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<document ID-DOI="10.11646/phytotaxa.567.2.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="7141712" checkinTime="1664861116526" checkinUser="plazi" docAuthor="Vieira-Uribe, Sebastián, Moreno, Juan Sebastián &amp; Espinosa, Felipe" docDate="2022" docId="377F8B2DD214FFCAFF27F193FE98FB31" docLanguage="en" docName="phytotaxa.567.2.3.pdf" docOrigin="Phytotaxa 567 (2)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Lepanthes hwangiae J. S. Moreno &amp; S. Vieira-Uribe 2022, sp. nov." docType="treatment" docVersion="1" lastPageNumber="159" masterDocId="CB46F355D213FFC0FFAFF61CFF8BFF99" masterDocTitle="Two new species of Lepanthes (Pleurothallidinae, Orchidaceae) from the Alto de Ventanas, Colombia" masterLastPageNumber="161" masterPageNumber="149" pageNumber="156" updateTime="1664906361310" updateUser="jonas" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of Lepanthes (Pleurothallidinae, Orchidaceae) from the Alto de Ventanas, Colombia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vieira-Uribe, Sebastián</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4325-1232</mods:nameIdentifier>
<mods:affiliation>Sociedad Colombiana de Orquideología, Carrera 52 # 73 - 298, Medellín, Colombia &amp; Grupo de Investigación en Biodiversidad Tropical - GIBIOT, Jardín Botánico de Medellín, Calle 73 # 51 D- 14, Medellín, Colombia &amp; Grupo de investigación Schultes, Fundación Ecotonos, Carrera 72 # 13 A- 56, Cali, Colombia &amp; utricseb @ gmail. com; https: // orcid. org / 0000 - 0003 - 4325 - 1232</mods:affiliation>
<mods:nameIdentifier type="email">utricseb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moreno, Juan Sebastián</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0875-9498</mods:nameIdentifier>
<mods:affiliation>Departamento de Biología, Universidad del Valle, Calle 13 # 100 - 00, Cali, Colombia &amp; Grupo de investigación Schultes, Fundación Ecotonos, Carrera 72 # 13 A- 56, Cali, Colombia &amp; semoreno 113 @ gmail. com; https: // orcid. org / 0000 - 0002 - 0875 - 9498</mods:affiliation>
<mods:nameIdentifier type="email">semoreno113@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Espinosa, Felipe</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-5932-7380</mods:nameIdentifier>
<mods:affiliation>Grupo de investigación Schultes, Fundación Ecotonos, Carrera 72 # 13 A- 56, Cali, Colombia &amp; felipeespinosam @ gmail. com; https: // orcid. org / 0000 - 0001 - 5932 - 7380</mods:affiliation>
<mods:nameIdentifier type="email">felipeespinosam@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-10-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>567</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>149</mods:start>
<mods:end>161</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.567.2.3</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7141712</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:377F8B2DD214FFCAFF27F193FE98FB31" httpUri="http://treatment.plazi.org/id/377F8B2DD214FFCAFF27F193FE98FB31" lastPageId="10" lastPageNumber="159" pageId="7" pageNumber="156">
<subSubSection box="[136,988,1934,1961]" pageId="7" pageNumber="156" type="nomenclature">
<paragraph blockId="7.[136,988,1934,1961]" box="[136,988,1934,1961]" pageId="7" pageNumber="156">
<taxonomicName authority="J.S.Moreno &amp; S.Vieira-Uribe" authorityName="J. S. Moreno &amp; S. Vieira-Uribe" authorityYear="2022" box="[136,706,1935,1961]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="156" phylum="Tracheophyta" rank="species" species="hwangiae" status="sp. nov.">
<emphasis box="[136,367,1935,1961]" italics="true" pageId="7" pageNumber="156">Lepanthes hwangiae</emphasis>
J.S.Moreno &amp; S.Vieira-Uribe
</taxonomicName>
,
<emphasis box="[717,802,1935,1960]" italics="true" pageId="7" pageNumber="156">
<taxonomicNameLabel box="[717,802,1935,1960]" pageId="7" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[816,924,1935,1961]" captionStart="FIGURE 4" captionStartId="6.[136,229,2014,2036]" captionTargetBox="[232,1394,439,1967]" captionTargetId="figure-153@6.[191,1397,425,1986]" captionTargetPageId="6" captionText="FIGURE 4. Drawing of Lepanthes hwangiae. A. Habit. B. Flower. C. Dissected perianth. D. Ovary, column and lip, lateral view.E. Ovary and column, dorsal and lateral views. F. Anther cap and pollinarium. Drawn by J.S.Moreno from the plant that served as type." figureDoi="http://doi.org/10.5281/zenodo.7141722" httpUri="https://zenodo.org/record/7141722/files/figure.png" pageId="7" pageNumber="156">Figures 4</figureCitation>
,
<figureCitation box="[936,952,1934,1961]" captionStart="FIGURE 5" captionStartId="7.[136,229,1794,1816]" captionTargetBox="[183,1404,190,1770]" captionTargetId="figure-18@7.[183,1404,190,1770]" captionTargetPageId="7" captionText="FIGURE 5. Lankester composite dissection plate (LCDP) of Lepanthes hwangiae. A. Habit. B. Flower. C. Dissected perianth, lip in expanded and natural position. D. Ovary, column and lip, lateral view. E. Ovary and column, dorsal and lateral views. F. Anther cap and pollinarium. Photographed by S.Vieira-Uribe and assembled by J.S.Moreno from the plant that served as type." figureDoi="http://doi.org/10.5281/zenodo.7141724" httpUri="https://zenodo.org/record/7141724/files/figure.png" pageId="7" pageNumber="156">5</figureCitation>
,
<figureCitation box="[964,979,1935,1961]" captionStart="FIGURE 7" captionStartId="9.[136,229,1794,1816]" captionTargetBox="[183,1404,190,1770]" captionTargetId="figure-18@9.[183,1404,190,1770]" captionTargetPageId="9" captionText="FIGURE 7. Side by side comparison of Lepanthes cyclochila and L.hwangiae. A. Flowers, 1. L. hwangiae, 2. L.cyclochila. B. Ovary, column and lip, lateral view. 1. L. hwangiae, 2. L.cyclochila. C. Expanded lip. 1. L. hwangiae, 2. L.cyclochila. D. Petal. 1. L. hwangiae, 2. L.cyclochila. Photographed by S.Vieira-Uribe and assembled by J.S.Moreno." figureDoi="http://doi.org/10.5281/zenodo.7141728" httpUri="https://zenodo.org/record/7141728/files/figure.png" pageId="7" pageNumber="156">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="156" type="materials_examined">
<paragraph blockId="7.[136,1452,2010,2068]" pageId="7" pageNumber="156">
<emphasis bold="true" box="[136,194,2010,2031]" pageId="7" pageNumber="156">Type:</emphasis>
<materialsCitation collectingDate="2022-01-31" collectorName="S. Vieira" country="Colombia" county="Municipio de Valdivia" elevation="2160" location="Cloud forest" municipality="Reserva Natural La Esperanza" pageId="7" pageNumber="156" specimenCount="1" stateProvince="Antioquia" typeStatus="holotype">
<collectingCountry box="[219,349,2010,2031]" name="Colombia" pageId="7" pageNumber="156">COLOMBIA</collectingCountry>
.
<collectingRegion box="[358,458,2010,2032]" country="Colombia" name="Antioquia" pageId="7" pageNumber="156">Antioquia</collectingRegion>
:
<collectingCounty box="[467,686,2010,2032]" pageId="7" pageNumber="156">Municipio de Valdivia</collectingCounty>
,
<collectingMunicipality box="[695,991,2010,2032]" pageId="7" pageNumber="156">Reserva Natural La Esperanza</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:377F8B2DD214FFCAFF27F193FE98FB31:BA096CE0D214FFC7FC46F1C6FBEFF869" box="[1001,1124,2010,2032]" country="Colombia" county="Municipio de Valdivia" municipality="Reserva Natural La Esperanza" name="Cloud forest" pageId="7" pageNumber="156" stateProvince="Antioquia">Cloud forest</location>
, epiphyte in thin branches next to the entrance to the reserve,
<quantity box="[453,528,2046,2068]" metricMagnitude="3" metricUnit="m" metricValue="2.16" pageId="7" pageNumber="156" unit="m" value="2160.0">
<elevation box="[453,528,2046,2068]" metricMagnitude="3" metricUnit="m" metricValue="2.16" pageId="7" pageNumber="156" unit="m" value="2160.0">2160 m</elevation>
</quantity>
,
<date box="[538,696,2046,2068]" pageId="7" pageNumber="156" value="2022-01-31">
<collectingDate box="[538,696,2046,2068]" pageId="7" pageNumber="156" value="2022-01-31">31 january 2022</collectingDate>
</date>
,
<emphasis box="[706,812,2046,2068]" italics="true" pageId="7" pageNumber="156">
<collectorName box="[706,782,2047,2068]" pageId="7" pageNumber="156">S.Vieira</collectorName>
30
</emphasis>
(
<typeStatus box="[825,914,2046,2068]" pageId="7" pageNumber="156">holotype</typeStatus>
: JAUM-Spirit!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="157" type="diagnosis">
<paragraph blockId="8.[136,1452,159,397]" pageId="8" pageNumber="157">
<taxonomicName authorityName="J. S. Moreno &amp; S. Vieira-Uribe" authorityYear="2022" box="[136,334,159,180]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="157" phylum="Tracheophyta" rank="species" species="hwangiae">
<emphasis box="[136,334,159,180]" italics="true" pageId="8" pageNumber="157">Lepanthes hwangiae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName authority="Luer &amp; R. Escobar ex Viveros &amp; W. E. Higgins (2007: 34)" authorityName="Luer &amp; R. Escobar ex Viveros &amp; W. E. Higgins" authorityPageNumber="34" authorityYear="2007" box="[511,1190,159,181]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="157" phylum="Tracheophyta" rank="species" species="cyclochila">
<emphasis box="[511,635,159,181]" italics="true" pageId="8" pageNumber="157">L. cyclochila</emphasis>
Luer &amp; R.Escobar ex Viveros &amp; W.E. Higgins (2007: 34)
</taxonomicName>
, but it can be distinguished by its transversally bilobed petals with the upper lobe bifurcate in unequal lobes, with the external lobe triangular, obtuse and the internal lobe oblong, obtuse (
<emphasis box="[474,501,232,253]" italics="true" pageId="8" pageNumber="157">vs.</emphasis>
the upper lobe obliquely truncate) and the lower lobe oblong and rounded (
<emphasis box="[1237,1257,232,253]" italics="true" pageId="8" pageNumber="157">vs</emphasis>
. lower lobe oblong, shallowly bifurcate into 2 triangular, narrowly rounded lobes) and the lip sub-circular, slightly emarginate, with the oblong and rounded basal lobes erect, embracing the column and touching each other well above the column (
<emphasis box="[1186,1206,304,325]" italics="true" pageId="8" pageNumber="157">vs</emphasis>
. sub-quadrate, markedly emarginate, with the shortly triangular and obtuse basal lobes of the lip erect on each side of the column without touching each other). (Figures 6,7).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7141726" ID-Zenodo-Dep="7141726" httpUri="https://zenodo.org/record/7141726/files/figure.png" pageId="8" pageNumber="157" startId="8.[136,229,1973,1995]" targetBox="[151,1436,423,1949]" targetPageId="8">
<paragraph blockId="8.[136,1452,1973,2068]" pageId="8" pageNumber="157">
<emphasis bold="true" box="[136,256,1973,1995]" pageId="8" pageNumber="157">FIGURE 6.</emphasis>
Lankester composite dissection plate (LCDP) of
<emphasis box="[752,966,1974,1995]" italics="true" pageId="8" pageNumber="157">
<taxonomicName authorityName="Luer &amp; R. Escobar ex Viveros &amp; W. E. Higgins" authorityYear="2007" box="[752,962,1974,1995]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="157" phylum="Tracheophyta" rank="species" species="cyclochila">Lepanthes cyclochila</taxonomicName>
.
</emphasis>
<emphasis bold="true" box="[973,997,1973,1994]" pageId="8" pageNumber="157">A.</emphasis>
Habit.
<emphasis bold="true" box="[1074,1096,1974,1995]" pageId="8" pageNumber="157">B.</emphasis>
Flower.
<emphasis bold="true" box="[1187,1210,1973,1995]" pageId="8" pageNumber="157">C.</emphasis>
Dissected perianth with lip expanded.
<emphasis bold="true" box="[276,300,2010,2031]" pageId="8" pageNumber="157">D.</emphasis>
Ovary, column and lip, lateral and dorsal view.
<emphasis bold="true" box="[782,805,2010,2031]" pageId="8" pageNumber="157">E.</emphasis>
Pollinarium and anther cap. Photographed by S.Vieira-Uribe and assembled by J.S.Moreno.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7141728" ID-Zenodo-Dep="7141728" httpUri="https://zenodo.org/record/7141728/files/figure.png" pageId="9" pageNumber="158" startId="9.[136,229,1794,1816]" targetBox="[183,1404,190,1770]" targetPageId="9">
<paragraph blockId="9.[136,1452,1794,1888]" pageId="9" pageNumber="158">
<emphasis bold="true" box="[136,255,1794,1816]" pageId="9" pageNumber="158">FIGURE 7.</emphasis>
Side by side comparison of
<taxonomicName authorityName="Luer &amp; R. Escobar ex Viveros &amp; W. E. Higgins" authorityYear="2007" box="[538,743,1794,1815]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="158" phylum="Tracheophyta" rank="species" species="cyclochila">
<emphasis box="[538,743,1794,1815]" italics="true" pageId="9" pageNumber="158">Lepanthes cyclochila</emphasis>
</taxonomicName>
and
<emphasis box="[793,912,1794,1815]" italics="true" pageId="9" pageNumber="158">
<taxonomicName authorityName="J. S. Moreno &amp; S. Vieira-Uribe" authorityYear="2022" box="[793,908,1794,1815]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="158" phylum="Tracheophyta" rank="species" species="hwangiae">L.hwangiae</taxonomicName>
.
</emphasis>
<emphasis bold="true" box="[919,943,1794,1815]" pageId="9" pageNumber="158">A.</emphasis>
Flowers, 1.
<emphasis box="[1066,1193,1794,1816]" italics="true" pageId="9" pageNumber="158">
<taxonomicName authorityName="J. S. Moreno &amp; S. Vieira-Uribe" authorityYear="2022" box="[1066,1189,1794,1816]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="158" phylum="Tracheophyta" rank="species" species="hwangiae">L. hwangiae</taxonomicName>
,
</emphasis>
2.
<emphasis box="[1226,1351,1794,1815]" italics="true" pageId="9" pageNumber="158">
<taxonomicName authorityName="Luer &amp; R. Escobar ex Viveros &amp; W. E. Higgins" authorityYear="2007" box="[1226,1347,1794,1815]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="158" phylum="Tracheophyta" rank="species" species="cyclochila">L.cyclochila</taxonomicName>
.
</emphasis>
<emphasis bold="true" box="[1357,1380,1794,1815]" pageId="9" pageNumber="158">B.</emphasis>
Ovary, column and lip, lateral view. 1.
<emphasis box="[436,561,1830,1852]" italics="true" pageId="9" pageNumber="158">
<taxonomicName authorityName="J. S. Moreno &amp; S. Vieira-Uribe" authorityYear="2022" box="[436,557,1830,1852]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="158" phylum="Tracheophyta" rank="species" species="hwangiae">L. hwangiae</taxonomicName>
,
</emphasis>
2.
<emphasis box="[589,714,1830,1851]" italics="true" pageId="9" pageNumber="158">
<taxonomicName authorityName="Luer &amp; R. Escobar ex Viveros &amp; W. E. Higgins" authorityYear="2007" box="[589,710,1830,1851]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="158" phylum="Tracheophyta" rank="species" species="cyclochila">L.cyclochila</taxonomicName>
.
</emphasis>
<emphasis bold="true" box="[719,742,1830,1852]" pageId="9" pageNumber="158">C.</emphasis>
Expanded lip. 1.
<emphasis box="[909,1034,1830,1852]" italics="true" pageId="9" pageNumber="158">
<taxonomicName authorityName="J. S. Moreno &amp; S. Vieira-Uribe" authorityYear="2022" box="[909,1030,1830,1852]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="158" phylum="Tracheophyta" rank="species" species="hwangiae">L. hwangiae</taxonomicName>
,
</emphasis>
2.
<emphasis box="[1062,1187,1830,1851]" italics="true" pageId="9" pageNumber="158">
<taxonomicName authorityName="Luer &amp; R. Escobar ex Viveros &amp; W. E. Higgins" authorityYear="2007" box="[1062,1184,1830,1851]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="158" phylum="Tracheophyta" rank="species" species="cyclochila">L.cyclochila</taxonomicName>
.
</emphasis>
<emphasis bold="true" box="[1192,1216,1830,1851]" pageId="9" pageNumber="158">D.</emphasis>
Petal. 1.
<emphasis box="[1304,1428,1830,1852]" italics="true" pageId="9" pageNumber="158">
<taxonomicName authorityName="J. S. Moreno &amp; S. Vieira-Uribe" authorityYear="2022" box="[1304,1425,1830,1852]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="158" phylum="Tracheophyta" rank="species" species="hwangiae">L. hwangiae</taxonomicName>
,
</emphasis>
2.
<emphasis box="[136,261,1866,1887]" italics="true" pageId="9" pageNumber="158">
<taxonomicName authorityName="Luer &amp; R. Escobar ex Viveros &amp; W. E. Higgins" authorityYear="2007" box="[136,257,1866,1887]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="158" phylum="Tracheophyta" rank="species" species="cyclochila">L.cyclochila</taxonomicName>
.
</emphasis>
Photographed by S.Vieira-Uribe and assembled by J.S.Moreno.
</paragraph>
</caption>
<subSubSection lastPageId="10" lastPageNumber="159" pageId="9" pageNumber="158" type="description">
<paragraph blockId="9.[136,1452,1934,2069]" lastBlockId="10.[136,1452,158,1192]" lastPageId="10" lastPageNumber="159" pageId="9" pageNumber="158">
<emphasis bold="true" box="[136,312,1934,1960]" pageId="9" pageNumber="158">Description:—</emphasis>
<emphasis box="[312,373,1935,1961]" italics="true" pageId="9" pageNumber="158">Plant</emphasis>
, epiphytic, caespitose, up to
<quantity box="[711,769,1935,1961]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="158" unit="cm" value="3.0">3 cm</quantity>
tall.
<emphasis box="[831,895,1935,1960]" italics="true" pageId="9" pageNumber="158">Roots</emphasis>
slender, flexuous, filiform,
<quantity box="[1218,1307,1935,1961]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="158" unit="mm" value="0.6">0.6 mm</quantity>
in diameter.
<emphasis box="[136,254,1971,1997]" italics="true" pageId="9" pageNumber="158">Ramicauls</emphasis>
slender, suberect,
<quantity box="[478,608,1971,1997]" metricMagnitude="-2" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="2.4" metricValueMin="1.2" pageId="9" pageNumber="158" unit="cm" value="1.7999999999999998" valueMax="2.4" valueMin="1.2">1.22.4 cm</quantity>
long, enclosed by 79 acuminate, ribbed, microscopically ciliate along the ribs lepanthiform sheaths, with ciliate dilated ostia.
<emphasis box="[783,861,2007,2032]" italics="true" pageId="9" pageNumber="158">Leaves</emphasis>
abaxially purple, elliptical, the apex acute, incised and tri-apiculate, 1.21.7 ×
<quantity box="[458,589,2042,2069]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="9" pageNumber="158" unit="cm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 cm</quantity>
, the cuneate base contracted into a
<emphasis box="[1005,1031,2043,2068]" italics="true" pageId="9" pageNumber="158">ca</emphasis>
.
<quantity box="[1049,1117,2043,2069]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="158" unit="mm" value="1.0">1 mm</quantity>
long petiole.
<emphasis box="[1279,1429,2043,2069]" italics="true" pageId="9" pageNumber="158">Inflorescence</emphasis>
a congested, distichous raceme successively many-flowered, up to 1/2 the length of the leaf,
<quantity box="[1156,1281,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="10" pageNumber="159" unit="cm" value="0.75" valueMax="0.9" valueMin="0.6">0.60.9 cm</quantity>
long including the peduncle, held on the abaxial surface of the leaf by a filiform, terete peduncle up to
<quantity box="[1106,1171,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="159" unit="mm" value="5.0">5 mm</quantity>
long borne near the apex of the ramicaul.
<emphasis box="[320,469,230,256]" italics="true" pageId="10" pageNumber="159">Floral bracts</emphasis>
conical, ciliate, acuminate, up to
<quantity box="[846,932,230,257]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="159" unit="mm" value="0.7">0.7 mm</quantity>
long;
<emphasis box="[1004,1096,230,256]" italics="true" pageId="10" pageNumber="159">pedicels</emphasis>
terete, verrucose, up to
<quantity box="[1366,1452,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="159" unit="mm" value="1.5">1.5 mm</quantity>
long.
<emphasis box="[202,272,267,293]" italics="true" pageId="10" pageNumber="159">Ovary</emphasis>
terete, costate, verrucose along the ribs, ca.
<quantity box="[787,876,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="159" unit="mm" value="1.1">1.1 mm</quantity>
long.
<emphasis box="[951,1043,266,292]" italics="true" pageId="10" pageNumber="159">Flowers</emphasis>
with the sepals translucent saffron; petals saffron with the base suffused with crimson; the lip magenta with fulvous-saffron margins, the column pale gray with the apex magenta.
<emphasis box="[403,546,338,364]" italics="true" pageId="10" pageNumber="159">Dorsal sepal</emphasis>
elliptic, with papillose margins towards the apex, acuminate, 3-veined, 2.3 ×
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="159" unit="mm" value="1.4">1.4 mm</quantity>
, connate at the base to the lateral sepals for
<emphasis box="[696,722,375,400]" italics="true" pageId="10" pageNumber="159">ca</emphasis>
.
<quantity box="[739,844,374,401]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="159" unit="mm" value="0.25">0.25 mm</quantity>
.
<emphasis box="[857,1018,374,400]" italics="true" pageId="10" pageNumber="159">Lateral sepals</emphasis>
ovate-elliptic, oblique, with papillose margins towards the apex, the apex acute, sub-acuminate, 2-veined, 1.92.0 ×
<quantity box="[1038,1177,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="10" pageNumber="159" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
, connate at the base for ca.
<quantity box="[175,279,446,473]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="159" unit="mm" value="0.25">0.25 mm</quantity>
.
<emphasis box="[290,360,446,472]" italics="true" pageId="10" pageNumber="159">Petals</emphasis>
transversally bilobed, microscopically pubescent, 0.8 ×
<quantity box="[994,1082,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="159" unit="mm" value="3.8">3.8 mm</quantity>
, 1 veined; the upper lobe oblong bifurcate in unequal lobes, the external lobe triangular, obtuse, the internal lobe oblong, obtuse,
<quantity box="[1251,1340,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="159" unit="mm" value="1.7">1.7 mm</quantity>
long; the lower lobe oblong, rounded, 2.0 mm long.
<emphasis box="[621,658,519,544]" italics="true" pageId="10" pageNumber="159">Lip</emphasis>
microscopically pubescent, sub-circular, bilobed, with the oblong and rounded basal lobes erect, embracing the column and touching each other well above the column, the apex rounded, emarginate, with a small apiculum in the sinus, 3-veined, 2.1 × 2.0 mm expanded, the base adnate to the base of the column.
<emphasis box="[234,323,626,652]" italics="true" pageId="10" pageNumber="159">Column</emphasis>
conical, terete,
<emphasis box="[500,533,627,652]" italics="true" pageId="10" pageNumber="159">ca.</emphasis>
<quantity box="[540,626,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="159" unit="mm" value="1.2">1.2 mm</quantity>
long including the rostellum, the anther dorsal, the stigma ventral.
<emphasis italics="true" pageId="10" pageNumber="159">Anther cap</emphasis>
purple, cordate, cucullate,
<quantity box="[479,565,662,689]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="159" unit="mm" value="0.5">0.5 mm</quantity>
long.
<taxonomicName box="[634,721,662,688]" class="Liliopsida" family="Poaceae" genus="Pollinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="10" pageNumber="151" phylum="Tracheophyta" rank="genus">Pollinia</taxonomicName>
2, yellow, narrowly obovoid,
<emphasis box="[1057,1083,663,688]" italics="true" pageId="10" pageNumber="159">ca</emphasis>
.
<quantity box="[1097,1183,662,689]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="159" unit="mm" value="0.6">0.6 mm</quantity>
long, attached to a drop like viscidium.
<emphasis box="[309,401,698,724]" italics="true" pageId="10" pageNumber="159">Capsule</emphasis>
not seen.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="159" type="distribution">
<paragraph blockId="10.[136,1452,158,1192]" pageId="10" pageNumber="159">
<emphasis bold="true" box="[189,488,734,760]" pageId="10" pageNumber="159">Distribution and ecology</emphasis>
:—
<taxonomicName authorityName="J. S. Moreno &amp; S. Vieira-Uribe" authorityYear="2022" box="[523,757,734,760]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="10" pageNumber="159" phylum="Tracheophyta" rank="species" species="hwangiae">
<emphasis box="[523,757,734,760]" italics="true" pageId="10" pageNumber="159">Lepanthes hwangiae</emphasis>
</taxonomicName>
is known from two localities in the Alto de Ventanas region in the north of the central Andes of
<collectingCountry box="[536,649,770,796]" name="Colombia" pageId="10" pageNumber="159">Colombia</collectingCountry>
, just a few kilometers apart and with the same elevation. It grows as an epiphyte in thin, moss-covered branches no more than
<quantity box="[757,843,806,833]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="159" unit="m" value="1.0">1 meter</quantity>
from the ground, in partially shaded and humid areas inside primary and secondary cloud forests.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="159" type="etymology">
<paragraph blockId="10.[136,1452,158,1192]" pageId="10" pageNumber="159">
<emphasis bold="true" box="[189,318,878,904]" pageId="10" pageNumber="159">Etymology</emphasis>
:—Named to honor Joanne Hwang Pemberton, wife of Robert W. Pemberton, generous donor to the expansion of the natural reserve that protects the habitat of the new species.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="159" type="discussion">
<paragraph blockId="10.[136,1452,158,1192]" pageId="10" pageNumber="159">
<emphasis bold="true" box="[189,458,950,976]" pageId="10" pageNumber="159">Taxonomic Discussion</emphasis>
:—
<taxonomicName authorityName="J. S. Moreno &amp; S. Vieira-Uribe" authorityYear="2022" box="[494,730,950,976]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="10" pageNumber="159" phylum="Tracheophyta" rank="species" species="hwangiae">
<emphasis box="[494,730,950,976]" italics="true" pageId="10" pageNumber="159">Lepanthes hwangiae</emphasis>
</taxonomicName>
is also similar to
<taxonomicName authorityName="Luer &amp; R. Escobar" authorityYear="1994" box="[953,1127,950,976]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="10" pageNumber="159" phylum="Tracheophyta" rank="species" species="petalopteryx">
<emphasis box="[953,1127,950,976]" italics="true" pageId="10" pageNumber="159">L. petalopteryx</emphasis>
</taxonomicName>
(
<figureCitation box="[1146,1268,950,976]" captionStart="FIGURE 3" captionStartId="5.[136,229,1794,1816]" captionTargetBox="[183,1404,190,1770]" captionTargetId="figure-18@5.[183,1404,190,1770]" captionTargetPageId="5" captionText="FIGURE 3. Comparison of the flower of Lepanthes pembertonii with similar species. A. L. anchicayae. B. L. pembertonii. C. L. petalopteryx. D. L. viahoensis. Photographs by J.S. Moreno (A) and S. Vieira-Uribe (B,C,D)." figureDoi="http://doi.org/10.5281/zenodo.7141720" httpUri="https://zenodo.org/record/7141720/files/figure.png" pageId="10" pageNumber="159">Figure 3C</figureCitation>
), but this latter species could be recognized by the petals with both lobes similar, obliquely subquadrate or oblong, oblique, with the inner angles shortly acuminate-obtuse, the outer angles subacute, the outer angles obtusely angled at the center (
<emphasis box="[1381,1404,1023,1048]" italics="true" pageId="10" pageNumber="159">vs</emphasis>
. the upper lobe oblong bifurcate in unequal lobes, the external lobe triangular, obtuse, the internal lobe oblong, obtuse and the lower lobe oblong, rounded). Finally, the lip in
<taxonomicName authorityName="Luer &amp; R. Escobar" authorityYear="1994" box="[710,880,1094,1120]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="10" pageNumber="159" phylum="Tracheophyta" rank="species" species="petalopteryx">
<emphasis box="[710,880,1094,1120]" italics="true" pageId="10" pageNumber="159">L. petalopteryx</emphasis>
</taxonomicName>
is reniform with the lobes surrounding the column without touching each other (
<emphasis box="[467,490,1131,1156]" italics="true" pageId="10" pageNumber="159">vs</emphasis>
. sub-circular, embracing the column and touching each other well above the column) (
<figureCitation box="[144,262,1166,1192]" captionStart="FIGURE 3" captionStartId="5.[136,229,1794,1816]" captionTargetBox="[183,1404,190,1770]" captionTargetId="figure-18@5.[183,1404,190,1770]" captionTargetPageId="5" captionText="FIGURE 3. Comparison of the flower of Lepanthes pembertonii with similar species. A. L. anchicayae. B. L. pembertonii. C. L. petalopteryx. D. L. viahoensis. Photographs by J.S. Moreno (A) and S. Vieira-Uribe (B,C,D)." figureDoi="http://doi.org/10.5281/zenodo.7141720" httpUri="https://zenodo.org/record/7141720/files/figure.png" pageId="10" pageNumber="159">Figure 3C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>