154 lines
18 KiB
XML
154 lines
18 KiB
XML
<document id="9A040B328F48B08CC1374EF23D8F14A3" ID-CLB-Dataset="31530" ID-DOI="10.5281/zenodo.1050060" ID-GBIF-Dataset="2d28bebf-e3c5-457b-9152-8a6877ded281" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510742923475" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="C063786AFFBEFFECFF0D15675E67971F" docLanguage="en" docName="zt03074p198.pdf.imf" docOrigin="Zootaxa 3074" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena punctilata Perkins, 2011, new species" docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="3C5A0012FFAEFFFDFF9A11335F219540" masterDocTitle="New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)" masterLastPageNumber="198" masterPageNumber="1" pageNumber="17" updateTime="1698485333395" updateUser="ExternalLinkService">
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<mods:mods id="0C9BF5DFF39C057A9233D4DB9D427266" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6A8CAB6501C3967474CCB33F553AFCC6">
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<mods:title id="AA909C2E8727315EFD79166CFBF4CE18">New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)</mods:title>
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<mods:name id="B9FD3AE359128BF4D8A14578A006C216" type="personal">
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<mods:roleTerm id="24146C69B566E711E45E5D8451B77982">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EA1A48A39EA71B5D2A4C3ACA8C5D65E3">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="0921012BB6995A726AAE417C45FA39FE">text</mods:typeOfResource>
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<mods:title id="367985C8F7367DEED4CBC6BEBCD19A4A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="42C38365E848F4A764F7A6C1A9693D9A">
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<mods:date id="CBFE2E5DC9AC19AF7B23D08A1DD5BDFA">2011</mods:date>
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<mods:detail id="CCC4AFCB645064D5B719ABC3063A0184" type="pubDate">
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<mods:number id="B09E1D1130A8B5BACF0AAC5635269B00">2011-10-28</mods:number>
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</mods:detail>
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<mods:detail id="F18D77C2215897ADC6273BD2A70E5BD9" type="volume">
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<mods:number id="E625B6353570B2AAB55AE2751227FFE9">3074</mods:number>
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<mods:classification id="4016EC486AB1745F88F86A0637B35CC4">journal article</mods:classification>
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<mods:identifier id="C12C14072845EC3C442C25D9DFF823C4" type="CLB-Dataset">31530</mods:identifier>
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<mods:identifier id="0E31213FC0682C73D98B4FCA099264F5" type="DOI">10.5281/zenodo.1050060</mods:identifier>
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<mods:identifier id="FDE4AEF475E8371B82A92C417FA5E6EB" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0CC6C4FA5DD4210718A9E4BAA209D0D6" type="Zenodo-Dep">1050060</mods:identifier>
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<treatment id="C063786AFFBEFFECFF0D15675E67971F" ID-DOI="http://doi.org/10.5281/zenodo.6184412" ID-GBIF-Taxon="137306947" ID-Zenodo-Dep="6184412" LSID="urn:lsid:plazi:treatment:C063786AFFBEFFECFF0D15675E67971F" httpUri="http://treatment.plazi.org/id/C063786AFFBEFFECFF0D15675E67971F" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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<subSubSection id="00D09AF7FFBEFFEDFF0D15675D5B91CE" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="4875C97CFFBEFFEDFF0D15675D0E912D" blockId="16.[151,634,1107,1166]" box="[151,559,1107,1134]" pageId="16" pageNumber="17">
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<heading id="133D7E10FFBEFFEDFF0D15675D0E912D" bold="true" box="[151,559,1107,1134]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<emphasis id="7ABE156EFFBEFFEDFF0D15675D0E912D" bold="true" box="[151,559,1107,1134]" pageId="16" pageNumber="17">
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<taxonomicName id="8FCAB2FFFFBEFFEDFF0D15675EB1912E" ID-CoL="3N2JJ" box="[151,400,1108,1134]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="punctilata" status="sp. nov.">
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<emphasis id="7ABE156EFFBEFFEDFF0D15675EB1912E" bold="true" box="[151,400,1108,1134]" italics="true" pageId="16" pageNumber="17">Hydraena punctilata</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="618DA815FFBEFFEDFE0515605D0E912D" box="[415,559,1107,1133]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4875C97CFFBEFFEDFF0D15455D5B91CE" blockId="16.[151,634,1107,1166]" box="[151,634,1142,1166]" pageId="16" pageNumber="17">
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<figureCitation id="D0F1D5F9FFBEFFEDFF0D15455FC291CE" box="[151,227,1142,1166]" captionStart="FIGURE 7" captionStartId="76.[151,250,1650,1673]" captionTargetBox="[285,1308,218,1613]" captionTargetId="figure@76.[268,1318,193,1629]" captionTargetPageId="76" captionText="FIGURE 7. Hydraena punctilata, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050074/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
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(habitus), 14 (aedeagus), 193 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FFBEFFEDFF0D15895CA090AC" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="4875C97CFFBEFFEDFF0D15895E7391B2" blockId="16.[151,1437,1209,2020]" pageId="16" pageNumber="17">
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<emphasis id="7ABE156EFFBEFFEDFF0D15895E699192" bold="true" box="[151,328,1209,1234]" pageId="16" pageNumber="17">
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<typeStatus id="977177DEFFBEFFEDFF0D15895FF39192" box="[151,210,1210,1234]" pageId="16" pageNumber="17">Type</typeStatus>
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Material.
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</emphasis>
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<typeStatus id="977177DEFFBEFFEDFECA158A5E999191" box="[336,440,1209,1233]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
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(male):
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<collectingCountry id="30DD89ECFFBEFFEDFDBA158A5D489192" box="[544,617,1209,1234]" name="Brazil" pageId="16" pageNumber="17">
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<emphasis id="7ABE156EFFBEFFEDFDBA158A5D489192" bold="true" box="[544,617,1209,1234]" pageId="16" pageNumber="17">Brazil</emphasis>
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</collectingCountry>
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, Amazonas, Est. [Estirao] Do
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<collectingCountry id="30DD89ECFFBEFFEDFC59158A5B039191" box="[963,1058,1209,1233]" name="Ecuador" pageId="16" pageNumber="17">Ecuador</collectingCountry>
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,
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<date id="3C74EFBCFFBEFFEDFBB4158A5BB59191" box="[1070,1172,1209,1233]" pageId="16" pageNumber="17" value="1973-10-01">1 x 1973</date>
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, M. Alvarenga. Deposited in the
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<collectionCode id="2EDB51B9FFBEFFEDFE9415E95E6E91B2" LSID="urn:lsid:biocol.org:col:33791" box="[270,335,1242,1266]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="16" pageNumber="17">MCZ</collectionCode>
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.
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</paragraph>
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<paragraph id="4875C97CFFBEFFEDFF5D15C85CA090AC" blockId="16.[151,1437,1209,2020]" pageId="16" pageNumber="17">
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<emphasis id="7ABE156EFFBEFFEDFF5D15C85EF29054" bold="true" box="[199,467,1275,1300]" pageId="16" pageNumber="17">Differential Diagnosis.</emphasis>
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Somewhat similar in dorsal habitus to
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<taxonomicName id="8FCAB2FFFFBEFFEDFC1715CE5B089054" box="[909,1065,1277,1300]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="paeminosa">
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<emphasis id="7ABE156EFFBEFFEDFC1715CE5B089054" box="[909,1065,1277,1300]" italics="true" pageId="16" pageNumber="17">H. paeminosa</emphasis>
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</taxonomicName>
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(
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<figureCitation id="D0F1D5F9FFBEFFEDFBA315CF5BA99054" box="[1081,1160,1276,1300]" captionStart="FIGURE 3" captionStartId="72.[151,250,1650,1673]" captionTargetBox="[227,1366,208,1616]" captionTargetId="figure@72.[208,1379,193,1629]" captionTargetPageId="72" captionText="FIGURE 3. Hydraena paeminosa, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050066/files/figure.png" pageId="16" pageNumber="17">Figs. 3</figureCitation>
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,
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<figureCitation id="D0F1D5F9FFBEFFEDFB0C15CF5B879054" box="[1174,1190,1276,1300]" captionStart="FIGURE 7" captionStartId="76.[151,250,1650,1673]" captionTargetBox="[285,1308,218,1613]" captionTargetId="figure@76.[268,1318,193,1629]" captionTargetPageId="76" captionText="FIGURE 7. Hydraena punctilata, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050074/files/figure.png" pageId="16" pageNumber="17">7</figureCitation>
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); differentiated therefrom by the slightly larger size (length ca.
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<quantity id="8F326499FFBEFFEDFDED142C5DFA9077" box="[631,731,1311,1335]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" pageId="16" pageNumber="17" unit="mm" value="1.76">1.76 mm</quantity>
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vs.
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<quantity id="8F326499FFBEFFEDFC90142C5C539077" box="[778,882,1311,1335]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="16" pageNumber="17" unit="mm" value="1.65">1.65 mm</quantity>
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), the more transverse pronotum (length/width ca. 0.69 vs. 0.82), the coarser pronotal punctation, and the greater degree of randomness of the elytra punctures. In addition, the hind tibiae of males are markedly arcuate in
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<taxonomicName id="8FCAB2FFFFBEFFEDFC81145A5C8F90C0" box="[795,942,1384,1408]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="punctilata">
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<emphasis id="7ABE156EFFBEFFEDFC81145A5C8F90C0" box="[795,942,1384,1408]" italics="true" pageId="16" pageNumber="17">H. punctilata</emphasis>
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</taxonomicName>
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, and straight in
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<taxonomicName id="8FCAB2FFFFBEFFEDFBF9145A5BDF90C0" box="[1123,1278,1385,1408]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="paeminosa">
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<emphasis id="7ABE156EFFBEFFEDFBF9145A5BDF90C0" box="[1123,1278,1385,1408]" italics="true" pageId="16" pageNumber="17">H. paeminosa</emphasis>
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</taxonomicName>
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. The pronotal punctation is as coarse as
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<taxonomicName id="8FCAB2FFFFBEFFEDFE2C14BE5D0D90E4" box="[438,556,1420,1444]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="neblina">
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<emphasis id="7ABE156EFFBEFFEDFE2C14BE5D0D90E4" box="[438,556,1420,1444]" italics="true" pageId="16" pageNumber="17">H. neblina</emphasis>
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</taxonomicName>
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(
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<figureCitation id="D0F1D5F9FFBEFFEDFDA114BF5D5F90E4" box="[571,638,1420,1444]" captionStart="FIGURE 8" captionStartId="77.[151,250,1612,1635]" captionTargetBox="[226,1357,193,1591]" captionTargetId="figure@77.[226,1360,193,1591]" captionTargetPageId="77" captionText="FIGURE 8. Hydraena neblina, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050076/files/figure.png" pageId="16" pageNumber="17">Fig. 8</figureCitation>
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), but the body form is much different in the two species. The male genitalia of the these three species markedly differ in many characters (
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<figureCitation id="D0F1D5F9FFBEFFEDFC14149C5B299087" box="[910,1032,1455,1479]" captionStart="FIGURES 9 – 11" captionStartId="78.[151,264,1606,1629]" captionText="FIGURES 9 – 11. Aedeagi of holotypes." pageId="16" pageNumber="17">Figs. 9, 10</figureCitation>
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,
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<figureCitation id="D0F1D5F9FFBEFFEDFB8E149C5B129087" box="[1044,1075,1455,1479]" captionStart="FIGURES 14 – 15" captionStartId="80.[151,264,1650,1673]" captionText="FIGURES 14 – 15. Aedeagi of holotypes." pageId="16" pageNumber="17">14</figureCitation>
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). The
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<typeStatus id="977177DEFFBEFFEDFBE0149C5BFD9087" box="[1146,1244,1455,1479]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
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is very fragmentary; as a result, the positions of the elytra in
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<figureCitation id="D0F1D5F9FFBEFFEDFD1514E75DF090AC" box="[655,721,1492,1516]" captionStart="FIGURE 7" captionStartId="76.[151,250,1650,1673]" captionTargetBox="[285,1308,218,1613]" captionTargetId="figure@76.[268,1318,193,1629]" captionTargetPageId="76" captionText="FIGURE 7. Hydraena punctilata, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050074/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
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are not natural.
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FFBEFFECFF5D14C45E82948F" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
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<paragraph id="4875C97CFFBEFFEDFF5D14C45E51933C" blockId="16.[151,1437,1209,2020]" pageId="16" pageNumber="17">
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<emphasis id="7ABE156EFFBEFFEDFF5D14C45E709350" bold="true" box="[199,337,1527,1552]" pageId="16" pageNumber="17">Description</emphasis>
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. Size:
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<typeStatus id="977177DEFFBEFFEDFE3714C45D2E934F" box="[429,527,1527,1551]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.76/0.87; head 0.26/0.41; pronotum 0.40/0.58, PA 0.46, PB 0.54; elytra 1.15/0.87. Dorsum of head with frons piceous, clypeus brown; pronotum dark brown to piceous, except grading to brown at margins; elytra dark brown; legs brown; maxillary palpi brown, tip missing.
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</paragraph>
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<paragraph id="4875C97CFFBEFFEDFF5D17BA5B2393A8" blockId="16.[151,1437,1209,2020]" pageId="16" pageNumber="17">Dorsum of head microreticulate, dull. Frons very coarsely densely punctate, punctures ca. 5–7xef, interstices ca. 0.5xpd to narrow walls. Clypeus punctures obsolete. Mentum and postmentum microreticulate, dull; postmentum rather coarsely punctate. Genae raised in posterior ridge, except medially.</paragraph>
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<paragraph id="4875C97CFFBEFFEDFF5D17C65D6092A4" blockId="16.[151,1437,1209,2020]" pageId="16" pageNumber="17">Pronotum and elytra coarsely densely punctate, subscabrous, each puncture with distinctive, short seta, interstices microreticulate on pronotum. Pronotum with median 1/3 of anterior margin emarginate, except middle are very slightly produced and with very narrow hyaline border; scintilla absent; PF1 absent; PF2 very shallow, almost imperceptible; PF3 and PF4 deep, confluent, separated from lateral margin by low irregular ridge. Elytra quite convex at summit of posterior declivity, slightly past midlength; lateral explanate margins wide; punctures random; interstices irregular surfaces, weakly shining. Apices in dorsal aspect weakly separately rounded, shape in posterior aspect uncertain (elytra are detached).</paragraph>
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<paragraph id="4875C97CFFBFFFECFF5D11A45E82948F" blockId="17.[151,1436,151,607]" pageId="17" pageNumber="18">
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Ratios of P2 width and plaque shape (P2/w/l/s) ca.
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<date id="3C74EFBCFFBFFFECFC9911A45C0895EF" box="[771,809,151,175]" bridgedPair="/" pageId="17" pageNumber="18">x/2</date>
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/8/4. P1 slightly wider than laminate; median carina sinuate in profile. P2 tapering to sharp point, tip contiguous with low median carina of metaventrite. Plaques moderately wide posteriorly, tapering to point anteriorly, narrowly separated, very slightly convergent anteriorly, almost parallel, weakly raised, located at sides of shallow median depression. Metaventrite with median carina in anterior 1/2, continued posteriorly to between anterior part of plaques, also with short point on each side, extended posteriorly from margin of each mesocoxal cavity. AIS slightly concave, width at straight posterior margin ca. equal plaque separation. Pro- and mesotibia very slightly arcuate, moderately slender. Metatibia distinctively arcuate in distal 1/2.
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<collectingCountry id="30DD89ECFFBFFFECFE9610A75E6794EC" box="[268,326,404,428]" name="Malta" pageId="17" pageNumber="18">Male</collectingCountry>
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abdominal apex symmetrical; last tergite with posterior 1/2 slightly upturned, free margin rounded. Females not yet known.
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FFBFFFECFF5D10EF5B5E94B4" box="[199,1151,476,500]" pageId="17" pageNumber="18" type="etymology">
|
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<paragraph id="4875C97CFFBFFFECFF5D10EF5B5E94B4" blockId="17.[151,1436,151,607]" box="[199,1151,476,500]" pageId="17" pageNumber="18">
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<emphasis id="7ABE156EFFBFFFECFF5D10EF5E6F94B4" bold="true" box="[199,334,476,500]" pageId="17" pageNumber="18">Etymology.</emphasis>
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Named in reference to the wide body form and coarse dorsal punctation.
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FFBFFFECFF5D10CC5E67971F" pageId="17" pageNumber="18" type="distribution">
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<paragraph id="4875C97CFFBFFFECFF5D10CC5CF49757" blockId="17.[151,1436,151,607]" box="[199,981,511,536]" pageId="17" pageNumber="18">
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<emphasis id="7ABE156EFFBFFFECFF5D10CC5E419758" bold="true" box="[199,352,511,536]" pageId="17" pageNumber="18">Distribution.</emphasis>
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Currently known only from the
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<typeStatus id="977177DEFFBFFFECFD5613335DDD9758" box="[716,764,512,536]" pageId="17" pageNumber="18">type</typeStatus>
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locality (
|
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<figureCitation id="D0F1D5F9FFBFFFECFCFC10CC5CE89757" box="[870,969,511,535]" captionStart="FIGURES 193 – 194. 193" captionStartId="187.[151,264,1650,1673]" captionTargetBox="[282,1304,193,1628]" captionTargetId="figure@187.[282,1305,193,1629]" captionTargetPageId="187" captionText="FIGURES 193 – 194. 193. Geographical distributions of species in the Paeminosa Group. 194. Geographical distributions of species in the Curvosa and Multispina Groups." httpUri="https://zenodo.org/record/1050236/files/figure.png" pageId="17" pageNumber="18">Fig. 193</figureCitation>
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).
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</paragraph>
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<paragraph id="4875C97CFFBFFFECFF5D13175E67971F" blockId="17.[151,1436,151,607]" pageId="17" pageNumber="18">
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<emphasis id="7ABE156EFFBFFFECFF5D13175E27977C" bold="true" box="[199,262,548,572]" pageId="17" pageNumber="18">Note.</emphasis>
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My attempts to glue the elytra to the body, in a natural position, before imaging were not entirely successful (
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<figureCitation id="D0F1D5F9FFBFFFECFF6E13745E19971F" box="[244,312,583,607]" captionStart="FIGURE 7" captionStartId="76.[151,250,1650,1673]" captionTargetBox="[285,1308,218,1613]" captionTargetId="figure@76.[268,1318,193,1629]" captionTargetPageId="76" captionText="FIGURE 7. Hydraena punctilata, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050074/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
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||
</document> |