286 lines
31 KiB
XML
286 lines
31 KiB
XML
<document id="E18EA9A9D411E998A35FB5791277E7AF" ID-DOI="10.5281/zenodo.199052" ID-GBIF-Dataset="a07f4b3c-e602-4b4e-95dc-87fef597d912" ID-ISSN="1175-5326" ID-Zenodo-Dep="199052" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460321171440" checkinUser="plazi" docAuthor="Park, Jong-Seok, Carlton, Christopher E. & Ferro, Michael L." docDate="2010" docId="03E2BD70FFFE8066FF5C295DFAE14D9D" docLanguage="en" docName="zt02662p027.pdf" docOrigin="Zootaxa 2662" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptusa (Adoxopisalia) smokyiensis Pace 1989" docType="treatment" docVersion="10" lastPageNumber="8" masterDocId="FFDBC508FFF98061FFCB282AFF954A52" masterDocTitle="Diversity and taxonomic review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species" masterLastPageNumber="27" masterPageNumber="1" pageNumber="8" updateTime="1698256666968" updateUser="plazi">
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<mods:title id="AB829BF438139644F73C5A8DC80D30DD">Diversity and taxonomic review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species</mods:title>
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<mods:namePart id="6EF937BA18795B205D9083698DB4973B">Park, Jong-Seok</mods:namePart>
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<mods:namePart id="0CF2EB7E57980473F22CF0B119DBAF2E">Carlton, Christopher E.</mods:namePart>
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<mods:namePart id="FA7F64558FF5ED7F7ADAA006AFE40784">Ferro, Michael L.</mods:namePart>
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<mods:typeOfResource id="731815C029F572B35867FDB6143A760F">text</mods:typeOfResource>
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<mods:title id="576CEB3F07F6184573F9FFA8C1501E3C">Zootaxa</mods:title>
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<mods:part id="003683240E848A7C027E67093F61F995">
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<mods:date id="A99E8F61AEB0C912E5A3300FCDD2844E">2010</mods:date>
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<mods:detail id="1EE2C1B5774EC28B7EFD9F75003F7334" type="volume">
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<mods:number id="1B52F53B3587F8ECB5332DABC2094008">2662</mods:number>
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<mods:identifier id="19CC1AB7EC0619265EF2E4ACE4A09CC2" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E2BD70FFFE8066FF5C295DFAE14D9D" ID-DOI="http://doi.org/10.5281/zenodo.5020583" ID-GBIF-Taxon="119402516" ID-Zenodo-Dep="5020583" LSID="urn:lsid:plazi:treatment:03E2BD70FFFE8066FF5C295DFAE14D9D" httpUri="http://treatment.plazi.org/id/03E2BD70FFFE8066FF5C295DFAE14D9D" lastPageNumber="8" pageId="7" pageNumber="8">
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<subSubSection id="C3515FEDFFFE8066FF5C295DFE3C4BE6" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8BF40C66FFFE8066FF5C295DFD474BC3" blockId="7.[151,722,375,436]" box="[151,722,375,401]" pageId="7" pageNumber="8">
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<heading id="D0BCBB0AFFFE8066FF5C295DFD474BC3" bold="true" box="[151,722,375,401]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<taxonomicName id="4C4B77E5FFFE8066FF5C295DFD474BC3" ID-CoL="6PPBZ" authority="Pace, 1989" authorityName="Pace" authorityYear="1989" box="[151,722,375,401]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="smokyiensis" subGenus="Adoxopisalia">
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<emphasis id="B93FD074FFFE8066FF5C295DFD474BC3" bold="true" box="[151,722,375,401]" pageId="7" pageNumber="8">
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<emphasis id="B93FD074FFFE8066FF5C295DFF624BC3" bold="true" box="[151,247,375,401]" italics="true" pageId="7" pageNumber="8">Leptusa</emphasis>
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(
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<emphasis id="B93FD074FFFE8066FEC3295DFE334BC3" bold="true" box="[264,422,375,401]" italics="true" pageId="7" pageNumber="8">Adoxopisalia</emphasis>
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)
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<emphasis id="B93FD074FFFE8066FE7C295DFDDD4BC3" bold="true" box="[439,584,375,401]" italics="true" pageId="7" pageNumber="8">smokyiensis</emphasis>
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<bibRefCitation id="EFDA7197FFFE8066FD84295DFD474BC3" author="Pace" box="[591,722,375,401]" pageId="7" pageNumber="8" refString="Pace, R. (1989) Monografia del genere Leptusa Kraatz (Coleoptera, Staphylinidae). Memorie del Museo Civico di Storia Naturale di Verona (II Serie), Sezione Scienze della vita (A: Biologica), 8, 1 - 307." type="journal article" year="1989">Pace, 1989</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BF40C66FFFE8066FF5C29B0FE3C4BE6" blockId="7.[151,722,375,436]" box="[151,425,410,436]" pageId="7" pageNumber="8">
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(
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<figureCitation id="137010E3FFFE8066FF5429B0FF7B4BE6" box="[159,238,410,436]" captionStart="FIGURE 1" captionStartId="3.[151,255,1170,1194]" captionTargetBox="[153,1432,194,1147]" captionTargetId="figure@3.[151,1436,194,1149]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Leptusa. a: Leptusa (Adoxopisalia) opaca Casey; b: L. (A.) pseudosmokyiensis sp. nov.; c: L. (A.) smokyiensis Pace; d: L. (A.) stocksi sp. nov.; e: L. (Dysleptusa) carolinensis Pace; f: L. (D.) gimmeli sp. nov.; g: L. (D.) pusio (Casey); h: L. (Eucryptusa) brevicollis Casey; i: L. (E.) ferroi sp. nov.; j: L. (Ulitusa) cribratula (Casey). Scale bars = 1 mm." httpUri="https://zenodo.org/record/199053/files/figure.png" pageId="7" pageNumber="8">Figs. 1</figureCitation>
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c, 4a
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<emphasis id="B93FD074FFFE8066FEE329B0FEA34BE1" box="[296,310,410,435]" italics="true" pageId="7" pageNumber="8">–</emphasis>
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h, Map 1)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3515FEDFFFE8066FF5C29CFFB9B4BAE" box="[151,1038,485,508]" pageId="7" pageNumber="8" type="reference_group">
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||
<paragraph id="8BF40C66FFFE8066FF5C29CFFB9B4BAE" blockId="7.[151,1038,485,508]" box="[151,1038,485,508]" pageId="7" pageNumber="8">
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<treatmentCitationGroup id="AB5B2B48FFFE8066FF5C29CFFB9B4BAE" box="[151,1038,485,508]" pageId="7" pageNumber="8">
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<taxonomicName id="4C4B77E5FFFE8066FF5C29CFFD5E4BAE" ID-CoL="6PPBZ" authority="Pace, 1989: 252" authorityName="Pace" authorityPageNumber="252" authorityYear="1989" box="[151,715,485,508]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="smokyiensis" subGenus="Adoxopisalia">
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<emphasis id="B93FD074FFFE8066FF5C29CFFF794BAE" box="[151,236,485,508]" italics="true" pageId="7" pageNumber="8">Leptusa</emphasis>
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(
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<emphasis id="B93FD074FFFE8066FF3029CFFE124BAE" box="[251,391,485,508]" italics="true" pageId="7" pageNumber="8">Adoxopisalia</emphasis>
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)
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<emphasis id="B93FD074FFFE8066FE5C29CFFD834BAE" box="[407,534,485,508]" italics="true" pageId="7" pageNumber="8">smokyiensis</emphasis>
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<treatmentCitation id="0AEA2A77FFFE8066FDD729CFFD5E4BAE" author="Pace" box="[540,715,485,508]" page="252" pageId="7" pageNumber="8" year="1989">
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<bibRefCitation id="EFDA7197FFFE8066FDD729CFFD004BAE" author="Pace" box="[540,661,485,508]" pageId="7" pageNumber="8" refString="Pace, R. (1989) Monografia del genere Leptusa Kraatz (Coleoptera, Staphylinidae). Memorie del Museo Civico di Storia Naturale di Verona (II Serie), Sezione Scienze della vita (A: Biologica), 8, 1 - 307." type="journal article" year="1989">Pace, 1989</bibRefCitation>
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: 252
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="0AEA2A77FFFE8066FD1D29CFFB9F4BAE" author="Klimaszewski" box="[726,1034,485,508]" page="9" pageId="7" pageNumber="8" year="2004">
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<bibRefCitation id="EFDA7197FFFE8066FD1D29CFFC7A4BAE" author="Klimaszewski" box="[726,1007,485,508]" pageId="7" pageNumber="8" refString="Klimaszewski, J., Pelletier, G. & Majka, C. (2004) A revision of Canadian Leptusa Kraatz (Col., Staphylinidae, Aleocharinae): new species, new distribution records, key and taxonomic considerations. Belgian Journal of Entomology, 6, 3 - 42." type="journal article" year="2004">
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Klimaszewski
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<emphasis id="B93FD074FFFE8066FCBF29CFFC3C4BAE" box="[884,937,485,508]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
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, 2004
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</bibRefCitation>
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: 9
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3515FEDFFFE8066FF5C2A00FE8249C6" pageId="7" pageNumber="8" type="materials_examined">
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||
<paragraph id="8BF40C66FFFE8066FF5C2A00FD9A48AD" blockId="7.[151,1437,554,1999]" pageId="7" pageNumber="8">
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||
<emphasis id="B93FD074FFFE8066FF5C2A00FC034816" bold="true" box="[151,918,554,580]" pageId="7" pageNumber="8">
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||
<typeStatus id="54F0B2C4FFFE8066FF5C2A00FF434816" box="[151,214,554,580]" pageId="7" pageNumber="8">Type</typeStatus>
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||
material examined.
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||
<typeStatus id="54F0B2C4FFFE8066FE162A00FDC14816" box="[477,596,554,580]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
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||
.
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||
<collectingCountry id="F35C4CF6FFFE8066FDA92A00FD2D4816" box="[610,696,554,580]" name="United States of America" pageId="7" pageNumber="8">U.S.A.</collectingCountry>
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: North Carolina:
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</emphasis>
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Ψ, glued on rectangular card, spermatheca dissected and mounted in balsam on a clear plastic card, “NC Gr. Sm. Mts. N. P. Clingmans Dome
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<quantity id="4CB3A183FFFE8066FAD32A7AFF3B48DD" metricMagnitude="3" metricUnit="m" metricValue="1.985" metricValueMax="2.02" metricValueMin="1.95" pageId="7" pageNumber="8" unit="m" value="1985.0" valueMax="2020.0" valueMin="1950.0">
|
||
1950
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||
<emphasis id="B93FD074FFFE8066FA982A7AFAF7483B" box="[1363,1378,592,617]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
2020 m
|
||
</quantity>
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||
<date id="FFF52AA6FFFE8066FF7E2A5FFE9A48DD" box="[181,271,629,655]" pageId="7" pageNumber="8" value="1986-06-02">2.VI.86</date>
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||
A. Smetana” [white label, printed]; “
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||
<typeStatus id="54F0B2C4FFFE8066FD742A5FFCE448DD" box="[703,881,629,655]" pageId="7" pageNumber="8" type="holotype">HOLOTYPUS</typeStatus>
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<taxonomicName id="4C4B77E5FFFE8066FCBC2A5FFBF248DC" box="[887,1127,629,654]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="smokyiensis">
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<emphasis id="B93FD074FFFE8066FCBC2A5FFBF248DC" box="[887,1127,629,654]" italics="true" pageId="7" pageNumber="8">Leptusa smokyiensis</emphasis>
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</taxonomicName>
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. Det.
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<bibRefCitation id="EFDA7197FFFE8066FB662A5FFACE48DD" author="PACE" box="[1197,1371,629,655]" pageId="7" pageNumber="8" refString="Pace, R. (1982) Leptusa raccolte dal Dr. Claude Besuchet nella Penisola Iberica, nella regione balcanica, in Turchia e in estremo oriente (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 89, 579 - 594." type="journal article" year="1982">
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R.
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<collectionCode id="ED5A94A3FFFE8066FB042A5FFA8248DD" box="[1231,1303,629,655]" pageId="7" pageNumber="8">PACE</collectionCode>
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1982
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</bibRefCitation>
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” [red label, handwritten]; “
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<taxonomicName id="4C4B77E5FFFE8066FE5A2AB0FD1748E1" box="[401,642,666,691]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="smokyiensis" status="sp. nov.">
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<emphasis id="B93FD074FFFE8066FE5A2AB0FD1748E1" box="[401,642,666,691]" italics="true" pageId="7" pageNumber="8">Leptusa smokyiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C6D0FFFFE8066FD412AB0FD5C48E6" box="[650,713,666,692]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
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||
det. R. Pace 1987” [white label, handwritten]; “
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<typeStatus id="54F0B2C4FFFE8066FB352AB0FA0948E6" box="[1278,1436,666,692]" pageId="7" pageNumber="8" type="holotype">
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<collectionCode id="ED5A94A3FFFE8066FB352AB0FA0948E6" box="[1278,1436,666,692]" pageId="7" pageNumber="8">HOLOTYPE</collectionCode>
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||
</typeStatus>
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||
<collectionCode id="ED5A94A3FFFE8066FF5C2AEAFF414888" box="[151,212,704,730]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="7" pageNumber="8">CNC</collectionCode>
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||
No. 20977” [red label, handwritten]. Deposited in Canadian National Collection of Insects, Arachnids and Nematodes (
|
||
<collectionCode id="ED5A94A3FFFE8066FE972ACFFE0B48AD" box="[348,414,741,767]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="7" pageNumber="8">CNC</collectionCode>
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), Ottawa.
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||
</paragraph>
|
||
<paragraph id="8BF40C66FFFE8066FF0D2B20FE8249C6" blockId="7.[151,1437,554,1999]" pageId="7" pageNumber="8">
|
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<emphasis id="B93FD074FFFE8066FF0D2B20FC134976" bold="true" box="[198,902,778,804]" pageId="7" pageNumber="8">
|
||
Additional material examined (n=2).
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||
<collectingCountry id="F35C4CF6FFFE8066FD562B20FD644976" box="[669,753,778,804]" name="United States of America" pageId="7" pageNumber="8">U.S.A.</collectingCountry>
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||
: Tennessee:
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||
</emphasis>
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||
Blount Co.: Ψ,
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||
<collectionCode id="ED5A94A3FFFE8066FB8C2B20FB394976" LSID="urn:lsid:biocol.org:col:13136" box="[1095,1196,778,804]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="7" pageNumber="8">GSMNP</collectionCode>
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||
, middle Prong Tr at 3944389 259358, 2200`, Berlese,
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<date id="FFF52AA6FFFE8066FDE92B1AFD544918" box="[546,705,816,842]" pageId="7" pageNumber="8" value="2006-06-17">17 June 2006</date>
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, C. Ware, C. Shapico, J. Hudson (
|
||
<collectionCode id="ED5A94A3FFFE8066FB802B1AFB364918" LSID="urn:lsid:biocol.org:col:33726" box="[1099,1187,816,842]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="7" pageNumber="8">LSAM</collectionCode>
|
||
); Sevier Co.: Ψ (slide mounted),
|
||
<collectionCode id="ED5A94A3FFFE8066FEDD2B7FFEEE493D" LSID="urn:lsid:biocol.org:col:13136" box="[278,379,853,879]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="7" pageNumber="8">GSMNP</collectionCode>
|
||
, Porters Creek
|
||
<geoCoordinate id="EE7F6AA1FFFE8066FDFB2B7FFD5D493D" box="[560,712,853,879]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="35.679832">N35°40.790’</geoCoordinate>
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<geoCoordinate id="EE7F6AA1FFFE8066FD1B2B7FFCE6493D" box="[720,883,853,879]" direction="west" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="-83.39758">W83°23.855’</geoCoordinate>
|
||
, SP2 Litter 3 of 3,
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||
<date id="FFF52AA6FFFE8066FB982B7FFA9B493D" box="[1107,1294,853,879]" pageId="7" pageNumber="8" value="2006-10-05">5 October 2006</date>
|
||
, M Gimmel (
|
||
<collectionCode id="ED5A94A3FFFE8066FF542B50FE9E49C6" LSID="urn:lsid:biocol.org:col:13136" box="[159,267,890,916]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="7" pageNumber="8">GSMNP</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3515FEDFFFE8066FF0D2BB5FE054CF6" pageId="7" pageNumber="8" type="description">
|
||
<paragraph id="8BF40C66FFFE8066FF0D2BB5FCB14E1D" blockId="7.[151,1437,554,1999]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93FD074FFFE8066FF0D2BB5FE6D49EB" bold="true" box="[198,504,927,953]" pageId="7" pageNumber="8">Redescription of female.</emphasis>
|
||
Length
|
||
<quantity id="4CB3A183FFFE8066FD972B8AFD7B49E8" box="[604,750,928,954]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="7" pageNumber="8" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">
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2.6
|
||
<emphasis id="B93FD074FFFE8066FD4B2B8AFD1A49EB" box="[640,655,928,953]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
2.8 mm
|
||
</quantity>
|
||
. Head and pronotum blackish brown; elytra and abdomen brown, abdominal tergites III
|
||
<emphasis id="B93FD074FFFE8066FE3E2BEFFD91498C" box="[501,516,965,990]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
V somewhat brighter; mouthparts, legs, and antennomeres 1
|
||
<emphasis id="B93FD074FFFE8066FB1D2BEFFB70498C" box="[1238,1253,965,990]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
3, 11 yellowish brown (
|
||
<figureCitation id="137010E3FFFE8066FF3D2BC0FEAB4E56" box="[246,318,1002,1028]" captionStart="FIGURE 1" captionStartId="3.[151,255,1170,1194]" captionTargetBox="[153,1432,194,1147]" captionTargetId="figure@3.[151,1436,194,1149]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Leptusa. a: Leptusa (Adoxopisalia) opaca Casey; b: L. (A.) pseudosmokyiensis sp. nov.; c: L. (A.) smokyiensis Pace; d: L. (A.) stocksi sp. nov.; e: L. (Dysleptusa) carolinensis Pace; f: L. (D.) gimmeli sp. nov.; g: L. (D.) pusio (Casey); h: L. (Eucryptusa) brevicollis Casey; i: L. (E.) ferroi sp. nov.; j: L. (Ulitusa) cribratula (Casey). Scale bars = 1 mm." httpUri="https://zenodo.org/record/199053/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
|
||
c). Head somewhat glossy, pubescent, with microsculpture. Antennomeres 1
|
||
<emphasis id="B93FD074FFFE8066FB252BC0FB684E51" box="[1262,1277,1002,1027]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
3 elongate, 4 longer than wide, 4 subquadrate, 6
|
||
<emphasis id="B93FD074FFFE8066FD892C3AFDC44E7B" box="[578,593,1040,1065]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
7 subquadrate, and 8
|
||
<emphasis id="B93FD074FFFE8066FC9A2C3AFCF54E7B" box="[849,864,1040,1065]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
10 transverse (
|
||
<figureCitation id="137010E3FFFE8066FBDE2C3AFBF74E78" box="[1045,1122,1040,1066]" captionStart="FIGURE 4" captionStartId="8.[151,256,1874,1898]" captionTargetBox="[158,1426,197,1845]" captionTargetId="figure@8.[151,1436,197,1853]" captionTargetPageId="8" captionText="FIGURE 4. Details of Leptusa (Adoxopisalia) smokyiensis Pace. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: female abdominal tergite VIII; g: female abdominal sternite VIII; h: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199056/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
a). Carina on each side of ventral surface of head complete, attaining gular suture.
|
||
</paragraph>
|
||
<paragraph id="8BF40C66FFFE8066FF0D2C71FC154F58" blockId="7.[151,1437,554,1999]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93FD074FFFE8066FF0D2C71FEDA4E26" box="[198,335,1115,1140]" italics="true" pageId="7" pageNumber="8">Mouthparts</emphasis>
|
||
. Labrum transverse, bearing 3 pairs of short setae, and 7 pairs of long setae, a-seta, b-seta, and many pores (
|
||
<figureCitation id="137010E3FFFE8066FEFB2CAAFEED4EC8" box="[304,376,1152,1178]" captionStart="FIGURE 4" captionStartId="8.[151,256,1874,1898]" captionTargetBox="[158,1426,197,1845]" captionTargetId="figure@8.[151,1436,197,1853]" captionTargetPageId="8" captionText="FIGURE 4. Details of Leptusa (Adoxopisalia) smokyiensis Pace. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: female abdominal tergite VIII; g: female abdominal sternite VIII; h: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199056/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
b). Labium with two distal setae in a longitudinal row, and several pores in median area; a pair of setal pores, 2 pairs of real pores and several pseudopores present in lateral area (
|
||
<figureCitation id="137010E3FFFE8066FB692C8FFB7E4EED" box="[1186,1259,1189,1215]" captionStart="FIGURE 4" captionStartId="8.[151,256,1874,1898]" captionTargetBox="[158,1426,197,1845]" captionTargetId="figure@8.[151,1436,197,1853]" captionTargetPageId="8" captionText="FIGURE 4. Details of Leptusa (Adoxopisalia) smokyiensis Pace. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: female abdominal tergite VIII; g: female abdominal sternite VIII; h: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199056/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
c). Labial palpi bearing 12 setae (a
|
||
<emphasis id="B93FD074FFFE8066FEB82CE1FE174EB6" box="[371,386,1227,1252]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
h, α
|
||
<emphasis id="B93FD074FFFE8066FE642CE1FE284EB6" box="[431,445,1227,1252]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
δ), γ-seta close to f-seta (
|
||
<figureCitation id="137010E3FFFE8066FD292CE0FCBF4EB6" box="[738,810,1226,1252]" captionStart="FIGURE 4" captionStartId="8.[151,256,1874,1898]" captionTargetBox="[158,1426,197,1845]" captionTargetId="figure@8.[151,1436,197,1853]" captionTargetPageId="8" captionText="FIGURE 4. Details of Leptusa (Adoxopisalia) smokyiensis Pace. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: female abdominal tergite VIII; g: female abdominal sternite VIII; h: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199056/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
d). Mentum trapezoidal, bearing 4 pairs of main setae (b, u, v, w), a pair of additional setae, and many pores (
|
||
<figureCitation id="137010E3FFFE8066FCD72CDAFCF14F58" box="[796,868,1264,1290]" captionStart="FIGURE 4" captionStartId="8.[151,256,1874,1898]" captionTargetBox="[158,1426,197,1845]" captionTargetId="figure@8.[151,1436,197,1853]" captionTargetPageId="8" captionText="FIGURE 4. Details of Leptusa (Adoxopisalia) smokyiensis Pace. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: female abdominal tergite VIII; g: female abdominal sternite VIII; h: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199056/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
<paragraph id="8BF40C66FFFE8066FF0D2D3FFE734F96" blockId="7.[151,1437,554,1999]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93FD074FFFE8066FF0D2D3FFEB64F7C" box="[198,291,1301,1326]" italics="true" pageId="7" pageNumber="8">Thorax.</emphasis>
|
||
Pronotum wider than head, and approximately 1.23 times wider than long; somewhat glossy, with microsculpture. Mesoventrite with distinct reticulate microsculpture. Metaventrite with setigerous punctures. Elytra as wide as pronotum, approximately 1.55 times longer than wide, and approximately 1.2 times longer than pronotum; latero-posterior margin emarginate, somewhat glossy, and pubescent; setigerous punctures present.
|
||
<collectingCountry id="F35C4CF6FFFE8066FF312D80FEA04F96" box="[250,309,1450,1476]" name="India" pageId="7" pageNumber="8">Hind</collectingCountry>
|
||
wings present.
|
||
</paragraph>
|
||
<paragraph id="8BF40C66FFFE8066FF0D2DFAFB044C08" blockId="7.[151,1437,554,1999]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93FD074FFFE8066FF0D2DFAFEA04FBB" box="[198,309,1488,1513]" italics="true" pageId="7" pageNumber="8">Abdomen</emphasis>
|
||
. Abdomen glossy, wider than elytra, widest at abdominal tergites V
|
||
<emphasis id="B93FD074FFFE8066FBB62DFAFB194FBB" box="[1149,1164,1488,1513]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
VI. Female abdominal tergites VI approximately 1.62 times wider than long; tergite VIII with posterior margin shallowly emarginate, bearing 4 pairs of main setae, and many additional short setae and pores (
|
||
<figureCitation id="137010E3FFFE8066FC302E30FBD74C66" box="[1019,1090,1562,1588]" captionStart="FIGURE 4" captionStartId="8.[151,256,1874,1898]" captionTargetBox="[158,1426,197,1845]" captionTargetId="figure@8.[151,1436,197,1853]" captionTargetPageId="8" captionText="FIGURE 4. Details of Leptusa (Adoxopisalia) smokyiensis Pace. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: female abdominal tergite VIII; g: female abdominal sternite VIII; h: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199056/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
f). Female abdominal sternite VIII bearing 5 pairs of main setae, and many additional short setae and pores (
|
||
<figureCitation id="137010E3FFFE8066FBE72E6AFBE14C08" box="[1068,1140,1600,1626]" captionStart="FIGURE 4" captionStartId="8.[151,256,1874,1898]" captionTargetBox="[158,1426,197,1845]" captionTargetId="figure@8.[151,1436,197,1853]" captionTargetPageId="8" captionText="FIGURE 4. Details of Leptusa (Adoxopisalia) smokyiensis Pace. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: female abdominal tergite VIII; g: female abdominal sternite VIII; h: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199056/files/figure.png" pageId="7" pageNumber="8">
|
||
Fig.
|
||
<quantity id="4CB3A183FFFE8066FBA82E6AFBE14C08" box="[1123,1140,1600,1626]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="7" pageNumber="8" unit="g" value="4.0">4</quantity>
|
||
</figureCitation>
|
||
g).
|
||
</paragraph>
|
||
<paragraph id="8BF40C66FFFE8066FF0D2E4FFDEE4C2D" blockId="7.[151,1437,554,1999]" box="[198,635,1637,1663]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93FD074FFFE8066FF0D2E4FFEA04C2C" box="[198,309,1637,1662]" italics="true" pageId="7" pageNumber="8">Genitalia</emphasis>
|
||
. Spermatheca as in
|
||
<figureCitation id="137010E3FFFE8066FDEB2E4FFDF34C2D" box="[544,614,1637,1663]" captionStart="FIGURE 4" captionStartId="8.[151,256,1874,1898]" captionTargetBox="[158,1426,197,1845]" captionTargetId="figure@8.[151,1436,197,1853]" captionTargetPageId="8" captionText="FIGURE 4. Details of Leptusa (Adoxopisalia) smokyiensis Pace. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: female abdominal tergite VIII; g: female abdominal sternite VIII; h: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199056/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
h.
|
||
</paragraph>
|
||
<paragraph id="8BF40C66FFFE8066FF0D2EA0FE054CF6" blockId="7.[151,1437,554,1999]" box="[198,400,1674,1700]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93FD074FFFE8066FF0D2EA0FE984CF6" bold="true" box="[198,269,1674,1700]" pageId="7" pageNumber="8">Male.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3515FEDFFFE8066FF0D2E85FD3E4C98" box="[198,683,1711,1738]" pageId="7" pageNumber="8" type="materials_examined">
|
||
<paragraph id="8BF40C66FFFE8066FF0D2E85FD3E4C98" blockId="7.[151,1437,554,1999]" box="[198,683,1711,1738]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93FD074FFFE8066FF0D2E85FEF94C9B" bold="true" box="[198,364,1711,1737]" pageId="7" pageNumber="8">
|
||
<typeStatus id="54F0B2C4FFFE8066FF0D2E85FE914C9B" box="[198,260,1711,1737]" pageId="7" pageNumber="8">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
North Carolina (
|
||
<collectionCode id="ED5A94A3FFFE8066FDF82E9AFD0A4C98" LSID="urn:lsid:biocol.org:col:13136" box="[563,671,1712,1738]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="7" pageNumber="8">GSMNP</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3515FEDFFFE8066FF0D2EFFFF694D0D" pageId="7" pageNumber="8" type="distribution">
|
||
<paragraph id="8BF40C66FFFE8066FF0D2EFFFE014D46" blockId="7.[151,1437,554,1999]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93FD074FFFE8066FF0D2EFFFEF24CBD" bold="true" box="[198,359,1749,1775]" pageId="7" pageNumber="8">Distribution.</emphasis>
|
||
<collectingCountry id="F35C4CF6FFFE8066FEA62EFFFE394CBD" box="[365,428,1749,1775]" name="United States of America" pageId="7" pageNumber="8">USA</collectingCountry>
|
||
: North Carolina, Tennessee. Locations where
|
||
<taxonomicName id="4C4B77E5FFFE8066FC0F2EFFFB374CBC" box="[964,1186,1749,1775]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="smokyiensis" subGenus="Adoxopisalia">
|
||
<emphasis id="B93FD074FFFE8066FC0F2EFFFC494CBC" box="[964,988,1749,1774]" italics="true" pageId="7" pageNumber="8">L.</emphasis>
|
||
(
|
||
<emphasis id="B93FD074FFFE8066FC272EFFFB904CBC" box="[1004,1029,1749,1774]" italics="true" pageId="7" pageNumber="8">A.</emphasis>
|
||
)
|
||
<emphasis id="B93FD074FFFE8066FBDD2EFFFB374CBC" box="[1046,1186,1749,1774]" italics="true" pageId="7" pageNumber="8">smokyiensis</emphasis>
|
||
</taxonomicName>
|
||
has been collected in GSMNP as in Map 1.
|
||
</paragraph>
|
||
<paragraph id="8BF40C66FFFE8066FF0D2F35FF694D0D" blockId="7.[151,1437,554,1999]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93FD074FFFE8066FF0D2F35FEBB4D6B" bold="true" box="[198,302,1823,1849]" pageId="7" pageNumber="8">Habitat.</emphasis>
|
||
Both specimens examined were collected from forest habitat by sifting leaf litter in mid to late summer.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3515FEDFFFE8066FF0D2F40FAE14D9D" pageId="7" pageNumber="8" type="discussion">
|
||
<paragraph id="8BF40C66FFFE8066FF0D2F40FAE14D9D" blockId="7.[151,1437,554,1999]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93FD074FFFE8066FF0D2F40FEC14DD6" bold="true" box="[198,340,1898,1924]" pageId="7" pageNumber="8">Comments.</emphasis>
|
||
<bibRefCitation id="EFDA7197FFFE8066FE952F40FE794DD6" author="Pace" box="[350,492,1898,1924]" pageId="7" pageNumber="8" refString="Pace, R. (1989) Monografia del genere Leptusa Kraatz (Coleoptera, Staphylinidae). Memorie del Museo Civico di Storia Naturale di Verona (II Serie), Sezione Scienze della vita (A: Biologica), 8, 1 - 307." type="journal article" year="1989">Pace (1989)</bibRefCitation>
|
||
mentioned that this species does not have hind wings, but our specimens have fully developed hind wings. We presume that the hind wings of the
|
||
<typeStatus id="54F0B2C4FFFE8066FC212FBAFBC34DF8" box="[1002,1110,1936,1962]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
may be damaged because wingless
|
||
<taxonomicName id="4C4B77E5FFFE8066FECF2F9FFEF44D9C" box="[260,353,1973,1998]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FD074FFFE8066FECF2F9FFEF44D9C" box="[260,353,1973,1998]" italics="true" pageId="7" pageNumber="8">Leptusa</emphasis>
|
||
</taxonomicName>
|
||
species usually have short and compact elytra, such as
|
||
<taxonomicName id="4C4B77E5FFFE8066FC272F9FFB354D9C" box="[1004,1184,1973,1999]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gimmeli" subGenus="Dysleptusa">
|
||
<emphasis id="B93FD074FFFE8066FC272F9FFC694D9C" box="[1004,1020,1973,1998]" italics="true" pageId="7" pageNumber="8">L</emphasis>
|
||
. (
|
||
<emphasis id="B93FD074FFFE8066FBDF2F9FFBA44D9C" box="[1044,1073,1973,1998]" italics="true" pageId="7" pageNumber="8">D.</emphasis>
|
||
)
|
||
<emphasis id="B93FD074FFFE8066FB892F9FFB354D9C" box="[1090,1184,1973,1998]" italics="true" pageId="7" pageNumber="8">gimmeli</emphasis>
|
||
</taxonomicName>
|
||
, described below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |