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Genus
<taxonomicName id="4C4B77E5FFF88060FF2529CDFE624853" ID-CoL="8VRSG" authority="Kraatz, 1856" authorityName="Kraatz" authorityYear="1856" box="[238,503,487,513]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B93FD074FFF88060FF2529CDFEDB4853" bold="true" box="[238,334,487,513]" italics="true" pageId="1" pageNumber="2">Leptusa</emphasis>
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<emphasis id="B93FD074FFF88060FF5C2A1AFF794815" box="[151,236,560,583]" italics="true" pageId="1" pageNumber="2">Leptusa</emphasis>
<treatmentCitation id="0AEA2A77FFF88060FF392A1AFE3C4815" author="Kraatz" box="[242,425,560,583]" page="60" pageId="1" pageNumber="2" year="1856">
<bibRefCitation id="EFDA7197FFF88060FF392A1AFE154815" author="Kraatz" box="[242,384,560,583]" pageId="1" pageNumber="2" refString="Kraatz, G. (1856) Naturgeschichte der Insecten Deutschlands, Erste Abteilung. Coleoptera. Bd. 2. Lief. 1 - 2. Verlag der Nicolaischen Buchhandlung, Berlin, 376 pp." type="book chapter" year="1856">Kraatz, 1856</bibRefCitation>
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<bibRefCitation id="EFDA7197FFF88060FE7F2A1AFDD24815" author="Fenyes" box="[436,583,560,583]" pageId="1" pageNumber="2" refString="Fenyes, A. (1920) Coleoptera Fam. Staphylinidae subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum. Fascicle 173 b. Louis Desmet-Verteneuil, Bruxelles, pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
: 120
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;
<treatmentCitation id="0AEA2A77FFF88060FD432A1AFB874815" author="Bernhauer" box="[648,1042,560,583]" page="553" pageId="1" pageNumber="2" year="1926">
<bibRefCitation id="EFDA7197FFF88060FD432A1AFC484815" author="Bernhauer" box="[648,989,560,583]" pageId="1" pageNumber="2" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 553
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;
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<bibRefCitation id="EFDA7197FFF88060FBD52A1AFB554815" author="Smetana" box="[1054,1216,560,583]" pageId="1" pageNumber="2" refString="Smetana, A. (1973) Die Leptusa - Arten der Tschechoslovakischen Republik einschliesslich Karpatorussland (Col., Staphylinidae). Stuttgarter Beitrage zur Naturkunde, Seria A (Biologie), 225, 1 - 46." type="journal article" year="1973">Smetana, 1973</bibRefCitation>
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;
<treatmentCitation id="0AEA2A77FFF88060FB2C2A1AFA0D4815" author="Lohse" box="[1255,1432,560,583]" page="42" pageId="1" pageNumber="2" year="1974">
<bibRefCitation id="EFDA7197FFF88060FB2C2A1AFAFA4815" author="Lohse" box="[1255,1391,560,583]" pageId="1" pageNumber="2" refString="Lohse, G. A. (1974) 22. U. F.: Aleocharinae. In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5. Staphylinidae II (Hypocyphtinae und Aleocharinae), Pselaphidae. Goecke &amp; Evers, Krefeld, pp. 11 - 72, 221 - 292." type="book chapter" year="1974">Lohse, 1974</bibRefCitation>
: 42
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;
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<bibRefCitation id="EFDA7197FFF88060FF0C2A7AFE5D4835" author="Moore" box="[199,456,592,615]" pageId="1" pageNumber="2" refString="Moore, I. M. &amp; Legner, E. F. (1975) A catalogue of the Staphylinidae of America north of Mexico (Coleoptera). University of California Division of Agricultural Sciences Special Publication No. 3015, 1 - 514." type="book chapter" year="1975">Moore &amp; Legner, 1975</bibRefCitation>
: 483
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;
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<bibRefCitation id="EFDA7197FFF88060FDC62A7AFD3D4835" author="Seevers" box="[525,680,592,615]" pageId="1" pageNumber="2" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, 1 - 289." type="journal article" year="1978">Seevers, 1978</bibRefCitation>
: 164
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;
<treatmentCitation id="0AEA2A77FFF88060FD262A7AFC064835" author="Pace" box="[749,915,592,615]" page="25" pageId="1" pageNumber="2" year="1989">
<bibRefCitation id="EFDA7197FFF88060FD262A7AFCFC4835" author="Pace" box="[749,873,592,615]" pageId="1" pageNumber="2" refString="Pace, R. (1989) Monografia del genere Leptusa Kraatz (Coleoptera, Staphylinidae). Memorie del Museo Civico di Storia Naturale di Verona (II Serie), Sezione Scienze della vita (A: Biologica), 8, 1 - 307." type="journal article" year="1989">Pace, 1989</bibRefCitation>
: 25
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;
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<bibRefCitation id="EFDA7197FFF88060FC6B2A7AFBB54835" author="Ashe" box="[928,1056,592,615]" pageId="1" pageNumber="2" refString="Ashe, J. S. (2001) XII. Subfamily Aleocharinae Fleming 1821. In: Thomas, M. C. &amp; Arnett, R. H. Jr. (Eds.), American Beetles, Volume 1, Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 358 - 374." type="journal article" year="2001">Ashe, 2001</bibRefCitation>
: 366
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;
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<bibRefCitation id="EFDA7197FFF88060FBAE2A7AFB624835" author="Assing" box="[1125,1271,592,615]" pageId="1" pageNumber="2" refString="Assing, V. (2002) New species and records of Leptusa Kraatz from the Palaearctic region (Coleoptera: Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 34, 971 - 1019." type="journal article" year="2002">Assing, 2002</bibRefCitation>
: 971
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; Gusarov, 2003: 113;
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, 2003b: 191;
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<bibRefCitation id="EFDA7197FFF88060FC8E2A5AFBF248D5" author="Klimaszewski" box="[837,1127,624,647]" pageId="1" pageNumber="2" refString="Klimaszewski, J., Pelletier, G. &amp; Majka, C. (2004) A revision of Canadian Leptusa Kraatz (Col., Staphylinidae, Aleocharinae): new species, new distribution records, key and taxonomic considerations. Belgian Journal of Entomology, 6, 3 - 42." type="journal article" year="2004">
Klimaszewski
<emphasis id="B93FD074FFF88060FC2E2A5AFB8A48D5" box="[997,1055,624,647]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
, 2004
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: 3
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;
<treatmentCitation id="0AEA2A77FFF88060FB582A5AFEBD48F5" author="Gouix" page="61" pageId="1" pageNumber="2" year="2007">
<bibRefCitation id="EFDA7197FFF88060FB582A5AFE9548F5" author="Gouix" pageId="1" pageNumber="2" refString="Gouix, N. &amp; Klimaszewski, J. (2007) Catalogue of Aleocharine Rove Beetles of Canada and Alaska (Coleoptera, Staphylinidae, Aleocharinae). Pensoft Publishers, Sofia - Moscow, 165 pp." type="book" year="2007">Gouix &amp; Klimaszewski, 2007</bibRefCitation>
: 61
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.
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</paragraph>
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<typeStatus id="54F0B2C4FFF88060FF5C2AFFFF4148BD" box="[151,212,725,751]" pageId="1" pageNumber="2">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C4B77E5FFF88060FE8B2AFFFCBE48BD" authority="Mannerheim, 1830" authorityName="Mannerheim" authorityYear="1830" box="[320,811,725,751]" class="Insecta" family="Staphylinidae" genus="Bolitochara" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="B93FD074FFF88060FE8B2AFFFDD448BC" box="[320,577,725,750]" italics="true" pageId="1" pageNumber="2">Bolitochara pulchella</emphasis>
<bibRefCitation id="EFDA7197FFF88060FD832AFFFCBE48BD" author="Mannerheim" box="[584,811,725,751]" pageId="1" pageNumber="2" refString="Mannerheim, C. G. (1830) Precis d'un nouvel arrangement de la famille des Brachelytres, de l'ordre des Insectes Coleopteres. St. Petersbourg, 87 pp." type="book" year="1830">Mannerheim, 1830</bibRefCitation>
</taxonomicName>
; designated by
<bibRefCitation id="EFDA7197FFF88060FC2B2AFFFAB848BD" author="Gusarov" box="[992,1325,725,751]" pageId="1" pageNumber="2" refString="Gusarov, V. I. &amp; Herman, L. H. (2003 a) Leptusa Kraatz, 1856 (Coleoptera: Staphylinidae: Aleocharinae): designation of the type species. Entomologische Blatter, 98 [2002], 115 - 119." type="book chapter" year="2003" yearSuffix="a">Gusarov &amp; Herman (2003a)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFF88060FF0D2AD0FE9D4E36" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BF40C66FFF88060FF0D2AD0FE9D4E36" blockId="1.[151,1437,725,1386]" pageId="1" pageNumber="2">
<emphasis id="B93FD074FFF88060FF0D2AD0FED34946" bold="true" box="[198,326,762,788]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
The genus
<taxonomicName id="4C4B77E5FFF88060FE1B2AD0FDB84941" box="[464,557,762,787]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B93FD074FFF88060FE1B2AD0FDB84941" box="[464,557,762,787]" italics="true" pageId="1" pageNumber="2">Leptusa</emphasis>
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can be recognized by the following combination of characters: body more or less convex, parallel-sided, pubescent; microsculpture present; hind wings often absent; eye size highly variable but usually small; antennae with 11 antennomeres, thickened toward apex; mandible short, right mandible with internal tooth; distal lobe of galea membranous and pubescent; maxillary palpomere 1 smallest, 2 moderately long, 3 largest, 4 subuliform; ligula narrow and long, not bifid, rounded apically; labial palpi with two apparent articles due to fusion of articles 1
<emphasis id="B93FD074FFF88060FCC52B9FFC88499C" box="[782,797,949,974]" italics="true" pageId="1" pageNumber="2"></emphasis>
2; pronotum usually widest at apical third or anterior half; mesocoxae narrowly separated; mesoventral process longer than metaventral process (approximately 2/3 length of mesocoxae); legs usually short, tarsal formula 4-4-5, first metatarsomere as long as following two articles or slightly longer. Abdominal tergites III
<emphasis id="B93FD074FFF88060FD1C2C0FFD734E6C" box="[727,742,1061,1086]" italics="true" pageId="1" pageNumber="2"></emphasis>
VI with setigerous punctures (
<bibRefCitation id="EFDA7197FFF88060FB872C0FFB5E4E6D" author="Pace" box="[1100,1227,1061,1087]" pageId="1" pageNumber="2" refString="Pace, R. (1989) Monografia del genere Leptusa Kraatz (Coleoptera, Staphylinidae). Memorie del Museo Civico di Storia Naturale di Verona (II Serie), Sezione Scienze della vita (A: Biologica), 8, 1 - 307." type="journal article" year="1989">Pace 1989</bibRefCitation>
,
<bibRefCitation id="EFDA7197FFF88060FB132C0FFF694E36" author="Klimaszewski" pageId="1" pageNumber="2" refString="Klimaszewski, J., Pelletier, G. &amp; Majka, C. (2004) A revision of Canadian Leptusa Kraatz (Col., Staphylinidae, Aleocharinae): new species, new distribution records, key and taxonomic considerations. Belgian Journal of Entomology, 6, 3 - 42." type="journal article" year="2004">
Klimaszewski
<emphasis id="B93FD074FFF88060FA4C2C0FFF204E36" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFF88060FF0D2C45FC324F38" pageId="1" pageNumber="2" type="discussion">
<paragraph id="8BF40C66FFF88060FF0D2C45FC324F38" blockId="1.[151,1437,725,1386]" pageId="1" pageNumber="2">
<emphasis id="B93FD074FFF88060FF0D2C45FEAA4EDB" bold="true" box="[198,319,1135,1161]" pageId="1" pageNumber="2">Remarks.</emphasis>
All species of
<taxonomicName id="4C4B77E5FFF88060FE352C5AFDCE4EDB" box="[510,603,1136,1161]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B93FD074FFF88060FE352C5AFDCE4EDB" box="[510,603,1136,1161]" italics="true" pageId="1" pageNumber="2">Leptusa</emphasis>
</taxonomicName>
collected in GSMNP have the following combination of characters: gular sutures widely separated, approximately 1/5 width of head; each side of ventral surface of head with carinae (fading before attaining gular suture or attaining gular suture); labrum bearing 3 pairs of short setae along anterior and lateral margin (
<figureCitation id="137010E3FFF88060FDF22CCAFD1B4EA8" box="[569,654,1248,1274]" captionStart="FIGURE 2" captionStartId="4.[151,259,1947,1971]" captionTargetBox="[188,1406,194,1927]" captionTargetId="figure@4.[178,1409,194,1933]" captionTargetPageId="4" captionText="FIGURE 2. Details of Leptusa (Adoxopisalia) opaca Casey. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: paramere; j: median lobe. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199054/files/figure.png" pageId="1" pageNumber="2">Figs. 2</figureCitation>
b
<emphasis id="B93FD074FFF88060FD562CCAFD394EAB" box="[669,684,1248,1273]" italics="true" pageId="1" pageNumber="2"></emphasis>
11b); median area of labium distinctly narrow or lateral areas meeting at middle, and a pair of distal setae present in a longitudinal row; mesoventrite carinate or not (if carinate, then not exceeding 1/2 length of mesoventrite). Abdominal sternites VII of males possess many pores occupying anywhere from anterior one-tenth to anterior half.
</paragraph>
</subSubSection>
</treatment>
</document>