143 lines
17 KiB
XML
143 lines
17 KiB
XML
<document id="F7F8A7AE20497AEEA71E61CA65C70D75" ID-CLB-Dataset="31530" ID-DOI="10.5281/zenodo.1050060" ID-GBIF-Dataset="2d28bebf-e3c5-457b-9152-8a6877ded281" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510742923475" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="C063786AFF9AFFC9FF0D13595CF792F5" docLanguage="en" docName="zt03074p198.pdf.imf" docOrigin="Zootaxa 3074" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena umbolenta Perkins, 2011, new species" docType="treatment" docVersion="7" lastPageNumber="53" masterDocId="3C5A0012FFAEFFFDFF9A11335F219540" masterDocTitle="New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)" masterLastPageNumber="198" masterPageNumber="1" pageNumber="53" updateTime="1698485333395" updateUser="ExternalLinkService">
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<mods:mods id="7D6B708ED104466E319DB212A3405F4F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4623274334EF60A9DDAE337258FFCD52">
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<mods:title id="425D426399DC712D8B1A32DE0653220C">New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="0B54AEF221FEE265028DB7C4E989C110" type="personal">
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<mods:roleTerm id="3C0D99149B7D3D572DD8CFCE29945CCC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BFDA42D56399DEE7883078BDBB35A547">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="BDDA88C8CAC41BFFDBE6CF9834706CEF">text</mods:typeOfResource>
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<mods:title id="0D331BE3ACB9960E0DDC2282E5414E84">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="22A8EBC0067B8839D66B19CC9E826B11">
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<mods:date id="3413335A7D35C4A61EF1FD8F66D72530">2011</mods:date>
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<mods:detail id="436021C3AC70AF3525231A5E946E4D8B" type="pubDate">
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<mods:number id="3A30D3953EAC4EDEC6053D45AEF484C0">2011-10-28</mods:number>
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</mods:detail>
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<mods:detail id="963FC4D145292B38944E3EC51425446C" type="volume">
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<mods:number id="CCC5C2D604CC37237A93D7265B915E5C">3074</mods:number>
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<mods:end id="68AE6733EA2D247E99812ACEFEFEB1B5">198</mods:end>
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<mods:classification id="88888CA7B9E14260131FF5F54E445FE4">journal article</mods:classification>
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<mods:identifier id="44A08FD78390C6FCA334F48461422065" type="CLB-Dataset">31530</mods:identifier>
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<mods:identifier id="6C959270832FCC8E130F6B1F8D1E8A8F" type="DOI">10.5281/zenodo.1050060</mods:identifier>
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<mods:identifier id="D5CA14755F3E2E16101DE7AA0640DA71" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7D1E83D5A601F478E1E09E162214180D" type="Zenodo-Dep">1050060</mods:identifier>
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<treatment id="C063786AFF9AFFC9FF0D13595CF792F5" ID-DOI="http://doi.org/10.5281/zenodo.6184542" ID-GBIF-Taxon="137307024" ID-Zenodo-Dep="6184542" LSID="urn:lsid:plazi:treatment:C063786AFF9AFFC9FF0D13595CF792F5" httpUri="http://treatment.plazi.org/id/C063786AFF9AFFC9FF0D13595CF792F5" lastPageNumber="53" pageId="52" pageNumber="53">
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<subSubSection id="00D09AF7FF9AFFC9FF0D13595C5E97E5" pageId="52" pageNumber="53" type="nomenclature">
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<paragraph id="4875C97CFF9AFFC9FF0D13595D1497C4" blockId="52.[151,895,618,677]" box="[151,565,618,644]" pageId="52" pageNumber="53">
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<heading id="133D7E10FF9AFFC9FF0D13595D1497C4" bold="true" box="[151,565,618,644]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
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<emphasis id="7ABE156EFF9AFFC9FF0D13595D1497C4" bold="true" box="[151,565,618,644]" pageId="52" pageNumber="53">
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<taxonomicName id="8FCAB2FFFF9AFFC9FF0D13595EB797C4" ID-CoL="6MJ44" box="[151,406,618,644]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="umbolenta" status="sp. nov.">
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<emphasis id="7ABE156EFF9AFFC9FF0D13595EB797C4" bold="true" box="[151,406,618,644]" italics="true" pageId="52" pageNumber="53">Hydraena umbolenta</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="618DA815FF9AFFC9FE3C13595D1497C4" box="[422,565,618,644]" pageId="52" pageNumber="53" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4875C97CFF9AFFC9FF0D13BE5C5E97E5" blockId="52.[151,895,618,677]" box="[151,895,653,677]" pageId="52" pageNumber="53">
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<figureCitation id="D0F1D5F9FF9AFFC9FF0D13BE5FDE97E5" box="[151,255,653,677]" captionStart="FIGURE 108" captionStartId="146.[151,250,1650,1673]" captionTargetBox="[243,1340,193,1625]" captionTargetId="figure@146.[243,1343,193,1629]" captionTargetPageId="146" captionText="FIGURE 108. Hydraena umbolenta, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050180/files/figure.png" pageId="52" pageNumber="53">Figs. 108</figureCitation>
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(habitus), 111 (aedeagus), 184 (spermatheca), 202 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FF9AFFC9FF0D13E65D429685" pageId="52" pageNumber="53" type="materials_examined">
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<paragraph id="4875C97CFF9AFFC9FF0D13E65A2D9651" blockId="52.[151,1437,725,1974]" pageId="52" pageNumber="53">
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<emphasis id="7ABE156EFF9AFFC9FF0D13E65E6997AE" bold="true" box="[151,328,725,750]" pageId="52" pageNumber="53">
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<typeStatus id="977177DEFF9AFFC9FF0D13E65FF397AD" box="[151,210,725,749]" pageId="52" pageNumber="53">Type</typeStatus>
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Material.
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</emphasis>
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<typeStatus id="977177DEFF9AFFC9FED413E65E9797AD" box="[334,438,725,749]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
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(male):
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<collectingCountry id="30DD89ECFF9AFFC9FD8613E65DAE97AD" box="[540,655,725,749]" name="Paraguay" pageId="52" pageNumber="53">
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<emphasis id="7ABE156EFF9AFFC9FD8613E65DAE97AD" bold="true" box="[540,655,725,749]" pageId="52" pageNumber="53">Paraguay</emphasis>
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</collectingCountry>
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: Rio Tebicuarymi, shore washing,
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<geoCoordinate id="2DFEAFBBFF9AFFC9FB8413E65BA497AD" box="[1054,1157,725,749]" direction="south" orientation="latitude" pageId="52" pageNumber="53" precision="925" value="-26.416666">26° 25' S</geoCoordinate>
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,
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<geoCoordinate id="2DFEAFBBFF9AFFC9FB0813E65A2297AD" box="[1170,1283,725,749]" direction="west" orientation="longitude" pageId="52" pageNumber="53" precision="925" value="-56.833332">56° 50' W</geoCoordinate>
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,
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<date id="3C74EFBCFF9AFFC9FA9413E65A5897AD" box="[1294,1401,725,749]" pageId="52" pageNumber="53" value="1991-01-20">20 i 1991</date>
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, S. Endrody-Younga (PAR-248). Deposited in the
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<collectionCode id="2EDB51B9FF9AFFC9FD3E13CA5DD79651" box="[676,758,761,785]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="52" pageNumber="53">TMSA</collectionCode>
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.
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<typeStatus id="977177DEFF9AFFC9FD6513C95C569652" box="[767,887,762,786]" pageId="52" pageNumber="53" type="paratype">
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<emphasis id="7ABE156EFF9AFFC9FD6513C95C569652" bold="true" box="[767,887,762,786]" pageId="52" pageNumber="53">Paratypes</emphasis>
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</typeStatus>
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: Same data as
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<typeStatus id="977177DEFF9AFFC9FBBC13CA5BA99651" box="[1062,1160,761,785]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
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(9
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<collectionCode id="2EDB51B9FF9AFFC9FB3713CA5A219651" box="[1197,1280,761,785]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="52" pageNumber="53">TMSA</collectionCode>
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).
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</paragraph>
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<paragraph id="4875C97CFF9AFFC9FF5D122E5D429685" blockId="52.[151,1437,725,1974]" pageId="52" pageNumber="53">
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<emphasis id="7ABE156EFF9AFFC9FF5D122E5EF29676" bold="true" box="[199,467,797,822]" pageId="52" pageNumber="53">Differential Diagnosis.</emphasis>
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Dorsal habitus very similar to
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<taxonomicName id="8FCAB2FFFF9AFFC9FCAE122D5CF69675" box="[820,983,797,821]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kellymilleri">
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<emphasis id="7ABE156EFF9AFFC9FCAE122D5CF69675" box="[820,983,797,821]" italics="true" pageId="52" pageNumber="53">H. kellymilleri</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="8FCAB2FFFF9AFFC9FB8A122D5BA09675" box="[1040,1153,797,821]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="loripes">
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<emphasis id="7ABE156EFF9AFFC9FB8A122D5BA09675" box="[1040,1153,797,821]" italics="true" pageId="52" pageNumber="53">H. loripes</emphasis>
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</taxonomicName>
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(
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<figureCitation id="D0F1D5F9FF9AFFC9FB0B122E5A169675" box="[1169,1335,797,821]" captionStart-0="FIGURE 108" captionStart-1="FIGURE 109" captionStart-2="FIGURE 110" captionStartId-0="146.[151,250,1650,1673]" captionStartId-1="147.[151,250,1650,1673]" captionStartId-2="148.[151,250,1650,1673]" captionTargetBox-0="[243,1340,193,1625]" captionTargetBox-1="[229,1352,206,1617]" captionTargetBox-2="[255,1324,211,1618]" captionTargetId-0="figure@146.[243,1343,193,1629]" captionTargetId-1="figure@147.[220,1368,193,1629]" captionTargetId-2="figure@148.[245,1342,193,1629]" captionTargetPageId-0="146" captionTargetPageId-1="147" captionTargetPageId-2="148" captionText-0="FIGURE 108. Hydraena umbolenta, holotype, dorsal and lateral habitus, and venter." captionText-1="FIGURE 109. Hydraena kellymilleri, holotype, dorsal and lateral habitus, and venter." captionText-2="FIGURE 110. Hydraena loripes, holotype, dorsal and lateral habitus, and venter." httpUri-0="https://zenodo.org/record/1050180/files/figure.png" httpUri-1="https://zenodo.org/record/1050182/files/figure.png" httpUri-2="https://zenodo.org/record/1050184/files/figure.png" pageId="52" pageNumber="53">Figs. 108–110</figureCitation>
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). In general,
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<taxonomicName id="8FCAB2FFFF9AFFC9FF4B12715E4A9619" box="[209,363,833,857]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="umbolenta">
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<emphasis id="7ABE156EFF9AFFC9FF4B12715E4A9619" box="[209,363,833,857]" italics="true" pageId="52" pageNumber="53">H. umbolenta</emphasis>
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</taxonomicName>
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is more coarsely punctate dorsally than the compared species, but the pronotum lacks foveae PF2, which are present in the two other species. In addition, males are differentiated by the thickened protibiae, and females by the non-serrate last tergite. The aedeagi of the three species, while having distinctively different details, also show a relationship (
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<figureCitation id="D0F1D5F9FF9AFFC9FE2E129E5D769685" box="[436,599,941,965]" captionStart="FIGURES 111 – 113" captionStartId="149.[151,264,1650,1673]" captionText="FIGURES 111 – 113. Aedeagi of holotypes." pageId="52" pageNumber="53">Figs. 111–113</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FF9AFFC9FF5D12E25BA4922D" pageId="52" pageNumber="53" type="description">
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<paragraph id="4875C97CFF9AFFC9FF5D12E25E159139" blockId="52.[151,1437,725,1974]" pageId="52" pageNumber="53">
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<emphasis id="7ABE156EFF9AFFC9FF5D12E25E7096AA" bold="true" box="[199,337,977,1002]" pageId="52" pageNumber="53">Description</emphasis>
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. Size:
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<typeStatus id="977177DEFF9AFFC9FE3712E25D2E96A9" box="[429,527,977,1001]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.25/0.54; head 0.22/0.32; pronotum 0.30/0.40, PA 0.36, PB 0.39; elytra 0.77/0.54. Dorsum of head piceous; pronotum testaceous in front of and behind piceous fascia, ratios of color bands, as measured in midline, ca.
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<date id="3C74EFBCFF9AFFC9FB8A152A5B7B9171" box="[1040,1114,1049,1073]" pageId="52" pageNumber="53" value="2010-05-05">5/10/5</date>
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, fascia laterally smaller and more lightly colored; elytra brown, columellae darker than interstices; legs testaceous; maxillary palpi testaceous, tip not darker.
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</paragraph>
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<paragraph id="4875C97CFF9AFFC9FF5D15B55C819049" blockId="52.[151,1437,725,1974]" pageId="52" pageNumber="53">Frons punctures ca. 1xef, not larger and denser near eyes; interstices effacedly microreticulate, weakly shining, 0.5–1xpd. Clypeus dull, microreticulate, very finely sparsely punctulate medially. Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate in median concavity, surrounding areas smooth, shining. Genae raised, shining, without posterior ridge.</paragraph>
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<paragraph id="4875C97CFF9AFFC9FF5D14255CBA9035" blockId="52.[151,1437,725,1974]" pageId="52" pageNumber="53">Pronotum weakly arcuate laterally; anterior margin straight behind eyes, emarginate behind frons, scintilla absent; punctures on disc larger and deeper than those on frons, interstices shining, 0.5–2xpd, punctures larger and denser at anterior and posterior; PF1, PF2 and PF4 absent; PF3 deep.</paragraph>
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<paragraph id="4875C97CFF9AFFC9FF5D14B25C2B9369" blockId="52.[151,1437,725,1974]" pageId="52" pageNumber="53">Elytra weakly arcuate laterally; summit of posterior declivity at ca. midlength; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming slightly smaller and more widely spaced toward posterior. Intervals not raised, shining, on basal 1/3 ca. 0.5–1xpd, as are interstices between punctures of a row, a few punctures subserial. Apices in dorsal aspect weakly separately rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
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<paragraph id="4875C97CFF9AFFC9FF5D17065BA4922D" blockId="52.[151,1437,725,1974]" pageId="52" pageNumber="53">
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Ratios of P2 width and plaque shape (P2/w/l/s) ca. 2.1/1/5/4. P1 laminate; median carina sinuate in profile. P2 l/w ca. 4/2.5, sides parallel, apex blunt, raised slightly above mesoventral intercoxal process. Plaques very narrow, subcarinate lines, straight, slightly converging toward one another anteriorly, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 1.5x P2. Protibia arcuate, moderately thick. Mesotibia straight. Metatibia arcuate, slender, very slightly widened subapically.
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<collectingCountry id="30DD89ECFF9AFFC9FC3317DA5CC29241" box="[937,995,1769,1793]" name="Malta" pageId="52" pageNumber="53">Male</collectingCountry>
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abdominal apex symmetrical; last tergite with apicomedian notch. Female abdominal apex: last tergite broadly rounded, without apicomedian notch, ca. 45 (n=1) hooked setae, tips sharply pointed; gonocoxite not midlongitudinally divided, apical margin sharply rounded, no transverse ridge evident at border of microreticulation. Spermatheca
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<typeStatus id="977177DEFF9AFFC9FBBC16655B77922E" box="[1062,1110,1878,1902]" pageId="52" pageNumber="53">type</typeStatus>
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C1.
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FF9AFFC9FF5D16495A7C92D1" box="[199,1373,1913,1938]" pageId="52" pageNumber="53" type="etymology">
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<paragraph id="4875C97CFF9AFFC9FF5D16495A7C92D1" blockId="52.[151,1437,725,1974]" box="[199,1373,1913,1938]" pageId="52" pageNumber="53">
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<emphasis id="7ABE156EFF9AFFC9FF5D16495E6F92D2" bold="true" box="[199,334,1914,1938]" pageId="52" pageNumber="53">Etymology.</emphasis>
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Named in reference to the many small bumps on the dorsal face of the aedeagal distal piece.
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FF9AFFC9FF5D16AE5CF792F5" box="[199,982,1949,1974]" pageId="52" pageNumber="53" type="distribution">
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||
<paragraph id="4875C97CFF9AFFC9FF5D16AE5CF792F5" blockId="52.[151,1437,725,1974]" box="[199,982,1949,1974]" pageId="52" pageNumber="53">
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||
<emphasis id="7ABE156EFF9AFFC9FF5D16AE5E4192F6" bold="true" box="[199,352,1949,1974]" pageId="52" pageNumber="53">Distribution.</emphasis>
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Currently known only from the
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<typeStatus id="977177DEFF9AFFC9FD5616AD5DDD92F6" box="[716,764,1950,1974]" pageId="52" pageNumber="53">type</typeStatus>
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locality (
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<figureCitation id="D0F1D5F9FF9AFFC9FCFC16AE5CE892F5" box="[870,969,1949,1973]" captionStart="FIGURE 202" captionStartId="195.[151,250,1650,1673]" captionText="FIGURE 202. Geographical distributions of Hydraena species." pageId="52" pageNumber="53">Fig. 202</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |