158 lines
18 KiB
XML
158 lines
18 KiB
XML
<document id="F37E0E0AF2833DD44CCAEA6079179B30" ID-CLB-Dataset="4537" ID-DOI="10.5252/g2011n4a4" ID-GBIF-Dataset="d3ae8332-0037-4484-83cf-4706b9b2315b" ID-ISSN="1638-9395" ID-Zenodo-Dep="5377989" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630283798029" checkinUser="marcus" docAuthor="Béthoux, Olivier, Schneider, Jörg W. & Klass, Klaus-Dieter" docDate="2011" docId="126F878EFFDDA24FFF7FBEC36544FD9E" docLanguage="en" docName="Geodiversitas.2011.33.4.625-635.pdf" docOrigin="Geodiversitas 33 (4)" docSource="http://dx.doi.org/10.5252/g2011n4a4" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Phyloblatta gaudryi" docType="treatment" docVersion="7" lastPageNumber="630" masterDocId="EE56FFF6FFDEA24AFFA9BB61647BFFB8" masterDocTitle="Redescription of the holotype of Phyloblatta gaudryi (Agnus, 1903) (Pennsylvanian; Commentry, France), an exceptionally wellpreserved stem-dictyopteran" masterLastPageNumber="635" masterPageNumber="625" pageNumber="628" updateTime="1699239604766" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods id="4E91B37CC97EFBA03DC5B6B6556F0827" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9A26CD2E4C39DBC3DDB1F3E1E170A82A">
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<mods:title id="EADBCAA885E952BD5332823AC49F246F">Redescription of the holotype of Phyloblatta gaudryi (Agnus, 1903) (Pennsylvanian; Commentry, France), an exceptionally wellpreserved stem-dictyopteran</mods:title>
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</mods:titleInfo>
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<mods:name id="583A56E90926B2728C69A18C634BE14C" type="personal">
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<mods:role id="56FB5B57BB4A6346065311C678555D88">
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<mods:roleTerm id="27222DD977B62FC33D5CEBA1425C6034">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DD45CD2D9918CC9A4091EC17E188CF57">Béthoux, Olivier</mods:namePart>
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</mods:name>
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<mods:name id="746D97D466F591BDEB04CA91376148F8" type="personal">
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<mods:roleTerm id="2E45F1B6CAC730DB4B8ACAAAD376E821">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5017E8287375281345524A7AD6B986A1">Schneider, Jörg W.</mods:namePart>
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</mods:name>
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<mods:name id="1508CB7D23EB3CCD69D819519F6C764D" type="personal">
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<mods:role id="0803D1B53FCE5B93417E1B13C84C649D">
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<mods:roleTerm id="E30289BF24BB70B3D2FF98CE0023568E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F49E1F5C8BECC446BB306A60957BEE49">Klass, Klaus-Dieter</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1B4E89714BB383C5390DC7B628EEE47D">text</mods:typeOfResource>
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<mods:titleInfo id="90DB02FC6CBA5B75AC3D88E1A4F126F8">
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<mods:title id="07F21B240CE8EE49305FB6C3EA34D746">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="F69329B562344401CB9618F1408CFCA2">
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<mods:date id="FBCDDA25EC1EFA0048A2F1F735F57650">2011</mods:date>
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<mods:detail id="09A644965ED1B4788DE2BE795BC89707" type="pubDate">
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<mods:number id="87754FCB71AB890A22323B965CD93C14">2011-12-31</mods:number>
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<mods:detail id="26FEBD5E9926633A841501ADEEC25984" type="volume">
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<mods:number id="6C1711C226D1BEDD4D2F24B4F00AB00F">33</mods:number>
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<mods:detail id="5CCE62F34A08AFE6B60D78A8C92270D6" type="issue">
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<mods:number id="CE58D9E9349AF243B0BCADBA32654A3D">4</mods:number>
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<mods:start id="8DF56418FD337CD1393C45EBDD5B7B1D">625</mods:start>
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<mods:end id="579487D13F702968ECBA21B26A34C241">635</mods:end>
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<mods:url id="2B97471CACC0ED71DC2FD827EF3016CD">http://dx.doi.org/10.5252/g2011n4a4</mods:url>
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</mods:location>
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<mods:classification id="C14D6249476B03657D7D30CACC4B06EF">journal article</mods:classification>
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<mods:identifier id="4E51EF5D323C77D25F2DC3CB6428012A" type="CLB-Dataset">4537</mods:identifier>
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<mods:identifier id="840BC77F4743156056C9CD0FBACAAC54" type="DOI">10.5252/g2011n4a4</mods:identifier>
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<mods:identifier id="6B096015E20C383F616716BA8DC77D4B" type="GBIF-Dataset">d3ae8332-0037-4484-83cf-4706b9b2315b</mods:identifier>
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<mods:identifier id="20FB2C8482F7CC1517D65B43043B0EC0" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="AEBD7A5388C95DCAD44D62CDD44C1CC6" type="Zenodo-Dep">5377989</mods:identifier>
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<treatment id="126F878EFFDDA24FFF7FBEC36544FD9E" ID-DOI="http://doi.org/10.5281/zenodo.5465287" ID-GBIF-Taxon="185366999" ID-Zenodo-Dep="5465287" LSID="urn:lsid:plazi:treatment:126F878EFFDDA24FFF7FBEC36544FD9E" httpUri="http://treatment.plazi.org/id/126F878EFFDDA24FFF7FBEC36544FD9E" lastPageId="5" lastPageNumber="630" pageId="3" pageNumber="628">
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<subSubSection id="D2DC6513FFDDA249FF7FBEC36634FA05" box="[214,591,1442,1469]" pageId="3" pageNumber="628" type="nomenclature">
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<paragraph id="9A793698FFDDA249FF7FBEC36634FA05" blockId="3.[214,591,1442,1469]" box="[214,591,1442,1469]" pageId="3" pageNumber="628">
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<heading id="C13181F4FFDDA249FF7FBEC36634FA05" box="[214,591,1442,1469]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="628" reason="2">
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<taxonomicName id="5DC64D1BFFDDA249FF7FBEC36634FA05" authority="(Agnus, 1903)" baseAuthorityName="Agnus" baseAuthorityYear="1903" box="[214,591,1442,1469]" class="Insecta" family="Archimylacridae" genus="Phyloblatta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="628" phylum="Arthropoda" rank="species" species="gaudryi">
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<emphasis id="A8B2EA8AFFDDA249FF7FBEC365D6FA05" bold="true" box="[214,429,1442,1469]" italics="true" pageId="3" pageNumber="628">Phyloblatta gaudryi</emphasis>
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(
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<bibRefCitation id="FE574B69FFDDA249FE12BEC2663CFA05" author="AGNUS A. N." box="[443,583,1443,1469]" pageId="3" pageNumber="628" pagination="291 - 294" refId="ref4668" refString="AGNUS A. N. 1903. - Deuxieme note sur les Blattidae paleozoiques (Orthopt. palaeoz.). Description d'une espece nouvelle. Bulletin de la Societe entomologique de France 17: 291 - 294." type="journal article" year="1903">Agnus, 1903</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D2DC6513FFDDA24EFF21BE876002FA33" lastPageId="4" lastPageNumber="629" pageId="3" pageNumber="628" type="description">
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<paragraph id="9A793698FFDDA24EFF21BE876002FA33" blockId="3.[134,669,1509,1722]" lastBlockId="4.[698,1233,1289,1419]" lastPageId="4" lastPageNumber="629" pageId="3" pageNumber="628">
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DESCRIPTION. — Negative imprint, right and left forewings, and left hind wing of a single individual, connected to remains of the pterothorax (wing venation described as if viewed from a positive imprint; numerical data: right
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<emphasis id="A8B2EA8AFFDDA249FF10BD3064C5F9D0" box="[185,190,1617,1640]" italics="true" pageId="3" pageNumber="628">/</emphasis>
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left side);
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<emphasis id="A8B2EA8AFFDDA249FEB2BD3365F8F9D2" bold="true" box="[283,387,1618,1642]" pageId="3" pageNumber="628">forewings:</emphasis>
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length 35.8/34.0 mm, width 14.3
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<emphasis id="A8B2EA8AFFDDA249FF1ABD0D64C2F93B" box="[179,185,1644,1667]" italics="true" pageId="3" pageNumber="628">/</emphasis>
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<quantity id="5D3E9B7DFFDDA249FF12BD0D6560F93B" box="[187,283,1644,1668]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="3" pageNumber="628" unit="mm" value="14.8">14.8 mm</quantity>
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; ScP reaching anterior wing margin at between 60 and 75% of the wing length; ScP concave, with numerous anterior branches; R concave, branched about 7.5
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<emphasis id="A8B2EA8AFFDDA249FCB0BE026764FAC2" box="[793,799,1379,1402]" italics="true" pageId="3" pageNumber="628">/</emphasis>
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<quantity id="5D3E9B7DFFDDA249FC88BE026714FAC2" box="[801,879,1379,1403]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="628" unit="mm" value="6.8">6.8 mm</quantity>
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distal to wing base, with no regular branching pattern; R with 13
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<emphasis id="A8B2EA8AFFDDA249FC67BE1F67AFFA2D" box="[974,980,1406,1429]" italics="true" pageId="3" pageNumber="628">/</emphasis>
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14 branches; M (excluding its branches associated with CuA) branched distal to R, concave, anteriorly pectinate in right forewing, irregularly branched in left forewing; M with 11
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<emphasis id="A8B2EA8AFFDDA249FB98BEAF604CFA5D" box="[1073,1079,1486,1509]" italics="true" pageId="3" pageNumber="628">/</emphasis>
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9 branches free of CuA; convex transverse to oblique, and somewhat irregular veinlets interconnecting M and CuA proximal to the branching of both, with a stronger basal veinlet and one
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<emphasis id="A8B2EA8AFFDDA249FCA0BD586774F9E8" box="[777,783,1593,1616]" italics="true" pageId="3" pageNumber="628">/</emphasis>
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several weaker veinlets; CuA convex; branches of M diverging from CuA + M
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<emphasis id="A8B2EA8AFFDDA249FC45BD356056F9D3" box="[1004,1069,1620,1643]" italics="true" pageId="3" pageNumber="628">partim</emphasis>
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concave; M
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<emphasis id="A8B2EA8AFFDDA249FB01BD3566A7F93E" italics="true" pageId="3" pageNumber="628">partim</emphasis>
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+ CuA with 17
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<emphasis id="A8B2EA8AFFDDA249FCDEBD0F6706F93D" box="[887,893,1646,1669]" italics="true" pageId="3" pageNumber="628">/</emphasis>
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18 branches; CuP simple, strongly curved, strongly concave, reaching the posterior wing margin near 40% of wing length; anal area with 16
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<emphasis id="A8B2EA8AFFDDA249FB19BDC560CDF903" box="[1200,1206,1700,1723]" italics="true" pageId="3" pageNumber="628">/</emphasis>
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14 AA branches, all reaching the posterior wing margin;
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<emphasis id="A8B2EA8AFFDAA24EFDD9BE6A64C1FA86" bold="true" pageId="4" pageNumber="629">hind wing</emphasis>
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: length
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<quantity id="5D3E9B7DFFDAA24EFEA4BE446517FA84" box="[269,364,1317,1341]" metricMagnitude="-2" metricUnit="m" metricValue="2.94" pageId="4" pageNumber="629" unit="mm" value="29.4">29.4 mm</quantity>
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, width of remigium
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<quantity id="5D3E9B7DFFDAA24EFD92BE4466E2FA84" box="[571,665,1317,1341]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="4" pageNumber="629" unit="mm" value="13.3">13.3 mm</quantity>
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; vannus folded over remigium (whether the vannus is provided with longitudinal folds, or not, cannot be observed); ScP concave, reaching the anterior wing margin at mid-length; R convex, branched
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<quantity id="5D3E9B7DFFDAA24EFDE2BEF066E7FA11" box="[587,668,1425,1449]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="4" pageNumber="629" unit="mm" value="5.7">5.7 mm</quantity>
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distal to wing base; RA convex, branched distally, with four branches; RP concave, branched
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<quantity id="5D3E9B7DFFDAA24EFDA4BEA96625FA58" box="[525,606,1480,1504]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="629" unit="mm" value="3.9">3.9 mm</quantity>
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distal to its origin, anteriorly pectinate, with 11 branches; M concave, weak at its origin, with three branches free of CuA; a simple branch of M diverges anteriorly from M
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<emphasis id="A8B2EA8AFFDAA24EFF01BD576490F9F5" box="[168,235,1590,1613]" italics="true" pageId="4" pageNumber="629">partim</emphasis>
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+ CuA; CuA located in a depression but concave, posteriorly pectinate, with six branches; CuP concave, sigmoid, weak, simple; AA(1?) (located on the remigium) with three branches reaching the posterior wing margin; basal to its main fork, AA(1?) emits two strong struts posteriorly; fold separating the remigium from the vannus crossing two veins (
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<figureCitation id="02FD2A1DFFDAA24EFBA7BE45603EFA84" box="[1038,1093,1316,1340]" captionStart="FIG" captionStartId="4.[136,147,1178,1195]" captionTargetBox="[207,1174,241,1132]" captionTargetId="figure-259@4.[207,1124,243,701]" captionTargetPageId="4" captionText="FIG. 3. — Phyloblatta gaudryi (Agnus, 1903); holotype specimen MNHN.F.R51454, left hind wing; reconstruction (continuous grey line indicates the remigium-vannus fold. *, indicates a branch of M diverging from CuA; °, indicates branches that cross the remigiumvannus fold) and photograph (negative imprint). See text for abbreviations. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5377995" httpUri="https://zenodo.org/record/5377995/files/figure.png" pageId="4" pageNumber="629">Fig. 3</figureCitation>
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[°]), and reaching the posterior wing margin at 38% of wing length; vannus moderately large, filled with numerous branches (belonging either to AA1 or AA2, see below).
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</paragraph>
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</subSubSection>
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<caption id="CEB96610FFDAA24EFF21BFFB67FCFB60" ID-DOI="http://doi.org/10.5281/zenodo.5377995" ID-Zenodo-Dep="5377995" httpUri="https://zenodo.org/record/5377995/files/figure.png" pageId="4" pageNumber="629" startId="4.[136,147,1178,1195]" targetBox="[207,1174,241,1132]" targetPageId="4">
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<paragraph id="9A793698FFDAA24EFF21BFFB67FCFB60" blockId="4.[136,1233,1178,1240]" pageId="4" pageNumber="629">
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FIG. 3. —
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<taxonomicName id="5DC64D1BFFDAA24EFF7EBFFB658BFB13" authority="(Agnus, 1903)" baseAuthorityName="Agnus" baseAuthorityYear="1903" box="[215,496,1178,1195]" class="Insecta" family="Archimylacridae" genus="Phyloblatta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="gaudryi">
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<emphasis id="A8B2EA8AFFDAA24EFF7EBFFB650DFB13" box="[215,374,1178,1195]" italics="true" pageId="4" pageNumber="629">Phyloblatta gaudryi</emphasis>
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(
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<bibRefCitation id="FE574B69FFDAA24EFED6BFFB6591FB13" author="AGNUS A. N." box="[383,490,1178,1195]" pageId="4" pageNumber="629" pagination="291 - 294" refId="ref4668" refString="AGNUS A. N. 1903. - Deuxieme note sur les Blattidae paleozoiques (Orthopt. palaeoz.). Description d'une espece nouvelle. Bulletin de la Societe entomologique de France 17: 291 - 294." type="journal article" year="1903">Agnus, 1903</bibRefCitation>
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)
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</taxonomicName>
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; holotype specimen MNHN.F.R51454, left hind wing; reconstruction (continuous grey line indicates the remigium-vannus fold.
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<emphasis id="A8B2EA8AFFDAA24EFE12BFD165B8FB79" bold="true" box="[443,451,1200,1217]" pageId="4" pageNumber="629">*</emphasis>
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, indicates a branch of M diverging from CuA;
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<emphasis id="A8B2EA8AFFDAA24EFCE3BFD1672AFB79" bold="true" box="[842,849,1200,1217]" pageId="4" pageNumber="629">°</emphasis>
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, indicates branches that cross the remigiumvannus fold) and photograph (negative imprint). See text for abbreviations. Scale bar: 5 mm.
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</paragraph>
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</caption>
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<subSubSection id="D2DC6513FFDAA24FFD12BEA26544FD9E" lastPageId="5" lastPageNumber="630" pageId="4" pageNumber="629" type="discussion">
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<paragraph id="9A793698FFDAA24EFD12BEA26764FA63" blockId="4.[697,1235,1475,1723]" box="[699,799,1475,1501]" pageId="4" pageNumber="629">REMARKS</paragraph>
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<paragraph id="9A793698FFDAA24EFD12BE8360FFF902" blockId="4.[697,1235,1475,1723]" pageId="4" pageNumber="629">
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The specimen most likely experienced moderate plastic deformation during fossilisation, as indicated by the forewings aspect ratio, differing in the right and left forewings. The difference observed in the fossil individual exceeds that observed between right and left forewings of
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<taxonomicName id="5DC64D1BFFDAA24EFC4CBDE1670EF903" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Blattidae" genus="Periplaneta" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="americana">
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<emphasis id="A8B2EA8AFFDAA24EFC4CBDE160ABF922" box="[997,1232,1664,1690]" italics="true" pageId="4" pageNumber="629">Periplaneta americana</emphasis>
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(Linnaeus, 1758)
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</taxonomicName>
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(
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<bibRefCitation id="FE574B69FFDAA24EFC2ABDC16049F902" author="SCHNEIDER J." box="[899,1074,1696,1722]" pageId="4" pageNumber="629" pagination="87 - 105" refId="ref5529" refString="SCHNEIDER J. 1977. - Zur Variabilitat der Flugel palaozoischer Blattodea (Insecta), Teil I. Freiberger Forschungshefte (C) 326: 87 - 105." type="journal article" year="1977">Schneider 1977</bibRefCitation>
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: fig. 2).
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</paragraph>
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<paragraph id="9A793698FFDBA24FFF36BB046567FFCE" blockId="5.[159,284,101,118]" box="[159,284,101,118]" pageId="5" pageNumber="630">
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Béthoux O.
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<emphasis id="A8B2EA8AFFDBA24FFF51BB046567FFCE" box="[248,284,101,118]" italics="true" pageId="5" pageNumber="630">et al.</emphasis>
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</paragraph>
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<paragraph id="9A793698FFDBA24FFF21BB8E6571FEBF" blockId="5.[136,671,239,550]" box="[136,266,239,265]" pageId="5" pageNumber="630">VARIABILITY</paragraph>
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<paragraph id="9A793698FFDBA24FFF21BA6F6544FD9E" blockId="5.[136,671,239,550]" pageId="5" pageNumber="630">
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As detailed by
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<bibRefCitation id="FE574B69FFDBA24FFE9BBA6F6666FE90" author="SCHNEIDER J." box="[306,541,270,296]" pageId="5" pageNumber="630" pagination="87 - 105" refId="ref5529" refString="SCHNEIDER J. 1977. - Zur Variabilitat der Flugel palaozoischer Blattodea (Insecta), Teil I. Freiberger Forschungshefte (C) 326: 87 - 105." type="journal article" year="1977">Schneider (1977: 81</bibRefCitation>
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, pl. 2), the forewing venation of
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<taxonomicName id="5DC64D1BFFDBA24FFED2BA4F6636FEF0" baseAuthorityName="Agnus" baseAuthorityYear="1903" box="[379,589,302,328]" class="Insecta" family="Archimylacridae" genus="Phyloblatta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="630" phylum="Arthropoda" rank="species" species="gaudryi">
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<emphasis id="A8B2EA8AFFDBA24FFED2BA4F6636FEF0" box="[379,589,302,328]" italics="true" pageId="5" pageNumber="630">Phyloblatta gaudryi</emphasis>
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</taxonomicName>
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is particularly variable with respect to other Palaeozoic stem-Dictyoptera. In particular the number of terminal branches and branching pattern of each vein (sector) varies considerably (e.g., the number of terminal branches of the anterior branch of R ranges from 1 to 9 (
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<bibRefCitation id="FE574B69FFDBA24FFEF3BA8D6673FDBE" author="SCHNEIDER J." box="[346,520,492,518]" pageId="5" pageNumber="630" pagination="87 - 105" refId="ref5529" refString="SCHNEIDER J. 1977. - Zur Variabilitat der Flugel palaozoischer Blattodea (Insecta), Teil I. Freiberger Forschungshefte (C) 326: 87 - 105." type="journal article" year="1977">Schneider 1977</bibRefCitation>
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: pl. 2, figs 12, 11, respectively).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |