFFBDFCC212565C84963D" box="[856,933,869,893]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="64" pageNumber="65">NMW</collectionCode>
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).
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</paragraph>
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<paragraph id="4875C97CFFEEFFBDFF5D12BA5D9991DD" blockId="64.[151,1437,833,2009]" pageId="64" pageNumber="65">
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<emphasis id="7ABE156EFFEEFFBDFF5D12BA5EF396E2" bold="true" box="[199,466,905,930]" pageId="64" pageNumber="65">Differential Diagnosis.</emphasis>
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A strongly shining species, with dorsal punctation very fine and sparse; the punctures on the pronotal disc are smaller than the largest punctures on the frons, an unusual condition. The elytra are unusual in that the punctures become slightly coarser posteriorly, and many of the punctures on the disc are random or only subserial. Similar in dorsal habitus to
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<taxonomicName id="8FCAB2FFFFEEFFBDFDD612C55C2E914D" authority="Perkins" authorityName="Perkins" box="[588,783,1013,1037]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="jivaro">
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<emphasis id="7ABE156EFFEEFFBDFDD612C55D93914D" box="[588,690,1014,1037]" italics="true" pageId="64" pageNumber="65">H. jivaro</emphasis>
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Perkins
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</taxonomicName>
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(
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<figureCitation id="D0F1D5F9FFEEFFBDFCBA12C65CAF914D" box="[800,910,1013,1037]" captionStart="FIGURE 102" captionStartId="142.[151,250,1650,1673]" captionTargetBox="[256,1333,213,1613]" captionTargetId="figure@142.[237,1350,193,1629]" captionTargetPageId="142" captionText="FIGURE 102. Hydraena jivaro, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050174/files/figure.png" pageId="64" pageNumber="65">Figs. 102</figureCitation>
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,
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<figureCitation id="D0F1D5F9FFEEFFBDFC0612C65CE8914D" box="[924,969,1013,1037]" captionStart="FIGURE 138" captionStartId="168.[151,250,1650,1673]" captionTargetBox="[252,1334,193,1626]" captionTargetId="figure@168.[252,1334,193,1629]" captionTargetPageId="168" captionText="FIGURE 138. Hydraena flagella, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050212/files/figure.png" pageId="64" pageNumber="65">138</figureCitation>
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); differentiated therefrom by the slightly finer pronotal punctation, the greater degree of randomness of the basal elytral punctures, the narrower P2, and the differently shaped plaques (arcuate and larger in
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<taxonomicName id="8FCAB2FFFFEEFFBDFD2D150D5C0F9115" box="[695,814,1085,1109]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="flagella">
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<emphasis id="7ABE156EFFEEFFBDFD2D150D5C0F9115" box="[695,814,1085,1109]" italics="true" pageId="64" pageNumber="65">H. flagella</emphasis>
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</taxonomicName>
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). The aedeagi of the two species are similar in the presence of a long gonopore-bearing flagellum, the presence of long setae on the dorsal surface, and general shape, but markedly differ in many details (
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<figureCitation id="D0F1D5F9FFEEFFBDFD9D15B65D5391DD" box="[519,626,1157,1181]" captionStart="FIGURES 105 – 107" captionStartId="145.[151,264,1650,1673]" captionText="FIGURES 105 – 107. Aedeagi of holotypes." pageId="64" pageNumber="65">Figs. 105</figureCitation>
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,
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<figureCitation id="D0F1D5F9FFEEFFBDFDE415B65D8D91DD" box="[638,684,1157,1181]" captionStart="FIGURES 140 – 141" captionStartId="170.[151,264,1650,1673]" captionText="FIGURES 140 – 141. Aedeagi of holotypes." pageId="64" pageNumber="65">140</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FFEEFFBCFF5D159A5CC194C7" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="description">
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<paragraph id="4875C97CFFEEFFBDFF5D159A5CA0906D" blockId="64.[151,1437,833,2009]" pageId="64" pageNumber="65">
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<emphasis id="7ABE156EFFEEFFBDFF5D159A5E709182" bold="true" box="[199,337,1193,1218]" pageId="64" pageNumber="65">Description</emphasis>
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. Size:
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<typeStatus id="977177DEFFEEFFBDFE37159A5D2E9181" box="[429,527,1193,1217]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.57/0.67; head 0.22/0.36; pronotum 0.42/0.52, PA 0.42, PB 0.46; elytra 0.93/0.67. Dorsum of head dark brown to piceous; pronotum testaceous in front of and behind piceous fascia, ratios of color bands, as measured in midline, ca.
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<date id="3C74EFBCFFEEFFBDFB5015C25A349049" box="[1226,1301,1265,1289]" pageId="64" pageNumber="65" value="2014-07-07">7/14/7</date>
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; elytra dark brown; legs light brown; maxillary palpi testaceous, tip not darker.
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</paragraph>
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<paragraph id="4875C97CFFEEFFBDFF5D14095DBE90A1" blockId="64.[151,1437,833,2009]" pageId="64" pageNumber="65">Frons punctures ca. 0.5xef, not larger, but slightly denser near eyes than medially; interstices strongly shining, ca. 1.5–2xpd laterally, 3–7xpd medially. Clypeus microreticulate, dull laterally, very strongly shining and very finely sparsely punctulate, or impunctulate, medially. Mentum very sparsely very finely punctulate, weakly shining; postmentum very finely densely micropunctulate in median concavity, surrounding areas smooth, shining. Genae raised, shining, without posterior ridge.</paragraph>
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<paragraph id="4875C97CFFEEFFBDFF5D14DD5A069331" blockId="64.[151,1437,833,2009]" pageId="64" pageNumber="65">Pronotum moderately arcuate laterally; anterior margin straight behind eyes, emarginate behind frons, scintilla absent, but this area virtually impunctate; punctation very fine and very sparse, punctures on disc smaller than largest punctures of frons, interstices strongly shining, ca. 4–10xpd, punctures larger and denser at anterior and posterior; PF1 absent; PF2 extremely shallow, almost imperceptible; PF3 deep; PF4 very shallow or obsolete.</paragraph>
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<paragraph id="4875C97CFFEEFFBDFF5D174E5CA19265" blockId="64.[151,1437,833,2009]" pageId="64" pageNumber="65">Elytra moderately convex, weakly arcuate laterally; summit of posterior declivity at ca. posterior 1/3; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming slightly coarser toward posterior, many punctures on disc random or subserial. Intervals not raised, shining, on basal 1/3 ca. 2–5xpd, as are interstices between punctures of a row. Apices in dorsal aspect truncate or subtruncate, in posterior aspect margins forming strong angle with one another.</paragraph>
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<paragraph id="4875C97CFFEEFFBDFF5D16025ABD9299" blockId="64.[151,1437,833,2009]" pageId="64" pageNumber="65">Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/2/8/5. P1 laminate; median carina sinuate in profile. P2 narrow, l/w ca. 5/1, sides parallel or slightly diverging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques large, wide in posterior 1/2, tapering to point anteriorly, arcuate, converging toward one another anteriorly, weakly raised, located at sides of deep median depression. Metaventrite with very small point on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca.</paragraph>
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<paragraph id="4875C97CFFEFFFBCFF0D11A45CC194C7" blockId="65.[151,1436,151,536]" pageId="65" pageNumber="66">
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<date id="3C74EFBCFFEFFFBCFF0D11A45F9295EF" box="[151,179,151,175]" pageId="65" pageNumber="66">4x</date>
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P2. Profemur (male) with small, sharply pointed tubercle on medial margin near trochanter. Protibia straight, gradually, moderately increasing in width from base to subapically, then excavate to apex. Mesotibia straight, slen- der. Metatrochanter with very small tubercle. Metatibia slender, straight. Abdominal apex slightly asymmetrical; last tergite cap-like, with very small apicomedian notch. Female (microslide mount, n=1): last tergite sharply rounded, without apicomedian notch, setae long, not hooked, ca. 18 on each side of median gap; gonocoxite not midlongitudinally divided, apical margin emarginate between sensilla clusters, no transverse ridge evident at bor- der of microreticulation; spermatheca
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<typeStatus id="977177DEFFEFFFBCFDD810435D5394C8" box="[578,626,368,392]" pageId="65" pageNumber="66">type</typeStatus>
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C1, accessory gland cup partial.
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FFEFFFBCFF5D10A75CB994EC" box="[199,920,404,428]" pageId="65" pageNumber="66" type="etymology">
|
||
<paragraph id="4875C97CFFEFFFBCFF5D10A75CB994EC" blockId="65.[151,1436,151,536]" box="[199,920,404,428]" pageId="65" pageNumber="66">
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<emphasis id="7ABE156EFFEFFFBCFF5D10A75E6F94EC" bold="true" box="[199,334,404,428]" pageId="65" pageNumber="66">Etymology.</emphasis>
|
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Named in reference to the large aedeagal flagellum.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="00D09AF7FFEFFFBCFF5D10845DCD9757" pageId="65" pageNumber="66" type="distribution">
|
||
<paragraph id="4875C97CFFEFFFBCFF5D10845CF7948F" blockId="65.[151,1436,151,536]" box="[199,982,439,464]" pageId="65" pageNumber="66">
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<emphasis id="7ABE156EFFEFFFBCFF5D10845E419490" bold="true" box="[199,352,439,464]" pageId="65" pageNumber="66">Distribution.</emphasis>
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Currently known only from the
|
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<typeStatus id="977177DEFFEFFFBCFD56108B5DDD9490" box="[716,764,440,464]" pageId="65" pageNumber="66">type</typeStatus>
|
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locality (
|
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<figureCitation id="D0F1D5F9FFEFFFBCFCFC10845CE8948F" box="[870,969,439,463]" captionStart="FIGURE 202" captionStartId="195.[151,250,1650,1673]" captionText="FIGURE 202. Geographical distributions of Hydraena species." pageId="65" pageNumber="66">Fig. 202</figureCitation>
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).
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</paragraph>
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<paragraph id="4875C97CFFEFFFBCFF5D10EF5DCD9757" blockId="65.[151,1436,151,536]" pageId="65" pageNumber="66">
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<emphasis id="7ABE156EFFEFFFBCFF5D10EF5E3094B4" bold="true" box="[199,273,476,500]" pageId="65" pageNumber="66">Notes.</emphasis>
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The aedeagus of
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<taxonomicName id="8FCAB2FFFFEFFFBCFE4E10EE5D6A94B4" box="[468,587,476,500]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="flagella">
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<emphasis id="7ABE156EFFEFFFBCFE4E10EE5D6A94B4" box="[468,587,476,500]" italics="true" pageId="65" pageNumber="66">H. flagella</emphasis>
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</taxonomicName>
|
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(
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<figureCitation id="D0F1D5F9FFEFFFBCFDC010EF5D9894B4" box="[602,697,476,500]" captionStart="FIGURES 140 – 141" captionStartId="170.[151,264,1650,1673]" captionText="FIGURES 140 – 141. Aedeagi of holotypes." pageId="65" pageNumber="66">Fig. 140</figureCitation>
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) has three long setae on the dorsal surface, perhaps indicating that this species is related to those in the
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<taxonomicName id="8FCAB2FFFFEFFFBCFDAA13325D569758" box="[560,631,513,536]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="143" phylum="Arthropoda" rank="species" species="jivaro">
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<emphasis id="7ABE156EFFEFFFBCFDAA13325D569758" box="[560,631,513,536]" italics="true" pageId="65" pageNumber="66">Jivaro</emphasis>
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</taxonomicName>
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Complex.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |