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<document id="68492203A684CDE1C54AE94A27A38626" ID-DOI="10.5281/zenodo.199052" ID-GBIF-Dataset="a07f4b3c-e602-4b4e-95dc-87fef597d912" ID-ISSN="1175-5326" ID-Zenodo-Dep="199052" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460321171440" checkinUser="plazi" docAuthor="Park, Jong-Seok, Carlton, Christopher E. &amp; Ferro, Michael L." docDate="2010" docId="03E2BD70FFF68070FF5C2A18FCBB4B8E" docLanguage="en" docName="zt02662p027.pdf" docOrigin="Zootaxa 2662" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptusa (Dysleptusa) pusio Casey 1906" docType="treatment" docVersion="10" lastPageNumber="18" masterDocId="FFDBC508FFF98061FFCB282AFF954A52" masterDocTitle="Diversity and taxonomic review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species" masterLastPageNumber="27" masterPageNumber="1" pageNumber="16" updateTime="1698256666968" updateUser="plazi">
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<mods:title id="B08E46E4F6CAA7B726B44B00BDBF043F">Diversity and taxonomic review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species</mods:title>
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<mods:namePart id="55A2A06C8745D488BE1E1BB85D865BE7">Carlton, Christopher E.</mods:namePart>
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<treatment id="03E2BD70FFF68070FF5C2A18FCBB4B8E" ID-DOI="http://doi.org/10.5281/zenodo.5020593" ID-GBIF-Taxon="119402522" ID-Zenodo-Dep="5020593" LSID="urn:lsid:plazi:treatment:03E2BD70FFF68070FF5C2A18FCBB4B8E" httpUri="http://treatment.plazi.org/id/03E2BD70FFF68070FF5C2A18FCBB4B8E" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
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<paragraph id="8BF40C66FFF6806EFF5C2A18FD1E481E" blockId="15.[151,651,562,623]" box="[151,651,562,588]" pageId="15" pageNumber="16">
<heading id="D0BCBB0AFFF6806EFF5C2A18FD1E481E" bold="true" box="[151,651,562,588]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="4C4B77E5FFF6806EFF5C2A18FD1E481E" ID-CoL="6PP8C" authority="Casey, 1906" authorityName="Casey" authorityYear="1906" box="[151,651,562,588]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pusio" subGenus="Dysleptusa">
<emphasis id="B93FD074FFF6806EFF5C2A18FD1E481E" bold="true" box="[151,651,562,588]" pageId="15" pageNumber="16">
<emphasis id="B93FD074FFF6806EFF5C2A18FF62481E" bold="true" box="[151,247,562,588]" italics="true" pageId="15" pageNumber="16">Leptusa</emphasis>
(
<emphasis id="B93FD074FFF6806EFEC32A18FE19481E" bold="true" box="[264,396,562,588]" italics="true" pageId="15" pageNumber="16">Dysleptusa</emphasis>
)
<emphasis id="B93FD074FFF6806EFE562A18FE4B481E" bold="true" box="[413,478,562,588]" italics="true" pageId="15" pageNumber="16">pusio</emphasis>
(
<bibRefCitation id="EFDA7197FFF6806EFE262A18FD16481E" author="Casey" box="[493,643,562,588]" pageId="15" pageNumber="16" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF40C66FFF6806EFF5C2A7FFE36483D" blockId="15.[151,651,562,623]" box="[151,419,597,623]" pageId="15" pageNumber="16">
(
<figureCitation id="137010E3FFF6806EFF542A7FFF7B483D" box="[159,238,597,623]" captionStart="FIGURE 1" captionStartId="3.[151,255,1170,1194]" captionTargetBox="[153,1432,194,1147]" captionTargetId="figure@3.[151,1436,194,1149]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Leptusa. a: Leptusa (Adoxopisalia) opaca Casey; b: L. (A.) pseudosmokyiensis sp. nov.; c: L. (A.) smokyiensis Pace; d: L. (A.) stocksi sp. nov.; e: L. (Dysleptusa) carolinensis Pace; f: L. (D.) gimmeli sp. nov.; g: L. (D.) pusio (Casey); h: L. (Eucryptusa) brevicollis Casey; i: L. (E.) ferroi sp. nov.; j: L. (Ulitusa) cribratula (Casey). Scale bars = 1 mm." httpUri="https://zenodo.org/record/199053/files/figure.png" pageId="15" pageNumber="16">
Figs.
<quantity id="4CB3A183FFF6806EFF292A7FFF7B483D" box="[226,238,597,623]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="15" pageNumber="16" unit="g" value="1.0">1</quantity>
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g, 8a
<emphasis id="B93FD074FFF6806EFEE22A7FFEAD483C" box="[297,312,597,622]" italics="true" pageId="15" pageNumber="16"></emphasis>
l, Map 1)
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFF6806EFF5C2AB0FE6B48A3" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8BF40C66FFF6806EFF5C2AB0FE7948E3" blockId="15.[151,1435,666,753]" box="[151,492,666,689]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB5B2B48FFF6806EFF5C2AB0FE7948E3" box="[151,492,666,689]" pageId="15" pageNumber="16">
<taxonomicName id="4C4B77E5FFF6806EFF5C2AB0FE7248E3" authority="Casey, 1906: 348" authorityName="Casey" authorityPageNumber="348" authorityYear="1906" box="[151,487,666,689]" class="Insecta" family="Staphylinidae" genus="Ulitusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="B93FD074FFF6806EFF5C2AB0FEB348E3" box="[151,294,666,689]" italics="true" pageId="15" pageNumber="16">Ulitusa pusio</emphasis>
<treatmentCitation id="0AEA2A77FFF6806EFEE72AB0FE7248E3" author="Casey" box="[300,487,666,689]" page="348" pageId="15" pageNumber="16" year="1906">
<bibRefCitation id="EFDA7197FFF6806EFEE72AB0FE2748E3" author="Casey" box="[300,434,666,689]" pageId="15" pageNumber="16" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
: 348
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8BF40C66FFF6806EFF5C2A90FE6B48A3" blockId="15.[151,1435,666,753]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB5B2B48FFF6806EFF5C2A90FE6B48A3" pageId="15" pageNumber="16">
<taxonomicName id="4C4B77E5FFF6806EFF5C2A90FD884883" ID-CoL="6PP8C" authority="Fenyes, 1920: 123" authorityName="Fenyes" authorityPageNumber="123" authorityYear="1920" box="[151,541,698,721]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="B93FD074FFF6806EFF5C2A90FEA64883" box="[151,307,698,721]" italics="true" pageId="15" pageNumber="16">Leptusa pusio</emphasis>
:
<treatmentCitation id="0AEA2A77FFF6806EFE832A90FD884883" author="Fenyes" box="[328,541,698,721]" page="123" pageId="15" pageNumber="16" year="1920">
<bibRefCitation id="EFDA7197FFF6806EFE832A90FE744883" author="Fenyes" box="[328,481,698,721]" pageId="15" pageNumber="16" refString="Fenyes, A. (1920) Coleoptera Fam. Staphylinidae subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum. Fascicle 173 b. Louis Desmet-Verteneuil, Bruxelles, pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
: 123
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AEA2A77FFF6806EFDE42A90FC404883" author="Bernhauer" box="[559,981,698,721]" page="556" pageId="15" pageNumber="16" year="1926">
<bibRefCitation id="EFDA7197FFF6806EFDE42A90FC0C4883" author="Bernhauer" box="[559,921,698,721]" pageId="15" pageNumber="16" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 556
</treatmentCitation>
;
<treatmentCitation id="0AEA2A77FFF6806EFC2C2A90FB364883" author="Pace" box="[999,1187,698,721]" page="107" pageId="15" pageNumber="16" year="1989">
<bibRefCitation id="EFDA7197FFF6806EFC2C2A90FBF24883" author="Pace" box="[999,1127,698,721]" pageId="15" pageNumber="16" refString="Pace, R. (1989) Monografia del genere Leptusa Kraatz (Coleoptera, Staphylinidae). Memorie del Museo Civico di Storia Naturale di Verona (II Serie), Sezione Scienze della vita (A: Biologica), 8, 1 - 307." type="journal article" year="1989">Pace, 1989</bibRefCitation>
: 107
</treatmentCitation>
; Gusarov, 2003: 115;
<treatmentCitation id="0AEA2A77FFF6806EFF0C2AF0FE6F48A3" author="Klimaszewski" box="[199,506,730,753]" page="9" pageId="15" pageNumber="16" year="2004">
<bibRefCitation id="EFDA7197FFF6806EFF0C2AF0FE7548A3" author="Klimaszewski" box="[199,480,730,753]" pageId="15" pageNumber="16" refString="Klimaszewski, J., Pelletier, G. &amp; Majka, C. (2004) A revision of Canadian Leptusa Kraatz (Col., Staphylinidae, Aleocharinae): new species, new distribution records, key and taxonomic considerations. Belgian Journal of Entomology, 6, 3 - 42." type="journal article" year="2004">
Klimaszewski
<emphasis id="B93FD074FFF6806EFEAF2AF0FE0F48A3" box="[356,410,730,753]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
, 2004
</bibRefCitation>
: 9
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFF6806EFF5C2B30FDEA4F41" pageId="15" pageNumber="16" type="materials_examined">
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<emphasis id="B93FD074FFF6806EFF5C2B30FD4C4966" bold="true" box="[151,729,794,820]" pageId="15" pageNumber="16">
Material examined (n=27).
<collectingCountry id="F35C4CF6FFF6806EFE242B30FDD14966" box="[495,580,794,820]" name="United States of America" pageId="15" pageNumber="16">U.S.A.</collectingCountry>
: Tennessee:
</emphasis>
Sevier Co.: 5ɗɗ, 7ΨΨ (Ψ slide mounted),
<collectionCode id="ED5A94A3FFF6806EFB1C2B30FAA94966" LSID="urn:lsid:biocol.org:col:13136" box="[1239,1340,794,820]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="15" pageNumber="16">GSMNP</collectionCode>
, Porters Creek,
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,
<date id="FFF52AA6FFF6806EFD812B15FD7E490B" box="[586,747,831,857]" pageId="15" pageNumber="16" value="2007-04-05">5 April 2007</date>
, SP2 Litter 2 of 3, M. Ferro (
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); ɗ, 2ΨΨ (ɗ slide mounted),
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, Porters Creek,
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,
<date id="FFF52AA6FFF6806EFC4B2B4EFB1A492C" box="[896,1167,868,894]" pageId="15" pageNumber="16" value="2006-04-12" valueMax="2006-05-18" valueMin="2006-04-12">
12 April
<emphasis id="B93FD074FFF6806EFC292B4EFC64492F" box="[994,1009,868,893]" italics="true" pageId="15" pageNumber="16"></emphasis>
18 May 2006
</date>
, SP25C -CWD Rear 1, M. Ferro (
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); ɗ,
<collectionCode id="ED5A94A3FFF6806EFE552BA3FD9649F1" LSID="urn:lsid:biocol.org:col:13136" box="[414,515,905,931]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="15" pageNumber="16">GSMNP</collectionCode>
, Porters Creek,
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<geoCoordinate id="EE7F6AA1FFF6806EFC962BA3FC6A49F1" box="[861,1023,905,931]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-83.39758">W83°23.855</geoCoordinate>
,
<date id="FFF52AA6FFF6806EFBC02BA3FB5049F1" box="[1035,1221,905,931]" pageId="15" pageNumber="16" value="2006-10-05">5 October 2006</date>
, SP2 Litter 1 of 3, M. Gimmel (
<collectionCode id="ED5A94A3FFF6806EFEFB2B87FE1D4995" LSID="urn:lsid:biocol.org:col:33726" box="[304,392,941,967]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="15" pageNumber="16">LSAM</collectionCode>
); ɗ,
<collectionCode id="ED5A94A3FFF6806EFE742B87FDB14995" LSID="urn:lsid:biocol.org:col:13136" box="[447,548,941,967]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="15" pageNumber="16">GSMNP</collectionCode>
, Porters Creek,
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<geoCoordinate id="EE7F6AA1FFF6806EFCB42B87FBB44995" box="[895,1057,941,967]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-83.39758">W83°23.855</geoCoordinate>
,
<date id="FFF52AA6FFF6806EFBE62B87FAAB4995" box="[1069,1342,941,967]" pageId="15" pageNumber="16" value="2007-04-12" valueMax="2007-05-18" valueMin="2007-04-12">
12 April
<emphasis id="B93FD074FFF6806EFB5B2B84FB0B4995" box="[1168,1182,942,967]" italics="true" pageId="15" pageNumber="16"></emphasis>
18 May 2007
</date>
, SP23B -CWD Rear 1, M. Ferro (
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); ɗ,
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, Porters Creek,
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<geoCoordinate id="EE7F6AA1FFF6806EFBD32BF8FB2F49BE" box="[1048,1210,978,1004]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-83.39758">W83°23.855</geoCoordinate>
,
<date id="FFF52AA6FFF6806EFB0D2BF8FAEA49BE" box="[1222,1407,978,1004]" pageId="15" pageNumber="16" value="2006-10-04">4 October 2006</date>
<emphasis id="B93FD074FFF6806EFAB52BF8FA1849B9" box="[1406,1421,978,1003]" italics="true" pageId="15" pageNumber="16"></emphasis>
<date id="FFF52AA6FFF6806EFA462BF8FE8E4E43" pageId="15" pageNumber="16" value="2007-04-01">1 April 2007</date>
, SP21B -CWD Rear 1, M. Ferro (
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); ɗ,
<collectionCode id="ED5A94A3FFF6806EFC8B2BDDFC334E43" LSID="urn:lsid:biocol.org:col:13136" box="[832,934,1015,1041]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="15" pageNumber="16">GSMNP</collectionCode>
, Sugarlands Quiet Walkway,
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,
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, SN32B -CWD Rear 1, M. Ferro (
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); Ψ,
<collectionCode id="ED5A94A3FFF6806EFB492C36FB724E64" LSID="urn:lsid:biocol.org:col:13136" box="[1154,1255,1052,1078]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="15" pageNumber="16">GSMNP</collectionCode>
, Porters Creek,
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<geoCoordinate id="EE7F6AA1FFF6806EFEF62C6BFE704E09" box="[317,485,1089,1115]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-83.39758">W83°23.855</geoCoordinate>
,
<date id="FFF52AA6FFF6806EFE3A2C6BFD264E09" box="[497,691,1089,1115]" pageId="15" pageNumber="16" value="2006-10-05">5 October 2006</date>
, SP2 Litter 2 of 3, M. Gimmel (
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); Ψ,
<collectionCode id="ED5A94A3FFF6806EFB1A2C6BFAAC4E09" LSID="urn:lsid:biocol.org:col:13136" box="[1233,1337,1089,1115]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="15" pageNumber="16">GSMNP</collectionCode>
, Porters Creek,
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<geoCoordinate id="EE7F6AA1FFF6806EFE522C4CFDD74ED2" box="[409,578,1126,1152]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-83.39758">W83°23.855</geoCoordinate>
,
<date id="FFF52AA6FFF6806EFD9B2C4CFCF34ED2" box="[592,870,1126,1152]" pageId="15" pageNumber="16" value="2006-05-18" valueMax="2006-06-24" valueMin="2006-05-18">
18 May
<emphasis id="B93FD074FFF6806EFD642C4CFD2B4E2D" box="[687,702,1126,1151]" italics="true" pageId="15" pageNumber="16"></emphasis>
24 June 2006
</date>
, SP24A -CWD Rear 1, M. Ferro (
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); Ψ,
<collectionCode id="ED5A94A3FFF6806EFF5C2CA0FF694EF6" LSID="urn:lsid:biocol.org:col:13136" box="[151,252,1162,1188]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="15" pageNumber="16">GSMNP</collectionCode>
, Porters Creek,
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<geoCoordinate id="EE7F6AA1FFF6806EFD9D2CA0FD6C4EF6" box="[598,761,1162,1188]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-83.39758">W83°23.855</geoCoordinate>
,
<date id="FFF52AA6FFF6806EFCCF2CA0FC2F4EF6" box="[772,954,1162,1188]" pageId="15" pageNumber="16" value="2006-10-04">4 October 2006</date>
<emphasis id="B93FD074FFF6806EFC702CA1FC5F4EF6" box="[955,970,1163,1188]" italics="true" pageId="15" pageNumber="16"></emphasis>
<date id="FFF52AA6FFF6806EFC022CA0FBF74EF6" box="[969,1122,1162,1188]" pageId="15" pageNumber="16" value="2007-04-01">1 April 2007</date>
, SP24A -CWD Rear 1, M. Ferro (
<collectionCode id="ED5A94A3FFF6806EFF2D2C85FEAB4E9B" LSID="urn:lsid:biocol.org:col:33726" box="[230,318,1199,1225]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="15" pageNumber="16">LSAM</collectionCode>
); Ψ,
<collectionCode id="ED5A94A3FFF6806EFEBF2C85FE4C4E9B" LSID="urn:lsid:biocol.org:col:13136" box="[372,473,1199,1225]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="15" pageNumber="16">GSMNP</collectionCode>
, Porters Creek,
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<geoCoordinate id="EE7F6AA1FFF6806EFCF82C85FC404E9B" box="[819,981,1199,1225]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-83.39758">W83°23.855</geoCoordinate>
,
<date id="FFF52AA6FFF6806EFC2A2C85FB7C4E9B" box="[993,1257,1199,1225]" pageId="15" pageNumber="16" value="2006-05-18" valueMax="2006-06-24" valueMin="2006-05-18">
18 May
<emphasis id="B93FD074FFF6806EFBF02C9AFBDF4E9B" box="[1083,1098,1200,1225]" italics="true" pageId="15" pageNumber="16"></emphasis>
24 June 2006
</date>
, SP24C -CWD Rear 1, M. Ferro (
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); Ψ,
<collectionCode id="ED5A94A3FFF6806EFE3F2CFEFDCC4EBC" LSID="urn:lsid:biocol.org:col:13136" box="[500,601,1236,1262]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="15" pageNumber="16">GSMNP</collectionCode>
, Greenbrier,
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<geoCoordinate id="EE7F6AA1FFF6806EFC442CFEFBA44EBC" box="[911,1073,1236,1262]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-83.38915">W83°23.349</geoCoordinate>
,
<date id="FFF52AA6FFF6806EFBF72CFEFB674EBC" box="[1084,1266,1236,1262]" pageId="15" pageNumber="16" value="2006-10-04">4 October 2006</date>
<emphasis id="B93FD074FFF6806EFB3A2CFEFA954EBF" box="[1265,1280,1236,1261]" italics="true" pageId="15" pageNumber="16"></emphasis>
<date id="FFF52AA6FFF6806EFACB2CFEFA024EBC" box="[1280,1431,1236,1262]" pageId="15" pageNumber="16" value="2007-04-01">1 April 2007</date>
, SP13A -CWD Rear 1, M. Ferro (
<collectionCode id="ED5A94A3FFF6806EFDD12CD3FDE74F41" LSID="urn:lsid:biocol.org:col:33726" box="[538,626,1273,1299]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="15" pageNumber="16">LSAM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFF68070FF0D2D37FB284AE0" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="8BF40C66FFF6806EFF0D2D37FE0D4F99" blockId="15.[151,1437,794,2035]" pageId="15" pageNumber="16">
<emphasis id="B93FD074FFF6806EFF0D2D37FEE94F65" bold="true" box="[198,380,1309,1335]" pageId="15" pageNumber="16">Redescription.</emphasis>
Length
<quantity id="4CB3A183FFF6806EFE162D34FDFB4F6A" box="[477,622,1310,1336]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="15" pageNumber="16" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">
1.9
<emphasis id="B93FD074FFF6806EFDC92D34FD854F65" box="[514,528,1310,1335]" italics="true" pageId="15" pageNumber="16"></emphasis>
2.2 mm
</quantity>
. Body brown, glossy; head and abdominal tergite VI blackish brown; antennomeres 1
<emphasis id="B93FD074FFF6806EFE992D69FEF44F0E" box="[338,353,1347,1372]" italics="true" pageId="15" pageNumber="16"></emphasis>
3, mouthparts and legs somewhat brighter (
<figureCitation id="137010E3FFF6806EFCA32D68FC394F0E" box="[872,940,1346,1372]" captionStart="FIGURE 1" captionStartId="3.[151,255,1170,1194]" captionTargetBox="[153,1432,194,1147]" captionTargetId="figure@3.[151,1436,194,1149]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Leptusa. a: Leptusa (Adoxopisalia) opaca Casey; b: L. (A.) pseudosmokyiensis sp. nov.; c: L. (A.) smokyiensis Pace; d: L. (A.) stocksi sp. nov.; e: L. (Dysleptusa) carolinensis Pace; f: L. (D.) gimmeli sp. nov.; g: L. (D.) pusio (Casey); h: L. (Eucryptusa) brevicollis Casey; i: L. (E.) ferroi sp. nov.; j: L. (Ulitusa) cribratula (Casey). Scale bars = 1 mm." httpUri="https://zenodo.org/record/199053/files/figure.png" pageId="15" pageNumber="16">
Fig.
<quantity id="4CB3A183FFF6806EFC6B2D68FC394F0E" box="[928,940,1346,1372]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="15" pageNumber="16" unit="g" value="1.0">1</quantity>
</figureCitation>
g). Head as wide as pronotum, with large setigerous punctures (distance between punctures more than their diameter) present. Antennomeres 1
<emphasis id="B93FD074FFF6806EFAB52D42FA184FD3" box="[1406,1421,1384,1409]" italics="true" pageId="15" pageNumber="16"></emphasis>
3 elongate, 4 subquadrate, 5
<emphasis id="B93FD074FFF6806EFE042DA6FE4B4FF7" box="[463,478,1420,1445]" italics="true" pageId="15" pageNumber="16"></emphasis>
10 transverse (
<figureCitation id="137010E3FFF6806EFD462DA6FD414FF4" box="[653,724,1420,1446]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
a). Carina on each side of ventral surface of head complete, attaining gular suture.
</paragraph>
<paragraph id="8BF40C66FFF6806EFF0D2DFCFB964CD1" blockId="15.[151,1437,794,2035]" pageId="15" pageNumber="16">
<emphasis id="B93FD074FFF6806EFF0D2DFCFEDA4FBD" box="[198,335,1494,1519]" italics="true" pageId="15" pageNumber="16">Mouthparts</emphasis>
. Labrum transverse, bearing 3 pairs of short setae, and 7 pairs of long setae, a-seta, b-seta, and pores (
<figureCitation id="137010E3FFF6806EFF2C2DD0FEBB4C46" box="[231,302,1530,1556]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
b). Labium bearing two distal setae in a longitudinal row in median area; a pair of setal pores, 2 pairs of real pores and several pseudopores present in lateral area (
<figureCitation id="137010E3FFF6806EFC552E35FC714C6B" box="[926,996,1567,1593]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
c). Labial palpi bearing 12 setae (a
<emphasis id="B93FD074FFF6806EFABC2E0AFA134C6B" box="[1399,1414,1568,1593]" italics="true" pageId="15" pageNumber="16"></emphasis>
h, α
<emphasis id="B93FD074FFF6806EFF6D2E6EFF204C0F" box="[166,181,1604,1629]" italics="true" pageId="15" pageNumber="16"></emphasis>
δ), γ-seta at least three times longer than α-, β-, δ-seta (
<figureCitation id="137010E3FFF6806EFC982E6EFC0E4C0C" box="[851,923,1604,1630]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
d). Mentum trapezoidal, bearing 4 pairs of main setae (b, u, v, w), a pair of additional setae, and many pores (
<figureCitation id="137010E3FFF6806EFC6B2E43FC734CD1" box="[928,998,1641,1667]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
e).
</paragraph>
<paragraph id="8BF40C66FFF6806EFF0D2EA4FB714D32" blockId="15.[151,1437,794,2035]" pageId="15" pageNumber="16">
<emphasis id="B93FD074FFF6806EFF0D2EA4FEB64CF5" box="[198,291,1678,1703]" italics="true" pageId="15" pageNumber="16">Thorax.</emphasis>
Pronotum approximately 1.32 times wider than long, with large setigerous punctures (distance between punctures more than their diameter) present. Mesoventrite with distinct reticulate microsculpture; incompletely carinate, carina extending approximately 1/3 length of mesoventrite. Metaventrite with large setigerous punctures present. Elytra wider than pronotum, approximately 1.46 times longer than wide, approximately 1.35 times longer than pronotum, latero-posterior margin emarginate, with large setigerous punctures (distance between punctures more than their diameter) present.
<collectingCountry id="F35C4CF6FFF6806EFC332F6CFBA64D32" box="[1016,1075,1862,1888]" name="India" pageId="15" pageNumber="16">Hind</collectingCountry>
wings present.
</paragraph>
<paragraph id="8BF40C66FFF68071FF0D2F41FE6F4B17" blockId="15.[151,1437,794,2035]" lastBlockId="16.[151,1437,152,325]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="B93FD074FFF6806EFF0D2F41FEA04DD6" box="[198,309,1899,1924]" italics="true" pageId="15" pageNumber="16">Abdomen</emphasis>
. Abdomen wider than elytra, widest at abdominal tergites VI
<emphasis id="B93FD074FFF6806EFB8F2F41FBC64DD6" box="[1092,1107,1899,1924]" italics="true" pageId="15" pageNumber="16"></emphasis>
VII. Abdominal tergite VI approximately 2.1 times wider than long. Male abdominal tergites VII
<emphasis id="B93FD074FFF6806EFC3D2FBAFB904DFB" box="[1014,1029,1936,1961]" italics="true" pageId="15" pageNumber="16"></emphasis>
VIII with single median elongate tubercle (
<figureCitation id="137010E3FFF6806EFECD2F9EFED94D9C" box="[262,332,1972,1998]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
f); tergite VIII bearing 4 pairs of main setae, many additional short setae and pores; posterior margin with several tubercles, and emarginate (
<figureCitation id="137010E3FFF6806EFD162FF3FCB34DA1" box="[733,806,2009,2035]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
f). Male abdominal sternite VII with many pores in anterior one-fourth; sternite VIII with posterior margin produced, bearing 9 pairs of main setae, and many additional short setae and pores (
<figureCitation id="137010E3FFE98071FDE12896FDE64A84" box="[554,627,188,214]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="16" pageNumber="17">
Fig.
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g). Female abdominal tergite VIII with posterior margin shallowly emarginate, bearing 4 pairs of main setae, and many additional short setae and pores (
<figureCitation id="137010E3FFE98071FB0728CBFA8D4AA9" box="[1228,1304,225,251]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="16" pageNumber="17">Fig. 8</figureCitation>
h). Female abdominal sternite VIII with posterior margin rounded, bearing 5 pairs of main setae, and many additional short setae and pores (
<figureCitation id="137010E3FFE98071FE572901FE764B17" box="[412,483,299,325]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="16" pageNumber="17">
Fig.
<date id="FFF52AA6FFE98071FE182901FE764B17" box="[467,483,299,325]" pageId="16" pageNumber="17">8</date>
</figureCitation>
i).
</paragraph>
<caption id="DF345CEEFFE98071FF5C2FBCFC704DA6" httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="16" pageNumber="17" targetBox="[163,1432,377,1919]" targetPageId="16">
<paragraph id="8BF40C66FFE98071FF5C2FBCFC704DA6" blockId="16.[151,1436,1942,2036]" pageId="16" pageNumber="17">
<emphasis id="B93FD074FFE98071FF5C2FBCFE8D4DFC" bold="true" box="[151,280,1942,1966]" pageId="16" pageNumber="17">FIGURE 8.</emphasis>
Details of
<taxonomicName id="4C4B77E5FFE98071FE452FBCFC9D4DFF" authority="Casey" authorityName="Casey" box="[398,776,1942,1965]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pusio" subGenus="Dysleptusa">
<emphasis id="B93FD074FFE98071FE452FBCFE774DFF" box="[398,482,1942,1965]" italics="true" pageId="16" pageNumber="17">Leptusa</emphasis>
(
<emphasis id="B93FD074FFE98071FE392FBCFDFD4DFF" box="[498,616,1942,1965]" italics="true" pageId="16" pageNumber="17">Dysleptusa</emphasis>
)
<emphasis id="B93FD074FFE98071FDB32FBCFD274DFF" box="[632,690,1942,1965]" italics="true" pageId="16" pageNumber="17">pusio</emphasis>
(Casey)
</taxonomicName>
. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BF40C66FFE88070FF0D28B2FB284AE0" blockId="17.[151,1436,152,476]" box="[198,1213,152,178]" pageId="17" pageNumber="18">
<emphasis id="B93FD074FFE88070FF0D28B2FEA04AE3" box="[198,309,152,177]" italics="true" pageId="17" pageNumber="18">Genitalia</emphasis>
. Paramere as in
<figureCitation id="137010E3FFE88070FE3C28B2FDA94AE0" box="[503,572,152,178]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
j. Median lobe as in
<figureCitation id="137010E3FFE88070FCE028B2FCFB4AE0" box="[811,878,152,178]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
k. Spermatheca as in
<figureCitation id="137010E3FFE88070FBA228B2FB3A4AE0" box="[1129,1199,152,178]" captionStart="FIGURE 8" captionStartId="16.[151,255,1942,1966]" captionTargetBox="[163,1432,377,1919]" captionTargetId="figure@16.[151,1436,365,1923]" captionTargetPageId="16" captionText="FIGURE 8. Details of Leptusa (Dysleptusa) pusio (Casey). a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199060/files/figure.png" pageId="17" pageNumber="18">
Fig.
<specimenCount id="9D4DC7EFFFE88070FB5428B2FB3A4AE0" box="[1183,1199,152,178]" pageId="17" pageNumber="18" type="generic">8</specimenCount>
</figureCitation>
l.
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFE88070FF0D2897FDDF4A85" box="[198,586,189,215]" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BF40C66FFE88070FF0D2897FDDF4A85" blockId="17.[151,1436,152,476]" box="[198,586,189,215]" pageId="17" pageNumber="18">
<emphasis id="B93FD074FFE88070FF0D2897FEF94A85" bold="true" box="[198,364,189,215]" pageId="17" pageNumber="18">
<typeStatus id="54F0B2C4FFE88070FF0D2897FE914A85" box="[198,260,189,215]" pageId="17" pageNumber="18">Type</typeStatus>
locality.
</emphasis>
Ohio (Cincinnati).
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFE88070FF0D28C8FB5C4B3E" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BF40C66FFE88070FF0D28C8FF7F4B70" blockId="17.[151,1436,152,476]" pageId="17" pageNumber="18">
<emphasis id="B93FD074FFE88070FF0D28C8FEF24AAE" bold="true" box="[198,359,226,252]" pageId="17" pageNumber="18">Distribution.</emphasis>
<collectingCountry id="F35C4CF6FFE88070FEA528C8FE384AAE" box="[366,429,226,252]" name="United States of America" pageId="17" pageNumber="18">USA</collectingCountry>
: Ohio, Tennessee. Locations where
<taxonomicName id="4C4B77E5FFE88070FC9C28C8FC7A4AA9" box="[855,1007,226,252]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pusio" subGenus="Dysleptusa">
<emphasis id="B93FD074FFE88070FC9C28C8FCFA4AA9" box="[855,879,226,251]" italics="true" pageId="17" pageNumber="18">L.</emphasis>
(
<emphasis id="B93FD074FFE88070FC4A28C8FC0B4AA9" box="[897,926,226,251]" italics="true" pageId="17" pageNumber="18">D.</emphasis>
)
<emphasis id="B93FD074FFE88070FC6428C8FC7A4AA9" box="[943,1007,226,251]" italics="true" pageId="17" pageNumber="18">pusio</emphasis>
</taxonomicName>
has been collected in GSMNP as in Map 1.
</paragraph>
<paragraph id="8BF40C66FFE88070FF0D2907FB5C4B3E" blockId="17.[151,1436,152,476]" pageId="17" pageNumber="18">
<emphasis id="B93FD074FFE88070FF0D2907FEBB4B15" bold="true" box="[198,302,301,327]" pageId="17" pageNumber="18">Habitat.</emphasis>
Specimens examined were collected from primary and secondary forest in GSMNP. Specimens were collected by sifting leaf litter and emergent from CWD decay classes I through III
<emphasis id="B93FD074FFE88070FB5C2978FB334B39" box="[1175,1190,338,363]" italics="true" pageId="17" pageNumber="18"></emphasis>
IV.
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFE88070FF0D295DFCBB4B8E" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BF40C66FFE88070FF0D295DFCBB4B8E" blockId="17.[151,1436,152,476]" pageId="17" pageNumber="18">
<emphasis id="B93FD074FFE88070FF0D295DFEC04BC3" bold="true" box="[198,341,375,401]" pageId="17" pageNumber="18">Comments.</emphasis>
Our conclusion that these specimens are conspecific with
<taxonomicName id="4C4B77E5FFE88070FBD32952FBEC4BC3" box="[1048,1145,376,401]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="B93FD074FFE88070FBD32952FBEC4BC3" box="[1048,1145,376,401]" italics="true" pageId="17" pageNumber="18">L. pusio</emphasis>
</taxonomicName>
is based on comparison with the genitalia, as illustrated by Gusarov (2003: figs 157-160). He designated the
<typeStatus id="54F0B2C4FFE88070FBB029B7FB724BE5" box="[1147,1255,413,439]" pageId="17" pageNumber="18" type="lectotype">lectotype</typeStatus>
of this species, and the illustrations were drawn using the
<typeStatus id="54F0B2C4FFE88070FD4329E8FCBF4B8E" box="[648,810,450,476]" pageId="17" pageNumber="18" type="paralectotype">paralectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>