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<document id="BB2ABFC40F5AF2F6F23714A81FB22AC8" ID-CLB-Dataset="7593" ID-DOI="10.11646/zootaxa.4950.1.12" ID-GBIF-Dataset="766e2d75-af43-43f7-a679-8c5969acb49f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4643348" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617002817976" checkinUser="plazi" docAuthor="Jiang, Ri-Xin, Ballerio, Alberto, Liu, Hao-Yi &amp; Wang, Shuo" docDate="2021" docId="126F87CFE840FFDEFEF5FC53FE441AC9" docLanguage="en" docName="zootaxa.4950.1.12.pdf" docOrigin="Zootaxa 4950 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterorthochaetes yunnanensis Ballerio 2014" docType="treatment" docVersion="5" lastPageNumber="198" masterDocId="EE56FFB7E840FFDDFE62FA36FFE11B06" masterDocTitle="Description of the male of Pterorthochaetes yunnanensis Ballerio, 2014 (Coleoptera Scarabaeoidea: Hybosoridae: Ceratocanthinae)" masterLastPageNumber="200" masterPageNumber="196" pageNumber="196" updateTime="1698935310677" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="33FB23FBC2E4866DDFD63E992D6E224D">
<mods:title id="B2347E70D8A305CC790D52A09794E31D">Description of the male of Pterorthochaetes yunnanensis Ballerio, 2014 (Coleoptera Scarabaeoidea: Hybosoridae: Ceratocanthinae)</mods:title>
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<mods:namePart id="97597B4F97A9210AB1791A0334D0E7EE">Jiang, Ri-Xin</mods:namePart>
<mods:nameIdentifier id="4AB3F1D3E84D9F3390D2A0F86872B3D4" type="ORCID">0000-0002-5339-853X</mods:nameIdentifier>
<mods:affiliation id="6384843435A627772B01A30989ECF625">College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, 53 Zhengzhou Road, Qing- dao, 266042, P. R. China. &amp; maoshuwuyouzhi @ 163. com; https: // orcid. org / 0000 - 0002 - 5339 - 853 X</mods:affiliation>
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<mods:namePart id="CD8ED2915B500E3993B3AE679CA88915">Ballerio, Alberto</mods:namePart>
<mods:affiliation id="8FE57D8E88C2F72598CBEB1DFF567761">Viale Venezia 45, Brescia, 2512, Italy.</mods:affiliation>
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</mods:role>
<mods:namePart id="DA53F325CCDF2FCAD7134C51B6B95750">Liu, Hao-Yi</mods:namePart>
<mods:nameIdentifier id="ADD74F8D9223190D0E88E0E0F016D043" type="ORCID">0000-0002-5600-1921</mods:nameIdentifier>
<mods:affiliation id="3E5763BEE790DADCA7EA78F3D285254A">Zhengzhou Foreign Language School-New Fengyang Campus, 66 Cuizhu Street, Zhengzhou, 450001, P. R. China. 3137068712 @ qq. com; https: // orcid. org / 0000 - 0002 - 5600 - 1921</mods:affiliation>
<mods:nameIdentifier id="7A1CC8924DE201C7AD47EE8FE2A0DB6E" type="email">3137068712@qq.com</mods:nameIdentifier>
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<mods:namePart id="D0DF622ADAFBF4FA39FC449F5B407B4F">Wang, Shuo</mods:namePart>
<mods:nameIdentifier id="6702ECA067B39C9F4A9C5294A29C3426" type="ORCID">0000-0003-0412-3799</mods:nameIdentifier>
<mods:affiliation id="45BFE6C99BF80814AF35D67D27522FDA">College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, 53 Zhengzhou Road, Qing- dao, 266042, P. R. China.</mods:affiliation>
<mods:nameIdentifier id="2263953C47BA5970282ED323B3FD10BE" type="email">shuowang@qust.edu.cn</mods:nameIdentifier>
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<mods:part id="43F49049678CA78B360BEB1E7A5CA3D2">
<mods:date id="8209D896534CA4DB9FE6448E87996264">2021</mods:date>
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<mods:identifier id="10D2C42D7214993DB8D1B36A8EBF01F1" type="DOI">10.11646/zootaxa.4950.1.12</mods:identifier>
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<treatment id="126F87CFE840FFDEFEF5FC53FE441AC9" ID-DOI="http://doi.org/10.5281/zenodo.4643352" ID-GBIF-Taxon="180776260" ID-Zenodo-Dep="4643352" LSID="urn:lsid:plazi:treatment:126F87CFE840FFDEFEF5FC53FE441AC9" httpUri="http://treatment.plazi.org/id/126F87CFE840FFDEFEF5FC53FE441AC9" lastPageId="3" lastPageNumber="198" pageId="0" pageNumber="196">
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<heading id="C13181B5E840FFDDFEF5FC53FC651D86" bold="true" box="[151,900,1637,1664]" fontSize="11" level="1" pageId="0" pageNumber="196" reason="1">
<emphasis id="A8B2EACBE840FFDDFEF5FC53FC651D86" bold="true" box="[151,900,1637,1664]" pageId="0" pageNumber="196">
<taxonomicName id="5DC64D5AE840FFDDFEF5FC53FD581D86" ID-CoL="4Q36H" authority="Ballerio, 2014" authorityName="Ballerio" authorityYear="2014" box="[151,697,1637,1664]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="A8B2EACBE840FFDDFEF5FC53FDE41D86" bold="true" box="[151,517,1637,1664]" italics="true" pageId="0" pageNumber="196">Pterorthochaetes yunnanensis</emphasis>
<bibRefCitation id="FE574B28E840FFDDFC6EFC50FD581D86" author="Ballerio, A." box="[524,697,1638,1664]" pageId="0" pageNumber="196" pagination="277 - 280" refId="ref2402" refString="Ballerio, A. (2014) New data on Palaearctic Ceratocanthinae with description of a new species from China (Coleoptera: Scarabaeoidea: Hybosoridae). Koleopterologische Rundschau, 84, 277 - 280." type="journal article" year="2014">Ballerio, 2014</bibRefCitation>
</taxonomicName>
(云南毛ńƙDz)
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<paragraph id="9A7936D9E840FFDDFEF5FCBAFEE71DA2" blockId="0.[151,900,1637,1700]" box="[151,262,1676,1700]" pageId="0" pageNumber="196">
(
<figureCitation id="02FD2A5CE840FFDDFEFDFCBAFF1D1DA2" box="[159,252,1676,1700]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1263,1287]" captionStartId-1="2.[151,250,1117,1141]" captionStartId-2="3.[151,250,1770,1794]" captionTargetBox-0="[176,1414,424,1234]" captionTargetBox-1="[151,1435,181,1093]" captionTargetBox-2="[151,1435,488,1746]" captionTargetId-0="figure-128@1.[171,1417,416,1240]" captionTargetId-1="figure-19@2.[151,1435,181,1093]" captionTargetId-2="figure-177@3.[151,1435,488,1746]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Male habitus of Pterorthochaetes yunnanensis Ballerio, 2014. A, dorsal view; B, enrolled, ventral view; C, enrolled, lateral view; D, enrolled, dorsal view. Scale bars: 1.0 mm." captionText-1="FIGURE 2. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, protibia, male, dorsal view; B, protibia, male, lateral view; C, mesotibia, male, ventral view; D, metatibia, male, ventral view; E, mesofemur, male, dorsal view; F, protibia, female, dorsal view; G, protibia, female, lateral view; H, mesotibia, female, ventral view; I, metatibia, female, ventral view. Scale bars: 0.5 mm." captionText-2="FIGURE 3. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, aedeagus, dorsal view; B, aedeagus, ventral view; C, parameres of aedeagus, dorsal view; D, left paramere; E, right paramere; F, internal sac of aedeagus; G, genital segment of male; H, bursal sclerites; I, spermatheca. Scale bars: 0.5 mm for A–G, 0.25 mm for H–I." figureDoi-0="http://doi.org/10.5281/zenodo.4643350" figureDoi-1="http://doi.org/10.5281/zenodo.4643354" figureDoi-2="http://doi.org/10.5281/zenodo.4643356" httpUri-0="https://zenodo.org/record/4643350/files/figure.png" httpUri-1="https://zenodo.org/record/4643354/files/figure.png" httpUri-2="https://zenodo.org/record/4643356/files/figure.png" pageId="0" pageNumber="196">Figs 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D2DC6552E840FFDDFEF5FCFFFEBC1C18" pageId="0" pageNumber="196" type="reference_group">
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<treatmentCitationGroup id="BAD611F7E840FFDDFEF5FCFFFEBC1C18" pageId="0" pageNumber="196">
<treatmentCitation id="1B6710C8E840FFDDFEF5FCFFFD7D1DE7" author="Ballerio, A." box="[151,668,1737,1761]" page="277" pageId="0" pageNumber="196" year="2014">
<taxonomicName id="5DC64D5AE840FFDDFEF5FCFFFD7D1DE7" ID-CoL="4Q36H" authority="Ballerio, 2014: 277" authorityName="Ballerio" authorityPageNumber="277" authorityYear="2014" box="[151,668,1737,1761]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="A8B2EACBE840FFDDFEF5FCFFFE2B1DE7" box="[151,458,1737,1761]" italics="true" pageId="0" pageNumber="196">Pterorthochaetes yunnanensis</emphasis>
<materialsCitation id="2AAE3C84E840FFDDFFB1FCFFFD7D1DE7" ID-GBIF-Occurrence="3061987301" box="[467,668,1737,1761]" collectingDate="1999-05-11" collectorName="Jach &amp; Naturhistorisches Museum" country="China" elevation="750" location="Wien" pageId="0" pageNumber="196" specimenCount="1" stateProvince="Wien">
<bibRefCitation id="FE574B28E840FFDDFFB1FCFFFD7D1DE7" author="Ballerio, A." box="[467,668,1737,1761]" pageId="0" pageNumber="196" pagination="277 - 280" refId="ref2402" refString="Ballerio, A. (2014) New data on Palaearctic Ceratocanthinae with description of a new species from China (Coleoptera: Scarabaeoidea: Hybosoridae). Koleopterologische Rundschau, 84, 277 - 280." type="journal article" year="2014">Ballerio, 2014: 277</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(
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locality: “
<collectingCountry id="E2D17649E840FFDDFD24FCFFFC691DE7" box="[838,904,1737,1761]" name="China" pageId="0" pageNumber="196">China</collectingCountry>
: Yunnan, Xishuangbanna (
<emphasis id="A8B2EACBE840FFDDFAFFFCFCFAE31DD9" bold="true" box="[1181,1282,1738,1759]" pageId="0" pageNumber="196">NJVhffi</emphasis>
), ca.
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w Menglun (
<emphasis id="A8B2EACBE840FFDDFF50FCDFFE841DF8" bold="true" box="[306,357,1769,1790]" pageId="0" pageNumber="196">勐仑</emphasis>
),
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<collectingDate id="FE3CE9F1E840FFDDFF19FCDEFE001C06" box="[379,481,1768,1792]" pageId="0" pageNumber="196" value="1999-05-11">5.11.1999</collectingDate>
</date>
, ca.
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<quantity id="5D3E9B3CE840FFDDFC72FCDEFD651DF9" box="[528,644,1768,1792]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="0" pageNumber="196" unit="m" value="750.0" valueMax="800.0" valueMin="700.0">700–800 m</quantity>
a.s.l.
</elevation>
, leg.
<collectorName id="3733530FE840FFDDFC96FCDEFCC51C06" box="[756,804,1768,1792]" pageId="0" pageNumber="196">Jäch</collectorName>
, et al.”, deposited in the
<collectorName id="3733530FE840FFDDFA40FCDEFAD21DF9" box="[1058,1331,1768,1792]" pageId="0" pageNumber="196">Naturhistorisches Museum</collectorName>
<collectingRegion id="5802F83BE840FFDDFB5BFCDEFA931C06" box="[1337,1394,1768,1792]" country="Austria" name="Wien" pageId="0" pageNumber="196">Wien</collectingRegion>
,
<collectingRegion id="5802F83BE840FFDDFB1EFCDEFF181C18" country="Austria" name="Wien" pageId="0" pageNumber="196">Vienna</collectingRegion>
,
<collectingCountry id="E2D17649E840FFDDFF60FD30FEB01C18" box="[258,337,1798,1822]" name="Austria" pageId="0" pageNumber="196">Austria</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D2DC6552E840FFDDFEF5FD78FBB61CA9" pageId="0" pageNumber="196" type="materials_examined">
<paragraph id="9A7936D9E840FFDDFEF5FD78FBB61CA9" blockId="0.[151,1437,1870,2038]" pageId="0" pageNumber="196">
<emphasis id="A8B2EACBE840FFDDFEF5FD78FE931C60" bold="true" box="[151,370,1870,1894]" pageId="0" pageNumber="196">Material examined</emphasis>
(
<specimenCount id="8CC0FD50E840FFDDFFE6FD79FDF01C60" box="[388,529,1870,1895]" pageId="0" pageNumber="196" type="generic">4 specimens</specimenCount>
):
<materialsCitation id="2AAE3C84E840FFDDFC44FD79FBB61CA9" ID-GBIF-Occurrence="3061987302" collectingDate="2020-08-10" collectionCode="R, MYNU, ABCB" country="China" county="Yingjiang County" elevation="930" location="Mangyun Village" municipality="Taiping Town" pageId="0" pageNumber="196" specimenCount="8" specimenCount-female="2" specimenCount-male="6" stateProvince="Yunnan">
<specimenCount id="8CC0FD50E840FFDDFC44FD79FD861C60" box="[550,615,1871,1895]" pageId="0" pageNumber="196" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="8CC0FD50E840FFDDFC14FD79FD401C60" box="[630,673,1871,1895]" pageId="0" pageNumber="196" type="female">1 ♀</specimenCount>
, labeled: “
<collectingCountry id="E2D17649E840FFDDFD79FD78FC801C60" box="[795,865,1870,1894]" name="China" pageId="0" pageNumber="196">China</collectingCountry>
:
<collectingRegion id="5802F83BE840FFDDFD0FFD79FBE21C60" box="[877,1027,1871,1894]" country="China" name="Yunnan" pageId="0" pageNumber="196">Yunnan Prov.</collectingRegion>
, Dehong Autonomous Prefecture (
<emphasis id="A8B2EACBE840FFDDFBE0FD78FE751C8E" bold="true" pageId="0" pageNumber="196">德 Ė傣tāüta治州›</emphasis>
,
<collectingCounty id="73184E55E840FFDDFFC1FD44FD841C8D" box="[419,613,1906,1931]" pageId="0" pageNumber="196">Yingjiang County</collectingCounty>
(
<emphasis id="A8B2EACBE840FFDDFC15FD44FD271C8E" bold="true" box="[631,710,1906,1928]" pageId="0" pageNumber="196">A江县</emphasis>
),
<collectingMunicipality id="7A1DACA3E840FFDDFCBDFD44FC941C8C" box="[735,885,1906,1930]" pageId="0" pageNumber="196">Taiping Town</collectingMunicipality>
(
<emphasis id="A8B2EACBE840FFDDFDE5FD44FC361C8E" bold="true" box="[903,983,1906,1928]" pageId="0" pageNumber="196">
<collectionCode id="FCD7AE1CE840FFDDFDE5FD44FC431C8E" box="[903,930,1906,1928]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="0" pageNumber="196">R</collectionCode>
平Þ
</emphasis>
),
<location id="9F196002E840FFDDFD92FD45FB4D1C8C" LSID="urn:lsid:plazi:treatment:126F87CFE840FFDEFEF5FC53FE441AC9:9F196002E840FFDDFD92FD45FB4D1C8C" box="[1008,1196,1906,1931]" country="China" county="Yingjiang County" municipality="Taiping Town" name="Mangyun Village" pageId="0" pageNumber="196" stateProvince="Yunnan">Mangyun Village</location>
(
<emphasis id="A8B2EACBE840FFDDFADCFD44FAFD1C8E" bold="true" box="[1214,1308,1906,1928]" pageId="0" pageNumber="196">K允ffl›</emphasis>
, H:
<quantity id="5D3E9B3CE840FFDDFB32FD45FA791C8C" box="[1360,1432,1907,1931]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="0" pageNumber="196" unit="m" value="930.0">
<elevation id="11EBD1EAE840FFDDFB32FD45FA791C8C" box="[1360,1432,1907,1931]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="0" pageNumber="196" unit="m" value="930.0">930 m</elevation>
</quantity>
,
<date id="EE781019E840FFDDFEF5FDA1FEC41CA9" box="[151,293,1943,1967]" pageId="0" pageNumber="196" value="2020-08-10">
<collectingDate id="FE3CE9F1E840FFDDFEF5FDA1FEC41CA9" box="[151,293,1943,1967]" pageId="0" pageNumber="196" value="2020-08-10">10.VIII.2020</collectingDate>
</date>
, in rottenwood, local people leg.” (
<specimenCount id="8CC0FD50E840FFDDFCFEFDA1FD381CA8" box="[668,729,1943,1967]" pageId="0" pageNumber="196" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="8CC0FD50E840FFDDFD6FFDA1FCD41CA8" box="[781,821,1943,1967]" pageId="0" pageNumber="196" type="female">1 ♀</specimenCount>
in
<collectionCode id="FCD7AE1CE840FFDDFD35FDA1FC4A1CA8" box="[855,939,1943,1966]" pageId="0" pageNumber="196">MYNU</collectionCode>
,
<specimenCount id="8CC0FD50E840FFDDFDD4FDA1FC3F1CA8" box="[950,990,1943,1967]" pageId="0" pageNumber="196" type="male">1 ♂</specimenCount>
in
<collectionCode id="FCD7AE1CE840FFDDFD9CFDA1FBAA1CA9" box="[1022,1099,1943,1967]" pageId="0" pageNumber="196">ABCB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D2DC6552E840FFDFFEA5FD8CFED51D96" lastPageId="2" lastPageNumber="198" pageId="0" pageNumber="196" type="description">
<paragraph id="9A7936D9E840FFDDFEA5FD8CFDD91CF0" blockId="0.[151,1437,1870,2038]" pageId="0" pageNumber="196">
<emphasis id="A8B2EACBE840FFDDFEA5FD8CFE511CD5" bold="true" box="[199,432,1978,2003]" inLineHeading="true" pageId="0" pageNumber="196" reason="1">Description of male.</emphasis>
Body (
<figureCitation id="02FD2A5CE840FFDDFC60FD8CFDA31CD5" box="[514,578,1978,2003]" captionStart="FIGURE 1" captionStartId="1.[151,250,1263,1287]" captionTargetBox="[176,1414,424,1234]" captionTargetId="figure-128@1.[171,1417,416,1240]" captionTargetPageId="1" captionText="FIGURE 1. Male habitus of Pterorthochaetes yunnanensis Ballerio, 2014. A, dorsal view; B, enrolled, ventral view; C, enrolled, lateral view; D, enrolled, dorsal view. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4643350" httpUri="https://zenodo.org/record/4643350/files/figure.png" pageId="0" pageNumber="196">Fig. 1</figureCitation>
) dark-brown to black, large sized, surface shiny, and covered with dense large punctures and short erect clavate setae.
</paragraph>
<paragraph id="9A7936D9E841FFDCFEA5FAA1FF261A81" blockId="1.[151,1437,151,391]" pageId="1" pageNumber="197">
Head (
<figureCitation id="02FD2A5CE841FFDCFF6DFAA1FE821BB6" box="[271,355,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,1263,1287]" captionTargetBox="[176,1414,424,1234]" captionTargetId="figure-128@1.[171,1417,416,1240]" captionTargetPageId="1" captionText="FIGURE 1. Male habitus of Pterorthochaetes yunnanensis Ballerio, 2014. A, dorsal view; B, enrolled, ventral view; C, enrolled, lateral view; D, enrolled, dorsal view. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4643350" httpUri="https://zenodo.org/record/4643350/files/figure.png" pageId="1" pageNumber="197">Fig. 1A</figureCitation>
), wider than long, subpentagonal, clypeus triangular, apex blunt, anterior margin serrate. Interocular distance about 9 times of the maximum width of dorsal ocular area. Genae obviously protruding outwards, dorsal ocular area large. Punctures of head strongly impressed and horseshoe-shaped on frons and clypeal disc, becoming irregular transverse lines near clypeal apex, all punctures on disc bearing an erect simple short seta. Antennae short, with nine antennomeres: scape securiform, pedicel about as long as wide and rounded; flagellomeres I about as long as wide, near trapezoidal, II-IV similar, strongly transverse, antennal club (flagellomeres V–VII) covered with dense short yellow hair.
</paragraph>
<caption id="CEB96651E841FFDCFEF5FED9FCCD1E2D" ID-DOI="http://doi.org/10.5281/zenodo.4643350" ID-Zenodo-Dep="4643350" httpUri="https://zenodo.org/record/4643350/files/figure.png" pageId="1" pageNumber="197" startId="1.[151,250,1263,1287]" targetBox="[176,1414,424,1234]" targetPageId="1">
<paragraph id="9A7936D9E841FFDCFEF5FED9FCCD1E2D" blockId="1.[151,1436,1263,1324]" pageId="1" pageNumber="197">
<emphasis id="A8B2EACBE841FFDCFEF5FED9FEF41E01" bold="true" box="[151,277,1263,1287]" pageId="1" pageNumber="197">FIGURE 1.</emphasis>
Male habitus of
<taxonomicName id="5DC64D5AE841FFDCFFAAFEC6FC781E0E" authority="Ballerio, 2014" authorityName="Ballerio" authorityYear="2014" box="[456,921,1264,1288]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="A8B2EACBE841FFDCFFAAFEC6FD1B1E01" box="[456,762,1264,1288]" italics="true" pageId="1" pageNumber="197">Pterorthochaetes yunnanensis</emphasis>
<bibRefCitation id="FE574B28E841FFDCFD61FEC6FC781E0E" author="Ballerio, A." box="[771,921,1264,1288]" pageId="1" pageNumber="197" pagination="277 - 280" refId="ref2402" refString="Ballerio, A. (2014) New data on Palaearctic Ceratocanthinae with description of a new species from China (Coleoptera: Scarabaeoidea: Hybosoridae). Koleopterologische Rundschau, 84, 277 - 280." type="journal article" year="2014">Ballerio, 2014</bibRefCitation>
</taxonomicName>
.
<emphasis id="A8B2EACBE841FFDCFDC6FED9FC5D1E01" bold="true" box="[932,956,1263,1287]" pageId="1" pageNumber="197">A,</emphasis>
dorsal view;
<emphasis id="A8B2EACBE841FFDCFA29FEC6FB831E0E" bold="true" box="[1099,1122,1264,1288]" pageId="1" pageNumber="197">B,</emphasis>
enrolled, ventral view;
<emphasis id="A8B2EACBE841FFDCFB39FED9FA951E01" bold="true" box="[1371,1396,1263,1287]" pageId="1" pageNumber="197">C,</emphasis>
enrolled, lateral view;
<emphasis id="A8B2EACBE841FFDCFF01FF22FE9D1E2A" bold="true" box="[355,380,1300,1324]" pageId="1" pageNumber="197">D,</emphasis>
enrolled, dorsal view. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="9A7936D9E841FFDCFEA5FF61FE871D24" blockId="1.[151,1437,1366,2038]" pageId="1" pageNumber="197">
Pronotum (
<figureCitation id="02FD2A5CE841FFDCFF5CFF60FE711E68" box="[318,400,1366,1390]" captionStart="FIGURE 1" captionStartId="1.[151,250,1263,1287]" captionTargetBox="[176,1414,424,1234]" captionTargetId="figure-128@1.[171,1417,416,1240]" captionTargetPageId="1" captionText="FIGURE 1. Male habitus of Pterorthochaetes yunnanensis Ballerio, 2014. A, dorsal view; B, enrolled, ventral view; C, enrolled, lateral view; D, enrolled, dorsal view. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4643350" httpUri="https://zenodo.org/record/4643350/files/figure.png" pageId="1" pageNumber="197">Fig. 1A</figureCitation>
) transverse, widest at middle. Anterior angles rounded, lateral margins curved, fringed with a row of short simple setae spaced out by an interval about their length. Margins of pronotum with thin but distinct groove. Disc of pronotum covered with dense ocellate punctures, larger ocellate punctures present also at discal sides then becoming larger horseshoe-shaped punctures, with opening directed outwards, punctures at sides bigger than on disc, each punctures with a pore in the middle and bearing a clavate erect short seta. Interpunctural distance shorter than (or rarely equal to) punctural diameter.
</paragraph>
<paragraph id="9A7936D9E841FFDCFEA5FC18FB5F1DB4" blockId="1.[151,1437,1366,2038]" pageId="1" pageNumber="197">
Scutellum (
<figureCitation id="02FD2A5CE841FFDCFF20FC18FE5C1D40" box="[322,445,1582,1606]" captionStart="FIGURE 1" captionStartId="1.[151,250,1263,1287]" captionTargetBox="[176,1414,424,1234]" captionTargetId="figure-128@1.[171,1417,416,1240]" captionTargetPageId="1" captionText="FIGURE 1. Male habitus of Pterorthochaetes yunnanensis Ballerio, 2014. A, dorsal view; B, enrolled, ventral view; C, enrolled, lateral view; D, enrolled, dorsal view. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4643350" httpUri="https://zenodo.org/record/4643350/files/figure.png" pageId="1" pageNumber="197">Figs 1A, D</figureCitation>
), slightly wider than long, lateral sides proximally subparallel and distinctly notched by apical portion of mesepisternum, base and sides smooth, apex elongate and acute and sides slightly curved inwards. Dorsal surface covered with dense horseshoe-shaped punctures with opening directed backwards, each with a pore in the middle and bearing a clavate erect short seta. Punctures on scutellum are larger at base and smaller at apex.
</paragraph>
<paragraph id="9A7936D9E841FFDCFEA5FC88FE871C60" blockId="1.[151,1437,1366,2038]" pageId="1" pageNumber="197">
Elytra (
<figureCitation id="02FD2A5CE841FFDCFF75FC88FE6E1DD0" box="[279,399,1726,1750]" captionStart="FIGURE 1" captionStartId="1.[151,250,1263,1287]" captionTargetBox="[176,1414,424,1234]" captionTargetId="figure-128@1.[171,1417,416,1240]" captionTargetPageId="1" captionText="FIGURE 1. Male habitus of Pterorthochaetes yunnanensis Ballerio, 2014. A, dorsal view; B, enrolled, ventral view; C, enrolled, lateral view; D, enrolled, dorsal view. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4643350" httpUri="https://zenodo.org/record/4643350/files/figure.png" pageId="1" pageNumber="197">Figs 1A, D</figureCitation>
) suboval in shape, strongly convex, longer than wide, surface, apart from base, which is covered by some transverse irregular short lines, covered with dense small horseshoe-shaped punctures, with opening directed backwards or outwards, each one with a pore in the middle bearing a clavate erect short seta, a few simple fine punctures mixed to horseshoe-shaped punctures. Interpunctural distance of horseshoe-shaped punctures equal to or shorter than punctural diameter.
</paragraph>
<paragraph id="9A7936D9E841FFDCFEA5FD45FB971CF0" blockId="1.[151,1437,1366,2038]" pageId="1" pageNumber="197">
Legs. Outer margin of protibiae (
<figureCitation id="02FD2A5CE841FFDCFC43FD44FD761C8C" box="[545,663,1906,1930]" captionStart="FIGURE 2" captionStartId="2.[151,250,1117,1141]" captionTargetBox="[151,1435,181,1093]" captionTargetId="figure-19@2.[151,1435,181,1093]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, protibia, male, dorsal view; B, protibia, male, lateral view; C, mesotibia, male, ventral view; D, metatibia, male, ventral view; E, mesofemur, male, dorsal view; F, protibia, female, dorsal view; G, protibia, female, lateral view; H, mesotibia, female, ventral view; I, metatibia, female, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4643354" httpUri="https://zenodo.org/record/4643354/files/figure.png" pageId="1" pageNumber="197">Figs 2A–B</figureCitation>
) with an irregular number of weak outer teeth and apex with two stronger outer teeth, apical spur abruptly hooked at apex. Mesofemora (
<figureCitation id="02FD2A5CE841FFDCFD4FFDA0FC9D1CA8" box="[813,892,1942,1966]" captionStart="FIGURE 2" captionStartId="2.[151,250,1117,1141]" captionTargetBox="[151,1435,181,1093]" captionTargetId="figure-19@2.[151,1435,181,1093]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, protibia, male, dorsal view; B, protibia, male, lateral view; C, mesotibia, male, ventral view; D, metatibia, male, ventral view; E, mesofemur, male, dorsal view; F, protibia, female, dorsal view; G, protibia, female, lateral view; H, mesotibia, female, ventral view; I, metatibia, female, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4643354" httpUri="https://zenodo.org/record/4643354/files/figure.png" pageId="1" pageNumber="197">Fig. 2E</figureCitation>
) obviously expanded near middle, mesotibiae (
<figureCitation id="02FD2A5CE841FFDCFB13FDA0FF591CD4" captionStart="FIGURE 2" captionStartId="2.[151,250,1117,1141]" captionTargetBox="[151,1435,181,1093]" captionTargetId="figure-19@2.[151,1435,181,1093]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, protibia, male, dorsal view; B, protibia, male, lateral view; C, mesotibia, male, ventral view; D, metatibia, male, ventral view; E, mesofemur, male, dorsal view; F, protibia, female, dorsal view; G, protibia, female, lateral view; H, mesotibia, female, ventral view; I, metatibia, female, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4643354" httpUri="https://zenodo.org/record/4643354/files/figure.png" pageId="1" pageNumber="197">Fig. 2C</figureCitation>
) slightly expanded basally, with a short inner apical spur bent inward at a right angle. Metatibiae (
<figureCitation id="02FD2A5CE841FFDCFAB8FD8CFACF1CD4" box="[1242,1326,1978,2002]" captionStart="FIGURE 2" captionStartId="2.[151,250,1117,1141]" captionTargetBox="[151,1435,181,1093]" captionTargetId="figure-19@2.[151,1435,181,1093]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, protibia, male, dorsal view; B, protibia, male, lateral view; C, mesotibia, male, ventral view; D, metatibia, male, ventral view; E, mesofemur, male, dorsal view; F, protibia, female, dorsal view; G, protibia, female, lateral view; H, mesotibia, female, ventral view; I, metatibia, female, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4643354" httpUri="https://zenodo.org/record/4643354/files/figure.png" pageId="1" pageNumber="197">Fig. 2D</figureCitation>
) subtriangular, strongly expanded apically, apex with two short spurs, the inner one longer and tortile.
</paragraph>
<caption id="CEB96651E842FFDFFEF5FE6BFE7C1FE6" ID-DOI="http://doi.org/10.5281/zenodo.4643354" ID-Zenodo-Dep="4643354" httpUri="https://zenodo.org/record/4643354/files/figure.png" pageId="2" pageNumber="198" startId="2.[151,250,1117,1141]" targetBox="[151,1435,181,1093]" targetPageId="2">
<paragraph id="9A7936D9E842FFDFFEF5FE6BFE7C1FE6" blockId="2.[151,1437,1117,1249]" pageId="2" pageNumber="198">
<emphasis id="A8B2EACBE842FFDFFEF5FE6BFEF21F73" bold="true" box="[151,275,1117,1141]" pageId="2" pageNumber="198">FIGURE 2.</emphasis>
Diagnostic characters of
<taxonomicName id="5DC64D5AE842FFDFFC75FE6BFC021F73" authority="Ballerio, 2014" authorityName="Ballerio" authorityYear="2014" box="[535,995,1117,1141]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="A8B2EACBE842FFDFFC75FE6BFCA91F73" box="[535,840,1117,1141]" italics="true" pageId="2" pageNumber="198">Pterorthochaetes yunnanensis</emphasis>
<bibRefCitation id="FE574B28E842FFDFFD2DFE6BFC021F73" author="Ballerio, A." box="[847,995,1117,1141]" pageId="2" pageNumber="198" pagination="277 - 280" refId="ref2402" refString="Ballerio, A. (2014) New data on Palaearctic Ceratocanthinae with description of a new species from China (Coleoptera: Scarabaeoidea: Hybosoridae). Koleopterologische Rundschau, 84, 277 - 280." type="journal article" year="2014">Ballerio, 2014</bibRefCitation>
</taxonomicName>
.
<emphasis id="A8B2EACBE842FFDFFD8FFE6BFBE41F73" bold="true" box="[1005,1029,1117,1141]" pageId="2" pageNumber="198">A,</emphasis>
protibia, male, dorsal view;
<emphasis id="A8B2EACBE842FFDFFB4BFE6BFAA11F73" bold="true" box="[1321,1344,1117,1141]" pageId="2" pageNumber="198">B,</emphasis>
protibia, male, lateral view;
<emphasis id="A8B2EACBE842FFDFFF02FEB7FE981F9F" bold="true" box="[352,377,1153,1177]" pageId="2" pageNumber="198">C,</emphasis>
mesotibia, male, ventral view;
<emphasis id="A8B2EACBE842FFDFFCA1FEB7FD3A1F9F" bold="true" box="[707,731,1153,1177]" pageId="2" pageNumber="198">D,</emphasis>
metatibia, male, ventral view;
<emphasis id="A8B2EACBE842FFDFFA43FEB7FBD91F9F" bold="true" box="[1057,1080,1153,1177]" pageId="2" pageNumber="198">E,</emphasis>
mesofemur, male, dorsal view;
<emphasis id="A8B2EACBE842FFDFFBEBFEB7FA7D1F9F" bold="true" box="[1417,1436,1153,1177]" pageId="2" pageNumber="198">F,</emphasis>
protibia, female, dorsal view;
<emphasis id="A8B2EACBE842FFDFFFA5FE93FE011FBB" bold="true" box="[455,480,1189,1213]" pageId="2" pageNumber="198">G,</emphasis>
protibia, female, lateral view;
<emphasis id="A8B2EACBE842FFDFFD7AFE93FCD01FBB" bold="true" box="[792,817,1189,1213]" pageId="2" pageNumber="198">H,</emphasis>
mesotibia, female, ventral view;
<emphasis id="A8B2EACBE842FFDFFAE1FE93FB721FBB" bold="true" box="[1155,1171,1189,1213]" pageId="2" pageNumber="198">I,</emphasis>
metatibia, female, ventral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="9A7936D9E842FFDFFEA5FF27FBE21E93" blockId="2.[151,1437,1297,2041]" pageId="2" pageNumber="198">
Male genitalia. Aedeagus (
<figureCitation id="02FD2A5CE842FFDFFF80FF27FDB81E2F" box="[482,601,1297,1321]" captionStart="FIGURE 3" captionStartId="3.[151,250,1770,1794]" captionTargetBox="[151,1435,488,1746]" captionTargetId="figure-177@3.[151,1435,488,1746]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, aedeagus, dorsal view; B, aedeagus, ventral view; C, parameres of aedeagus, dorsal view; D, left paramere; E, right paramere; F, internal sac of aedeagus; G, genital segment of male; H, bursal sclerites; I, spermatheca. Scale bars: 0.5 mm for A–G, 0.25 mm for H–I." figureDoi="http://doi.org/10.5281/zenodo.4643356" httpUri="https://zenodo.org/record/4643356/files/figure.png" pageId="2" pageNumber="198">Figs 3A–E</figureCitation>
), asymmetrical and tortile, basal piece about 2.5 times the length of parameres. Parameres as in
<figureCitation id="02FD2A5CE842FFDFFF25FF03FE241E4B" box="[327,453,1333,1357]" captionStart="FIGURE 3" captionStartId="3.[151,250,1770,1794]" captionTargetBox="[151,1435,488,1746]" captionTargetId="figure-177@3.[151,1435,488,1746]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, aedeagus, dorsal view; B, aedeagus, ventral view; C, parameres of aedeagus, dorsal view; D, left paramere; E, right paramere; F, internal sac of aedeagus; G, genital segment of male; H, bursal sclerites; I, spermatheca. Scale bars: 0.5 mm for A–G, 0.25 mm for H–I." figureDoi="http://doi.org/10.5281/zenodo.4643356" httpUri="https://zenodo.org/record/4643356/files/figure.png" pageId="2" pageNumber="198">Figs. 3A–E</figureCitation>
. Internal sac with a distinctive weak sclerotized plate irregularly S-shaped occupying about half the length of the sac and with a transverse thick short sclerification in the middle (
<figureCitation id="02FD2A5CE842FFDFFA46FF6FFB941E77" box="[1060,1141,1369,1393]" captionStart="FIGURE 3" captionStartId="3.[151,250,1770,1794]" captionTargetBox="[151,1435,488,1746]" captionTargetId="figure-177@3.[151,1435,488,1746]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, aedeagus, dorsal view; B, aedeagus, ventral view; C, parameres of aedeagus, dorsal view; D, left paramere; E, right paramere; F, internal sac of aedeagus; G, genital segment of male; H, bursal sclerites; I, spermatheca. Scale bars: 0.5 mm for A–G, 0.25 mm for H–I." figureDoi="http://doi.org/10.5281/zenodo.4643356" httpUri="https://zenodo.org/record/4643356/files/figure.png" pageId="2" pageNumber="198">Fig. 3F</figureCitation>
). Genital segment (
<figureCitation id="02FD2A5CE842FFDFFB20FF6FFA751E77" box="[1346,1428,1369,1393]" captionStart="FIGURE 3" captionStartId="3.[151,250,1770,1794]" captionTargetBox="[151,1435,488,1746]" captionTargetId="figure-177@3.[151,1435,488,1746]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, aedeagus, dorsal view; B, aedeagus, ventral view; C, parameres of aedeagus, dorsal view; D, left paramere; E, right paramere; F, internal sac of aedeagus; G, genital segment of male; H, bursal sclerites; I, spermatheca. Scale bars: 0.5 mm for A–G, 0.25 mm for H–I." figureDoi="http://doi.org/10.5281/zenodo.4643356" httpUri="https://zenodo.org/record/4643356/files/figure.png" pageId="2" pageNumber="198">Fig. 3G</figureCitation>
) asymmetrical and curved with branches apically fused and with short manubrium.
</paragraph>
<paragraph id="9A7936D9E842FFDFFEA5FF97FD111EDA" blockId="2.[151,1437,1297,2041]" pageId="2" pageNumber="198">
Measurements of male: BL:
<quantity id="5D3E9B3CE842FFDFFC6AFF97FD4B1EBE" box="[520,682,1441,1465]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="5.95" metricValueMin="5.65" pageId="2" pageNumber="198" unit="mm" value="5.8" valueMax="5.95" valueMin="5.65">5.65–5.95 mm</quantity>
; HL: 0.95–1.00 mm, HW:
<quantity id="5D3E9B3CE842FFDFFDBEFF97FB9F1EBE" box="[988,1150,1441,1465]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="1.85" metricValueMin="1.75" pageId="2" pageNumber="198" unit="mm" value="1.8" valueMax="1.85" valueMin="1.75">1.75–1.85 mm</quantity>
; PL:
<quantity id="5D3E9B3CE842FFDFFADCFF97FABE1EBE" box="[1214,1375,1441,1465]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="2" pageNumber="198" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.70–1.80 mm</quantity>
, PW:
<quantity id="5D3E9B3CE842FFDFFEF5FFF3FED71EDA" box="[151,310,1477,1501]" metricMagnitude="-3" metricUnit="m" metricValue="2.875" metricValueMax="2.95" metricValueMin="2.8" pageId="2" pageNumber="198" unit="mm" value="2.875" valueMax="2.95" valueMin="2.8">2.80–2.95 mm</quantity>
; EL: 3.00–
<quantity id="5D3E9B3CE842FFDFFFCEFFF3FDED1EDA" box="[428,524,1477,1501]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" pageId="2" pageNumber="198" unit="mm" value="3.15">3.15 mm</quantity>
, EW:
<quantity id="5D3E9B3CE842FFDFFC2FFFF3FD0D1EDA" box="[589,748,1477,1501]" metricMagnitude="-3" metricUnit="m" metricValue="2.725" metricValueMax="2.75" metricValueMin="2.7" pageId="2" pageNumber="198" unit="mm" value="2.725" valueMax="2.75" valueMin="2.7">2.70–2.75 mm</quantity>
.
</paragraph>
<paragraph id="9A7936D9E842FFDFFEA5FFDFFA8D1D4F" blockId="2.[151,1437,1297,2041]" pageId="2" pageNumber="198">
<emphasis id="A8B2EACBE842FFDFFEA5FFDFFB9D1D07" box="[199,1148,1513,1537]" inLineHeading="true" pageId="2" pageNumber="198" reason="1">
<emphasis id="A8B2EACBE842FFDFFEA5FFDFFD8E1D07" bold="true" box="[199,623,1513,1537]" pageId="2" pageNumber="198">Supplementary description of female</emphasis>
(based on the specimen from Mangyun village).
</emphasis>
Generally similar to male, apical spur of protibiae (
<figureCitation id="02FD2A5CE842FFDFFFF9FC3BFDEE1D23" box="[411,527,1549,1573]" captionStart="FIGURE 2" captionStartId="2.[151,250,1117,1141]" captionTargetBox="[151,1435,181,1093]" captionTargetId="figure-19@2.[151,1435,181,1093]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, protibia, male, dorsal view; B, protibia, male, lateral view; C, mesotibia, male, ventral view; D, metatibia, male, ventral view; E, mesofemur, male, dorsal view; F, protibia, female, dorsal view; G, protibia, female, lateral view; H, mesotibia, female, ventral view; I, metatibia, female, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4643354" httpUri="https://zenodo.org/record/4643354/files/figure.png" pageId="2" pageNumber="198">Figs 2F–G</figureCitation>
) thinner and straighter; apical spur of mesotibiae (
<figureCitation id="02FD2A5CE842FFDFFA45FC3BFB9B1D23" box="[1063,1146,1549,1573]" captionStart="FIGURE 2" captionStartId="2.[151,250,1117,1141]" captionTargetBox="[151,1435,181,1093]" captionTargetId="figure-19@2.[151,1435,181,1093]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, protibia, male, dorsal view; B, protibia, male, lateral view; C, mesotibia, male, ventral view; D, metatibia, male, ventral view; E, mesofemur, male, dorsal view; F, protibia, female, dorsal view; G, protibia, female, lateral view; H, mesotibia, female, ventral view; I, metatibia, female, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4643354" httpUri="https://zenodo.org/record/4643354/files/figure.png" pageId="2" pageNumber="198">Fig. 2H</figureCitation>
) straight. Outer apical spur of metatibiae (
<figureCitation id="02FD2A5CE842FFDFFF50FC07FE9B1D4F" box="[306,378,1585,1609]" captionStart="FIGURE 2" captionStartId="2.[151,250,1117,1141]" captionTargetBox="[151,1435,181,1093]" captionTargetId="figure-19@2.[151,1435,181,1093]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, protibia, male, dorsal view; B, protibia, male, lateral view; C, mesotibia, male, ventral view; D, metatibia, male, ventral view; E, mesofemur, male, dorsal view; F, protibia, female, dorsal view; G, protibia, female, lateral view; H, mesotibia, female, ventral view; I, metatibia, female, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4643354" httpUri="https://zenodo.org/record/4643354/files/figure.png" pageId="2" pageNumber="198">Fig. 2I</figureCitation>
) straight and sharp. Spermatheca (
<figureCitation id="02FD2A5CE842FFDFFC8FFC07FCD41D4F" box="[749,821,1585,1609]" captionStart="FIGURE 3" captionStartId="3.[151,250,1770,1794]" captionTargetBox="[151,1435,488,1746]" captionTargetId="figure-177@3.[151,1435,488,1746]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic characters of Pterorthochaetes yunnanensis Ballerio, 2014. A, aedeagus, dorsal view; B, aedeagus, ventral view; C, parameres of aedeagus, dorsal view; D, left paramere; E, right paramere; F, internal sac of aedeagus; G, genital segment of male; H, bursal sclerites; I, spermatheca. Scale bars: 0.5 mm for A–G, 0.25 mm for H–I." figureDoi="http://doi.org/10.5281/zenodo.4643356" httpUri="https://zenodo.org/record/4643356/files/figure.png" pageId="2" pageNumber="198">Fig. 3I</figureCitation>
) C-shaped, both extremities flattened in lateral view.
</paragraph>
<paragraph id="9A7936D9E842FFDFFEA5FC63FED51D96" blockId="2.[151,1437,1297,2041]" pageId="2" pageNumber="198">
Measurements of female: BL:
<quantity id="5D3E9B3CE842FFDFFC72FC63FD921D6A" box="[528,627,1621,1645]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" pageId="2" pageNumber="198" unit="mm" value="5.75">5.75 mm</quantity>
; HL: 1.00 mm, HW:
<quantity id="5D3E9B3CE842FFDFFD3AFC63FC5D1D6A" box="[856,956,1621,1645]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="198" unit="mm" value="1.8">1.80 mm</quantity>
; PL:
<quantity id="5D3E9B3CE842FFDFFD97FC63FBB61D6A" box="[1013,1111,1621,1645]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="2" pageNumber="198" unit="mm" value="1.75">1.75 mm</quantity>
, PW:
<quantity id="5D3E9B3CE842FFDFFAFAFC63FB1A1D6A" box="[1176,1275,1621,1645]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" pageId="2" pageNumber="198" unit="mm" value="2.85">2.85 mm</quantity>
; EL: 3.00 mm, EW:
<quantity id="5D3E9B3CE842FFDFFEACFC4FFED11D96" box="[206,304,1657,1681]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" pageId="2" pageNumber="198" unit="mm" value="2.65">2.65 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="D2DC6552E842FFDFFEA5FCABFF3C1C8B" pageId="2" pageNumber="198" type="distribution">
<paragraph id="9A7936D9E842FFDFFEA5FCABFDED1DDF" blockId="2.[151,1437,1297,2041]" pageId="2" pageNumber="198">
<emphasis id="A8B2EACBE842FFDFFEA5FCABFEB81DB2" box="[199,345,1693,1717]" inLineHeading="true" pageId="2" pageNumber="198" reason="1">
<emphasis id="A8B2EACBE842FFDFFEA5FCABFEB31DB3" bold="true" box="[199,338,1693,1717]" pageId="2" pageNumber="198">Distribution</emphasis>
.
</emphasis>
<collectingCountry id="E2D17649E842FFDFFF03FCABFE441DB3" box="[353,421,1693,1717]" name="China" pageId="2" pageNumber="198">China</collectingCountry>
:
<collectingRegion id="5802F83BE842FFDFFFCCFCABFD8B1DB3" box="[430,618,1693,1717]" country="China" name="Yunnan" pageId="2" pageNumber="198">Yunnan Province</collectingRegion>
: Dai Autonomous Prefecture of Xishuangbanna (
<typeStatus id="457D887BE842FFDFFA18FCABFB481DB3" box="[1146,1193,1693,1717]" pageId="2" pageNumber="198">type</typeStatus>
locality), Dehong Autonomous Prefecture (new record).
</paragraph>
<paragraph id="9A7936D9E842FFDFFEA5FCD3FF3C1C8B" blockId="2.[151,1437,1297,2041]" pageId="2" pageNumber="198">
<emphasis id="A8B2EACBE842FFDFFEA5FCD3FE4F1DFB" bold="true" box="[199,430,1765,1789]" inLineHeading="true" pageId="2" pageNumber="198" reason="1">Habitat and biology.</emphasis>
The adults of
<taxonomicName id="5DC64D5AE842FFDFFC21FCD0FD011DFB" authorityName="Ballerio" authorityYear="2014" box="[579,736,1766,1789]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="A8B2EACBE842FFDFFC21FCD0FD011DFB" box="[579,736,1766,1789]" italics="true" pageId="2" pageNumber="198">P. yunnanensis</emphasis>
</taxonomicName>
can be collected from rotten wood. The locality of new specimens in this study is in the tropical rainforest climate and characterized by the presence of a moist broadleaf evergreen primary forest (
<bibRefCitation id="FE574B28E842FFDFFE80FD1BFE811C43" author="Zeng, J. - M." box="[226,352,1837,1861]" pageId="2" pageNumber="198" pagination="1 - 18" refId="ref2647" refString="Zeng, J. - M. (2018) The Classification System of Natural Forests and Its Geographic Distribution in Yunnan. Journal of South- west Forestry University, 38 (6), 1 - 18." type="journal article" year="2018">Zeng, 2018</bibRefCitation>
). They were also found associated with adults and larvae of
<taxonomicName id="5DC64D5AE842FFDFFDBEFD1BFBAA1C43" authorityName="Leach" authorityYear="1815" box="[988,1099,1837,1861]" class="Insecta" family="Passalidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
species inside rotten wood, this latter observation confirms what has already been reported for the genus
<taxonomicName id="5DC64D5AE842FFDFFDC6FD67FBBA1C6F" authorityName="Gestro" authorityYear="1898" box="[932,1115,1873,1897]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="A8B2EACBE842FFDFFDC6FD67FBBA1C6F" box="[932,1115,1873,1897]" italics="true" pageId="2" pageNumber="198">Pterorthochaetes</emphasis>
</taxonomicName>
in
<collectingCountry id="E2D17649E842FFDFFA1FFD67FB3E1C6F" box="[1149,1247,1873,1897]" name="Malaysia" pageId="2" pageNumber="198">Malaysia</collectingCountry>
(
<bibRefCitation id="FE574B28E842FFDFFA8FFD67FA791C6F" author="Kon, M. &amp; Johki, Y. &amp; Ballerio, A." box="[1261,1432,1873,1897]" pageId="2" pageNumber="198" pagination="93 - 95" refId="ref2439" refString="Kon, M., Johki, Y. &amp; Ballerio, A. (2010) The ceratocanthid beetle Pterorthochaetes haroldi (Coleoptera, Ceratocanthidae), collected from a gallery of the passalid beetle, Leptaulax planus (Coleoptera, Passalidae) in Sabah, Malaysia. Kogane, 11, 93 - 95." type="journal article" year="2010">
Kon
<emphasis id="A8B2EACBE842FFDFFB46FD64FABB1C6F" box="[1316,1370,1873,1897]" italics="true" pageId="2" pageNumber="198">et al.</emphasis>
2010
</bibRefCitation>
,
<bibRefCitation id="FE574B28E842FFDFFEF5FD43FF301C8B" author="Kon, M. &amp; Ballerio, A. &amp; Pittino, R &amp; Johki, Y." box="[151,209,1909,1933]" pageId="2" pageNumber="198" pagination="99 - 102" refId="ref2496" refString="Kon, M., Ballerio, A., Pittino, R, &amp; Johki, Y. (2014) A new record of Leptaulax eschscholtzi Kaup (Coleoptera, Passalidae) from the Malay Peninsula, with notes on the microhabitat and possible colony composition of L. eschscholtzi and the scarabguests in its gallery. Kogane, 16, 99 - 102." type="journal article" year="2014">2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D2DC6552E842FFDEFEA5FDAFFE441AC9" lastPageId="3" lastPageNumber="199" pageId="2" pageNumber="198" type="discussion">
<paragraph id="9A7936D9E842FFDEFEA5FDAFFE441AC9" blockId="2.[151,1437,1297,2041]" lastBlockId="3.[151,1437,151,463]" lastPageId="3" lastPageNumber="199" pageId="2" pageNumber="198">
<emphasis id="A8B2EACBE842FFDFFEA5FDAFFEA71CB6" box="[199,326,1945,1969]" inLineHeading="true" pageId="2" pageNumber="198" reason="1">
<emphasis id="A8B2EACBE842FFDFFEA5FDAFFEDE1CB7" bold="true" box="[199,319,1945,1969]" pageId="2" pageNumber="198">Discussion</emphasis>
.
</emphasis>
The discovery of more specimens of
<taxonomicName id="5DC64D5AE842FFDFFC83FDACFC601CB7" authorityName="Ballerio" authorityYear="2014" box="[737,897,1946,1969]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="A8B2EACBE842FFDFFC83FDACFC601CB7" box="[737,897,1946,1969]" italics="true" pageId="2" pageNumber="198">P. yunnanensis</emphasis>
</taxonomicName>
allows us to better define the species. The distinguishing characters listed by
<bibRefCitation id="FE574B28E842FFDFFFB7FD8BFD9B1CD3" author="Ballerio, A." box="[469,634,1981,2005]" pageId="2" pageNumber="198" pagination="277 - 280" refId="ref2402" refString="Ballerio, A. (2014) New data on Palaearctic Ceratocanthinae with description of a new species from China (Coleoptera: Scarabaeoidea: Hybosoridae). Koleopterologische Rundschau, 84, 277 - 280." type="journal article" year="2014">Ballerio (2014)</bibRefCitation>
are confirmed and can be better defined. The species is characterized by: a) the presence of small impressed ocellate punctures on pronotal disc, present also at discal sides, although larger, then becoming horseshoe-shaped and distinctly larger at pronotal sides; b) pronotal interpunctural distance smaller than punctural diameter, c) pronotal lateral margins fringed with a row of long simple setae, spaced out by an interval about their length or longer, d) elytral punctation made of dense medium sized impressed horseshoe-shaped punctures with opening directed backwards, spaced out by an interval subequal to (or shorter than) their diameter, mixed with simple impressed punctures irregularly distributed, e) shape of bursal sclerites, f) shape of endophallites.
<taxonomicName id="5DC64D5AE843FFDEFA00FB11FA611A39" authorityName="Gestro" authorityYear="1898" box="[1122,1408,295,319]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="A8B2EACBE843FFDEFA00FB11FA611A39" box="[1122,1408,295,319]" italics="true" pageId="3" pageNumber="199">Pterorthochaetes insularis</emphasis>
</taxonomicName>
is confirmed as the species closest to it and the differences listed by
<bibRefCitation id="FE574B28E843FFDEFD2DFB7DFC131A62" author="Ballerio, A." box="[847,1010,331,356]" pageId="3" pageNumber="199" pagination="277 - 280" refId="ref2402" refString="Ballerio, A. (2014) New data on Palaearctic Ceratocanthinae with description of a new species from China (Coleoptera: Scarabaeoidea: Hybosoridae). Koleopterologische Rundschau, 84, 277 - 280." type="journal article" year="2014">Ballerio (2014)</bibRefCitation>
are also confirmed apart from the periscutellar punctation, which is basically the same with some transverse irregular lines. The endophallites are different from the ones present in
<taxonomicName id="5DC64D5AE843FFDEFF09FBA2FE051AAD" authorityName="Gestro" authorityYear="1898" box="[363,484,403,427]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="A8B2EACBE843FFDEFF09FBA2FE051AAD" box="[363,484,403,427]" italics="true" pageId="3" pageNumber="199">P. insularis</emphasis>
</taxonomicName>
, a species which lacks the weak sclerotized plate irregularly S-shaped occupying about half the length of the sac.
</paragraph>
</subSubSection>
</treatment>
</document>
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