179 lines
20 KiB
XML
179 lines
20 KiB
XML
<document id="7FC3AA84F81EDDD01FFD8E0D8B4FDBFF" ID-CLB-Dataset="31530" ID-DOI="10.5281/zenodo.1050060" ID-GBIF-Dataset="2d28bebf-e3c5-457b-9152-8a6877ded281" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510742923475" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="C063786AFFECFFBFFF0D11A45C3C93FA" docLanguage="en" docName="zt03074p198.pdf.imf" docOrigin="Zootaxa 3074" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena venezuela Perkins, 2011, new species" docType="treatment" docVersion="7" lastPageNumber="67" masterDocId="3C5A0012FFAEFFFDFF9A11335F219540" masterDocTitle="New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)" masterLastPageNumber="198" masterPageNumber="1" pageNumber="67" updateTime="1698485333395" updateUser="ExternalLinkService">
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<mods:titleInfo id="8246A592F1781B0E526060BF4341A95B">
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<mods:title id="1560BA8FD8D05566679B83C318670944">New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)</mods:title>
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<mods:name id="B344D4E61BBB1108CC13B42C66C2246E" type="personal">
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<mods:roleTerm id="C3E70A42397CC7D73AD09A8C78001F7F">Author</mods:roleTerm>
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<mods:namePart id="88AD15593B9CBD541F8A686F43F74734">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="E7C853CD21CBF023954B8C6545BFAFFD">text</mods:typeOfResource>
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<mods:title id="B4C2DD3F00654EE87A932A7AAA43C43F">Zootaxa</mods:title>
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<mods:part id="1053E61B4A614DBDE16E759BE45101C2">
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<mods:date id="CE7F281E767B475EC52D867ED375AA7C">2011</mods:date>
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<mods:detail id="827066D04DDCB7A0E6899F422BA2E4E2" type="pubDate">
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<mods:number id="F390811E71C5E6EB152B3F27E0FD2FE0">2011-10-28</mods:number>
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<mods:detail id="FDF3FC6C94859797835C50CC5683B062" type="volume">
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<mods:number id="E7EF902377863677E3D16FD2690B819C">3074</mods:number>
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<mods:classification id="7F3426351CCC9D47667131BEE0C58ED2">journal article</mods:classification>
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<mods:identifier id="1FE653888E92FE11C9AF799ECF7565DE" type="CLB-Dataset">31530</mods:identifier>
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<mods:identifier id="5859883F60643E604FFC5492629FEACE" type="DOI">10.5281/zenodo.1050060</mods:identifier>
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<mods:identifier id="FA66D65AE0C30978E37849E7C0A1788A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A85EEDE77BB2676BF5B02EAB372600B8" type="Zenodo-Dep">1050060</mods:identifier>
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<treatment id="C063786AFFECFFBFFF0D11A45C3C93FA" ID-DOI="http://doi.org/10.5281/zenodo.6184586" ID-GBIF-Taxon="137306936" ID-Zenodo-Dep="6184586" LSID="urn:lsid:plazi:treatment:C063786AFFECFFBFFF0D11A45C3C93FA" httpUri="http://treatment.plazi.org/id/C063786AFFECFFBFFF0D11A45C3C93FA" lastPageNumber="67" pageId="66" pageNumber="67">
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<subSubSection id="00D09AF7FFECFFBFFF0D11A45D859592" pageId="66" pageNumber="67" type="nomenclature">
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<paragraph id="4875C97CFFECFFBFFF0D11A45D0D95F1" blockId="66.[151,676,151,210]" box="[151,556,151,177]" pageId="66" pageNumber="67">
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<heading id="133D7E10FFECFFBFFF0D11A45D0D95F1" bold="true" box="[151,556,151,177]" fontSize="11" level="1" pageId="66" pageNumber="67" reason="1">
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<emphasis id="7ABE156EFFECFFBFFF0D11A45D0D95F1" bold="true" box="[151,556,151,177]" pageId="66" pageNumber="67">
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<taxonomicName id="8FCAB2FFFFECFFBFFF0D11A45EAC95F1" ID-CoL="3N3KL" box="[151,397,151,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="venezuela" status="sp. nov.">
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<emphasis id="7ABE156EFFECFFBFFF0D11A45EAC95F1" bold="true" box="[151,397,151,177]" italics="true" pageId="66" pageNumber="67">Hydraena venezuela</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="618DA815FFECFFBFFE0611A45D0D95F1" box="[412,556,151,177]" pageId="66" pageNumber="67" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="4875C97CFFECFFBFFF0D11895D859592" blockId="66.[151,676,151,210]" box="[151,676,186,210]" pageId="66" pageNumber="67">
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<figureCitation id="D0F1D5F9FFECFFBFFF0D11895FDE9592" box="[151,255,186,210]" captionStart="FIGURE 142" captionStartId="171.[151,250,1650,1673]" captionTargetBox="[220,1366,193,1625]" captionTargetId="figure@171.[220,1366,193,1629]" captionTargetPageId="171" captionText="FIGURE 142. Hydraena venezuela, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050216/files/figure.png" pageId="66" pageNumber="67">Figs. 142</figureCitation>
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(habitus), 144 (aedeagus), 199 (map)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="00D09AF7FFECFFBFFF0D10305C9A977A" pageId="66" pageNumber="67" type="materials_examined">
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<paragraph id="4875C97CFFECFFBFFF0D10305FD594EA" blockId="66.[151,1437,258,1723]" pageId="66" pageNumber="67">
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<emphasis id="7ABE156EFFECFFBFFF0D10305E66945B" bold="true" box="[151,327,258,283]" pageId="66" pageNumber="67">
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<typeStatus id="977177DEFFECFFBFFF0D10305FF3945B" box="[151,210,259,283]" pageId="66" pageNumber="67">Type</typeStatus>
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Material.
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</emphasis>
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<typeStatus id="977177DEFFECFFBFFED710315E94945A" box="[333,437,258,282]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
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(male):
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<taxonomicName id="8FCAB2FFFFECFFBFFD8010305DB3945B" box="[538,658,259,283]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="172" phylum="Arthropoda" rank="species" species="venezuela">
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<collectingCountry id="30DD89ECFFECFFBFFD8010305DB3945B" box="[538,658,259,283]" name="Venezuela" pageId="66" pageNumber="67">
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||
<emphasis id="7ABE156EFFECFFBFFD8010305DB3945B" bold="true" box="[538,658,259,283]" pageId="66" pageNumber="67">Venezuela</emphasis>
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</collectingCountry>
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</taxonomicName>
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: Bolivar, Los Pijiguaos: outcrop/morichal, detrital pools, elev.
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<quantity id="8F326499FFECFFBFFAC710315AB6945A" box="[1373,1431,258,282]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="66" pageNumber="67" unit="m" value="80.0">80 m</quantity>
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,
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<geoCoordinate id="2DFEAFBBFFECFFBFFF0D10155E09947E" box="[151,296,294,318]" direction="north" orientation="latitude" pageId="66" pageNumber="67" precision="1" value="6.5936165">6° 35.617' N</geoCoordinate>
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,
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<geoCoordinate id="2DFEAFBBFFECFFBFFEAE10155EF9947E" box="[308,472,294,318]" direction="west" orientation="longitude" pageId="66" pageNumber="67" precision="1" value="-66.82063">66° 49.238' W</geoCoordinate>
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,
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<date id="3C74EFBCFFECFFBFFE7910155D71947E" box="[483,592,294,318]" pageId="66" pageNumber="67" value="2009-01-12">12 i 2009</date>
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, Short, Miller (VZ09-0112-01C). Deposited in the
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<collectionCode id="2EDB51B9FFECFFBFFB0A10145BFA947F" LSID="urn:lsid:biocol.org:col:33834" box="[1168,1243,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="66" pageNumber="67">MIZA</collectionCode>
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.
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<typeStatus id="977177DEFFECFFBFFB7C10145A7F947F" box="[1254,1374,295,319]" pageId="66" pageNumber="67" type="paratype">
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<emphasis id="7ABE156EFFECFFBFFB7C10145A7F947F" bold="true" box="[1254,1374,295,319]" pageId="66" pageNumber="67">Paratypes</emphasis>
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</typeStatus>
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(29):
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<taxonomicName id="8FCAB2FFFFECFFBFFF0D10785E2E9423" box="[151,271,331,355]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="172" phylum="Arthropoda" rank="species" species="venezuela">
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<collectingCountry id="30DD89ECFFECFFBFFF0D10785E2E9423" box="[151,271,331,355]" name="Venezuela" pageId="66" pageNumber="67">
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<emphasis id="7ABE156EFFECFFBFFF0D10785E2E9423" bold="true" box="[151,271,331,355]" pageId="66" pageNumber="67">Venezuela</emphasis>
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</collectingCountry>
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</taxonomicName>
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: Bolivar, Same data as
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<typeStatus id="977177DEFFECFFBFFDB510795DB09422" box="[559,657,330,354]" pageId="66" pageNumber="67" type="holotype">holotype</typeStatus>
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(26
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<collectionCode id="2EDB51B9FFECFFBFFD5310795C3B9422" box="[713,794,330,354]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="66" pageNumber="67">SEMC</collectionCode>
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); Los Pijiguaos: outcrop/morichal, margin of morichal, elev.
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<quantity id="8F326499FFECFFBFFF4F105D5E3394C6" box="[213,274,366,390]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="66" pageNumber="67" unit="m" value="80.0">80 m</quantity>
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,
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<geoCoordinate id="2DFEAFBBFFECFFBFFEBB105D5E9694C6" box="[289,439,366,390]" direction="north" orientation="latitude" pageId="66" pageNumber="67" precision="1" value="6.5936165">6° 35.617' N</geoCoordinate>
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,
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<geoCoordinate id="2DFEAFBBFFECFFBFFE5F105D5D4C94C6" box="[453,621,366,390]" direction="west" orientation="longitude" pageId="66" pageNumber="67" precision="1" value="-66.82063">66° 49.238' W</geoCoordinate>
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,
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<date id="3C74EFBCFFECFFBFFDE1105D5DCC94C6" box="[635,749,366,390]" pageId="66" pageNumber="67" value="2009-01-12">12 i 2009</date>
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, Short, Garcia, Camacho, Miller, Joly (VZ09-0112-01A) (3
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<collectionCode id="2EDB51B9FFECFFBFFF0D10A15FC894EA" box="[151,233,402,426]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="66" pageNumber="67">SEMC</collectionCode>
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||
).
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</paragraph>
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<paragraph id="4875C97CFFECFFBFFF5D10855C9A977A" blockId="66.[151,1437,258,1723]" pageId="66" pageNumber="67">
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||
<emphasis id="7ABE156EFFECFFBFFF5D10855EF5948F" bold="true" box="[199,468,438,463]" pageId="66" pageNumber="67">Differential Diagnosis.</emphasis>
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A finely sparsely punctate, strongly shining species with distinctive pronotal fascia. Somewhat similar in dorsal habitus to
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<taxonomicName id="8FCAB2FFFFECFFBFFDDF10EF5D8994B3" box="[581,680,475,499]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="shorti">
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<emphasis id="7ABE156EFFECFFBFFDDF10EF5D8994B3" box="[581,680,475,499]" italics="true" pageId="66" pageNumber="67">H. shorti</emphasis>
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</taxonomicName>
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(
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<figureCitation id="D0F1D5F9FFECFFBFFD2D10E95C3494B2" box="[695,789,474,498]" captionStart="FIGURE 68" captionStartId="120.[151,250,1650,1673]" captionTargetBox="[235,1350,193,1627]" captionTargetId="figure@120.[235,1351,193,1629]" captionTargetPageId="120" captionText="FIGURE 68. Hydraena shorti, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050136/files/figure.png" pageId="66" pageNumber="67">Figs. 68</figureCitation>
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,
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<figureCitation id="D0F1D5F9FFECFFBFFCBB10E95C6E94B2" box="[801,847,474,498]" captionStart="FIGURE 142" captionStartId="171.[151,250,1650,1673]" captionTargetBox="[220,1366,193,1625]" captionTargetId="figure@171.[220,1366,193,1629]" captionTargetPageId="171" captionText="FIGURE 142. Hydraena venezuela, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050216/files/figure.png" pageId="66" pageNumber="67">142</figureCitation>
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); differentiated therefrom by the larger size (ca. 1.38 vs.
|
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<quantity id="8F326499FFECFFBFFF5A10CD5E0B9756" box="[192,298,510,534]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="66" pageNumber="67" unit="mm" value="1.25">1.25 mm</quantity>
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), the wider pronotal fascia (color band ratios ca.
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<date id="3C74EFBCFFECFFBFFCC510CD5C879756" box="[863,934,510,534]" pageId="66" pageNumber="67" value="2015-05-05">5/15/5</date>
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vs.
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<date id="3C74EFBCFFECFFBFFC4210CD5B029756" box="[984,1059,510,534]" pageId="66" pageNumber="67" value="2010-06-06">6/10/6</date>
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), the narrower P2, and the wider plaques. The aedeagi of the two species markedly differ (
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<figureCitation id="D0F1D5F9FFECFFBFFC8D13115C54977A" box="[791,885,546,570]" captionStart="FIGURES 71 – 73" captionStartId="123.[151,264,1582,1605]" captionText="FIGURES 71 – 73. Aedeagi of holotypes." pageId="66" pageNumber="67">Figs. 72</figureCitation>
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,
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<figureCitation id="D0F1D5F9FFECFFBFFC1B13115C8E977A" box="[897,943,546,570]" captionStart="FIGURES 144 – 145" captionStartId="173.[151,264,1650,1673]" captionText="FIGURES 144 – 145. Aedeagi of holotypes." pageId="66" pageNumber="67">144</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FFECFFBFFF5D13755FF7936A" pageId="66" pageNumber="67" type="description">
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<paragraph id="4875C97CFFECFFBFFF5D13755D47978A" blockId="66.[151,1437,258,1723]" pageId="66" pageNumber="67">
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||
<emphasis id="7ABE156EFFECFFBFFF5D13755E70971F" bold="true" box="[199,337,582,607]" pageId="66" pageNumber="67">Description</emphasis>
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. Size:
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<typeStatus id="977177DEFFECFFBFFE3713755D2E971E" box="[429,527,582,606]" pageId="66" pageNumber="67" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.38/0.58; head 0.22/0.35; pronotum 0.35/0.46, PA 0.38, PB 0.43; elytra 0.80/0.58. Dorsum of head piceous; pronotum testaceous in front of and behind piceous fascia, ratios of color bands, as measured in midline, ca.
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<date id="3C74EFBCFFECFFBFFC7413BD5B1897E6" box="[1006,1081,654,678]" pageId="66" pageNumber="67" value="2015-05-05">5/15/5</date>
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; elytra dark brown; legs brown; maxillary palpi testaceous, tip not darker.
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</paragraph>
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<paragraph id="4875C97CFFECFFBFFF5D13E45C81961A" blockId="66.[151,1437,258,1723]" pageId="66" pageNumber="67">Frons punctures ca. 1xef laterally, smaller and sparser medially; interstices shining, 1–2xpd laterally, 3–5xpd medially. Clypeus microreticulate laterally, very finely sparsely punctulate medially. Mentum finely, moderately sparsely punctulate, shining; postmentum very finely densely micropunctulate in small median area, surrounding areas smooth, shining. Genae raised, shining, without posterior ridge.</paragraph>
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<paragraph id="4875C97CFFECFFBFFF5D12545FDD96AA" blockId="66.[151,1437,258,1723]" pageId="66" pageNumber="67">Pronotum weakly arcuate laterally; anterior margin straight behind eyes, emarginate behind frons, scintilla absent; finely sparsely punctate, punctures on disc ca. 1xpd frons punctures, interstices strongly shining, 4–8xpd, punctures slightly larger and denser at anterior and posterior; PF1 and PF2 absent; PF3 deep; PF4 very shallow or obsolete.</paragraph>
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<paragraph id="4875C97CFFECFFBFFF5D12C55D9B91DE" blockId="66.[151,1437,258,1723]" pageId="66" pageNumber="67">Elytra weakly convex; weakly arcuate laterally; summit of posterior declivity at about midlength; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming finer and more widely spaced toward posterior. Intervals not raised, shining, on basal 1/3 ca. 2–3xpd, as are interstices between punctures of a row, a few punctures subserial. Apices in dorsal aspect subtruncate, in posterior aspect margins forming rather large angle with one another.</paragraph>
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<paragraph id="4875C97CFFECFFBFFF5D15995FF7936A" blockId="66.[151,1437,258,1723]" pageId="66" pageNumber="67">
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Ratios of P2 width and plaque shape (P2/w/l/s) ca. 2.5/2/6/4. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately wide, straight, slightly converging toward one another anteriorly, weakly raised, located at sides of deep median depression. Metaventrite with very small point on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 1.5x P2. Protibia very slightly arcuate, widest at ca. distal 1/4, where located tiny sharp tooth. Meso- and metatibia slender, extremely slightly, if at all, arcuate.
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<collectingCountry id="30DD89ECFFECFFBFFF6214B15E1090DA" box="[248,305,1410,1434]" name="Malta" pageId="66" pageNumber="67">Male</collectingCountry>
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abdominal apex very slightly, if at all, asymmetrical; last tergite with apicomedian notch. Female (microslide mount, n=1): last tergite sharply rounded, without apicomedian notch, ca. 28 long, closely spaced, some contiguous, moderately hooked setae; gonocoxite not midlongitudinally divided, apical margin sharply rounded, no transverse ridge evident at border of microreticulation; spermatheca
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<typeStatus id="977177DEFFECFFBFFBDC14DC5B579347" box="[1094,1142,1519,1543]" pageId="66" pageNumber="67">type</typeStatus>
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C1, accessory gland cup large.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00D09AF7FFECFFBFFF5D17045CBF930E" box="[199,926,1590,1615]" pageId="66" pageNumber="67" type="etymology">
|
||
<paragraph id="4875C97CFFECFFBFFF5D17045CBF930E" blockId="66.[151,1437,258,1723]" box="[199,926,1590,1615]" pageId="66" pageNumber="67">
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||
<emphasis id="7ABE156EFFECFFBFFF5D17045E6F930F" bold="true" box="[199,334,1591,1615]" pageId="66" pageNumber="67">Etymology.</emphasis>
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Named in reference to the geographical distribution.
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FFECFFBFFF5D17695C3C93FA" pageId="66" pageNumber="67" type="distribution">
|
||
<paragraph id="4875C97CFFECFFBFFF5D17695B5D9332" blockId="66.[151,1437,258,1723]" box="[199,1148,1626,1651]" pageId="66" pageNumber="67">
|
||
<emphasis id="7ABE156EFFECFFBFFF5D17695E419333" bold="true" box="[199,352,1626,1651]" pageId="66" pageNumber="67">Distribution.</emphasis>
|
||
Currently known from the
|
||
<typeStatus id="977177DEFFECFFBFFD0917685DE29333" box="[659,707,1627,1651]" pageId="66" pageNumber="67">type</typeStatus>
|
||
locality and one nearby site (
|
||
<figureCitation id="D0F1D5F9FFECFFBFFB9717695B519332" box="[1037,1136,1626,1650]" captionStart="FIGURE 199" captionStartId="192.[151,250,1650,1673]" captionText="FIGURE 199. Geographical distributions of Hydraena species." pageId="66" pageNumber="67">Fig. 199</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4875C97CFFECFFBFFF5D174C5C3C93FA" blockId="66.[151,1437,258,1723]" pageId="66" pageNumber="67">
|
||
<emphasis id="7ABE156EFFECFFBFFF5D174C5E3093D7" bold="true" box="[199,273,1663,1687]" pageId="66" pageNumber="67">Notes.</emphasis>
|
||
The aedeagus of
|
||
<taxonomicName id="8FCAB2FFFFECFFBFFE4717B35D4E93D7" box="[477,623,1663,1687]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="venezuela">
|
||
<emphasis id="7ABE156EFFECFFBFFE4717B35D4E93D7" box="[477,623,1663,1687]" italics="true" pageId="66" pageNumber="67">H. venezuela</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="D0F1D5F9FFECFFBFFD1B174D5DC393D6" box="[641,738,1662,1686]" captionStart="FIGURES 144 – 145" captionStartId="173.[151,264,1650,1673]" captionText="FIGURES 144 – 145. Aedeagi of holotypes." pageId="66" pageNumber="67">Fig. 144</figureCitation>
|
||
) has two long setae on the dorsal surface, perhaps indicating that this species is related to those in the
|
||
<taxonomicName id="8FCAB2FFFFECFFBFFDFB17975D8693FB" box="[609,679,1700,1723]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="143" phylum="Arthropoda" rank="species" species="jivaro">
|
||
<emphasis id="7ABE156EFFECFFBFFDFB17975D8693FB" box="[609,679,1700,1723]" italics="true" pageId="66" pageNumber="67">Jivaro</emphasis>
|
||
</taxonomicName>
|
||
Complex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |