155 lines
18 KiB
XML
155 lines
18 KiB
XML
<document id="C221E02C0448D0662D3DC25987A668A0" ID-CLB-Dataset="31530" ID-DOI="10.5281/zenodo.1050060" ID-GBIF-Dataset="2d28bebf-e3c5-457b-9152-8a6877ded281" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510742923475" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="C063786AFF83FFD3FF0D144D5B829677" docLanguage="en" docName="zt03074p198.pdf.imf" docOrigin="Zootaxa 3074" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena pantanalensis Perkins, 2011, new species" docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="3C5A0012FFAEFFFDFF9A11335F219540" masterDocTitle="New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)" masterLastPageNumber="198" masterPageNumber="1" pageNumber="46" updateTime="1698485333395" updateUser="ExternalLinkService">
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<mods:title id="A29A3F4F8E37B635393CCCE586D2DDA7">New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)</mods:title>
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<mods:name id="56E631761D2BBD7BAAD09EC9C23ACEA3" type="personal">
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<mods:namePart id="0E03060C3DFC3A2DCA5D822A714DFB31">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="C342EC2D7605EBBBA828A9499C64FB8A">text</mods:typeOfResource>
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<mods:title id="70461ADBE8D7E165223DAFB8788F5BD2">Zootaxa</mods:title>
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<mods:part id="3E09D9A84FA528CF715DF8B2D5CFE3BC">
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<mods:date id="3872D29FFC5DC2577565494BA6275B83">2011</mods:date>
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<mods:detail id="3E743EB9A42BB57E74EDB4D89FD3E8F5" type="pubDate">
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<mods:number id="3107B203B278D136FCC04EE5B296DD9B">2011-10-28</mods:number>
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<mods:detail id="7C5FB0968035B15FB5E598CFFBFCE5BF" type="volume">
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<mods:number id="1442D0685490CA97677A8BDB05FE4424">3074</mods:number>
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<mods:classification id="D91015816CB4DCDA9C00599D13DBA0EA">journal article</mods:classification>
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<mods:identifier id="EEA2B33EE03326FC75B283663589F648" type="CLB-Dataset">31530</mods:identifier>
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<mods:identifier id="4FFBC8C1C822175F523C5CD855E30CC8" type="DOI">10.5281/zenodo.1050060</mods:identifier>
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<mods:identifier id="A58C480DA77459984C910CF945C481D3" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D834D45E4F32F82127A84D961155214E" type="Zenodo-Dep">1050060</mods:identifier>
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<treatment id="C063786AFF83FFD3FF0D144D5B829677" ID-DOI="http://doi.org/10.5281/zenodo.6184512" ID-GBIF-Taxon="137306989" ID-Zenodo-Dep="6184512" LSID="urn:lsid:plazi:treatment:C063786AFF83FFD3FF0D144D5B829677" httpUri="http://treatment.plazi.org/id/C063786AFF83FFD3FF0D144D5B829677" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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<subSubSection id="00D09AF7FF83FFD0FF0D144D5DA990F9" pageId="45" pageNumber="46" type="nomenclature">
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<paragraph id="4875C97CFF83FFD0FF0D144D5D7D90D8" blockId="45.[151,648,1406,1465]" box="[151,604,1406,1432]" pageId="45" pageNumber="46">
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<heading id="133D7E10FF83FFD0FF0D144D5D7D90D8" bold="true" box="[151,604,1406,1432]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
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<emphasis id="7ABE156EFF83FFD0FF0D144D5D7D90D8" bold="true" box="[151,604,1406,1432]" pageId="45" pageNumber="46">
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<taxonomicName id="8FCAB2FFFF83FFD0FF0D144D5E9C90D8" ID-CoL="6MJC3" box="[151,445,1406,1432]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pantanalensis" status="sp. nov.">
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<emphasis id="7ABE156EFF83FFD0FF0D144D5E9C90D8" bold="true" box="[151,445,1406,1432]" italics="true" pageId="45" pageNumber="46">Hydraena pantanalensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="618DA815FF83FFD0FE57144D5D7D90D8" box="[461,604,1406,1432]" pageId="45" pageNumber="46" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4875C97CFF83FFD0FF0D14925DA990F9" blockId="45.[151,648,1406,1465]" box="[151,648,1441,1465]" pageId="45" pageNumber="46">
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<figureCitation id="D0F1D5F9FF83FFD0FF0D14925FD090F9" box="[151,241,1441,1465]" captionStart="FIGURE 85" captionStartId="132.[151,250,1650,1673]" captionTargetBox="[222,1365,193,1626]" captionTargetId="figure@132.[222,1365,193,1629]" captionTargetPageId="132" captionText="FIGURE 85. Hydraena pantanalensis, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050156/files/figure.png" pageId="45" pageNumber="46">Figs. 85</figureCitation>
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(habitus), 89 (aedeagus), 200 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FF83FFD0FF0D14D75B189394" pageId="45" pageNumber="46" type="materials_examined">
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<paragraph id="4875C97CFF83FFD0FF0D14D75BA3935F" blockId="45.[151,1437,1507,2036]" pageId="45" pageNumber="46">
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<emphasis id="7ABE156EFF83FFD0FF0D14D75E6990BC" bold="true" box="[151,328,1507,1532]" pageId="45" pageNumber="46">
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<typeStatus id="977177DEFF83FFD0FF0D14D75FF390BC" box="[151,210,1508,1532]" pageId="45" pageNumber="46">Type</typeStatus>
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Material.
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</emphasis>
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<typeStatus id="977177DEFF83FFD0FED414D75E9790BC" box="[334,438,1508,1532]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
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(male):
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<collectingCountry id="30DD89ECFF83FFD0FD8614D05D4490BC" box="[540,613,1507,1532]" name="Brazil" pageId="45" pageNumber="46">
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<emphasis id="7ABE156EFF83FFD0FD8614D05D4490BC" bold="true" box="[540,613,1507,1532]" pageId="45" pageNumber="46">Brazil</emphasis>
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</collectingCountry>
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: Matto Grosso, Rio Bento Gomes (Pantanal); Quelle, Quellbach, Campo Allegre, II,
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<geoCoordinate id="2DFEAFBBFF83FFD0FE8317345EA0935F" box="[281,385,1543,1567]" direction="south" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="-15.75">15° 45' S</geoCoordinate>
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,
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<geoCoordinate id="2DFEAFBBFF83FFD0FE1717345EDE935F" box="[397,511,1543,1567]" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-56.55">56° 33' W</geoCoordinate>
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,
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<date id="3C74EFBCFF83FFD0FD9317345D44935F" box="[521,613,1543,1567]" pageId="45" pageNumber="46" value="1993-01-01">1 i 1993</date>
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–
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<date id="3C74EFBCFF83FFD0FD1B17345C25935F" box="[641,772,1543,1567]" pageId="45" pageNumber="46" value="1994-12-30">30 xii 1994</date>
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, E. Stuhr. Deposited in the
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<collectionCode id="2EDB51B9FF83FFD0FBAF173B5B5F935F" box="[1077,1150,1544,1567]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="45" pageNumber="46">NMW</collectionCode>
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.
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</paragraph>
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<paragraph id="4875C97CFF83FFD0FF5D17185B189394" blockId="45.[151,1437,1507,2036]" pageId="45" pageNumber="46">
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<emphasis id="7ABE156EFF83FFD0FF5D17185EF39304" bold="true" box="[199,466,1579,1604]" pageId="45" pageNumber="46">Differential Diagnosis.</emphasis>
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This species has rather large and dense elytral punctures, the interstices only 0.5–1xpd on the basal 1/3; the elytra are rather convex at the summit of the posterior declivity. Similar in dorsal habitus to
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<taxonomicName id="8FCAB2FFFF83FFD0FA1B17625E3B93CC" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="tucumanica">
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<emphasis id="7ABE156EFF83FFD0FA1B17625E3B93CC" italics="true" pageId="45" pageNumber="46">H. tucumanica</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="8FCAB2FFFF83FFD0FEC817465EE393CC" box="[338,450,1652,1676]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="diffusa">
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<emphasis id="7ABE156EFF83FFD0FEC817465EE393CC" box="[338,450,1652,1676]" italics="true" pageId="45" pageNumber="46">H. diffusa</emphasis>
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</taxonomicName>
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(
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<figureCitation id="D0F1D5F9FF83FFD0FE4817475D1093CC" box="[466,561,1652,1676]" captionStart="FIGURE 74" captionStartId="124.[151,250,1607,1630]" captionTargetBox="[226,1357,193,1585]" captionTargetId="figure@124.[226,1360,193,1586]" captionTargetPageId="124" captionText="FIGURE 74. Hydraena tucumanica, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050142/files/figure.png" pageId="45" pageNumber="46">Figs. 74</figureCitation>
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,
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<figureCitation id="D0F1D5F9FF83FFD0FDA717475D7A93CC" box="[573,603,1652,1676]" captionStart="FIGURE 85" captionStartId="132.[151,250,1650,1673]" captionTargetBox="[222,1365,193,1626]" captionTargetId="figure@132.[222,1365,193,1629]" captionTargetPageId="132" captionText="FIGURE 85. Hydraena pantanalensis, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050156/files/figure.png" pageId="45" pageNumber="46">85</figureCitation>
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,
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<figureCitation id="D0F1D5F9FF83FFD0FDF217475DB793CC" box="[616,662,1652,1676]" captionStart="FIGURE 139" captionStartId="169.[151,250,1650,1673]" captionTargetBox="[256,1331,202,1616]" captionTargetId="figure@169.[247,1340,193,1629]" captionTargetPageId="169" captionText="FIGURE 139. Hydraena diffusa, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050214/files/figure.png" pageId="45" pageNumber="46">139</figureCitation>
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); differing from both species by the smaller size (ca.
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<quantity id="8F326499FF83FFD0FB6E17475A7C93CC" box="[1268,1373,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="45" pageNumber="46" unit="mm" value="1.22">1.22 mm</quantity>
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). The aedeagus has some resemblance in basic shape to that of
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<taxonomicName id="8FCAB2FFFF83FFD0FC8717AA5CA993F0" box="[797,904,1688,1712]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="orcula">
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<emphasis id="7ABE156EFF83FFD0FC8717AA5CA993F0" box="[797,904,1688,1712]" italics="true" pageId="45" pageNumber="46">H. orcula</emphasis>
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</taxonomicName>
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. Reliable determinations will require examination of the very dissimilar male genitalia of the four species (
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<figureCitation id="D0F1D5F9FF83FFD0FCDB178F5CBE9394" box="[833,927,1724,1748]" captionStart="FIGURES 78 – 81" captionStartId="128.[151,264,1650,1673]" captionText="FIGURES 78 – 81. Aedeagi of holotypes." pageId="45" pageNumber="46">Figs. 78</figureCitation>
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,
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<figureCitation id="D0F1D5F9FF83FFD0FC31178F5CD29394" box="[939,1011,1724,1748]" captionStart-0="FIGURE 84" captionStart-1="FIGURE 85" captionStartId-0="131.[151,250,1650,1673]" captionStartId-1="132.[151,250,1650,1673]" captionTargetBox-0="[258,1339,222,1606]" captionTargetBox-1="[222,1365,193,1626]" captionTargetId-0="figure@131.[227,1360,193,1629]" captionTargetId-1="figure@132.[222,1365,193,1629]" captionTargetPageId-0="131" captionTargetPageId-1="132" captionText-0="FIGURE 84. Hydraena orcula, holotype, dorsal and lateral habitus, and venter." captionText-1="FIGURE 85. Hydraena pantanalensis, holotype, dorsal and lateral habitus, and venter." httpUri-0="https://zenodo.org/record/1050154/files/figure.png" httpUri-1="https://zenodo.org/record/1050156/files/figure.png" pageId="45" pageNumber="46">84–85</figureCitation>
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,
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<figureCitation id="D0F1D5F9FF83FFD0FC64178F5B0A9394" box="[1022,1067,1724,1748]" captionStart="FIGURES 140 – 141" captionStartId="170.[151,264,1650,1673]" captionText="FIGURES 140 – 141. Aedeagi of holotypes." pageId="45" pageNumber="46">141</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FF83FFD3FF5D17EC5D7A97E7" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
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<paragraph id="4875C97CFF83FFD0FF5D17EC5ABC9224" blockId="45.[151,1437,1507,2036]" pageId="45" pageNumber="46">
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<emphasis id="7ABE156EFF83FFD0FF5D17EC5E7093B8" bold="true" box="[199,337,1759,1784]" pageId="45" pageNumber="46">Description</emphasis>
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. Size:
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<typeStatus id="977177DEFF83FFD0FE3717EC5D2E93B7" box="[429,527,1759,1783]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.22/0.49; head 0.19/0.31; pronotum 0.29/0.39, PA 0.33, PB 0.36; elytra 0.74/0.49. The
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<typeStatus id="977177DEFF83FFD0FCCF16375C96925C" box="[853,951,1796,1820]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
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is teneral; following is a prediction of the mature color. Dorsum of head piceous; pronotum testaceous in front of and behind piceous macula, ratios of color bands, as measured in midline, ca.
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<date id="3C74EFBCFF83FFD0FD86167F5D479224" box="[540,614,1868,1892]" pageId="45" pageNumber="46" value="2010-05-05">5/10/5</date>
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; elytra dark brown; legs brown; maxillary palpi testaceous, tip not darker.
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</paragraph>
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<paragraph id="4875C97CFF83FFD0FF5D16425A5692B4" blockId="45.[151,1437,1507,2036]" pageId="45" pageNumber="46">Frons punctures ca. 1–2xef, larger and denser near eyes than medially; interstices 0.5–1xpd, microreticulate, dull laterally, 1–3xpd and shining medially. Clypeus microreticulate laterally, very finely sparsely punctulate medially. Mentum microreticulate, dull, very sparsely very finely punctulate; postmentum dull, very finely densely micropunctulate. Genae concave and dull laterally, raised and weakly shining medially, without posterior ridge.</paragraph>
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<paragraph id="4875C97CFF80FFD3FF5D11AB5BF095B7" blockId="46.[151,1437,151,824]" pageId="46" pageNumber="47">Pronotum weakly arcuate laterally; anterior margin straight behind eyes, emarginate behind frons, scintilla absent; punctures on disc slightly larger and deeper than those of frons, interstices shining, 1–3xpd, punctures larger and denser at anterior and posterior; PF1 absent; PF2 very shallow; PF3 deep; PF4 absent.</paragraph>
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<paragraph id="4875C97CFF80FFD3FF5D10375DEA94EC" blockId="46.[151,1437,151,824]" pageId="46" pageNumber="47">Elytra weakly arcuate laterally; summit of posterior declivity at ca. posterior 1/3; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming finer and more widely spaced toward posterior. Intervals not raised, shining, on basal 1/3 ca. 0.5–1xpd, as are interstices between punctures of a row, a few punctures subserial. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very shallow angle with one another.</paragraph>
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<paragraph id="4875C97CFF80FFD3FF5D10845D7A97E7" blockId="46.[151,1437,151,824]" pageId="46" pageNumber="47">
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Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1.5/1.5/5/3. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2.5/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately wide, straight, parallel or slightly converging toward one another anteriorly, weakly raised, located at sides of deep median depression. Metaventrite with very small point on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca.
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<date id="3C74EFBCFF80FFD3FB1C13745B83971F" box="[1158,1186,583,607]" pageId="46" pageNumber="47">2x</date>
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P2. All tibiae slender, straight. Abdominal apex symmetrical; last tergite concave on each side of median longitudinal ridge, free margin sharply rounded. Female not yet known.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="00D09AF7FF80FFD3FF5D13875CBF978C" box="[199,926,692,716]" pageId="46" pageNumber="47" type="etymology">
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<paragraph id="4875C97CFF80FFD3FF5D13875CBF978C" blockId="46.[151,1437,151,824]" box="[199,926,692,716]" pageId="46" pageNumber="47">
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||
<emphasis id="7ABE156EFF80FFD3FF5D13875E6F978C" bold="true" box="[199,334,692,716]" pageId="46" pageNumber="47">Etymology.</emphasis>
|
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Named in reference to the geographical distribution.
|
||
</paragraph>
|
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</subSubSection>
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<subSubSection id="00D09AF7FF80FFD3FF5D13E45B829677" pageId="46" pageNumber="47" type="distribution">
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<paragraph id="4875C97CFF80FFD3FF5D13E45CF797AF" blockId="46.[151,1437,151,824]" box="[199,982,727,752]" pageId="46" pageNumber="47">
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<emphasis id="7ABE156EFF80FFD3FF5D13E45E4197B0" bold="true" box="[199,352,727,752]" pageId="46" pageNumber="47">Distribution.</emphasis>
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Currently known only from the
|
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<typeStatus id="977177DEFF80FFD3FD5613EB5DDD97B0" box="[716,764,728,752]" pageId="46" pageNumber="47">type</typeStatus>
|
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locality (
|
||
<figureCitation id="D0F1D5F9FF80FFD3FCFC13E45CEB97AF" box="[870,970,727,751]" captionStart="FIGURE 200" captionStartId="193.[151,250,1650,1673]" captionText="FIGURE 200. Geographical distributions of Hydraena species." pageId="46" pageNumber="47">Fig. 200</figureCitation>
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||
).
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</paragraph>
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<paragraph id="4875C97CFF80FFD3FF5D13CF5B829677" blockId="46.[151,1437,151,824]" pageId="46" pageNumber="47">
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<emphasis id="7ABE156EFF80FFD3FF5D13CF5E309654" bold="true" box="[199,273,764,788]" pageId="46" pageNumber="47">Notes.</emphasis>
|
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The
|
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<typeStatus id="977177DEFF80FFD3FED413CF5E919654" box="[334,432,764,788]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
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has a very narrow transverse groove across the posterior margin of the frons (
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<figureCitation id="D0F1D5F9FF80FFD3FAA213CF5AAF9654" box="[1336,1422,764,788]" captionStart="FIGURE 85" captionStartId="132.[151,250,1650,1673]" captionTargetBox="[222,1365,193,1626]" captionTargetId="figure@132.[222,1365,193,1629]" captionTargetPageId="132" captionText="FIGURE 85. Hydraena pantanalensis, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050156/files/figure.png" pageId="46" pageNumber="47">Fig. 85</figureCitation>
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); more specimens are needed to verify if this is constant in the species, or a variable character.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |