686 lines
80 KiB
XML
686 lines
80 KiB
XML
<document id="1AB85253D85422638110ECD7AECF94AE" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C24C6021EB72BF9FBA92005" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heteropsis oberthueri Lees, sp. nov." docType="treatment" docVersion="9" lastPageNumber="85" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="83" updateTime="1698662715872" updateUser="ExternalLinkService">
|
||
<mods:mods id="DD0A955A96A1613AD9298475BDA125A0" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="02A352C15DF1DCAE63B943EA3B34D344">
|
||
<mods:title id="8CDA67AF8AA6A78CA76F4DDC63D459B5">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="0EDB61BA1DDE2F8D8B9D7168B3E003BE" type="personal">
|
||
<mods:role id="156A98CEF0A7631DD2194C7DC68B6DA5">
|
||
<mods:roleTerm id="743BC917C220EA82AC05CDF2EE1CFEC6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CB437FFD1204E143387A3A9FD4D32402">C, Lees David</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="B9B33194AE5A691AD3663EF97EBCC55C">text</mods:typeOfResource>
|
||
<mods:relatedItem id="7F7F5632383AA3C6DA2984EEF61FA958" type="host">
|
||
<mods:titleInfo id="419F81FDA8EF1D84F2B7BC0EC54D78EB">
|
||
<mods:title id="C07802970DE87619F14E003C21FEDF09">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="130E314E472F22A130021160578DCF2E">
|
||
<mods:date id="E4EF5270916E768F8AB1DCF38EA3566A">2016</mods:date>
|
||
<mods:detail id="C8018CA409A3BE9F392670518A457ACA" type="volume">
|
||
<mods:number id="669D3D6B261C1C977073891D53429529">4118</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C59BB226555FFF1A0BCF9F3B146E41BD" type="issue">
|
||
<mods:number id="5144175090F884FB7C7F72199B407BF9">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="234941BFB778D7B9E79BD95309A8BA85" unit="page">
|
||
<mods:start id="CB2DD017EE0F8D26A7ADD6EF4739E879">1</mods:start>
|
||
<mods:end id="E5D604CE1F7F6C575F7FAFB1A90CE90C">97</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="FC18A361EB6886676A9D1C4D18DAEB68">journal article</mods:classification>
|
||
<mods:identifier id="737F0DA7FE1C6FA9BF557C7BBD4F345F" type="CLB-Dataset">38930</mods:identifier>
|
||
<mods:identifier id="D7A9D852D6828A4345415323C6892C1B" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
|
||
<mods:identifier id="65EF100D74B5BF6F3AA239358B386E22" type="GBIF-Dataset">a4e408c4-2575-4b5c-878b-e1b9010ec619</mods:identifier>
|
||
<mods:identifier id="903C72D74965B8BE821DE2E393EC5231" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="1F38F37B032BA7259994C9398B6CE4CD" type="Zenodo-Dep">264597</mods:identifier>
|
||
<mods:identifier id="B73DF747844DE9B38A3E886D4C398E93" type="ZooBank">CFA586DA-10EE-468B-80EE-35351E3845FD</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="038747324C24C6021EB72BF9FBA92005" ID-DOI="http://doi.org/10.5281/zenodo.6086458" ID-GBIF-Taxon="120817150" ID-Zenodo-Dep="6086458" LSID="urn:lsid:plazi:treatment:038747324C24C6021EB72BF9FBA92005" httpUri="http://treatment.plazi.org/id/038747324C24C6021EB72BF9FBA92005" lastPageId="85" lastPageNumber="85" pageId="82" pageNumber="83">
|
||
<subSubSection id="C334A5AF4C24C6051EB72BF9FBF52664" pageId="82" pageNumber="83" type="nomenclature">
|
||
<paragraph id="8B91F6244C24C6051EB72BF9FD99279A" blockId="82.[151,1062,1748,1808]" box="[151,586,1748,1774]" pageId="82" pageNumber="83">
|
||
<heading id="D0D941484C24C6051EB72BF9FD99279A" bold="true" box="[151,586,1748,1774]" fontSize="11" level="1" pageId="82" pageNumber="83" reason="1">
|
||
<emphasis id="B95A2A364C24C6051EB72BF9FD99279A" bold="true" box="[151,586,1748,1774]" pageId="82" pageNumber="83">
|
||
<taxonomicName id="4C2E8DA74C24C6051EB72BF9FE33279A" authority="Lees" authorityName="Lees" box="[151,480,1748,1774]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="oberthueri" status="sp. nov.">
|
||
<emphasis id="B95A2A364C24C6051EB72BF9FE4D279A" bold="true" box="[151,414,1748,1774]" italics="true" pageId="82" pageNumber="83">Heteropsis oberthueri</emphasis>
|
||
Lees
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A269974D4C24C6051FCD2BF9FD99279A" box="[493,586,1748,1774]" pageId="82" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C24C6051EB72BDAFBF52664" blockId="82.[151,1062,1748,1808]" box="[151,1062,1783,1808]" pageId="82" pageNumber="83">LSID: urn:lsid:zoobank.org:act:FEE8CA6F-A53B-43A5-8C9F-E5D69AD0E468</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C24C6051EB72A12FEEB260F" pageId="82" pageNumber="83" type="description">
|
||
<paragraph id="8B91F6244C24C6051EB72A12FEEB260F" blockId="82.[151,1436,1855,2029]" pageId="82" pageNumber="83">
|
||
<emphasis id="B95A2A364C24C6051EB72A12FE88262C" bold="true" box="[151,347,1855,1880]" pageId="82" pageNumber="83">Prior references</emphasis>
|
||
: sp. 89, sp. 87 (on some envelopes). “KA518_Heteopsis_
|
||
<taxonomicName id="4C2E8DA74C24C6051A252A12FB81262C" box="[1029,1106,1855,1880]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="anceps">anceps</taxonomicName>
|
||
” [
|
||
<emphasis id="B95A2A364C24C6051A502A6DFB5C2623" box="[1136,1167,1856,1879]" italics="true" pageId="82" pageNumber="83">sic</emphasis>
|
||
] in Aduse-Poku
|
||
<emphasis id="B95A2A364C24C6051B792A6DFA4F262C" box="[1369,1436,1855,1880]" italics="true" pageId="82" pageNumber="83">et al.,</emphasis>
|
||
(2015,
|
||
<figureCitation id="1315EAA14C24C6051EC62A4EFEF4260F" box="[230,295,1891,1916]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="82" pageNumber="83">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C24C6041EE72AA4FE9A23E3" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83" type="materials_examined">
|
||
<paragraph id="8B91F6244C24C6051EE72AA4FE9A2699" blockId="82.[151,1436,1855,2029]" pageId="82" pageNumber="83">
|
||
<emphasis id="B95A2A364C24C6051EE72AA4FD2026D6" bold="true" box="[199,755,1929,1954]" pageId="82" pageNumber="83">
|
||
<typeStatus id="549548864C24C6051EE72AA4FED126D6" box="[199,258,1929,1954]" pageId="82" pageNumber="83">Type</typeStatus>
|
||
material. Deposition
|
||
<collectionCode id="ED3F6EE14C24C6051C302AA4FDB826D6" box="[528,619,1929,1954]" pageId="82" pageNumber="83">BMNH</collectionCode>
|
||
:
|
||
<typeStatus id="549548864C24C6051C5D2AA4FD3D26D6" box="[637,750,1929,1954]" pageId="82" pageNumber="83" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
♂ (
|
||
<figureCitation id="1315EAA14C24C6051D072AA4FCAD26D6" box="[807,894,1929,1954]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="82" pageNumber="83">Fig. 24</figureCitation>
|
||
D),
|
||
<collectingCountry id="F339B6B44C24C6051D8C2AA4FBE026D6" box="[940,1075,1929,1954]" name="Madagascar" pageId="82" pageNumber="83">Madagascar</collectingCountry>
|
||
E, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C24C6051B022AA4FA4526D6" box="[1314,1430,1926,1954]" direction="south" orientation="latitude" pageId="82" pageNumber="83" precision="5" value="-17.5131">17.5131o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C24C6051EB72A9DFED926BC" box="[151,266,1965,1993]" direction="east" orientation="longitude" pageId="82" pageNumber="83" precision="5" value="48.7269">48.7269o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C24C6051F1B2A9DFE5F26BD" box="[315,396,1968,1993]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="82" pageNumber="83" unit="km" value="1.5">1.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C24C6051FB82A9DFDEF26BC" box="[408,572,1968,1993]" metricMagnitude="3" metricUnit="m" metricValue="1.064" metricValueMax="1.089" metricValueMin="1.039" pageId="82" pageNumber="83" unit="m" value="1064.0" valueMax="1089.0" valueMin="1039.0">1064 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C24C6051C682A9DFD6826BD" box="[584,699,1968,1993]" pageId="82" pageNumber="83" value="2006-11-05">5/11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C24C6051D682A9DFCBC26BD" box="[840,879,1968,1993]" pageId="82" pageNumber="83">DL</collectionCode>
|
||
06-064, G04 [leg for
|
||
<collectionCode id="ED3F6EE14C24C6051A7F2A9CFB4C26BD" box="[1119,1183,1969,1993]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="82" pageNumber="83">DNA</collectionCode>
|
||
],
|
||
<collectionCode id="ED3F6EE14C24C6051A942A9DFACD26BD" box="[1204,1310,1968,1993]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="82" pageNumber="83">NHMUK</collectionCode>
|
||
010289159 [
|
||
<collectingCountry id="F339B6B44C24C6051EBD2AF9FF0B2698" box="[157,216,2004,2028]" name="Qatar" pageId="82" pageNumber="83">QTR</collectingCountry>
|
||
barcode].
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C25C6041EE72DB7FE9A23E3" blockId="83.[151,1437,151,2030]" pageId="83" pageNumber="84">
|
||
<typeStatus id="549548864C25C6041EE72DB7FEEC21C7" box="[199,319,154,179]" pageId="83" pageNumber="84" type="paratype">
|
||
<emphasis id="B95A2A364C25C6041EE72DB7FEEC21C7" bold="true" box="[199,319,154,179]" pageId="83" pageNumber="84">Paratypes</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<emphasis id="B95A2A364C25C6041F6D2DB7FDE621C7" bold="true" box="[333,565,154,179]" pageId="83" pageNumber="84">
|
||
Deposition
|
||
<collectionCode id="ED3F6EE14C25C6041FF42DB7FDFC21C7" box="[468,559,154,179]" pageId="83" pageNumber="84">BMHN</collectionCode>
|
||
:
|
||
</emphasis>
|
||
♂,
|
||
<collectingCountry id="F339B6B44C25C6041C7F2DB7FD3521C7" box="[607,742,154,179]" name="Madagascar" pageId="83" pageNumber="84">Madagascar</collectingCountry>
|
||
E, Ambavala camp, Zahamena,
|
||
<geoCoordinate id="EE1A90E34C25C6041A752DB7FB1A21C7" box="[1109,1225,151,179]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5451">17.5451o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041AF52DB6FA9A21C6" box="[1237,1353,151,179]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="5" value="48.7237">48.7237o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041B592DB7FF6D21A3" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="83" pageNumber="84" unit="km" value="0.5">0.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041EE92D92FEBF21A2" box="[201,364,190,215]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.35" metricValueMin="1.25" pageId="83" pageNumber="84" unit="m" value="1300.0" valueMax="1350.0" valueMin="1250.0">1300 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C25C6041F582D93FE3F21A3" box="[376,492,190,215]" pageId="83" pageNumber="84" value="2006-11-06">6/11/2006</date>
|
||
, 13:10, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C25C6041CE52D93FD3F21A3" box="[709,748,190,215]" pageId="83" pageNumber="84">DL</collectionCode>
|
||
06-130A,
|
||
<collectionCode id="ED3F6EE14C25C6041D7C2D93FC1521A3" box="[860,966,190,215]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="83" pageNumber="84">NHMUK</collectionCode>
|
||
010289160 [
|
||
<collectingCountry id="F339B6B44C25C6041A722D92FB5E21A3" box="[1106,1165,191,215]" name="Qatar" pageId="83" pageNumber="84">QTR</collectingCountry>
|
||
barcode]; ♀ (
|
||
<figureCitation id="1315EAA14C25C6041B0A2D93FAAF21A3" box="[1322,1404,190,215]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="83" pageNumber="84">Fig. 24</figureCitation>
|
||
E), E, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C25C6041F4F2DC9FE302189" box="[367,483,225,253]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5376">17.5376o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041FD02DC9FDA12189" box="[496,626,225,253]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="1" value="48.71965">48.71965o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041C832DC8FCD02189" box="[675,771,228,253]" metricMagnitude="2" metricUnit="m" metricValue="9.399999999999999" pageId="83" pageNumber="84" unit="km" value="0.94">0.94 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041D302DC8FC112189" box="[784,962,228,253]" metricMagnitude="3" metricUnit="m" metricValue="1.1800000000000002" metricValueMax="1.3050000000000002" metricValueMin="1.0550000000000002" pageId="83" pageNumber="84" unit="m" value="1180.0" valueMax="1305.0" valueMin="1055.0">1180 +/- 125 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C25C6041DEF2DC9FB902189" box="[975,1091,228,253]" pageId="83" pageNumber="84" value="2006-11-06">6/11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C25C6041AF12DC9FB2A2189" box="[1233,1273,228,253]" pageId="83" pageNumber="84">DL</collectionCode>
|
||
06-130, IA585 [isotope voucher],
|
||
<collectionCode id="ED3F6EE14C25C6041F492C26FE072050" box="[361,468,267,292]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="83" pageNumber="84">NHMUK</collectionCode>
|
||
010289161 [
|
||
<collectingCountry id="F339B6B44C25C6041C7E2C21FD4A2050" box="[606,665,268,292]" name="Qatar" pageId="83" pageNumber="84">QTR</collectingCountry>
|
||
barcode]; ♂, E, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C25C6041A282C26FBAF2050" box="[1032,1148,264,292]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5376">17.5376o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041AA92C26FAD92057" box="[1161,1290,264,292]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="1" value="48.71965">48.71965o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041B192C21FA4B2050" box="[1337,1432,267,292]" metricMagnitude="2" metricUnit="m" metricValue="9.399999999999999" pageId="83" pageNumber="84" unit="km" value="0.94">0.94 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041EB72C02FE8B2033" box="[151,344,303,328]" metricMagnitude="3" metricUnit="m" metricValue="1.1800000000000002" metricValueMax="1.3050000000000002" metricValueMin="1.0550000000000002" pageId="83" pageNumber="84" unit="m" value="1180.0" valueMax="1305.0" valueMin="1055.0">1180 +/- 125 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C25C6041F4A2C02FE0D203C" box="[362,478,303,328]" pageId="83" pageNumber="84" value="2006-11-06">6/11/2006</date>
|
||
, D.C. Lees: 371
|
||
<collectionCode id="ED3F6EE14C25C6041C852C02FD1E2033" box="[677,717,303,327]" pageId="83" pageNumber="84">DL</collectionCode>
|
||
,
|
||
<collectionCode id="ED3F6EE14C25C6041CFE2C02FC9B2033" box="[734,840,303,327]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="83" pageNumber="84">NHMUK</collectionCode>
|
||
010289192 [
|
||
<collectingCountry id="F339B6B44C25C6041DF92C02FBC72033" box="[985,1044,303,327]" name="Qatar" pageId="83" pageNumber="84">QTR</collectingCountry>
|
||
barcode]; ♂, E, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C25C6041EB72C78FF3D201A" box="[151,238,338,366]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="555" value="-17.53">17.53o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041F212C78FEA6201A" box="[257,373,338,366]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="5" value="48.7269">48.7269o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041F8F2C78FDD5201A" box="[431,518,341,366]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="83" pageNumber="84" unit="km" value="1.5">1.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041C372C78FD18201A" box="[535,715,341,366]" metricMagnitude="3" metricUnit="m" metricValue="1.064" metricValueMax="1.089" metricValueMin="1.039" pageId="83" pageNumber="84" unit="m" value="1064.0" valueMax="1089.0" valueMin="1039.0">1064 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C25C6041CFD2C78FC83201A" box="[733,848,341,366]" pageId="83" pageNumber="84" value="2006-11-06">6/11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C25C6041DCD2C78FBC6201A" box="[1005,1045,341,366]" pageId="83" pageNumber="84">DL</collectionCode>
|
||
06-086, IA586 [isotope voucher],
|
||
<collectionCode id="ED3F6EE14C25C6041EB72C51FED220E1" box="[151,257,380,405]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="83" pageNumber="84">NHMUK</collectionCode>
|
||
010289162 [
|
||
<collectingCountry id="F339B6B44C25C6041FAC2C50FE1420E1" box="[396,455,381,405]" name="Qatar" pageId="83" pageNumber="84">QTR</collectingCountry>
|
||
barcode]; ♂, E, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C25C6041D132C51FC7420E1" box="[819,935,377,405]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5376">17.5376o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041D932C51FBE620E0" box="[947,1077,377,405]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="1" value="48.71965">48.71965o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041A432C50FB1220E1" box="[1123,1217,380,405]" metricMagnitude="2" metricUnit="m" metricValue="9.399999999999999" pageId="83" pageNumber="84" unit="km" value="0.94">0.94 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041AEC2C50FAAA20E0" box="[1228,1401,380,405]" metricMagnitude="3" metricUnit="m" metricValue="1.1800000000000002" metricValueMax="1.3050000000000002" metricValueMin="1.0550000000000002" pageId="83" pageNumber="84" unit="m" value="1180.0" valueMax="1305.0" valueMin="1055.0">1180 +/- 125 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C25C6041BA42C51FED920CD" pageId="83" pageNumber="84" value="2006-11-06" valueMax="2006-11-07" valueMin="2006-11-06">6- 7/11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C25C6041F942C8DFE0E20CD" box="[436,477,416,441]" pageId="83" pageNumber="84">DL</collectionCode>
|
||
06-1001, IA324 [isotope voucher],
|
||
<collectionCode id="ED3F6EE14C25C6041DAF2C8DFC2A20CD" box="[911,1017,416,441]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="83" pageNumber="84">NHMUK</collectionCode>
|
||
010289163 [
|
||
<collectingCountry id="F339B6B44C25C6041AAF2C8DFB1920CC" box="[1167,1226,416,440]" name="Qatar" pageId="83" pageNumber="84">QTR</collectingCountry>
|
||
barcode]; ♀, E, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C25C6041F742CEBFE1420AB" box="[340,455,451,479]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5376">17.5376o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041FF72CEBFD8A20AA" box="[471,601,451,479]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="1" value="48.71965">48.71965o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041CAF2CEAFD2220AB" box="[655,753,454,479]" metricMagnitude="2" metricUnit="m" metricValue="9.399999999999999" pageId="83" pageNumber="84" unit="km" value="0.94">0.94 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041D202CEAFC6A20AA" box="[768,953,454,479]" metricMagnitude="3" metricUnit="m" metricValue="1.1800000000000002" metricValueMax="1.3050000000000002" metricValueMin="1.0550000000000002" pageId="83" pageNumber="84" unit="m" value="1180.0" valueMax="1305.0" valueMin="1055.0">1180 +/- 125 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C25C6041DE82CEBFBEF20AB" box="[968,1084,454,479]" pageId="83" pageNumber="84" value="2006-11-06">6/11/2006</date>
|
||
, 11:30, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C25C6041B012CEBFA9A20AB" box="[1313,1353,454,479]" pageId="83" pageNumber="84">DL</collectionCode>
|
||
06-112, IA587 [isotope voucher],
|
||
<collectionCode id="ED3F6EE14C25C6041FEE2CC7FDEB2377" box="[462,568,490,515]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="83" pageNumber="84">NHMUK</collectionCode>
|
||
010289164 [
|
||
<collectingCountry id="F339B6B44C25C6041CEB2CC6FCD52377" box="[715,774,491,515]" name="Qatar" pageId="83" pageNumber="84">QTR</collectingCountry>
|
||
barcode]; ♀, E, Towards Ambavala camp, Zahamena,
|
||
<geoCoordinate id="EE1A90E34C25C6041EB72F3DFED9235D" box="[151,266,525,553]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5376">17.5376o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041F382F3DFE49235C" box="[280,410,525,553]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="1" value="48.71965">48.71965o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041FEC2F3CFDFF235D" box="[460,556,528,553]" metricMagnitude="2" metricUnit="m" metricValue="9.399999999999999" pageId="83" pageNumber="84" unit="km" value="0.94">0.94 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041C182F3CFD3F235C" box="[568,748,528,553]" metricMagnitude="3" metricUnit="m" metricValue="1.1800000000000002" metricValueMax="1.3050000000000002" metricValueMin="1.0550000000000002" pageId="83" pageNumber="84" unit="m" value="1180.0" valueMax="1305.0" valueMin="1055.0">1180 +/- 125 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C25C6041CD92F3DFCBF235D" box="[761,876,528,553]" pageId="83" pageNumber="84" value="2006-11-06">6/11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C25C6041DDB2F3DFBF7235D" box="[1019,1060,528,553]" pageId="83" pageNumber="84">DL</collectionCode>
|
||
06-141, IA588 [isotope voucher],
|
||
<collectionCode id="ED3F6EE14C25C6041EB72F1AFED22324" box="[151,257,567,592]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="83" pageNumber="84">NHMUK</collectionCode>
|
||
010289165 [
|
||
<collectingCountry id="F339B6B44C25C6041FB02F15FE182324" box="[400,459,568,592]" name="Qatar" pageId="83" pageNumber="84">QTR</collectingCountry>
|
||
barcode]; ♀, E, returning to Bemoara camp, Zahamena,
|
||
<geoCoordinate id="EE1A90E34C25C6041A4A2F1AFB0D2324" box="[1130,1246,564,592]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5356">17.5356o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041ACE2F1AFAA3233B" box="[1262,1392,564,592]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="1" value="48.71875">48.71875o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041EB72F76FF252300" box="[151,246,603,628]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="83" pageNumber="84" unit="km" value="0.7">0.70 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041F222F76FE602307" box="[258,435,603,627]" metricMagnitude="3" metricUnit="m" metricValue="1.18" metricValueMax="1.28" metricValueMin="1.08" pageId="83" pageNumber="84" unit="m" value="1180.0" valueMax="1280.0" valueMin="1080.0">1180 +/- 100 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C25C6041F9F2F76FDE12300" box="[447,562,603,628]" pageId="83" pageNumber="84" value="2006-11-05">5/11/2006</date>
|
||
, 14:59, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C25C6041D2E2F76FCE52300" box="[782,822,603,628]" pageId="83" pageNumber="84">DL</collectionCode>
|
||
06-059, IA589 [isotope voucher],
|
||
<collectionCode id="ED3F6EE14C25C6041A942F76FACD2300" box="[1204,1310,603,628]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="83" pageNumber="84">NHMUK</collectionCode>
|
||
010289166 [
|
||
<collectingCountry id="F339B6B44C25C6041EBD2F52FF0B23E3" box="[157,216,639,663]" name="Qatar" pageId="83" pageNumber="84">QTR</collectingCountry>
|
||
barcode];
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C25C6041EE72F88FB1D223B" pageId="83" pageNumber="84" type="description">
|
||
<paragraph id="8B91F6244C25C6041EE72F88FB3A2396" blockId="83.[151,1437,151,2030]" pageId="83" pageNumber="84">
|
||
<emphasis id="B95A2A364C25C6041EE72F88FE7C23CA" bold="true" box="[199,431,677,702]" pageId="83" pageNumber="84">Deposition MNHN:</emphasis>
|
||
♂,
|
||
<collectingCountry id="F339B6B44C25C6041FF82F88FD8C23CA" box="[472,607,677,702]" name="Madagascar" pageId="83" pageNumber="84">Madagascar</collectingCountry>
|
||
E, Ambavala, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C25C6041D9A2F88FBFD23CA" box="[954,1070,674,702]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5451">17.5451o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041A1B2F8BFB7D23C9" box="[1083,1198,674,702]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="5" value="48.7237">48.7237o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041AFC2F88FAE823CA" box="[1244,1339,677,702]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="83" pageNumber="84" unit="km" value="0.25">0.25 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041B662F88FF312395" metricMagnitude="3" metricUnit="m" metricValue="1.325" metricValueMax="1.375" metricValueMin="1.275" pageId="83" pageNumber="84" unit="m" value="1325.0" valueMax="1375.0" valueMin="1275.0">1325 +/ - 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C25C6041ECE2FE4FEB22396" box="[238,353,713,738]" pageId="83" pageNumber="84" value="2006-11-06">6/11/2006</date>
|
||
, 14:11, D.C. Lees: DL06-117, IA584 [isotope voucher], DSC03096.jpg [image];
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C25C6041EE72FC2FE282258" blockId="83.[151,1437,151,2030]" pageId="83" pageNumber="84">
|
||
<emphasis id="B95A2A364C25C6041EE72FC2FE4E227C" bold="true" box="[199,413,751,776]" pageId="83" pageNumber="84">Deposition ABRI:</emphasis>
|
||
♂,
|
||
<collectingCountry id="F339B6B44C25C6041FE82FC2FD9C227C" box="[456,591,751,776]" name="Madagascar" pageId="83" pageNumber="84">Madagascar</collectingCountry>
|
||
E, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C25C6041D0D2FC2FC73227C" box="[813,928,748,776]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5376">17.5376o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041D8D2FC2FBFC2273" box="[941,1071,748,776]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="1" value="48.71965">48.71965o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041A7F2FDDFB6D227C" box="[1119,1214,751,776]" metricMagnitude="2" metricUnit="m" metricValue="9.399999999999999" pageId="83" pageNumber="84" unit="km" value="0.94">0.94 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041AEA2FDDFAA92273" box="[1226,1402,751,776]" metricMagnitude="3" metricUnit="m" metricValue="1.1800000000000002" metricValueMax="1.3050000000000002" metricValueMin="1.0550000000000002" pageId="83" pageNumber="84" unit="m" value="1180.0" valueMax="1305.0" valueMin="1055.0">1180 +/- 125 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C25C6041BA62FC2FF272258" pageId="83" pageNumber="84" value="2006-11-06">6/ 11/2006</date>
|
||
, D.C. Lees: DL06-121.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C25C6041EE72E1AFB1D223B" blockId="83.[151,1437,151,2030]" box="[199,1230,823,848]" pageId="83" pageNumber="84">
|
||
<emphasis id="B95A2A364C25C6041EE72E1AFE6D2224" bold="true" box="[199,446,823,848]" pageId="83" pageNumber="84">Deposition summary</emphasis>
|
||
: BMNH (HT ♂, 4 PT ♂♂, 4 PT ♀♀), MNHN (PT ♂), ABRI (PT ♂).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C25C6041EE72E70FB992201" box="[199,1098,858,886]" pageId="83" pageNumber="84" type="materials_examined">
|
||
<paragraph id="8B91F6244C25C6041EE72E70FB992201" blockId="83.[151,1437,151,2030]" box="[199,1098,858,886]" pageId="83" pageNumber="84">
|
||
<emphasis id="B95A2A364C25C6041EE72E70FEB62202" bold="true" box="[199,357,861,886]" pageId="83" pageNumber="84">
|
||
<typeStatus id="549548864C25C6041EE72E70FED12202" box="[199,258,861,886]" pageId="83" pageNumber="84">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
NW Zahamena,
|
||
<geoCoordinate id="EE1A90E34C25C6041C042E70FD4B2202" box="[548,664,858,886]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5131">17.5131o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C25C6041C842E70FCCB2201" box="[676,792,858,886]" direction="east" orientation="longitude" pageId="83" pageNumber="84" precision="5" value="48.7269">48.7269o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C25C6041D672E70FC4B2202" box="[839,920,861,886]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="83" pageNumber="84" unit="km" value="1.5">1.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C25C6041D832E70FB952201" box="[931,1094,861,886]" metricMagnitude="3" metricUnit="m" metricValue="1.064" metricValueMax="1.089" metricValueMin="1.039" pageId="83" pageNumber="84" unit="m" value="1064.0" valueMax="1089.0" valueMin="1039.0">
|
||
1064 +/-
|
||
<specimenCount id="9D283DAD4C25C6041A2A2E70FB952201" box="[1034,1094,861,886]" pageId="83" pageNumber="84" type="generic">25 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C25C6041EE72EACFEEF24A2" pageId="83" pageNumber="84" type="diagnosis">
|
||
<paragraph id="8B91F6244C25C6041EE72EACFEEF24A2" blockId="83.[151,1437,151,2030]" pageId="83" pageNumber="84">
|
||
<emphasis id="B95A2A364C25C6041EE72EACFE9222EE" bold="true" box="[199,321,897,922]" pageId="83" pageNumber="84">Diagnosis.</emphasis>
|
||
The species is particularly similar to
|
||
<taxonomicName id="4C2E8DA74C25C6041D002EAFFC4B22ED" box="[800,920,898,921]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||
<emphasis id="B95A2A364C25C6041D002EAFFC4B22ED" box="[800,920,898,921]" italics="true" pageId="83" pageNumber="84">Ht. tianae</emphasis>
|
||
</taxonomicName>
|
||
described above and in fact few obvious distinguishing features are apparent that are likely to distinguish all examples, but that species is however allopatric, only known from the Angavo massif, and on average, the HWV Mb of
|
||
<taxonomicName id="4C2E8DA74C25C6041A452EE7FAD92295" box="[1125,1290,968,993]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="oberthueri">
|
||
<emphasis id="B95A2A364C25C6041A452EE7FAD92295" box="[1125,1290,968,993]" italics="true" pageId="83" pageNumber="84">Ht. oberthueri</emphasis>
|
||
</taxonomicName>
|
||
is relatively straight and there is usually no space-M3 ocellus expressed on the HWD (
|
||
<figureCitation id="1315EAA14C25C6041DFD2EC1FBFD2571" box="[989,1070,1004,1029]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="83" pageNumber="84">Fig. 24</figureCitation>
|
||
D–E). The wings have a slightly greyer cast.
|
||
<taxonomicName id="4C2E8DA74C25C6041F06293CFE19255D" box="[294,458,1040,1065]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="oberthueri">
|
||
<emphasis id="B95A2A364C25C6041F06293CFE19255D" box="[294,458,1040,1065]" italics="true" pageId="83" pageNumber="84">Ht. oberthueri</emphasis>
|
||
</taxonomicName>
|
||
also strongly resembles
|
||
<taxonomicName id="4C2E8DA74C25C6041CCD293CFC53255D" box="[749,896,1040,1065]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="andasibe">
|
||
<emphasis id="B95A2A364C25C6041CCD293CFC53255D" box="[749,896,1040,1065]" italics="true" pageId="83" pageNumber="84">Ht. andasibe</emphasis>
|
||
</taxonomicName>
|
||
. It can be distinguished from that last species (which has slightly more conspicuous white spots on the ventral wing surfaces) by the less straight, usually more evenly outangled HWV Mb at space M
|
||
<quantity id="4CD65BC14C25C6041C6F2975FDA92504" box="[591,634,1111,1136]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="83" pageNumber="84" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="4C2E8DA74C25C6041CA02974FCDC2504" box="[640,783,1111,1136]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="andasibe">
|
||
<emphasis id="B95A2A364C25C6041CA02974FCDC2504" box="[640,783,1111,1136]" italics="true" pageId="83" pageNumber="84">Ht. andasibe</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1315EAA14C25C6041D3E297AFCA22504" box="[798,881,1111,1136]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="83" pageNumber="84">Fig. 22</figureCitation>
|
||
C), and from more yellowish lowland
|
||
<taxonomicName id="4C2E8DA74C25C6041B3B2974FA4F2504" box="[1307,1436,1111,1136]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="strigula">
|
||
<emphasis id="B95A2A364C25C6041B3B2974FA4F2504" box="[1307,1436,1111,1136]" italics="true" pageId="83" pageNumber="84">Ht. strigula</emphasis>
|
||
</taxonomicName>
|
||
which have a conspicuous orange crescent on the HWD, and from
|
||
<taxonomicName id="4C2E8DA74C25C6041DBF2950FBCF25E0" box="[927,1052,1147,1172]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="ankova">
|
||
<emphasis id="B95A2A364C25C6041DBF2950FBCF25E0" box="[927,1052,1147,1172]" italics="true" pageId="83" pageNumber="84">Ht. ankova</emphasis>
|
||
</taxonomicName>
|
||
which is smaller and has ocellus space-M3 as well as in space-CuA1 expressed on the HWD (
|
||
<figureCitation id="1315EAA14C25C6041D7129B2FC7725C3" box="[849,932,1183,1208]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="83" pageNumber="84">Fig. 19</figureCitation>
|
||
C). The ventrally very similar
|
||
<taxonomicName id="4C2E8DA74C25C6041ADE298DFA4F25C3" box="[1278,1436,1184,1207]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="roussettae">
|
||
<emphasis id="B95A2A364C25C6041ADE298DFA4F25C3" box="[1278,1436,1184,1207]" italics="true" pageId="83" pageNumber="84">Ht. roussettae</emphasis>
|
||
</taxonomicName>
|
||
has a strong tendency to expression of a small ocellus M3 on the HWD (
|
||
<figureCitation id="1315EAA14C25C6041D9C29EEFBDF25AF" box="[956,1036,1219,1244]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="83" pageNumber="84">Fig. 22</figureCitation>
|
||
A–B). The potentially sympatric (in ‘Antsianaka’ region)
|
||
<taxonomicName id="4C2E8DA74C25C6041FAA29C5FDD0258B" box="[394,515,1256,1279]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="anceps">
|
||
<emphasis id="B95A2A364C25C6041FAA29C5FDD0258B" box="[394,515,1256,1279]" italics="true" pageId="83" pageNumber="84">Ht. anceps</emphasis>
|
||
</taxonomicName>
|
||
has a relatively straight HWV Mb and a yellowish ventral cast with sparse light brown irroration and a distinct black ventral androconial patch on the ♂ sternal abdomen (
|
||
<figureCitation id="1315EAA14C25C6041AEB2827FAF02457" box="[1227,1315,1290,1315]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="83" pageNumber="84">Fig. 22</figureCitation>
|
||
D). The ♂ genitalia are similar to those of
|
||
<taxonomicName id="4C2E8DA74C25C6041FDB2802FDB82432" box="[507,619,1327,1350]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||
<emphasis id="B95A2A364C25C6041FDB2802FDB82432" box="[507,619,1327,1350]" italics="true" pageId="83" pageNumber="84">Ht. tianae</emphasis>
|
||
</taxonomicName>
|
||
, differing by the more strongly inpointed subterminal spine on the valve and the terminal head covered in spinoid setae being enlarged. They also differ from the potentially sympatric
|
||
<taxonomicName id="4C2E8DA74C25C6041B59287EFE7D24FA" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="anceps">
|
||
<emphasis id="B95A2A364C25C6041B59287EFF3624F9" italics="true" pageId="83" pageNumber="84">Ht. anceps</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBF8BD54C25C6041ED52858FE7424FA" author="Oberthur" box="[245,423,1397,1422]" pageId="83" pageNumber="84" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, being smaller and with a narrower distal valve tip (
|
||
<bibRefCitation id="EFBF8BD54C25C6041A28285BFB1624FA" author="Lees" box="[1032,1221,1398,1422]" pageId="83" pageNumber="84" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 107</bibRefCitation>
|
||
). DNA differences separating
|
||
<taxonomicName id="4C2E8DA74C25C6041F3128B7FE6324C6" box="[273,432,1433,1458]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="oberthueri">
|
||
<emphasis id="B95A2A364C25C6041F3128B7FE6324C6" box="[273,432,1433,1458]" italics="true" pageId="83" pageNumber="84">Ht. oberthueri</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C2E8DA74C25C6041FD428B7FDB724C5" box="[500,612,1434,1457]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||
<emphasis id="B95A2A364C25C6041FD428B7FDB724C5" box="[500,612,1434,1457]" italics="true" pageId="83" pageNumber="84">Ht. tianae</emphasis>
|
||
</taxonomicName>
|
||
are indicated by
|
||
<bibRefCitation id="EFBF8BD54C25C6041D0428B4FBED24C6" author="Aduse-Poku" box="[804,1086,1433,1458]" pageId="83" pageNumber="84" refString="Aduse-Poku, K., Lees, D. C., Brattstrom, O., Kodandaramaiah, U., Collins, S. & Brakefield, P. M., (2016) Molecular phylogeny and generic-level taxonomy of the widespread palaeotropical ' Heteropsis clade' (Nymphalidae: Satyrinae: Mycalesina). Systematic Entomology, in press. http: // dx. doi. org / syen. 12183" type="book" year="2016">
|
||
Aduse-Poku
|
||
<emphasis id="B95A2A364C25C6041D9528B7FC2724C6" box="[949,1012,1433,1458]" italics="true" pageId="83" pageNumber="84">et al.,</emphasis>
|
||
(2016
|
||
</bibRefCitation>
|
||
, in press). See below regarding male genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C25C6031EE728CDFECB2292" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="description">
|
||
<paragraph id="8B91F6244C25C6031EE728CDFB4E21A0" blockId="83.[151,1437,151,2030]" lastBlockId="84.[151,1437,151,2013]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
|
||
<emphasis id="B95A2A364C25C6041EE728CDFE7C248D" bold="true" box="[199,431,1504,1529]" pageId="83" pageNumber="84">Description. Wings</emphasis>
|
||
: description is similar to that of
|
||
<taxonomicName id="4C2E8DA74C25C6041D1A28CFFC7D248D" box="[826,942,1506,1529]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||
<emphasis id="B95A2A364C25C6041D1A28CFFC7D248D" box="[826,942,1506,1529]" italics="true" pageId="83" pageNumber="84">Ht. tianae</emphasis>
|
||
</taxonomicName>
|
||
. Upperside uniform mid brown, with FW space-CuA1 ocellus featuring an orange ocellus ‘ring’ which is sometimes rather ‘squashed’ along the diagonal from mid-costa to tergal angle. FWD space-M1 ocellus small with narrow orange ring. HWD space-CuA1 ocellus is the only one expressed there and is somewhat elliptic, with a narrow orange ring. Darkish brown, diffuse, slightly wavy and uprecurved hair-brush emanates mainly from below vein 1A+2A, as far as mid-vein. Underside greyish brown, similar to
|
||
<taxonomicName id="4C2E8DA74C25C6041FE52BB8FDEA27D8" box="[453,569,1685,1708]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||
<emphasis id="B95A2A364C25C6041FE52BB8FDEA27D8" box="[453,569,1685,1708]" italics="true" pageId="83" pageNumber="84">Ht. tianae</emphasis>
|
||
</taxonomicName>
|
||
, rather more uniform and less strongly irrorated than in many
|
||
<taxonomicName id="4C2E8DA74C25C6041B3C2BB8FA4727D8" box="[1308,1428,1685,1708]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A2A364C25C6041B3C2BB8FA4727D8" box="[1308,1428,1685,1708]" italics="true" pageId="83" pageNumber="84">Heteropsis</emphasis>
|
||
</taxonomicName>
|
||
, rather uniformly irrorated brown and ochreous brown. Space-CuA1 ocellus with orange ring yellowing proximad, following darkish brown concave curve of Mb before it bends back to mid-costa. Space-M1 ocellus FWV a white pupil in HT. On HWV, space-CuA1 ocellus small and slightly elliptic with narrow black iris, also without orange ring trace. HWV space-Rs-M2 ocelli as white pupil-points and space-CuA2 ocellus hardly expressed in HT. HWV Mb darkish brown and fairly straight but minutely irregular, only gently curving, with very small yellow Mf distad of it in space-M2. FWV cell area with four darker brown rather equally spaced transverse Cbs. Basal areas not much darker than areas distad of Mb, and darker brown PMb weakly represented.
|
||
<emphasis id="B95A2A364C25C6041A622AA0FB6426D2" bold="true" box="[1090,1207,1933,1958]" pageId="83" pageNumber="84">Variation.</emphasis>
|
||
♂♂ similar to each other, but seasonal variation unknown as the entire
|
||
<typeStatus id="549548864C25C6041CFB2A9FFCD826BE" box="[731,779,1970,1994]" pageId="83" pageNumber="84">type</typeStatus>
|
||
series was taken in a few days in early November. Space- M1 ocellus FWV sometimes features a narrow black iris and space-M2 sometimes just as a white pupil without narrow black iris nor trace of orange ring. Space-CuA2 ocellus HWV slightly expressed in some specimens. ♀♀ similar in pattern, but larger and somewhat lighter dorsally with a more rounded HW shape.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE72DF2FDCA204B" blockId="84.[151,1437,151,2013]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72DF2FEBF218C" bold="true" box="[199,364,223,248]" pageId="84" pageNumber="85">Wingspan/fwl</emphasis>
|
||
: range
|
||
<quantity id="4CD65BC14C22C6031FE62DF2FDBD218C" box="[454,622,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="3.335" metricValueMax="3.55" metricValueMin="3.12" pageId="84" pageNumber="85" unit="mm" value="33.35" valueMax="35.5" valueMin="31.2">31.2–35.5 mm</quantity>
|
||
./17.0–
|
||
<quantity id="4CD65BC14C22C6031C9A2DCDFCFD2183" box="[698,814,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.879" pageId="84" pageNumber="85" unit="mm" value="18.79">18.79 mm</quantity>
|
||
(n=6 ♂♂), mean= 33.8+/ 1.44 SD/17.87 +/- 0.54 SD mm (n= 6 ♂♂), including HT ♂ 32.4/
|
||
<quantity id="4CD65BC14C22C6031C632C2EFD65206F" box="[579,694,259,283]" metricMagnitude="-2" metricUnit="m" metricValue="1.879" pageId="84" pageNumber="85" unit="mm" value="18.79">18.79 mm</quantity>
|
||
. Range 34.0–39.7/17.8–20.0 mm (n=4 ♀♀), mean = 36.2 +/- 3.03 SD/18.75 +/- 1 SD mm (n=4 ♀♀).
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE72C67FE4B2086" blockId="84.[151,1437,151,2013]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72C67FE872017" bold="true" box="[199,340,330,355]" pageId="84" pageNumber="85">Androconia</emphasis>
|
||
: HWD vein 1A+2A has a narrowly tapered inflation and 3A is much more swollen distad, from base to mid-vein, covered in thin grey-brown scales. The length of HW veins M2, M3, CuA1 and CuA2 are also narrowly inflated, and have specialized scales on the dorsal surface. Abdominal black androconia are just visible ventro-laterally around A4–A5. HWD discocellular brush dark brown. Sdp HWD grey, small, lenticular, composed of narrow grey scales).
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE72CD0FE672329" blockId="84.[151,1437,151,2013]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72CD0FEDB2362" bold="true" box="[199,264,509,534]" pageId="84" pageNumber="85">Palps</emphasis>
|
||
: penultimate segment with narrow brown medial strip, flanked by yellow more towards where labial palp potentially wipes compound eye and fringed by dark brown scales, with yellow scales mainly on mesad face, brown inside towards tip.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE72F45FECB2292" blockId="84.[151,1437,151,2013]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72F45FE9823F5" bold="true" box="[199,331,616,641]" pageId="84" pageNumber="85">♂ genitalia</emphasis>
|
||
: 371 DL (
|
||
<figureCitation id="1315EAA14C22C6031FE42F45FDCB23F5" box="[452,536,616,641]" captionStart="FIGURE 25" captionStartId="80.[151,250,1045,1067]" captionTargetBox="[254,1341,201,1005]" captionTargetId="figure@80.[231,1356,193,1023]" captionTargetPageId="80" captionText="FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264622/files/figure.png" pageId="84" pageNumber="85">Fig. 25</figureCitation>
|
||
B, PT): description more or less as for
|
||
<taxonomicName id="4C2E8DA74C22C6031DF42F44FB9523F4" box="[980,1094,617,640]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||
<emphasis id="B95A2A364C22C6031DF42F44FB9523F4" box="[980,1094,617,640]" italics="true" pageId="84" pageNumber="85">Ht. tianae</emphasis>
|
||
</taxonomicName>
|
||
: from LV, uncus very slightly proud of dorsal curve of tegumen (which has a small proximad notch from DV) and slightly longer than it, fairly straight and parallel-sided to hook without a distinct ‘head’ towards tip (inflated from DV before tip), fairly narrow at hinge with vinculum. Valve without prominent dorsal ‘shoulder’. Valve arm with fairly short ‘neck’ and slight club at tip covered in spinoid setae (its crest particularly raised from DV), with distinct ‘beak’ oriented towards uncus and slightly mesad, with none of valve tip proud of uncus. Gnathos from rather small base with deepened (although not ear-like structure) near base, recurved downwards at tapered tip (appearing sinuate from DV and slightly inrecurved at pointed tip). Saccus not very long and parallel sided, aedeagus slightly shorter than valve and quite recurved, both towards and away from ostium, also proximally uprecurved. ♂ genitalia different from those of potentially sympatric (
|
||
<taxonomicName id="4C2E8DA74C22C6031F992E86FCDB22B7" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[441,776,938,963]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="anceps">
|
||
<emphasis id="B95A2A364C22C6031F992E86FDE022B6" box="[441,563,939,962]" italics="true" pageId="84" pageNumber="85">Ht. anceps</emphasis>
|
||
<bibRefCitation id="EFBF8BD54C22C6031C1D2E87FCDB22B7" author="Oberthur" box="[573,776,938,963]" pageId="84" pageNumber="85" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">(Oberthür, 1916))</bibRefCitation>
|
||
</taxonomicName>
|
||
, which are larger and with broader distal valve tip (
|
||
<bibRefCitation id="EFBF8BD54C22C6031B7F2E87FED82292" author="Lees" pageId="84" pageNumber="85" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 107</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C22C6031EE72EDCFE47255A" pageId="84" pageNumber="85" type="etymology">
|
||
<paragraph id="8B91F6244C22C6031EE72EDCFE47255A" blockId="84.[151,1437,151,2013]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72EDCFE9D257E" bold="true" box="[199,334,1009,1034]" pageId="84" pageNumber="85">Etymology.</emphasis>
|
||
after Charles Oberthür, who in 1916 produced one of the first treatments, a seminal work on the
|
||
<collectingCountry id="F339B6B44C22C6031EB72938FED7255A" box="[151,260,1045,1070]" name="Madagascar" pageId="84" pageNumber="85">Malagasy</collectingCountry>
|
||
<taxonomicName id="4C2E8DA74C22C6031F2B2938FE43255A" box="[267,400,1045,1070]" class="Insecta" family="Nymphalidae" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="subTribe" subTribe="Mycalesina">Mycalesina</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C22C6031EE72914FEFE245C" pageId="84" pageNumber="85" type="discussion">
|
||
<paragraph id="8B91F6244C22C6031EE72914FA5B25C9" blockId="84.[151,1437,151,2013]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72914FE9F2526" bold="true" box="[199,332,1081,1106]" pageId="84" pageNumber="85">Discussion.</emphasis>
|
||
No historical material has been found in museums. The species was first found in the field during an expedition to northwest RNI Zahamena in
|
||
<date id="FF90D0E44C22C6031C812971FC882501" box="[673,859,1116,1141]" pageId="84" pageNumber="85" value="2006-11">November 2006</date>
|
||
, where it was hoped to rediscover
|
||
<taxonomicName id="4C2E8DA74C22C6031AC82973FAB22501" box="[1256,1377,1118,1141]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="anceps">
|
||
<emphasis id="B95A2A364C22C6031AC82973FAB22501" box="[1256,1377,1118,1141]" italics="true" pageId="84" pageNumber="85">Ht. anceps</emphasis>
|
||
</taxonomicName>
|
||
. The STs of the similar species
|
||
<taxonomicName id="4C2E8DA74C22C6031FF229ADFC5325ED" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[466,896,1152,1177]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="antsianakana">
|
||
<emphasis id="B95A2A364C22C6031FF229ADFD1525ED" box="[466,710,1152,1177]" italics="true" pageId="84" pageNumber="85">Culapa antsianakana</emphasis>
|
||
<bibRefCitation id="EFBF8BD54C22C6031CF229ADFC5325ED" author="Oberthur" box="[722,896,1152,1177]" pageId="84" pageNumber="85" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||
</taxonomicName>
|
||
and of
|
||
<taxonomicName id="4C2E8DA74C22C6031DC229ADFA9425ED" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[994,1351,1152,1177]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="anceps">
|
||
<emphasis id="B95A2A364C22C6031DC229ADFB5D25EC" box="[994,1166,1152,1177]" italics="true" pageId="84" pageNumber="85">Culapa anceps</emphasis>
|
||
<bibRefCitation id="EFBF8BD54C22C6031AB929ADFA9425ED" author="Oberthur" box="[1177,1351,1152,1177]" pageId="84" pageNumber="85" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||
</taxonomicName>
|
||
(LT ♂ designated above under
|
||
<taxonomicName id="4C2E8DA74C22C6031F862988FD9125C8" box="[422,578,1189,1212]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="roussettae">
|
||
<emphasis id="B95A2A364C22C6031F862988FD9125C8" box="[422,578,1189,1212]" italics="true" pageId="84" pageNumber="85">Ht. roussettae</emphasis>
|
||
</taxonomicName>
|
||
) were examined but the species strongly resembles
|
||
<taxonomicName id="4C2E8DA74C22C6031AA92988FB2A25C8" box="[1161,1273,1189,1212]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||
<emphasis id="B95A2A364C22C6031AA92988FB2A25C8" box="[1161,1273,1189,1212]" italics="true" pageId="84" pageNumber="85">Ht. tianae</emphasis>
|
||
</taxonomicName>
|
||
(see above).
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE729EAFEFE245C" blockId="84.[151,1437,151,2013]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE729EAFD202594" bold="true" box="[199,755,1223,1248]" pageId="84" pageNumber="85">Additional information. DNA divergences:</emphasis>
|
||
COI-5P cluster number BOLD:ACW4996 (exemplar BMAD242-15, DL06-11), about 2.58% divergent to
|
||
<taxonomicName id="4C2E8DA74C22C6031D3229C0FC592470" box="[786,906,1261,1284]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||
<emphasis id="B95A2A364C22C6031D3229C0FC592470" box="[786,906,1261,1284]" italics="true" pageId="84" pageNumber="85">Ht. tianae</emphasis>
|
||
</taxonomicName>
|
||
(BOLD:AAE4112, exemplar BMAD200-15, DL14Z-051).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C22C6031EE7281EFE042732" pageId="84" pageNumber="85" type="description">
|
||
<paragraph id="8B91F6244C22C6031EE7281EFE8624E7" blockId="84.[151,1437,151,2013]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE7281EFE362438" bold="true" box="[199,485,1331,1356]" pageId="84" pageNumber="85">Phylogeny/sister species</emphasis>
|
||
: this species was not known at the time of
|
||
<bibRefCitation id="EFBF8BD54C22C6031DE92819FB80243F" author="Lees" box="[969,1107,1331,1355]" pageId="84" pageNumber="85" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997)</bibRefCitation>
|
||
. Closely related to
|
||
<taxonomicName id="4C2E8DA74C22C6031B0C2819FA4F243F" box="[1324,1436,1332,1355]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||
<emphasis id="B95A2A364C22C6031B0C2819FA4F243F" box="[1324,1436,1332,1355]" italics="true" pageId="84" pageNumber="85">Ht. tianae</emphasis>
|
||
</taxonomicName>
|
||
(sister in
|
||
<bibRefCitation id="EFBF8BD54C22C6031F20287BFDC6241B" author="Aduse-Poku" box="[256,533,1366,1391]" pageId="84" pageNumber="85" refString="Aduse-Poku, K., Lees, D. C., Brattstrom, O., Kodandaramaiah, U., Collins, S. & Brakefield, P. M., (2016) Molecular phylogeny and generic-level taxonomy of the widespread palaeotropical ' Heteropsis clade' (Nymphalidae: Satyrinae: Mycalesina). Systematic Entomology, in press. http: // dx. doi. org / syen. 12183" type="book" year="2016">
|
||
Aduse-Poku
|
||
<emphasis id="B95A2A364C22C6031FB22875FE18241B" box="[402,459,1366,1391]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
|
||
, 2016
|
||
</bibRefCitation>
|
||
, in press). which it closely resembles, and based on the close relationship of the COI-5P barcode.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE728B3FE2524C3" blockId="84.[151,1437,151,2013]" box="[199,502,1438,1463]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE728B3FE2524C3" bold="true" box="[199,502,1438,1463]" pageId="84" pageNumber="85">Ecology and distribution.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE728EFFDD724AF" blockId="84.[151,1437,151,2013]" box="[199,516,1474,1499]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE728EFFEF024AF" bold="true" box="[199,291,1474,1499]" pageId="84" pageNumber="85">Habitat</emphasis>
|
||
: primary rainforest.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE728C8FE6A248A" blockId="84.[151,1437,151,2013]" box="[199,441,1509,1534]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE728C8FE90248A" bold="true" box="[199,323,1509,1534]" pageId="84" pageNumber="85">Behaviour</emphasis>
|
||
: flies low.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE72B24FCB12756" blockId="84.[151,1437,151,2013]" box="[199,866,1545,1570]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72B24FEEE2756" bold="true" box="[199,317,1545,1570]" pageId="84" pageNumber="85">Hostplant</emphasis>
|
||
: unknown, but likely to be low-growing grasses.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE72B00FE042732" blockId="84.[151,1437,151,2013]" box="[199,471,1581,1606]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72B00FE8B2732" bold="true" box="[199,344,1581,1606]" pageId="84" pageNumber="85">Early stages</emphasis>
|
||
: unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C22C6031EE72B7CFC2427F9" pageId="84" pageNumber="85" type="distribution">
|
||
<paragraph id="8B91F6244C22C6031EE72B7CFBA8271E" blockId="84.[151,1437,151,2013]" box="[199,1147,1617,1642]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72B7CFE8A271E" bold="true" box="[199,345,1617,1642]" pageId="84" pageNumber="85">Distribution</emphasis>
|
||
: as far as is known, endemic to RNI Zahamena (
|
||
<figureCitation id="1315EAA14C22C6031D5C2B7CFC1C271D" box="[892,975,1617,1642]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="84" pageNumber="85">Fig. 30</figureCitation>
|
||
C, brown dots).
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C22C6031EE72B59FC2427F9" blockId="84.[151,1437,151,2013]" box="[199,1015,1652,1677]" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72B59FE4A27F9" bold="true" box="[199,409,1652,1677]" pageId="84" pageNumber="85">Elevational range</emphasis>
|
||
:
|
||
<quantity id="4CD65BC14C22C6031F882B58FD9627F8" box="[424,581,1653,1677]" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" metricValueMax="1.31" metricValueMin="1.18" pageId="84" pageNumber="85" unit="m" value="1245.0" valueMax="1310.0" valueMin="1180.0">1180–1310 m</quantity>
|
||
. (n=27, including referred specimens).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C22C6021EE72BB6FBA92005" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="materials_examined">
|
||
<paragraph id="8B91F6244C22C6021EE72BB6FBA92005" blockId="84.[151,1437,151,2013]" lastBlockId="85.[151,1436,151,369]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
|
||
<emphasis id="B95A2A364C22C6031EE72BB6FE6F27C0" bold="true" box="[199,444,1691,1716]" pageId="84" pageNumber="85">Referred specimens.</emphasis>
|
||
♀,
|
||
<collectingCountry id="F339B6B44C22C6031FCA2BB6FDA227C0" box="[490,625,1691,1716]" name="Madagascar" pageId="84" pageNumber="85">Madagascar</collectingCountry>
|
||
E, returning to Bemoara camp, Zahamena,
|
||
<geoCoordinate id="EE1A90E34C22C6031A4D2BB6FB3227C0" box="[1133,1249,1688,1716]" direction="south" orientation="latitude" pageId="84" pageNumber="85" precision="5" value="-17.5376">17.5376o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C22C6031AD02BB6FAA227C7" box="[1264,1393,1688,1716]" direction="east" orientation="longitude" pageId="84" pageNumber="85" precision="1" value="48.71965">48.71965o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C22C6031EB72B92FF2A27AC" box="[151,249,1727,1752]" metricMagnitude="2" metricUnit="m" metricValue="9.399999999999999" pageId="84" pageNumber="85" unit="km" value="0.94">0.94 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C22C6031F272B92FE6C27A3" box="[263,447,1727,1752]" metricMagnitude="3" metricUnit="m" metricValue="1.1800000000000002" metricValueMax="1.3050000000000002" metricValueMin="1.0550000000000002" pageId="84" pageNumber="85" unit="m" value="1180.0" valueMax="1305.0" valueMin="1055.0">1180 +/- 125 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C22C6031FEE2B92FD9227AC" box="[462,577,1727,1752]" pageId="84" pageNumber="85" value="2006-11-05">5/11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C22C6031CF62B92FD2E27AC" box="[726,765,1727,1752]" pageId="84" pageNumber="85">DL</collectionCode>
|
||
06-060, G03 [leg for
|
||
<collectionCode id="ED3F6EE14C22C6031DD72BEDFBE927AC" box="[1015,1082,1728,1752]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="84" pageNumber="85">DNA</collectionCode>
|
||
]; specimen, data as above but:
|
||
<collectionCode id="ED3F6EE14C22C6031EB72BC8FF6C278A" box="[151,191,1765,1790]" pageId="84" pageNumber="85">DL</collectionCode>
|
||
06-066; specimen, data as above but:
|
||
<collectionCode id="ED3F6EE14C22C6031C452BC8FD5E278A" box="[613,653,1765,1790]" pageId="84" pageNumber="85">DL</collectionCode>
|
||
06-067; ♀, E, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C22C6031DC32BC8FB85278A" box="[995,1110,1762,1790]" direction="south" orientation="latitude" pageId="84" pageNumber="85" precision="5" value="-17.5131">17.5131o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C22C6031A442BC8FB042789" box="[1124,1239,1762,1790]" direction="east" orientation="longitude" pageId="84" pageNumber="85" precision="5" value="48.7269">48.7269o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C22C6031B272BC8FA8A278A" box="[1287,1369,1765,1790]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="84" pageNumber="85" unit="km" value="1.5">1.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C22C6031B442BC8FF2A2650" metricMagnitude="3" metricUnit="m" metricValue="1.064" metricValueMax="1.089" metricValueMin="1.039" pageId="84" pageNumber="85" unit="m" value="1064.0" valueMax="1089.0" valueMin="1039.0">1064 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C22C6031F242A21FEA42651" box="[260,375,1804,1829]" pageId="84" pageNumber="85" value="2006-11-05">5/11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C22C6031C212A21FDFA2651" box="[513,553,1804,1829]" pageId="84" pageNumber="85">DL</collectionCode>
|
||
06-061; ♂, E, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C22C6031D5A2A21FC022650" box="[890,977,1800,1829]" direction="south" orientation="latitude" pageId="84" pageNumber="85" precision="555" value="-17.53">17.53o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C22C6031DFD2A21FB822650" box="[989,1105,1800,1829]" direction="east" orientation="longitude" pageId="84" pageNumber="85" precision="5" value="48.7269">48.7269o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C22C6031A5F2A21FB1C2651" box="[1151,1231,1804,1829]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="84" pageNumber="85" unit="km" value="1.5">1.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C22C6031AFA2A21FAA82650" box="[1242,1403,1804,1829]" metricMagnitude="3" metricUnit="m" metricValue="1.064" metricValueMax="1.089" metricValueMin="1.039" pageId="84" pageNumber="85" unit="m" value="1064.0" valueMax="1089.0" valueMin="1039.0">1064 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C22C6031BA62A21FF27263C" pageId="84" pageNumber="85" value="2006-11-06">6/ 11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C22C6031FA12A02FE7A263C" box="[385,425,1839,1864]" pageId="84" pageNumber="85">DL</collectionCode>
|
||
06-070; ♂, data as above but:
|
||
<collectionCode id="ED3F6EE14C22C6031CDA2A02FCF1263C" box="[762,802,1839,1864]" pageId="84" pageNumber="85">DL</collectionCode>
|
||
06-118,
|
||
<collectionCode id="ED3F6EE14C22C6031D5D2A02FC07263C" box="[893,980,1839,1864]" pageId="84" pageNumber="85">BMAD</collectionCode>
|
||
242-15 [
|
||
<collectionCode id="ED3F6EE14C22C6031A0E2A1DFBBD263C" box="[1070,1134,1840,1864]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="84" pageNumber="85">DNA</collectionCode>
|
||
barcode]; ♀, E, just above cascade, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C22C6031F912A7BFDF6261B" box="[433,549,1875,1903]" direction="south" orientation="latitude" pageId="84" pageNumber="85" precision="5" value="-17.5376">17.5376o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C22C6031C112A7BFD60261A" box="[561,691,1875,1903]" direction="east" orientation="longitude" pageId="84" pageNumber="85" precision="1" value="48.71965">48.71965o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C22C6031CC22A7AFC93261B" box="[738,832,1878,1903]" metricMagnitude="2" metricUnit="m" metricValue="9.399999999999999" pageId="84" pageNumber="85" unit="km" value="0.94">0.94 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C22C6031D6B2A7AFC29261A" box="[843,1018,1878,1903]" metricMagnitude="3" metricUnit="m" metricValue="1.1800000000000002" metricValueMax="1.3050000000000002" metricValueMin="1.0550000000000002" pageId="84" pageNumber="85" unit="m" value="1180.0" valueMax="1305.0" valueMin="1055.0">1180 +/- 125 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C22C6031A252A7BFBA9261B" box="[1029,1146,1878,1903]" pageId="84" pageNumber="85" value="2006-11-06">6/11/2006</date>
|
||
: 15:29, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C22C6031B712A7BFAAB261B" box="[1361,1400,1878,1903]" pageId="84" pageNumber="85">DL</collectionCode>
|
||
06- 084; ♂, data as above but [without time]:
|
||
<collectionCode id="ED3F6EE14C22C6031C512A57FD4A26E7" box="[625,665,1914,1939]" pageId="84" pageNumber="85">DL</collectionCode>
|
||
06-132,
|
||
<collectionCode id="ED3F6EE14C22C6031CD52A57FCF826E7" box="[757,811,1914,1939]" country="USA" httpUri="http://grbio.org/cool/s9x3-awht" name="Dicty Stock Center" pageId="84" pageNumber="85">DSC</collectionCode>
|
||
_0825 [photo]; ♀, data as above but:
|
||
<collectionCode id="ED3F6EE14C22C6031AF22A57FB2926E7" box="[1234,1274,1914,1939]" pageId="84" pageNumber="85">DL</collectionCode>
|
||
06-119, IA432 [isotope voucher], KAP518 [=KA-P518; extract number, sequenced specimen in Aduse-Poku
|
||
<emphasis id="B95A2A364C22C6031B272AB2FA9426C2" box="[1287,1351,1949,1974]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
|
||
, 2015, mislabelled as
|
||
<taxonomicName id="4C2E8DA74C22C6031F1D2AE8FDD826A8" box="[317,523,1989,2012]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="anceps">
|
||
<emphasis id="B95A2A364C22C6031F1D2AE8FDD826A8" box="[317,523,1989,2012]" italics="true" pageId="84" pageNumber="85">Heteropsis anceps</emphasis>
|
||
</taxonomicName>
|
||
); ♀, data as above but:
|
||
<collectionCode id="ED3F6EE14C22C6031D362AE9FCEC26A9" box="[790,831,1988,2013]" pageId="84" pageNumber="85">DL</collectionCode>
|
||
06-124; ♂, E, Ambavala camp, Zahamena,
|
||
<geoCoordinate id="EE1A90E34C22C6031B032AE9FA4526A9" box="[1315,1430,1985,2013]" direction="south" orientation="latitude" pageId="84" pageNumber="85" precision="5" value="-17.5451">17.5451o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C23C6021EB72DB6FED921C6" box="[151,266,151,179]" direction="east" orientation="longitude" pageId="85" pageNumber="86" precision="5" value="48.7237">48.7237o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C23C6021F1B2DB7FE5F21C7" box="[315,396,154,179]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="85" pageNumber="86" unit="km" value="0.5">0.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C23C6021FB92DB6FDED21C6" box="[409,574,154,179]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.35" metricValueMin="1.25" pageId="85" pageNumber="86" unit="m" value="1300.0" valueMax="1350.0" valueMin="1250.0">1300 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C23C6021C6B2DB7FD6D21C7" box="[587,702,154,179]" pageId="85" pageNumber="86" value="2006-11-06">6/11/2006</date>
|
||
, 13:23, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C23C6021DBB2DB7FC1721C7" box="[923,964,154,179]" pageId="85" pageNumber="86">DL</collectionCode>
|
||
06-126; ♂, data as above but: 13:16, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C23C6021EFA2DECFED021AE" box="[218,259,193,218]" pageId="85" pageNumber="86">DL</collectionCode>
|
||
06-127; ♂, E, Ankosy summit, Zahamena,
|
||
<geoCoordinate id="EE1A90E34C23C6021CC02DECFC8721AD" box="[736,852,190,218]" direction="south" orientation="latitude" pageId="85" pageNumber="86" precision="5" value="-17.4946">17.4946o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C23C6021D402DECFC1621AD" box="[864,965,190,217]" direction="east" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="48.733">48.733o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C23C6021DD32DECFBFC21AE" box="[1011,1071,193,218]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="85" pageNumber="86" unit="km" value="1.0">1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C23C6021A192DECFB4321AD" box="[1081,1168,193,217]" metricMagnitude="3" metricUnit="m" metricValue="1.321" pageId="85" pageNumber="86" unit="m" value="1321.0">1321 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C23C6021ABB2DECFADC21AE" box="[1179,1295,193,218]" pageId="85" pageNumber="86" value="2006-11-07">7/11/2006</date>
|
||
, 10:37, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C23C6021EFB2DCAFED72074" box="[219,260,231,256]" pageId="85" pageNumber="86">DL</collectionCode>
|
||
06-156; ♂, E, near Ankosy summit, Zahamena,
|
||
<geoCoordinate id="EE1A90E34C23C6021D3E2DCAFC4C2074" box="[798,927,228,256]" direction="south" orientation="latitude" pageId="85" pageNumber="86" precision="1" value="-17.49405">17.49405o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C23C6021D8C2DCAFBFD218B" box="[940,1070,228,256]" direction="east" orientation="longitude" pageId="85" pageNumber="86" precision="1" value="48.73325">48.73325o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C23C6021A7E2DCAFB192074" box="[1118,1226,231,256]" metricMagnitude="1" metricUnit="m" metricValue="6.7" pageId="85" pageNumber="86" unit="km" value="0.067">0.067 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C23C6021AF72DC5FAA9218B" box="[1239,1402,231,256]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.325" metricValueMin="1.275" pageId="85" pageNumber="86" unit="m" value="1300.0" valueMax="1325.0" valueMin="1275.0">1300 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C23C6021BA62DC5FF272053" pageId="85" pageNumber="86" value="2006-11-07">7/ 11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C23C6021F5D2C23FE752053" box="[381,422,270,295]" pageId="85" pageNumber="86">DL</collectionCode>
|
||
06-188; ♀, E, Zahamena,
|
||
<geoCoordinate id="EE1A90E34C23C6021CE32C23FCE52053" box="[707,822,267,295]" direction="south" orientation="latitude" pageId="85" pageNumber="86" precision="5" value="-17.5131">17.5131o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C23C6021D622C23FC652052" box="[834,950,267,295]" direction="east" orientation="longitude" pageId="85" pageNumber="86" precision="5" value="48.7269">48.7269o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C23C6021DC32C23FBE72053" box="[995,1076,270,295]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="85" pageNumber="86" unit="km" value="1.5">1.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C23C6021A1E2C23FB0D2052" box="[1086,1246,270,295]" metricMagnitude="3" metricUnit="m" metricValue="1.064" metricValueMax="1.089" metricValueMin="1.039" pageId="85" pageNumber="86" unit="m" value="1064.0" valueMax="1089.0" valueMin="1039.0">1064 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C23C6021AC92C23FA8E2053" box="[1257,1373,270,295]" pageId="85" pageNumber="86" value="2006-11-06">6/11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C23C6021EFA2C19FED12039" box="[218,258,308,333]" pageId="85" pageNumber="86">DL</collectionCode>
|
||
06-67, IA69 [isotope voucher]; ♂, E, Zahamena NW,
|
||
<geoCoordinate id="EE1A90E34C23C6021D7B2C19FC1D2039" box="[859,974,305,333]" direction="south" orientation="latitude" pageId="85" pageNumber="86" precision="5" value="-17.4946">17.4946o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C23C6021DFB2C18FB932039" box="[987,1088,305,333]" direction="east" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="48.733">48.733o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C23C6021A4F2C18FB782039" box="[1135,1195,308,333]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="85" pageNumber="86" unit="km" value="1.0">1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C23C6021A962C18FA8B2039" box="[1206,1368,308,333]" metricMagnitude="3" metricUnit="m" metricValue="1.321" metricValueMax="1.3459999999999999" metricValueMin="1.2959999999999998" pageId="85" pageNumber="86" unit="m" value="1321.0" valueMax="1346.0" valueMin="1296.0">1321 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C23C6021B432C19FF012005" pageId="85" pageNumber="86" value="2006-11-06">6/11/ 2006</date>
|
||
, D.C. Lees: 364
|
||
<collectionCode id="ED3F6EE14C23C6021FA62C75FE7F2005" box="[390,428,344,369]" pageId="85" pageNumber="86">DL</collectionCode>
|
||
[genitalia voucher], IA29 [isotope voucher], leg sample [KAP].
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |