treatments-xml/data/D1/FC/D9/D1FCD9D1AC07541DB881DFE02A1C1BD7.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1012.57047" ID-GBIF-Dataset="96b35ac5-ed73-465c-9e88-76f169714a83" ID-PMC="PMC7854560" ID-Pensoft-Pub="1313-2970-1012-21" ID-Pensoft-UUID="1AC67CC9000A5FCF86DECCE264464FD8" ID-PubMed="33584107" ID-ZooBank="4B5ACAB053224893BC53B4A48F8DC20C" ModsDocID="1313-2970-1012-21" checkinTime="1611749546682" checkinUser="pensoft" docAuthor="Rivera-Quiroz, Francisco Andres, Petcharad, Booppa &amp; Miller, Jeremy A." docDate="2021" docId="D1FCD9D1AC07541DB881DFE02A1C1BD7" docLanguage="en" docName="ZooKeys 1012: 21-53" docOrigin="ZooKeys 1012" docPubDate="2021-01-26" docSource="http://dx.doi.org/10.3897/zookeys.1012.57047" docTitle="Patu shiluensis Lin &amp; Li 2009" docType="treatment" docVersion="4" id="1AC67CC9000A5FCF86DECCE264464FD8" lastPageNumber="21" masterDocId="1AC67CC9000A5FCF86DECCE264464FD8" masterDocTitle="First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995" masterLastPageNumber="53" masterPageNumber="21" pageNumber="21" updateTime="1668149733872" updateUser="ExternalLinkService">
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<mods:title>First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rivera-Quiroz, Francisco Andres</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8748-4930</mods:nameIdentifier>
<mods:affiliation>Department of Terrestrial Zoology, Understanding Evolution group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands &amp; Institute for Biology Leiden (IBL), Leiden University, Sylviusweg 72, 2333 BE Leiden, the Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">andres.riveraquiroz@naturalis.nl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Petcharad, Booppa</mods:namePart>
<mods:affiliation>Institute for Biology Leiden (IBL), Leiden University, Sylviusweg 72, 2333 BE Leiden, the Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Jeremy A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8918-9775</mods:nameIdentifier>
<mods:affiliation>Department of Terrestrial Zoology, Understanding Evolution group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands</mods:affiliation>
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<mods:date>2021</mods:date>
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<mods:number>2021-01-26</mods:number>
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<mods:number>1012</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1012.57047</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1012.57047</mods:identifier>
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<treatment ID-GBIF-Taxon="177132022" LSID="urn:lsid:plazi:treatment:D1FCD9D1AC07541DB881DFE02A1C1BD7" httpUri="http://treatment.plazi.org/id/D1FCD9D1AC07541DB881DFE02A1C1BD7" lastPageNumber="21" pageId="0" pageNumber="21">
<subSubSection pageId="0" pageNumber="21" type="nomenclature">
<paragraph pageId="0" pageNumber="21">
<taxonomicName LSID="D1FCD9D1-AC07-541D-B881-DFE02A1C1BD7" authority="Lin &amp; Li, 2009" authorityName="Lin &amp; Li" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu shiluensis" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="shiluensis">Patu shiluensis Lin &amp; Li, 2009</taxonomicName>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">Figures 14</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. a, c Patu shiluensis Lin &amp; Li, 2009 b, d Crassignatha seedam sp. nov. Epigynum, cleared: a, b dorsal view c, d ventral view. Scale bars: 0.03 mm (a, c); 0.05 mm (b, d)." figureDoi="10.3897/zookeys.1012.57047.figure15" httpUri="https://binary.pensoft.net/fig/501623" pageId="0" pageNumber="21">, 15a, c</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="reference_group">
<paragraph pageId="0" pageNumber="21">
<taxonomicName authorityName="Lin &amp; Li" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu shiluensis" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="shiluensis">Patu shiluensis</taxonomicName>
Lin &amp; Li, 2009: 59, figs 11A, B, 12A, B, 13A-D.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="materials_examined">
<paragraph pageId="0" pageNumber="21">Collected material.</paragraph>
<paragraph pageId="0" pageNumber="21">
Thailand • 4♀; Phuket Province, Siray Island. Mixed tropical forest; 
<geoCoordinate degrees="7" direction="north" minutes="53.355" orientation="latitude" precision="1" value="7.88925">7°53.355'N</geoCoordinate>
, 
<geoCoordinate degrees="98" direction="east" minutes="26.083" orientation="longitude" precision="1" value="98.434715">98°26.083'E</geoCoordinate>
. 132 m; August 02-06, 2018; Booppa Petcharad, Jeremy Miller, F. Andres Rivera-Quiroz leg.; Winkler extractor; RMNH.5106642.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="distribution">
<paragraph pageId="0" pageNumber="21">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="21">Known only from its type locality, Shilu Town, Hainan Province, China and the specimens collected for the present work.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="morphological remarks">
<paragraph pageId="0" pageNumber="21">Morphological remarks.</paragraph>
<paragraph pageId="0" pageNumber="21">
Carapace pale yellow with black margin, smooth texture (Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14b</figureCitation>
). Legs black and semi-transparent. Abdomen oval, longer than wide (Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14a, b</figureCitation>
). Ventrally same color as carapace, dorsally, darker with pale yellow patches.
</paragraph>
<caption doi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" pageId="0" pageNumber="21">Figure 14.</emphasis>
<taxonomicName authorityName="Lin &amp; Li" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu shiluensis" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="shiluensis">
<emphasis italics="true" pageId="0" pageNumber="21">Patu shiluensis</emphasis>
</taxonomicName>
Lin &amp; Li, 2009 female: Habitus: 
<emphasis bold="true" pageId="0" pageNumber="21">a</emphasis>
ventral view 
<emphasis bold="true" pageId="0" pageNumber="21">b</emphasis>
dorsal view. Epigynum: 
<emphasis bold="true" pageId="0" pageNumber="21">c</emphasis>
ventral view 
<emphasis bold="true" pageId="0" pageNumber="21">d</emphasis>
dorsal view, cleared. Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="21">a, b</emphasis>
); 0.06 mm (
<emphasis bold="true" pageId="0" pageNumber="21">c</emphasis>
); 0.03 mm (
<emphasis bold="true" pageId="0" pageNumber="21">d</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Vulva</emphasis>
: Epigynum weakly sclerotized, semi-transparent (Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14c</figureCitation>
). Atrium semi-circular slightly wider than inner distance between S (Figs 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14c</figureCitation>
, 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. a, c Patu shiluensis Lin &amp; Li, 2009 b, d Crassignatha seedam sp. nov. Epigynum, cleared: a, b dorsal view c, d ventral view. Scale bars: 0.03 mm (a, c); 0.05 mm (b, d)." figureDoi="10.3897/zookeys.1012.57047.figure15" httpUri="https://binary.pensoft.net/fig/501623" pageId="0" pageNumber="21">15c</figureCitation>
). Spermathecae spherical slightly more sclerotized than epigynum, separated by 0.5 
<normalizedToken originalValue="×">x</normalizedToken>
their diameter (Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14d</figureCitation>
). Copulatory ducts spring-like, spiraling 3 
<normalizedToken originalValue="×">x</normalizedToken>
over themselves. Fertilization ducts as long as S width, projecting posteriorly (Figs 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14d</figureCitation>
, 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. a, c Patu shiluensis Lin &amp; Li, 2009 b, d Crassignatha seedam sp. nov. Epigynum, cleared: a, b dorsal view c, d ventral view. Scale bars: 0.03 mm (a, c); 0.05 mm (b, d)." figureDoi="10.3897/zookeys.1012.57047.figure15" httpUri="https://binary.pensoft.net/fig/501623" pageId="0" pageNumber="21">15a, c</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1012.57047.figure15" httpUri="https://binary.pensoft.net/fig/501623" pageId="0" pageNumber="21" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" pageId="0" pageNumber="21">Figure 15.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="21">a, c</emphasis>
<taxonomicName authorityName="Lin &amp; Li" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu shiluensis" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="shiluensis">
<emphasis italics="true" pageId="0" pageNumber="21">Patu shiluensis</emphasis>
</taxonomicName>
Lin &amp; Li, 2009 
<emphasis bold="true" pageId="0" pageNumber="21">b, d</emphasis>
<taxonomicName authorityName="Rivera-Quiroz &amp; Petcharad &amp; Miller" authorityYear="2021" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha seedam" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="seedam">
<emphasis italics="true" pageId="0" pageNumber="21">Crassignatha seedam</emphasis>
</taxonomicName>
sp. nov. Epigynum, cleared: 
<emphasis bold="true" pageId="0" pageNumber="21">a, b</emphasis>
dorsal view 
<emphasis bold="true" pageId="0" pageNumber="21">c, d</emphasis>
ventral view. Scale bars: 0.03 mm (
<emphasis bold="true" pageId="0" pageNumber="21">a, c</emphasis>
); 0.05 mm (
<emphasis bold="true" pageId="0" pageNumber="21">b, d</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" pageId="0" pageNumber="21">Female</emphasis>
: Total length 0.52, carapace 0.21 long, 0.2 wide; clypeus 0.04; Chelicera 0.07 long, 0.05 wide; Leg I: femur 0.15, patella 0.07, tibia 0.1, metatarsus 0.07, tarsus 0.1; leg formula I-II-IV-III; abdomen 0.34 long, 0.28 wide.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="notes">
<paragraph pageId="0" pageNumber="21">Notes.</paragraph>
<paragraph pageId="0" pageNumber="21">
Small somatic variations can be seen between the specimen we collected in Thailand and the ones previously described from China (compare Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14b</figureCitation>
to 
<bibRefCitation author="Lin, Y" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="47 - 68" refId="B23" refString="Lin, Y, Li, S, 2009. First described Patu spiders (Araneae, Symphytognathidae) from Asia. Zootaxa 2154: 47 - 68" title="First described Patu spiders (Araneae, Symphytognathidae) from Asia." volume="2154" year="2009">Lin and Li 2009</bibRefCitation>
: fig. 11). However, we did not find any objective differences in the female genitalia.
</paragraph>
<paragraph pageId="0" pageNumber="21">
Secretory ampullae (Figs 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14d</figureCitation>
, 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. a, c Patu shiluensis Lin &amp; Li, 2009 b, d Crassignatha seedam sp. nov. Epigynum, cleared: a, b dorsal view c, d ventral view. Scale bars: 0.03 mm (a, c); 0.05 mm (b, d)." figureDoi="10.3897/zookeys.1012.57047.figure15" httpUri="https://binary.pensoft.net/fig/501623" pageId="0" pageNumber="21">15a</figureCitation>
) were very evident in our specimens; these glandular structures might be homologous to the accessory glands in 
<bibRefCitation DOI="https://doi.org/10.1111/zoj.12199" author="Lopardo, L" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="21" pagination="527 - 786" refId="B27" refString="Lopardo, L, Hormiga, G, 2015. Out of the twilight zone: Phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea). Zoological Journal of the Linnean Society 173 (3): 527 - 786, DOI: https://doi.org/10.1111/zoj.12199" title="Out of the twilight zone: Phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea)." url="https://doi.org/10.1111/zoj.12199" volume="173" year="2015">Lopardo and Hormiga (2015)</bibRefCitation>
. These structures were found in one anapid (
<taxonomicName authorityName="Kobayashi" authorityYear="1940" class="Trilobita" family="Hystricuridae" genus="Tasmanaspis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tasmanaspis" order="Aulacopleurida" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Tasmanaspis</emphasis>
</taxonomicName>
) and several mysmenids, but scored as absent or unknown for all the symphytognathids.
</paragraph>
<paragraph pageId="0" pageNumber="21">
The authors of this species mentioned it to be close to 
<taxonomicName authorityName="Saaristo" authorityYear="1996" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu silho" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="silho">
<emphasis italics="true" pageId="0" pageNumber="21">Patu silho</emphasis>
</taxonomicName>
Saaristo, 1996 from Seychelles. The possibility of 
<taxonomicName lsidName="P. silho" pageId="0" pageNumber="21" rank="species" species="silho">
<emphasis italics="true" pageId="0" pageNumber="21">P. silho</emphasis>
</taxonomicName>
not being a true 
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Patu</emphasis>
</taxonomicName>
was discussed by its author (
<bibRefCitation author="Saaristo, MI" journalOrPublisher="Phelsuma" pageId="0" pageNumber="21" pagination="53 - 56" refId="B43" refString="Saaristo, MI, 1996. Symphytognathidae (Arachnida, Araneae), a new spider family for the granitic islands of Seychelles. Phelsuma 4: 53 - 56" title="Symphytognathidae (Arachnida, Araneae), a new spider family for the granitic islands of Seychelles." volume="4" year="1996">Saaristo 1996</bibRefCitation>
; 
<bibRefCitation author="Saaristo, MI" journalOrPublisher="Press Manchester, UK" pageId="0" pageNumber="21" refId="B44" refString="Saaristo, MI, 2010. Araneae. In: Gerlach J, Marusik Y (Eds) Arachnida and Myriapoda of the Seychelles islands. Press Manchester, UK" title="Araneae. In: Gerlach J, Marusik Y (Eds) Arachnida and Myriapoda of the Seychelles islands." year="2010">2010</bibRefCitation>
) mentioning evident differences on somatic and sexual characters between 
<taxonomicName lsidName="P. silho" pageId="0" pageNumber="21" rank="species" species="silho">
<emphasis italics="true" pageId="0" pageNumber="21">P. silho</emphasis>
</taxonomicName>
and other 
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Patu</emphasis>
</taxonomicName>
species. Nevertheless, the author deemed appropriate to place it in this genus. We also consider this species might be misplaced in 
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Patu</emphasis>
</taxonomicName>
but would need further and more detailed analysis out of the scope of this work to clarify it (see discussion on 
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Patu</emphasis>
</taxonomicName>
relationships below).
</paragraph>
</subSubSection>
</treatment>
</document>