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<document id="DC2863CE4D013EA635D7299BA3DA1B7B" ID-CLB-Dataset="25362" ID-DOI="10.11646/zootaxa.4674.5.6" ID-GBIF-Dataset="534da4e0-2064-4098-93c2-1f614aef9d2e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3465101" ID-ZooBank="250CE8E7-B7B5-4503-873C-5E79827F6D3C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569830604473" checkinUser="plazi" docAuthor="Shear, William A. &amp; Crawford, Rodney L." docDate="2019" docId="03D17B73FFDF6D73D9CD6FBB811748CA" docLanguage="en" docName="zootaxa.4674.5.6.pdf" docOrigin="Zootaxa 4674 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Opiona catorycha Shear &amp; Crawford 2019, new species" docType="treatment" docVersion="11" lastPageNumber="578" masterDocId="FFE8030BFFDB6D74D95A6D1684364C0D" masterDocTitle="Cave millipedes of the United States. XVI. Two new species from Oregon Caves National Monument, Oregon (Chordeumatida, Conotylidae and Caseyidae)" masterLastPageNumber="580" masterPageNumber="571" pageNumber="575" updateTime="1698756535735" updateUser="ExternalLinkService">
<mods:mods id="372647E5D309FC12085E692401A7332F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="196936AC228330817D0C51908F0CEF52">Cave millipedes of the United States. XVI. Two new species from Oregon Caves National Monument, Oregon (Chordeumatida, Conotylidae and Caseyidae)</mods:title>
</mods:titleInfo>
<mods:name id="5B1C131F16717ECBE2CF31F509066E0C" type="personal">
<mods:role id="AA3463361D9D1296571047120F2B37E7">
<mods:roleTerm id="DA4A3050CB3D0C1F9212AE90D7C7ECC6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F85B621D1D84EA7E107130789D1CAA80">Shear, William A.</mods:namePart>
</mods:name>
<mods:name id="F87FC29D6B09247598A25DBBC0CE67E9" type="personal">
<mods:role id="B11ABCB2D419894BAA2EF09A3D8E68A2">
<mods:roleTerm id="BB685907044DAECDD2271DF04FEBFF02">Author</mods:roleTerm>
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<mods:namePart id="326BE560741D73D61594B28096BA778B">Crawford, Rodney L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="92A562A8E54B69043FBD4BC6EA7837B4">text</mods:typeOfResource>
<mods:relatedItem id="2E3A4E52885DC37FE7ADB7FBD290D012" type="host">
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<mods:title id="BB6BBB651EC3FDD403C353D1A0126FB2">Zootaxa</mods:title>
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<mods:part id="A1745B1A27C8A6A11C6834CE63853275">
<mods:date id="F1B028522C6A642029F4F86288E8FBB3">2019</mods:date>
<mods:detail id="6AE136DAFE6996B1C1F14629B2354C67" type="pubDate">
<mods:number id="6D38DC274F9D7BD796DC83DA1029C77F">2019-09-27</mods:number>
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<mods:detail id="9AF4BCD524496115A3026413177F4896" type="volume">
<mods:number id="572F830154CCD0E8215432DF4F504F11">4674</mods:number>
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<mods:number id="D0136701237532F68344E2C7066E768D">5</mods:number>
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<mods:classification id="17F95DFB0F9EC2A2AEE41FEE8D3FE845">journal article</mods:classification>
<mods:identifier id="4B9D4D3ED49190446C3E3A4C3E436EA4" type="CLB-Dataset">25362</mods:identifier>
<mods:identifier id="172386022AEF95708A222C69E0771BF3" type="DOI">10.11646/zootaxa.4674.5.6</mods:identifier>
<mods:identifier id="B2F0707558AE45A1265AA0EE2E6A11E2" type="GBIF-Dataset">534da4e0-2064-4098-93c2-1f614aef9d2e</mods:identifier>
<mods:identifier id="D70072208CDC26476F22F945C97735A0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="150E56FB8E7B8AC70410A0C6125FC36E" type="Zenodo-Dep">3465101</mods:identifier>
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<treatment id="03D17B73FFDF6D73D9CD6FBB811748CA" ID-DOI="http://doi.org/10.5281/zenodo.5940781" ID-GBIF-Taxon="159609126" ID-Zenodo-Dep="5940781" LSID="urn:lsid:plazi:treatment:03D17B73FFDF6D73D9CD6FBB811748CA" httpUri="http://treatment.plazi.org/id/03D17B73FFDF6D73D9CD6FBB811748CA" lastPageId="7" lastPageNumber="578" pageId="4" pageNumber="575">
<subSubSection id="C36299EEFFDF6D70D9CD6FBB87374ECA" box="[151,769,685,712]" pageId="4" pageNumber="575" type="nomenclature">
<paragraph id="8BC7CA65FFDF6D70D9CD6FBB87374ECA" blockId="4.[151,769,685,747]" box="[151,769,685,712]" pageId="4" pageNumber="575">
<heading id="D08F7D09FFDF6D70D9CD6FBB87374ECA" bold="true" box="[151,769,685,712]" fontSize="11" level="1" pageId="4" pageNumber="575" reason="1">
<emphasis id="B90C1677FFDF6D70D9CD6FBB87374ECA" bold="true" box="[151,769,685,712]" pageId="4" pageNumber="575">
<taxonomicName id="4C78B1E6FFDF6D70D9CD6FBB86544ECA" authority="Shear &amp; Crawford" authorityName="Shear &amp; Crawford" authorityYear="2019" box="[151,610,685,712]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="575" phylum="Arthropoda" rank="species" species="catorycha" status="sp. nov.">
<emphasis id="B90C1677FFDF6D70D9CD6FBB85594EC5" bold="true" box="[151,367,685,712]" italics="true" pageId="4" pageNumber="575">Opiona catorycha</emphasis>
Shear &amp; Crawford
</taxonomicName>
,
<taxonomicNameLabel id="A23FAB0CFFDF6D70DB2B6FB887374ECA" box="[625,769,685,712]" pageId="4" pageNumber="575" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFDF6D70D9CD6FC785384EE6" box="[151,270,721,747]" pageId="4" pageNumber="575" type="description">
<paragraph id="8BC7CA65FFDF6D70D9CD6FC785384EE6" blockId="4.[151,769,685,747]" box="[151,270,721,747]" pageId="4" pageNumber="575">
<figureCitation id="1343D6E0FFDF6D70D9CD6FC785384EE6" box="[151,270,721,747]" captionStart-0="FIGURE 6" captionStart-1="FIGURES 7–10" captionStart-2="FIGURES 11, 12" captionStart-3="FIGURES 13, 14" captionStartId-0="4.[151,250,1978,2003]" captionStartId-1="5.[151,264,1297,1322]" captionStartId-2="6.[151,264,1931,1956]" captionStartId-3="7.[151,264,1815,1840]" captionTargetBox-0="[170,1407,1014,1945]" captionTargetBox-1="[152,1436,174,1278]" captionTargetBox-2="[167,1422,166,1902]" captionTargetBox-3="[152,1437,1238,1795]" captionTargetId-0="figure@4.[170,1416,1014,1953]" captionTargetId-1="figure@5.[151,1436,174,1278]" captionTargetId-2="figure@6.[165,1422,166,1911]" captionTargetId-3="figure@7.[151,1437,1237,1795]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 6. Opiona catorycha, new species. Freshly preserved male from No Name Cave, lateral view; g, gonopod; L1–L3, legs 1–3, respectively; R4, ring 4; t9, telopodite of leg 9; tL3, telopodite of leg 3 (small object just to right of label)." captionText-1="FIGURES 7–10. Opiona catorycha, new species. Fig. 7. Head and collum, frontal view. Fig. 8. Legpair 1, anterior view. Fig. 9. Legpairs 2 and 3, anterior view; cx3, coxa of leg 3; pf3, prefemur of leg 3; S3, sternum of legpair 3; sp2, spermatic process of coxa 2; t2, telopodite of leg 2; t3, telopodite of leg 3. Fig. 10. Tips of legpair 3 coxae, anterior view." captionText-2="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." captionText-3="FIGURES 13, 14. Opiona catorycha, new species. Fig. 13. Left leg 9, anterior view. Fig. 14. Legpair 10, anterior view." figureDoi-0="http://doi.org/10.5281/zenodo.3467972" figureDoi-1="http://doi.org/10.5281/zenodo.3467974" figureDoi-2="http://doi.org/10.5281/zenodo.3467976" figureDoi-3="http://doi.org/10.5281/zenodo.3467978" httpUri-0="https://zenodo.org/record/3467972/files/figure.png" httpUri-1="https://zenodo.org/record/3467974/files/figure.png" httpUri-2="https://zenodo.org/record/3467976/files/figure.png" httpUri-3="https://zenodo.org/record/3467978/files/figure.png" pageId="4" pageNumber="575">Figs. 6–14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFDF6D70D9CD6E0480CD4FEC" pageId="4" pageNumber="575" type="materials_examined">
<paragraph id="8BC7CA65FFDF6D70D9CD6E0480CD4FEC" blockId="4.[151,1437,786,993]" pageId="4" pageNumber="575">
<emphasis id="B90C1677FFDF6D70D9CD6E04854F4F20" bold="true" box="[151,377,786,813]" pageId="4" pageNumber="575">Material examined</emphasis>
:
<materialsCitation id="3B10C038FFDF6D70D8DD6E0585604F94" ID-GBIF-Occurrence="2423596450" collectingDate="1992-08-31" collectingDateMax="2018-11-11" collectingDateMin="1992-08-31" collectionCode="UWBM" county="Josephine Co." latitude="42.288" location="Oregon Cave, Oregon Caves National Monument" pageId="4" pageNumber="575" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oregon" typeStatus="holotype">
Type
<specimenCount id="9D7E01ECFFDF6D70D89F6E05860C4F20" box="[453,570,787,813]" pageId="4" pageNumber="575" type="generic">specimens</specimenCount>
.
<specimenCount id="9D7E01ECFFDF6D70DB126E0586B44F20" box="[584,642,787,813]" pageId="4" pageNumber="575" type="male">Male</specimenCount>
<typeStatus id="54C374C7FFDF6D70DBD26E0586DC4F20" box="[648,746,787,813]" pageId="4" pageNumber="575" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED6952A0FFDF6D70DBA26E0587604F20" box="[760,854,787,813]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="4" pageNumber="575" type="Museum">UWBM</collectionCode>
) from
<location id="8EA79CBEFFDF6D70DAFB6E0584E44F5C" LSID="urn:lsid:plazi:treatment:03D17B73FFDF6D73D9CD6FBB811748CA:8EA79CBEFFDF6D70DAFB6E0584E44F5C" county="Josephine Co." latitude="42.288" name="Oregon Cave, Oregon Caves National Monument" pageId="4" pageNumber="575" stateProvince="Oregon">Oregon Cave, Oregon Caves National Monument</location>
,
<collectingCounty id="62A6B2E9FFDF6D70D9876E21854B4F5C" box="[221,381,823,849]" pageId="4" pageNumber="575">Josephine Co.</collectingCounty>
,
<collectingRegion id="49BC0487FFDF6D70D8D36E2185EB4F5C" box="[393,477,823,849]" country="United States of America" name="Oregon" pageId="4" pageNumber="575">Oregon</collectingRegion>
(
<geoCoordinate id="EE4CACA2FFDF6D70D8B66E2186464F5C" box="[492,624,823,849]" degrees="42.0974" direction="north" orientation="latitude" pageId="4" pageNumber="575" precision="5" value="42.0974">42.0974º N</geoCoordinate>
,
<geoCoordinate id="EE4CACA2FFDF6D70DB216E21872D4F5C" box="[635,795,823,849]" pageId="4" pageNumber="575">-123.4099º W</geoCoordinate>
), collected
<date id="FFC6ECA5FFDF6D70DAC66E2180784F5C" box="[924,1102,823,849]" pageId="4" pageNumber="575" value="1992-08-31">
<collectingDate id="EF82154DFFDF6D70DAC66E2180784F5C" box="[924,1102,823,849]" pageId="4" pageNumber="575" value="1992-08-31">31 August 1992</collectingDate>
</date>
by a drift net intercepting the inflow of the main cave stream “River Styx” where it flows from an impenetrable opening into a passage along the cave’s tour route
</materialsCitation>
.
<materialsCitation id="3B10C038FFDF6D70D83B6E6980CD4FEC" ID-GBIF-Occurrence="2423596449" collectingDate="1992-08-31" collectingDateMax="2018-11-11" collectingDateMin="1992-08-31" collectionCode="UWBM, CASC" county="Josephine Co." latitude="42.288" location="River Styx" pageId="4" pageNumber="575" specimenCount="5" specimenCount-female="1" specimenCount-male="2" stateProvince="Oregon" typeStatus="paratype">
<specimenCount id="9D7E01ECFFDF6D70D83B6E6985AD4F94" box="[353,411,895,921]" pageId="4" pageNumber="575" type="male">Male</specimenCount>
and
<specimenCount id="9D7E01ECFFDF6D70D88B6E69862B4F94" box="[465,541,895,921]" pageId="4" pageNumber="575" type="female">female</specimenCount>
<typeStatus id="54C374C7FFDF6D70DB7F6E6986A64F94" box="[549,656,895,921]" pageId="4" pageNumber="575" type="paratype">paratypes</typeStatus>
from
<location id="8EA79CBEFFDF6D70DB8F6E6987B34F94" LSID="urn:lsid:plazi:treatment:03D17B73FFDF6D73D9CD6FBB811748CA:8EA79CBEFFDF6D70DB8F6E6987B34F94" box="[725,901,895,921]" county="Josephine Co." latitude="42.288" name="No Name Cave" pageId="4" pageNumber="575" stateProvince="Oregon">No Name Cave</location>
,
<quantity id="4C806780FFDF6D70DACB6E6987DB4F94" box="[913,1005,895,921]" metricMagnitude="4" metricUnit="m" metricValue="1.78" pageId="4" pageNumber="575" unit="km" value="17.8">
<locationDeviation id="9A75D347FFDF6D70DACB6E6987DB4F94" box="[913,1005,895,921]" metricMagnitude="4" metricUnit="m" metricValue="1.78" pageId="4" pageNumber="575" unit="km" value="17.8">17.8 km</locationDeviation>
</quantity>
south-southwest of
<location id="8EA79CBEFFDF6D70DD8A6E6981604F94" LSID="urn:lsid:plazi:treatment:03D17B73FFDF6D73D9CD6FBB811748CA:8EA79CBEFFDF6D70DD8A6E6981604F94" box="[1232,1366,895,921]" county="Josephine Co." latitude="42.288" name="Grants Pass" pageId="4" pageNumber="575" stateProvince="Oregon">Grants Pass</location>
,
<collectingCounty id="62A6B2E9FFDF6D70DC396E6985304FB0" pageId="4" pageNumber="575">Josephine Co.</collectingCounty>
,
<collectingRegion id="49BC0487FFDF6D70D8486EB585504FB0" box="[274,358,931,957]" country="United States of America" name="Oregon" pageId="4" pageNumber="575">Oregon</collectingRegion>
(
<geoCoordinate id="EE4CACA2FFDF6D70D82F6EB585C94FB0" box="[373,511,931,957]" degrees="42.2880" direction="north" orientation="latitude" pageId="4" pageNumber="575" precision="5" value="42.288">42.2880 º N</geoCoordinate>
,
<geoCoordinate id="EE4CACA2FFDF6D70DB516EB486874FB0" box="[523,689,930,957]" pageId="4" pageNumber="575">-123.2519 º W</geoCoordinate>
), collected
<date id="FFC6ECA5FFDF6D70DA6B6EB5803D4FB0" box="[817,1035,931,957]" pageId="4" pageNumber="575" value="2018-11-11">
<collectingDate id="EF82154DFFDF6D70DA6B6EB5803D4FB0" box="[817,1035,931,957]" pageId="4" pageNumber="575" value="2018-11-11">11 November 2018</collectingDate>
</date>
, by
<location id="8EA79CBEFFDF6D70DD636EB581344FB0" LSID="urn:lsid:plazi:treatment:03D17B73FFDF6D73D9CD6FBB811748CA:8EA79CBEFFDF6D70DD636EB581344FB0" box="[1081,1282,931,957]" county="Josephine Co." latitude="42.288" name="Neil Marchington" pageId="4" pageNumber="575" stateProvince="Oregon">Neil Marchington</location>
and
<date id="FFC6ECA5FFDF6D70DC626EB584E74FEC" pageId="4" pageNumber="575" value="2011-08-08">
<collectingDate id="EF82154DFFDF6D70DC626EB584E74FEC" pageId="4" pageNumber="575" value="2011-08-08">8 August 2011</collectingDate>
</date>
, by
<location id="8EA79CBEFFDF6D70D9A56ED185AF4FEC" LSID="urn:lsid:plazi:treatment:03D17B73FFDF6D73D9CD6FBB811748CA:8EA79CBEFFDF6D70D9A56ED185AF4FEC" box="[255,409,967,993]" county="Josephine Co." latitude="42.288" name="Darrell Ubick" pageId="4" pageNumber="575" stateProvince="Oregon">Darrell Ubick</location>
(
<location id="8EA79CBEFFDF6D70D8F26ED185D84FEC" LSID="urn:lsid:plazi:treatment:03D17B73FFDF6D73D9CD6FBB811748CA:8EA79CBEFFDF6D70D8F26ED185D84FEC" box="[424,494,967,993]" county="Josephine Co." latitude="42.288" name="Ubick" pageId="4" pageNumber="575" stateProvince="Oregon">Ubick</location>
<specimenCount id="9D7E01ECFFDF6D70D8AC6ED1865D4FEC" box="[502,619,967,993]" pageId="4" pageNumber="575" type="generic">specimens</specimenCount>
deposited in the
<collectingRegion id="49BC0487FFDF6D70DA706ED187AA4FEC" box="[810,924,967,993]" country="United States of America" name="California" pageId="4" pageNumber="575">California</collectingRegion>
<location id="8EA79CBEFFDF6D70DAFB6ED180A04FEC" LSID="urn:lsid:plazi:treatment:03D17B73FFDF6D73D9CD6FBB811748CA:8EA79CBEFFDF6D70DAFB6ED180A04FEC" box="[929,1174,967,993]" county="Josephine Co." latitude="42.288" name="Academy of Sciences" pageId="4" pageNumber="575" stateProvince="Oregon">Academy of Sciences</location>
,
<collectionCode id="ED6952A0FFDF6D70DDFB6ED180D94FEC" box="[1185,1263,967,993]" pageId="4" pageNumber="575">CASC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF079AEDFFDF6D70D9CD6AAC81074BFA" ID-DOI="http://doi.org/10.5281/zenodo.3467972" ID-Zenodo-Dep="3467972" httpUri="https://zenodo.org/record/3467972/files/figure.png" pageId="4" pageNumber="575" startId="4.[151,250,1978,2003]" targetBox="[170,1407,1014,1945]" targetPageId="4">
<paragraph id="8BC7CA65FFDF6D70D9CD6AAC81074BFA" blockId="4.[151,1436,1978,2039]" pageId="4" pageNumber="575">
<emphasis id="B90C1677FFDF6D70D9CD6AAC85224BDE" bold="true" box="[151,276,1978,2003]" pageId="4" pageNumber="575">FIGURE 6.</emphasis>
<emphasis id="B90C1677FFDF6D70D8466AAA85ED4BDE" box="[284,475,1979,2003]" italics="true" pageId="4" pageNumber="575">
<taxonomicName id="4C78B1E6FFDF6D70D8466AAA85E14BDE" authority="Shear &amp; Crawford, 2019" authorityName="Shear &amp; Crawford" authorityYear="2019" box="[284,471,1979,2003]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="575" phylum="Arthropoda" rank="species" species="catorycha" status="sp. nov.">Opiona catorycha</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A23FAB0CFFDF6D70D8B86AAA86564BDE" box="[482,608,1978,2003]" pageId="4" pageNumber="575" rank="species">new species</taxonomicNameLabel>
. Freshly preserved male from No Name Cave, lateral view;
<emphasis id="B90C1677FFDF6D70DD966AAD80EF4BDE" bold="true" box="[1228,1241,1979,2003]" pageId="4" pageNumber="575">g</emphasis>
, gonopod;
<emphasis id="B90C1677FFDF6D70DC146AAC81A04BDE" bold="true" box="[1358,1430,1978,2003]" pageId="4" pageNumber="575">L1–L3</emphasis>
, legs 1–3, respectively;
<emphasis id="B90C1677FFDF6D70D8DB6AC885A94BFA" bold="true" box="[385,415,2014,2039]" pageId="4" pageNumber="575">R4</emphasis>
, ring 4;
<emphasis id="B90C1677FFDF6D70D8AC6AC886274BFA" bold="true" box="[502,529,2014,2039]" pageId="4" pageNumber="575">t9,</emphasis>
telopodite of leg 9;
<emphasis id="B90C1677FFDF6D70DB846AC8873C4BFA" bold="true" box="[734,778,2014,2039]" pageId="4" pageNumber="575">tL3,</emphasis>
telopodite of leg 3 (small object just to right of label).
</paragraph>
</caption>
<caption id="DF079AEDFFDE6D71D9CD6807809E497F" ID-DOI="http://doi.org/10.5281/zenodo.3467974" ID-Zenodo-Dep="3467974" httpUri="https://zenodo.org/record/3467974/files/figure.png" pageId="5" pageNumber="576" startId="5.[151,264,1297,1322]" targetBox="[152,1436,174,1278]" targetPageId="5">
<paragraph id="8BC7CA65FFDE6D71D9CD6807809E497F" blockId="5.[151,1437,1297,1394]" pageId="5" pageNumber="576">
<emphasis id="B90C1677FFDE6D71D9CD6807857E4927" bold="true" box="[151,328,1297,1322]" pageId="5" pageNumber="576">FIGURES 7–10.</emphasis>
<emphasis id="B90C1677FFDE6D71D815680486384927" box="[335,526,1298,1322]" italics="true" pageId="5" pageNumber="576">
<taxonomicName id="4C78B1E6FFDE6D71D8156804863C4927" authority="Shear &amp; Crawford, 2019" authorityName="Shear &amp; Crawford" authorityYear="2019" box="[335,522,1298,1322]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="catorycha" status="sp. nov.">Opiona catorycha</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A23FAB0CFFDE6D71DB4E680486A44927" box="[532,658,1297,1322]" pageId="5" pageNumber="576" rank="species">new species</taxonomicNameLabel>
. Fig. 7. Head and collum, frontal view. Fig. 8. Legpair 1, anterior view. Fig. 9. Legpairs 2 and 3, anterior view;
<emphasis id="B90C1677FFDE6D71D8A76823861E4943" bold="true" box="[509,552,1333,1358]" pageId="5" pageNumber="576">cx3,</emphasis>
coxa of leg 3;
<emphasis id="B90C1677FFDE6D71DB98682386D34943" bold="true" box="[706,741,1333,1358]" pageId="5" pageNumber="576">pf3</emphasis>
, prefemur of leg 3;
<emphasis id="B90C1677FFDE6D71DAE8682387FA4943" bold="true" box="[946,972,1333,1358]" pageId="5" pageNumber="576">S3</emphasis>
, sternum of legpair 3;
<emphasis id="B90C1677FFDE6D71DDEF682380D64943" bold="true" box="[1205,1248,1333,1358]" pageId="5" pageNumber="576">sp2,</emphasis>
spermatic process of coxa 2;
<emphasis id="B90C1677FFDE6D71D859684F852E497F" bold="true" box="[259,280,1369,1394]" pageId="5" pageNumber="576">t2</emphasis>
, telopodite of leg 2;
<emphasis id="B90C1677FFDE6D71D8B1684F8637497F" bold="true" box="[491,513,1369,1394]" pageId="5" pageNumber="576">t3</emphasis>
, telopodite of leg 3. Fig. 10. Tips of legpair 3 coxae, anterior view.
</paragraph>
</caption>
<subSubSection id="C36299EEFFDE6D71D99D68B686B54A0F" pageId="5" pageNumber="576" type="diagnosis">
<paragraph id="8BC7CA65FFDE6D71D99D68B686B54A0F" blockId="5.[151,1437,1440,2043]" pageId="5" pageNumber="576">
<emphasis id="B90C1677FFDE6D71D99D68B6850C49B6" bold="true" box="[199,314,1440,1467]" pageId="5" pageNumber="576">Diagnosis</emphasis>
.
<taxonomicName id="4C78B1E6FFDE6D71D81D68B7862449B7" authorityName="Shear &amp; Crawford" authorityYear="2019" box="[327,530,1440,1467]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="catorycha">
<emphasis id="B90C1677FFDE6D71D81D68B7862449B7" box="[327,530,1440,1467]" italics="true" pageId="5" pageNumber="576">Opiona catorycha</emphasis>
</taxonomicName>
can be confused with no other
<taxonomicName id="4C78B1E6FFDE6D71DA3068B7878849B6" authorityName="Chamberlin" authorityYear="1951" box="[874,958,1441,1467]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="576" phylum="Arthropoda" rank="genus">
<emphasis id="B90C1677FFDE6D71DA3068B7878849B6" box="[874,958,1441,1467]" italics="true" pageId="5" pageNumber="576">Opiona</emphasis>
</taxonomicName>
species, due to the extraordinary complexity of the anterior gonopods, featuring numerous flagelliform branches of at least five distinct
<typeStatus id="54C374C7FFDE6D71DDE968D380D849D2" box="[1203,1262,1477,1503]" pageId="5" pageNumber="576">types</typeStatus>
combined with its troglomorphic adaptations (
<figureCitation id="1343D6E0FFDE6D71D8B568FE86404A0F" box="[495,630,1512,1538]" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="5" pageNumber="576">Figs. 11, 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFDE6D73D99D6B1A81A74D86" lastPageId="7" lastPageNumber="578" pageId="5" pageNumber="576" type="description">
<paragraph id="8BC7CA65FFDE6D73D99D6B1A86404D4E" blockId="5.[151,1437,1440,2043]" lastBlockId="7.[151,1437,153,1224]" lastPageId="7" lastPageNumber="578" pageId="5" pageNumber="576">
<emphasis id="B90C1677FFDE6D71D99D6B1A85674A2A" bold="true" box="[199,337,1548,1575]" pageId="5" pageNumber="576">Description</emphasis>
. Male
<typeStatus id="54C374C7FFDE6D71D8FD6B1A863B4A2B" box="[423,525,1548,1574]" pageId="5" pageNumber="576" type="holotype">holotype</typeStatus>
. Length about
<quantity id="4C806780FFDE6D71DBE76B1A87244A2A" box="[701,786,1548,1575]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="576" unit="mm" value="13.0">13 mm</quantity>
, width 1.0 mm. Eyes absent (
<figureCitation id="1343D6E0FFDE6D71DD2B6B1A808F4A2A" box="[1137,1209,1548,1575]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1297,1322]" captionTargetBox="[152,1436,174,1278]" captionTargetId="figure@5.[151,1436,174,1278]" captionTargetPageId="5" captionText="FIGURES 7–10. Opiona catorycha, new species. Fig. 7. Head and collum, frontal view. Fig. 8. Legpair 1, anterior view. Fig. 9. Legpairs 2 and 3, anterior view; cx3, coxa of leg 3; pf3, prefemur of leg 3; S3, sternum of legpair 3; sp2, spermatic process of coxa 2; t2, telopodite of leg 2; t3, telopodite of leg 3. Fig. 10. Tips of legpair 3 coxae, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467974" httpUri="https://zenodo.org/record/3467974/files/figure.png" pageId="5" pageNumber="576">Fig. 7</figureCitation>
). Rings cylindrical. Metazonital setae short, slightly clavate. Fourth ring enlarged, trunk flexed at fourth ring (
<figureCitation id="1343D6E0FFDE6D71DDFF6B2680DA4A46" box="[1189,1260,1584,1611]" captionStart="FIGURE 6" captionStartId="4.[151,250,1978,2003]" captionTargetBox="[170,1407,1014,1945]" captionTargetId="figure@4.[170,1416,1014,1953]" captionTargetPageId="4" captionText="FIGURE 6. Opiona catorycha, new species. Freshly preserved male from No Name Cave, lateral view; g, gonopod; L1–L3, legs 1–3, respectively; R4, ring 4; t9, telopodite of leg 9; tL3, telopodite of leg 3 (small object just to right of label)." figureDoi="http://doi.org/10.5281/zenodo.3467972" httpUri="https://zenodo.org/record/3467972/files/figure.png" pageId="5" pageNumber="576">Fig. 6</figureCitation>
). Unpigmented (both preserved and when alive). First legpair incrassate, podomeres with distinctive ensiform setae ventrally, coxae with large posterior lobes bearing five or six peg-like setae (
<figureCitation id="1343D6E0FFDE6D71DA6B6B6E87434A9F" box="[817,885,1656,1682]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1297,1322]" captionTargetBox="[152,1436,174,1278]" captionTargetId="figure@5.[151,1436,174,1278]" captionTargetPageId="5" captionText="FIGURES 7–10. Opiona catorycha, new species. Fig. 7. Head and collum, frontal view. Fig. 8. Legpair 1, anterior view. Fig. 9. Legpairs 2 and 3, anterior view; cx3, coxa of leg 3; pf3, prefemur of leg 3; S3, sternum of legpair 3; sp2, spermatic process of coxa 2; t2, telopodite of leg 2; t3, telopodite of leg 3. Fig. 10. Tips of legpair 3 coxae, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467974" httpUri="https://zenodo.org/record/3467974/files/figure.png" pageId="5" pageNumber="576">Fig. 8</figureCitation>
). Second legpair reduced to six podomeres, coxal gonapophyses about as long as prefemora, with fine, tightly appressed setae; distal three podomeres with curled setae (
<figureCitation id="1343D6E0FFDE6D71D99D6BD6853C4AD7" box="[199,266,1728,1754]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1297,1322]" captionTargetBox="[152,1436,174,1278]" captionTargetId="figure@5.[151,1436,174,1278]" captionTargetPageId="5" captionText="FIGURES 7–10. Opiona catorycha, new species. Fig. 7. Head and collum, frontal view. Fig. 8. Legpair 1, anterior view. Fig. 9. Legpairs 2 and 3, anterior view; cx3, coxa of leg 3; pf3, prefemur of leg 3; S3, sternum of legpair 3; sp2, spermatic process of coxa 2; t2, telopodite of leg 2; t3, telopodite of leg 3. Fig. 10. Tips of legpair 3 coxae, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467974" httpUri="https://zenodo.org/record/3467974/files/figure.png" pageId="5" pageNumber="576">Fig. 9</figureCitation>
). Third legpair with coxae greatly extended ventrally into subclavate lobes with apical tufts of setae, prefemora greatly shortened, flattened, ventrolaterally expanded; femora, postfemora, tibiae and tarsi much reduced, and tarsus without claw (
<figureCitation id="1343D6E0FFDE6D71D8EF6A1E86074B2F" box="[437,561,1800,1826]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1297,1322]" captionTargetBox="[152,1436,174,1278]" captionTargetId="figure@5.[151,1436,174,1278]" captionTargetPageId="5" captionText="FIGURES 7–10. Opiona catorycha, new species. Fig. 7. Head and collum, frontal view. Fig. 8. Legpair 1, anterior view. Fig. 9. Legpairs 2 and 3, anterior view; cx3, coxa of leg 3; pf3, prefemur of leg 3; S3, sternum of legpair 3; sp2, spermatic process of coxa 2; t2, telopodite of leg 2; t3, telopodite of leg 3. Fig. 10. Tips of legpair 3 coxae, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467974" httpUri="https://zenodo.org/record/3467974/files/figure.png" pageId="5" pageNumber="576">Figs. 9, 10</figureCitation>
). Anterior gonopods (
<figureCitation id="1343D6E0FFDE6D71DA706A1E87864B2F" box="[810,944,1800,1826]" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="5" pageNumber="576">Figs. 11–12</figureCitation>
) extraordinarily complex, sternum (
<emphasis id="B90C1677FFDE6D71DC116A1F816A4B2E" bold="true" box="[1355,1372,1801,1827]" pageId="5" pageNumber="576">s,</emphasis>
<figureCitation id="1343D6E0FFDE6D71DC3F6A1E84E94B4B" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="5" pageNumber="576">Figs. 11, 12</figureCitation>
) well sclerotized anteriorly and posteriorly, with sigmoidially curved, fimbriate lateral processes. Anterior angiocoxite (
<emphasis id="B90C1677FFDE6D71D8706A47856C4B66" bold="true" box="[298,346,1873,1899]" pageId="5" pageNumber="576">aac,</emphasis>
<figureCitation id="1343D6E0FFDE6D71D83A6A4685D24B67" box="[352,484,1872,1898]" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="5" pageNumber="576">Figs. 11, 12</figureCitation>
) in anterior view with two relatively broader branches: the lateralmost shorter, acute; the more medial longer, broader, with subapical tooth—medial to these is a series of curved, flattened, blade-like pseudoflagella (
<emphasis id="B90C1677FFDE6D71D8146A8E85584BBE" bold="true" box="[334,366,1944,1971]" pageId="5" pageNumber="576">pf,</emphasis>
<figureCitation id="1343D6E0FFDE6D71D8236A8E86304BBF" box="[377,518,1944,1970]" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="5" pageNumber="576">Figs. 11, 12</figureCitation>
), and more medial still a few fine, fimbriate pseudoflagella. In posterior view, additional pseudoflagella are seen, these with knobbed tips, knobs set with fine filaments (
<figureCitation id="1343D6E0FFDE6D71DDF06AAA81364BDB" box="[1194,1280,1980,2006]" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="5" pageNumber="576">Fig. 12</figureCitation>
). Near lateral base of angiocoxite on each side, a single thin filament (
<emphasis id="B90C1677FFDE6D71DA4E6AF687124BF6" bold="true" box="[788,804,2016,2043]" pageId="5" pageNumber="576">f,</emphasis>
<figureCitation id="1343D6E0FFDE6D71DA766AF6874A4BF7" box="[812,892,2016,2042]" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="5" pageNumber="576">Fig. 12</figureCitation>
) with fine branches. Posterior angiocoxite (
<emphasis id="B90C1677FFDE6D71DC316AF781AB4BF6" bold="true" box="[1387,1437,2017,2043]" pageId="5" pageNumber="576">pac,</emphasis>
<figureCitation id="1343D6E0FFDC6D73D9CD6D8F84DE4CBE" box="[151,232,153,179]" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="7" pageNumber="578">Fig. 12</figureCitation>
) with posteriorly curved, fimbriate branch, more anterior, erect, flattened branch; at least 3 strong pseudoflagella with blunt, slightly bifurcate tips arise behind posterior colpocoxite. Colpocoxites (
<emphasis id="B90C1677FFDC6D73DDFD6DAB80F14CDA" bold="true" box="[1191,1223,189,215]" pageId="7" pageNumber="578">cc,</emphasis>
<figureCitation id="1343D6E0FFDC6D73DD956DAB81174CDA" box="[1231,1313,189,215]" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="7" pageNumber="578">Fig. 12</figureCitation>
) lobe-like, poorly sclerotized. Ninth legpair with three-branched coxal processes, branches curved, acute. Telopodite article typically swollen, flattened, cordate in lateral view (
<figureCitation id="1343D6E0FFDC6D73DB846C1387074D12" box="[734,817,261,287]" captionStart="FIGURES 13, 14" captionStartId="7.[151,264,1815,1840]" captionTargetBox="[152,1437,1238,1795]" captionTargetId="figure@7.[151,1437,1237,1795]" captionTargetPageId="7" captionText="FIGURES 13, 14. Opiona catorycha, new species. Fig. 13. Left leg 9, anterior view. Fig. 14. Legpair 10, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467978" httpUri="https://zenodo.org/record/3467978/files/figure.png" pageId="7" pageNumber="578">Fig. 13</figureCitation>
). Tenth legpair typical of genus, coxae with large, deep glands, posterior hooked process (
<figureCitation id="1343D6E0FFDC6D73DB4F6C3F86524D4E" box="[533,612,297,323]" captionStart="FIGURES 13, 14" captionStartId="7.[151,264,1815,1840]" captionTargetBox="[152,1437,1238,1795]" captionTargetId="figure@7.[151,1437,1237,1795]" captionTargetPageId="7" captionText="FIGURES 13, 14. Opiona catorycha, new species. Fig. 13. Left leg 9, anterior view. Fig. 14. Legpair 10, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467978" httpUri="https://zenodo.org/record/3467978/files/figure.png" pageId="7" pageNumber="578">Fig 14</figureCitation>
.).
</paragraph>
<caption id="DF079AEDFFDD6D72D9CD6A9D85DF4BE1" ID-DOI="http://doi.org/10.5281/zenodo.3467976" ID-Zenodo-Dep="3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="6" pageNumber="577" startId="6.[151,264,1931,1956]" targetBox="[167,1422,166,1902]" targetPageId="6">
<paragraph id="8BC7CA65FFDD6D72D9CD6A9D85DF4BE1" blockId="6.[151,1437,1931,2028]" pageId="6" pageNumber="577">
<emphasis id="B90C1677FFDD6D72D9CD6A9D85614BA9" bold="true" box="[151,343,1931,1956]" pageId="6" pageNumber="577">FIGURES 11, 12.</emphasis>
<emphasis id="B90C1677FFDD6D72D83A6A9A86164BAE" box="[352,544,1931,1955]" italics="true" pageId="6" pageNumber="577">
<taxonomicName id="4C78B1E6FFDD6D72D83A6A9A862A4BAE" authority="Shear &amp; Crawford, 2019" authorityName="Shear &amp; Crawford" authorityYear="2019" box="[352,540,1931,1955]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="catorycha" status="sp. nov.">Opiona catorycha</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A23FAB0CFFDD6D72DB726A9A86914BA9" box="[552,679,1931,1956]" pageId="6" pageNumber="577" rank="species">new species</taxonomicNameLabel>
. Fig. 11. Gonopods, posterior view (scanning electron micrograph);
<emphasis id="B90C1677FFDD6D72DC2B6A9D81AA4BAE" bold="true" box="[1393,1436,1931,1955]" pageId="6" pageNumber="577">aac,</emphasis>
anterior angiocoxite;
<emphasis id="B90C1677FFDD6D72D8346AB985BD4BCA" bold="true" box="[366,395,1967,1991]" pageId="6" pageNumber="577">cc,</emphasis>
colpocoxite;
<emphasis id="B90C1677FFDD6D72DB496AB986764BCA" bold="true" box="[531,576,1967,1991]" pageId="6" pageNumber="577">pac,</emphasis>
posterior angiocoxite;
<emphasis id="B90C1677FFDD6D72DA726AB9870E4BCA" bold="true" box="[808,824,1967,1991]" pageId="6" pageNumber="577">s,</emphasis>
gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11,
<emphasis id="B90C1677FFDD6D72D9E66AC584EF4BE1" bold="true" box="[188,217,2003,2028]" pageId="6" pageNumber="577">pf,</emphasis>
pseudoflagellar branches..
</paragraph>
</caption>
<paragraph id="8BC7CA65FFDC6D73D99D6C5B81A74D86" blockId="7.[151,1437,153,1224]" pageId="7" pageNumber="578">
Female
<typeStatus id="54C374C7FFDC6D73D87A6C5B85B24D6A" box="[288,388,333,359]" pageId="7" pageNumber="578" type="paratype">paratype</typeStatus>
. Length about
<quantity id="4C806780FFDC6D73DB726C5B86B84D6A" box="[552,654,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="7" pageNumber="578" unit="mm" value="13.2">13.2 mm</quantity>
, width 1.0 mm. Similar to male in nonsexual characters. First and second legpairs reduced. Third legpair with sternum strongly modified, cupped anteriorly, partially enclosing vulvae.
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFDC6D73D99D6C8287C64DFA" pageId="7" pageNumber="578" type="distribution">
<paragraph id="8BC7CA65FFDC6D73D99D6C8287C64DFA" blockId="7.[151,1437,153,1224]" pageId="7" pageNumber="578">
<emphasis id="B90C1677FFDC6D73D99D6C82856F4DA2" bold="true" box="[199,345,404,431]" pageId="7" pageNumber="578">Distribution</emphasis>
. Known only from the type and
<typeStatus id="54C374C7FFDC6D73DB916C83871D4DA2" box="[715,811,405,431]" pageId="7" pageNumber="578" type="paratype">paratype</typeStatus>
localities, but possibly occurring in other marble caves in the region. Numerous small lenses of marble are scattered through the area between
<collectingRegion id="49BC0487FFDC6D73DD3F6CAF808F4DDE" box="[1125,1209,441,467]" country="United States of America" name="Oregon" pageId="7" pageNumber="578">Oregon</collectingRegion>
Cave and No Name Cave, many of them with small caves (N. Marchington, pers. com. to WAS).
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFDC6D73D99D6F1781A748AE" pageId="7" pageNumber="578" type="discussion">
<paragraph id="8BC7CA65FFDC6D73D99D6F17862E4F52" blockId="7.[151,1437,153,1224]" pageId="7" pageNumber="578">
<emphasis id="B90C1677FFDC6D73D99D6F17853C4E16" bold="true" box="[199,266,513,539]" pageId="7" pageNumber="578">Notes</emphasis>
. The differences between the scanning electron micrograph of the gonopods (
<figureCitation id="1343D6E0FFDC6D73DDD86F1780E54E16" box="[1154,1235,513,539]" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="7" pageNumber="578">Fig. 11</figureCitation>
) and the drawing (
<figureCitation id="1343D6E0FFDC6D73D9FA6F3384C54E32" box="[160,243,549,575]" captionStart="FIGURES 11, 12" captionStartId="6.[151,264,1931,1956]" captionTargetBox="[167,1422,166,1902]" captionTargetId="figure@6.[165,1422,166,1911]" captionTargetPageId="6" captionText="FIGURES 11, 12. Opiona catorycha, new species. Fig. 11. Gonopods, posterior view (scanning electron micrograph); aac, anterior angiocoxite; cc, colpocoxite; pac, posterior angiocoxite; s, gonopod sternum. Fig. 12 Drawing of same; labels as in Fig. 11, pf, pseudoflagellar branches.." figureDoi="http://doi.org/10.5281/zenodo.3467976" httpUri="https://zenodo.org/record/3467976/files/figure.png" pageId="7" pageNumber="578">Fig. 12</figureCitation>
) are instructive. The drawing of the anterior gonopods was made from material temporarily mounted in glycerine on a microscope slide and the gonopods were flattened from their normal position by the cover glass, expanding and pushing apart the tightly clustered pseudoflagella, thereby allowing the several
<typeStatus id="54C374C7FFDC6D73DD956F7B813C4E8A" box="[1231,1290,621,647]" pageId="7" pageNumber="578">types</typeStatus>
to be distinguished. The pressure of the cover glass also changed the orientation of a number of features of the gonopods. Shear (2011) discussed in detail the gonopod structure of species of
<emphasis id="B90C1677FFDC6D73DA126FA387954EC2" box="[840,931,693,719]" italics="true" pageId="7" pageNumber="578">
<taxonomicName id="4C78B1E6FFDC6D73DA126FA387A94EC2" authorityName="Chamberlin" authorityYear="1951" box="[840,927,693,719]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">Opiona</taxonomicName>
,
</emphasis>
recognizing two groups, but found the gonopods of
<taxonomicName id="4C78B1E6FFDC6D73D9AA6FCF85494EFE" authorityName="Shear" authorityYear="2011" box="[240,383,729,755]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="graeningi">
<emphasis id="B90C1677FFDC6D73D9AA6FCF85494EFE" box="[240,383,729,755]" italics="true" pageId="7" pageNumber="578">O. graeningi</emphasis>
</taxonomicName>
to be somewhat intermediate. The gonopods of
<taxonomicName id="4C78B1E6FFDC6D73DACC6FCF801E4EFE" authorityName="Shear &amp; Crawford" authorityYear="2019" box="[918,1064,729,755]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="catorycha">
<emphasis id="B90C1677FFDC6D73DACC6FCF801E4EFE" box="[918,1064,729,755]" italics="true" pageId="7" pageNumber="578">O. catorycha</emphasis>
</taxonomicName>
have a shape similar to that of
<emphasis id="B90C1677FFDC6D73DCDB6FCF853D4F1A" italics="true" pageId="7" pageNumber="578">
<taxonomicName id="4C78B1E6FFDC6D73DCDB6FCF85314F1A" authorityName="Shear" authorityYear="2011" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="graeningi">O. graeningi</taxonomicName>
,
</emphasis>
the main differences being the deeply divided anterior angiocoxites of the former, and the very numerous pseudoflagella. The coxal process of the posterior gonopods of
<taxonomicName id="4C78B1E6FFDC6D73DA396E3787C54F36" authorityName="Shear" authorityYear="2011" box="[867,1011,801,827]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="graeningi">
<emphasis id="B90C1677FFDC6D73DA396E3787C54F36" box="[867,1011,801,827]" italics="true" pageId="7" pageNumber="578">O. graeningi</emphasis>
</taxonomicName>
is flattened and blade-like, while that of
<taxonomicName id="4C78B1E6FFDC6D73D9EC6E53857E4F52" authorityName="Shear &amp; Crawford" authorityYear="2019" box="[182,328,837,863]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="catorycha">
<emphasis id="B90C1677FFDC6D73D9EC6E53857E4F52" box="[182,328,837,863]" italics="true" pageId="7" pageNumber="578">O. catorycha</emphasis>
</taxonomicName>
is three-branched.
</paragraph>
<paragraph id="8BC7CA65FFDC6D73D99D6E7F86904FE2" blockId="7.[151,1437,153,1224]" pageId="7" pageNumber="578">
<figureCitation id="1343D6E0FFDC6D73D99D6E7F85104F8E" box="[199,294,873,899]" captionStart="FIGURE 6" captionStartId="4.[151,250,1978,2003]" captionTargetBox="[170,1407,1014,1945]" captionTargetId="figure@4.[170,1416,1014,1953]" captionTargetPageId="4" captionText="FIGURE 6. Opiona catorycha, new species. Freshly preserved male from No Name Cave, lateral view; g, gonopod; L1–L3, legs 1–3, respectively; R4, ring 4; t9, telopodite of leg 9; tL3, telopodite of leg 3 (small object just to right of label)." figureDoi="http://doi.org/10.5281/zenodo.3467972" httpUri="https://zenodo.org/record/3467972/files/figure.png" pageId="7" pageNumber="578">Figure 6</figureCitation>
depicts a freshly preserved specimen from No Name Cave. As in other species of
<emphasis id="B90C1677FFDC6D73DD916E7F81104F8E" box="[1227,1318,873,899]" italics="true" pageId="7" pageNumber="578">
<taxonomicName id="4C78B1E6FFDC6D73DD916E7F81144F8E" authorityName="Chamberlin" authorityYear="1951" box="[1227,1314,873,899]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">Opiona</taxonomicName>
,
</emphasis>
the fourth ring is enlarged to accommodate powerful muscles attached to the third legpair and the trunk flexes at almost a right angle at this point. It seems safe to assume that the male third legpair is involved somehow in mating, but there have been no observations of mating in any caseyid.
</paragraph>
<paragraph id="8BC7CA65FFDC6D73D99D6EEF81A748AE" blockId="7.[151,1437,153,1224]" pageId="7" pageNumber="578">
Although the
<typeStatus id="54C374C7FFDC6D73D83C6EEF85FE481E" box="[358,456,1017,1043]" pageId="7" pageNumber="578" type="holotype">holotype</typeStatus>
was intercepted in stream flow, we assume that the species is terrestrial and that the specimen was washed downstream from a cave passage not accessible to humans. No specimens of this species (or any caseyid) were taken at other times in the year-long baited pitfall trapping throughout
<collectingRegion id="49BC0487FFDC6D73DDD1695780E94856" box="[1163,1247,1089,1115]" country="United States of America" name="Oregon" pageId="7" pageNumber="578">Oregon</collectingRegion>
Cave. No Name Cave is also a
<typeStatus id="54C374C7FFDC6D73D864697385A84872" box="[318,414,1125,1151]" pageId="7" pageNumber="578" type="paratype">paratype</typeStatus>
locality for the unique relict spider
<taxonomicName id="4C78B1E6FFDC6D73DA606973853948AE" authority="Griswold, Audisio &amp; Ledford, 2012" authorityName="Griswold, Audisio &amp; Ledford" authorityYear="2012" class="Arachnida" family="Trogloraptoridae" genus="Trogloraptor" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="marchingtoni">
<emphasis id="B90C1677FFDC6D73DA606973805C4872" box="[826,1130,1125,1151]" italics="true" pageId="7" pageNumber="578">Trogloraptor marchingtoni</emphasis>
<bibRefCitation id="EFE9B794FFDC6D73DD286973853948AE" author="Griswold, C. E. &amp; Audisio, T. &amp; Ledford, J. M." pageId="7" pageNumber="578" pagination="77 - 102" refId="ref4113" refString="Griswold, C. E., Audisio, T. &amp; Ledford, J. M. (2012) An extraordinary new family of spiders from caves in the Pacific Northwest (Araneae, Trogloraptoridae, new family). ZooKeys, 215, 77 - 102. https: // doi. org / 10.3897 / zookeys. 215.3547" type="journal article" year="2012">Griswold, Audisio &amp; Ledford, 2012</bibRefCitation>
</taxonomicName>
, which is the only described species of the family
<taxonomicName id="4C78B1E6FFDC6D73DA18699F81BB48AE" authority="Griswold, Audisio &amp; Leford, 2012" authorityName="Griswold, Audisio &amp; Leford" authorityYear="2012" box="[834,1421,1161,1187]" class="Arachnida" family="Trogloraptoridae" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="578" phylum="Arthropoda" rank="family">Trogloraptoridae Griswold, Audisio &amp; Leford, 2012</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFDC6D73D99D69BA811748CA" box="[199,1313,1196,1224]" pageId="7" pageNumber="578" type="etymology">
<paragraph id="8BC7CA65FFDC6D73D99D69BA811748CA" blockId="7.[151,1437,153,1224]" box="[199,1313,1196,1224]" pageId="7" pageNumber="578">
<emphasis id="B90C1677FFDC6D73D99D69BA857E48CA" bold="true" box="[199,328,1196,1223]" italics="true" pageId="7" pageNumber="578">Etymology,</emphasis>
The species epithet (Latinized Greek) is an adjective meaning “underground, in caves.”
</paragraph>
</subSubSection>
</treatment>
</document>
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