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<document id="8F62CA372B984EAD5474AF25D089BF07" ID-CLB-Dataset="107636" ID-DOI="10.5852/ejt.2022.833.1879" ID-GBIF-Dataset="69674de4-93c1-463c-9ae1-2a9939cfc966" ID-ISSN="2118-9773" ID-Zenodo-Dep="6916920" ID-ZooBank="EADA40BF-86E5-4D5B-8D43-3F6FB8F2C687" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658962515064" checkinUser="felipe" docAuthor="Wichard, Wilfried &amp; Espeland, Marianne" docDate="2022" docId="1A5A8785FF81156AC9BFFB420056E7F7" docLanguage="en" docName="ejt-833_1-11.pdf.imf" docOrigin="European Journal of Taxonomy 833" docSource="http://zoobank.org/eada40bf-86e5-4d5b-8d43-3f6fb8f2c687" docStyle="DocumentStyle:8DBBCD0342F59C7B93E4D60327A07408.3:EJT.2018-.journal_article.type2" docStyleId="8DBBCD0342F59C7B93E4D60327A07408" docStyleName="EJT.2018-.journal_article.type2" docStyleVersion="3" docTitle="Cretapsyche kachini Wichard &amp; Espeland 2022, sp. nov." docType="treatment" docUuid="7B10F380-13D1-4885-9A19-0414DE8F4975" docUuidSource="ZooBank" docVersion="7" lastPageNumber="7" masterDocId="E663FFFDFF85156CCB2CFFBE0526E53A" masterDocTitle="The family Cretapsychidae (Insecta, Trichoptera) from mid- Cretaceous Burmese amber, with descriptions of two new species" masterLastPageNumber="11" masterPageNumber="1" pageNumber="5" updateTime="1699362387126" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9FB3C7A758705EACD613D75F1A6D174C">The family Cretapsychidae (Insecta, Trichoptera) from mid- Cretaceous Burmese amber, with descriptions of two new species</mods:title>
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<mods:namePart id="BDD4095555433B93FBDD08800E4B369D">Wichard, Wilfried</mods:namePart>
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<mods:affiliation id="3BE6B9828076AF183093D276120F5FE1">Institute of Biology, University of Cologne, Herbert Lewinstrasse 2, 50931 Köln, Germany.</mods:affiliation>
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<mods:namePart id="91AC58C1F916A6FE2FEF98F016983C00">Espeland, Marianne</mods:namePart>
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<mods:affiliation id="80F5B9994E2B41C5FDD05D19D1C0992A">Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauerallee 127, 53113 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="57EC6CFBB98D1638EDF5136A9D8A297F" type="email">m.espeland@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:date id="722C68327718608262CED826F57AC80E">2022</mods:date>
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<subSubSection id="DAE96518FF811568C9BFFB4206CAE02D" box="[659,1004,1276,1303]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="924C3693FF811568C9BFFB4206CAE02D" blockId="4.[415,1248,1276,1374]" box="[659,1004,1276,1303]" pageId="4" pageNumber="5">
<heading id="C90481FFFF811568C9BFFB4206CAE02D" box="[659,1004,1276,1303]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="55F34D10FF811568C9BFFB4206ACE02D" authority="Wichard &amp; Espeland, 2022" authorityName="Wichard &amp; Espeland" authorityYear="2022" box="[659,906,1276,1303]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kachini" status="sp. nov.">
<emphasis id="A087EA81FF811568C9BFFB4206ACE02D" bold="true" box="[659,906,1276,1303]" italics="true" pageId="4" pageNumber="5">Cretapsyche kachini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBB457FAFF811568C8BEFB4306CAE02D" box="[914,1004,1277,1303]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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urn:lsid:zoobank.org:act:
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<figureCitation id="0AC82A16FF811568C831FAFA0644E064" box="[797,866,1348,1374]" captionStart="Fig" captionStartId="5.[227,270,1964,1990]" captionTargetBox="[263,1401,302,1912]" captionTargetId="figure-52@5.[263,1401,302,1912]" captionTargetPageId="5" captionText="Fig. 3. Cretapsyche kachini sp. nov., holotype, ♂ (ZFMK-TRI000829). A. Dorsal view. B. Drawing of a forewing. C. Maxillary and labial palps. D. Drawing of the maxillary palp. E. Male genitalia in ventrolateral view, distinctly flattened and partially distorted. F. Drawing of male genitalia, left half." figureDoi="http://doi.org/10.5281/zenodo.6916926" httpUri="https://zenodo.org/record/6916926/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
</paragraph>
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<subSubSection id="DAE96518FF811568CBCFFA3B044BE303" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="924C3693FF811568CBCFFA3B047AE0A5" blockId="4.[227,348,1413,1439]" box="[227,348,1413,1439]" pageId="4" pageNumber="5">
<heading id="C90481FFFF811568CBCFFA3B047AE0A5" bold="true" box="[227,348,1413,1439]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="A087EA81FF811568CBCFFA3B047AE0A5" bold="true" box="[227,348,1413,1439]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="924C3693FF811568CBCFFA0A044BE303" blockId="4.[227,1438,1460,1593]" pageId="4" pageNumber="5">
<taxonomicName id="55F34D10FF811568CBCFFA0A04C5E0F4" authority="Wichard &amp; Espeland, 2022" authorityName="Wichard &amp; Espeland" authorityYear="2022" box="[227,483,1460,1486]" class="Insecta" family="Cretapsychidae" genus="Creatapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kachini" status="sp. nov.">
<emphasis id="A087EA81FF811568CBCFFA0A04C5E0F4" box="[227,483,1460,1486]" italics="true" pageId="4" pageNumber="5">Creatapsyche kachini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBB457FAFF811568CAC3FA0B076DE0F4" box="[495,587,1461,1487]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from all other species in the genus by having Rs branching before M in the forewing. In all other species, Rs and M branch at about the same level, or M branches before Rs. Additionally, in the male genitalia the harpago is two-thirds the length of the coxopodite.
</paragraph>
</subSubSection>
<subSubSection id="DAE96518FF811568CBCFF9DF0416E3F7" pageId="4" pageNumber="5" type="etymology">
<paragraph id="924C3693FF811568CBCFF9DF044CE341" blockId="4.[227,362,1633,1659]" box="[227,362,1633,1659]" pageId="4" pageNumber="5">
<heading id="C90481FFFF811568CBCFF9DF044CE341" bold="true" box="[227,362,1633,1659]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="A087EA81FF811568CBCFF9DF044CE341" bold="true" box="[227,362,1633,1659]" pageId="4" pageNumber="5">Etymology</emphasis>
</heading>
</paragraph>
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<taxonomicName id="55F34D10FF811568CBCFF92E04F5E390" authority="Wichard &amp; Espeland, 2022" authorityName="Wichard &amp; Espeland" authorityYear="2022" box="[227,467,1680,1706]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kachini" status="sp. nov.">
<emphasis id="A087EA81FF811568CBCFF92E04F5E390" box="[227,467,1680,1706]" italics="true" pageId="4" pageNumber="5">Cretapsyche kachini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBB457FAFF811568CAF7F92E0711E390" box="[475,567,1680,1706]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
is named after
<collectingRegion id="5037F871FF811568C9C2F92E06A0E390" box="[750,902,1680,1706]" country="Myanmar" name="Kachin" pageId="4" pageNumber="5">Kachin State</collectingRegion>
in
<collectingCountry id="EAE47603FF811568C881F92E0107E390" box="[941,1057,1680,1706]" name="Myanmar" pageId="4" pageNumber="5">Myanmar</collectingCountry>
, where the amber inclusion was found.
</paragraph>
</subSubSection>
<subSubSection id="DAE96518FF811568CBCFF94B061CED34" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="924C3693FF811568CBCFF94B04F6E235" blockId="4.[227,464,1781,1807]" box="[227,464,1781,1807]" pageId="4" pageNumber="5">
<heading id="C90481FFFF811568CBCFF94B04F6E235" bold="true" box="[227,464,1781,1807]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="A087EA81FF811568CBCFF94B04F6E235" bold="true" box="[227,464,1781,1807]" pageId="4" pageNumber="5">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="924C3693FF811568CA3CF89A04EBE293" blockId="4.[227,1437,1828,1961]" pageId="4" pageNumber="5">
<materialsCitation id="229B3CCEFF811568CA3CF89A04EBE293" ID-GBIF-Occurrence="3866793301" collectionCode="ZFMK" country="Myanmar" location="Kachin State" pageId="4" pageNumber="5" specimenCode="ZFMK-TRI000829" specimenCount="1" specimenCount-male="1" stateProvince="Kachin State" typeStatus="holotype">
<emphasis id="A087EA81FF811568CA3CF89A04A4E204" bold="true" box="[272,386,1828,1854]" pageId="4" pageNumber="5">
<typeStatus id="4D488831FF811568CA3CF89A04A4E204" box="[272,386,1828,1854]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="EAE47603FF811568CBCFF8F6045AE258" box="[227,380,1864,1890]" name="Myanmar" pageId="4" pageNumber="5">MYANMAR</collectingCountry>
<emphasis id="A087EA81FF811568CAB0F8F90776E258" bold="true" box="[412,592,1863,1890]" pageId="4" pageNumber="5">
<collectingRegion id="5037F871FF811568CAB0F8F90719E25B" box="[412,575,1863,1889]" country="Myanmar" name="Kachin" pageId="4" pageNumber="5">Kachin State</collectingRegion>
</emphasis>
<specimenCount id="84F5FD1AFF811568C974F8F60757E258" box="[600,625,1864,1890]" pageId="4" pageNumber="5" type="male"></specimenCount>
; exact locality unknown;
<geologicalTimeScale id="E0D3B638FF811568C88EF8F600BEE258" box="[930,1432,1864,1890]" pageId="4" pageNumber="5">Mid-Cretaceous Burmese amber inclusion</geologicalTimeScale>
; deposited in the amber collection of the
<collectionCode id="F4E2AE56FF811568C9D7F8D20676E2BC" box="[763,848,1900,1926]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="5" type="Museum">ZFMK</collectionCode>
; former Patrick Müller Collection, BUB 540;
<specimenCode id="C2559EE8FF811568CBCFF83104EFE293" box="[227,457,1935,1961]" collectionCode="ZFMK-TRI" pageId="4" pageNumber="5">ZFMK-TRI000829</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="924C3693FF811568CBCFF86E061CED34" blockId="4.[227,1436,2000,2063]" pageId="4" pageNumber="5">Body well preserved; forewing in dorsal view, hindwing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view.</paragraph>
</subSubSection>
<caption id="C68C661BFF801569CBCFF812005DED37" ID-DOI="http://doi.org/10.5281/zenodo.6916926" ID-Zenodo-Dep="6916926" httpUri="https://zenodo.org/record/6916926/files/figure.png" pageId="5" pageNumber="6" startId="5.[227,270,1964,1990]" targetBox="[263,1401,302,1912]" targetPageId="5">
<paragraph id="924C3693FF801569CBCFF812005DED37" blockId="5.[227,1437,1964,2062]" pageId="5" pageNumber="6">
<emphasis id="A087EA81FF801569CBCFF8120415E2FD" bold="true" box="[227,307,1964,1991]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="55F34D10FF801569CA11F8130709E2FD" authority="Wichard &amp; Espeland, 2022" authorityName="Wichard &amp; Espeland" authorityYear="2022" box="[317,559,1965,1991]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kachini" status="sp. nov.">
<emphasis id="A087EA81FF801569CA11F8130709E2FD" box="[317,559,1965,1991]" italics="true" pageId="5" pageNumber="6">Cretapsyche kachini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBB457FAFF801569C916F81307BCE2FD" box="[570,666,1965,1991]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ZFMK-TRI000829).
<emphasis id="A087EA81FF801569CF65F8120178E2FC" bold="true" box="[1097,1118,1964,1990]" pageId="5" pageNumber="6">A</emphasis>
. Dorsal view.
<emphasis id="A087EA81FF801569CE3CF8130002E2FD" bold="true" box="[1296,1316,1965,1991]" pageId="5" pageNumber="6">B</emphasis>
. Drawing of a forewing.
<emphasis id="A087EA81FF801569CAB5F86E048AE2D0" bold="true" box="[409,428,2000,2026]" pageId="5" pageNumber="6">C</emphasis>
. Maxillary and labial palps.
<emphasis id="A087EA81FF801569C827F86E0638E2D0" bold="true" box="[779,798,2000,2026]" pageId="5" pageNumber="6">D</emphasis>
. Drawing of the maxillary palp.
<emphasis id="A087EA81FF801569CF82F86E0199E2D0" bold="true" box="[1198,1215,2000,2026]" pageId="5" pageNumber="6">E</emphasis>
. Male genitalia in ventrolateral view, distinctly flattened and partially distorted.
<emphasis id="A087EA81FF801569C899F84D06E1ED37" bold="true" box="[949,967,2035,2061]" pageId="5" pageNumber="6">F</emphasis>
. Drawing of male genitalia, left half.
</paragraph>
</caption>
<subSubSection id="DAE96518FF83156ACBCFFE900056E7F7" pageId="6" pageNumber="7" type="description">
<paragraph id="924C3693FF83156ACBCFFE900452E472" blockId="6.[227,372,302,328]" box="[227,372,302,328]" pageId="6" pageNumber="7">
<heading id="C90481FFFF83156ACBCFFE900452E472" bold="true" box="[227,372,302,328]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="A087EA81FF83156ACBCFFE900452E472" bold="true" box="[227,372,302,328]" pageId="6" pageNumber="7">Description</emphasis>
</heading>
</paragraph>
<paragraph id="924C3693FF83156ACA3CFEDE0476E440" blockId="6.[225,1439,352,718]" box="[272,336,352,378]" pageId="6" pageNumber="7">
<emphasis id="A087EA81FF83156ACA3CFEDE0476E440" bold="true" box="[272,336,352,378]" pageId="6" pageNumber="7">Male</emphasis>
</paragraph>
<paragraph id="924C3693FF83156ACBCFFE3B0056E7F7" blockId="6.[225,1439,352,718]" pageId="6" pageNumber="7">
Ocelli absent. Antennae probably about as long as forewings (
<typeStatus id="4D488831FF83156AC8E5FE3B0114E4A5" box="[969,1074,389,415]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
antennae incomplete). Scapus about as long as head. Maxillary palps 5-segmented, 2
<superScript id="65869BDBFF83156AC878FE140643E482" attach="left" box="[852,869,426,440]" fontSize="6" pageId="6" pageNumber="7">nd</superScript>
segment longest, following segments decreasing in length, 3
<superScript id="65869BDBFF83156ACA40FE6E045CE4E4" attach="left" box="[364,378,464,478]" fontSize="6" pageId="6" pageNumber="7">rd</superScript>
segment basally barrel-shaped and extended laterad to form pointed spur corresponding in length to 4
<superScript id="65869BDBFF83156ACAAAFE4804B5E73E" attach="left" box="[390,403,502,516]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
and 5
<superScript id="65869BDBFF83156ACACCFE4804CBE73E" attach="left" box="[480,493,502,516]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
segments. Body and wings light brown. Forewing with length
<quantity id="550B9B76FF83156ACFC7FE49006EE72B" box="[1259,1352,503,529]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="7" unit="mm" value="2.8">2.8 mm</quantity>
longer than small hindwing. In forewings, apical forks II, III and V present, discoidal cell absent, thyridial cell present, Cu2 running continuously to Cu1b. In hindwings, apical forks I, III and V present (difficult to see,
<typeStatus id="4D488831FF83156ACA35FDD604A6E7B8" box="[281,384,616,642]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
forewing hides hindwings in dorsal view). Tibial spur formula 2/2/4. In male genitalia, two-segmented inferior appendages characterized by basally broad coxopodite, tapering and curving mesad. Harpago elongate and tapering continuously towards apex, two-thirds of coxopodite length.
</paragraph>
</subSubSection>
</treatment>
</document>