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<document id="140845C6CFD0BF1291D07338D648033A" ID-CLB-Dataset="45826" ID-DOI="10.5281/zenodo.200703" ID-GBIF-Dataset="c3116728-c150-4a0c-835a-f7705577eb87" ID-ISSN="1175-5326" ID-Zenodo-Dep="200703" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460417722364" checkinUser="plazi" docAuthor="Lotzkat, Sebastian, Bienentreu, Joe-Felix, Hertz, Andreas &amp; Köhler, Gunther" docDate="2011" docId="F77D87AE7F415321FF3FF970FBB8FF4F" docLanguage="en" docName="zt03125p021.pdf" docOrigin="Zootaxa 3125" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anolis benedikti Lotzkat, Bienentreu, Hertz &amp; Köhler, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="18" masterDocId="0B44FFD67F465330FFA8FFECFFC8FF9B" masterDocTitle="A new species of Anolis (Squamata: Iguania: Dactyloidae) formerly referred to as A. pachypus from the Cordillera de Talamanca of western Panama and adjacent Costa Rica" masterLastPageNumber="21" masterPageNumber="1" pageNumber="8" updateTime="1698329135495" updateUser="ExternalLinkService">
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<mods:title id="71AB48ECED8CF9987832E948071DBCA8">A new species of Anolis (Squamata: Iguania: Dactyloidae) formerly referred to as A. pachypus from the Cordillera de Talamanca of western Panama and adjacent Costa Rica</mods:title>
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<mods:namePart id="6B65F4FED8BD6AE1E04918B5759F1803">Lotzkat, Sebastian</mods:namePart>
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<mods:namePart id="2380F8B505D8B2173C75C2E4833467E0">Bienentreu, Joe-Felix</mods:namePart>
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<mods:namePart id="9B302577E3C0D135392B2439F49A42B6">Hertz, Andreas</mods:namePart>
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<mods:namePart id="B401D758DB750227B7B05136C14172D2">Köhler, Gunther</mods:namePart>
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<mods:date id="3A29CCCC2D4A12F1FB684723F6436ED4">2011</mods:date>
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<treatment id="F77D87AE7F415321FF3FF970FBB8FF4F" ID-DOI="http://doi.org/10.5281/zenodo.5617003" ID-GBIF-Taxon="119425603" ID-Zenodo-Dep="5617003" LSID="urn:lsid:plazi:treatment:F77D87AE7F415321FF3FF970FBB8FF4F" httpUri="http://treatment.plazi.org/id/F77D87AE7F415321FF3FF970FBB8FF4F" lastPageId="17" lastPageNumber="18" pageId="7" pageNumber="8">
<subSubSection id="37CE65337F415337FF3FF970FEA1F943" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="7F6B36B87F415337FF3FF970FE7BF92E" blockId="7.[151,435,1692,1752]" box="[151,435,1692,1717]" pageId="7" pageNumber="8">
<heading id="242381D47F415337FF3FF970FE7BF92E" bold="true" box="[151,435,1692,1717]" fontSize="10" level="2" pageId="7" pageNumber="8" reason="2">
<emphasis id="4DA0EAAA7F415337FF3FF970FE7BF92E" bold="true" box="[151,435,1692,1717]" pageId="7" pageNumber="8">
<taxonomicName id="B8D44D3B7F415337FF3FF970FE9AF92E" box="[151,338,1692,1717]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="benedikti" status="sp. nov.">
<emphasis id="4DA0EAAA7F415337FF3FF970FE9AF92E" bold="true" box="[151,338,1692,1717]" italics="true" pageId="7" pageNumber="8">Anolis benedikti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="569357D17F415337FEF1F971FE7BF92E" box="[345,435,1693,1717]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7F6B36B87F415337FF3FF92CFEA1F943" blockId="7.[151,435,1692,1752]" box="[151,361,1728,1752]" pageId="7" pageNumber="8">
<figureCitation id="E7EF2A3D7F415337FF3FF92CFF33F943" box="[151,251,1728,1752]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1851]" captionTargetBox="[305,1272,193,1803]" captionTargetId="figure@3.[305,1272,193,1804]" captionTargetPageId="3" captionText="FIGURE 1. Dewlaps of anoles related to Anolis pachypus from Costa Rica and western Panama, representative of the five OTUs (width of tweezers 3 mm): (A) OTU A (SMF 89729, Sendero La Cascada, Bajo Mono, Chiriquí, Panama); (B) OTU B (GK 2297, road from San José de la Montaña to Birrí, Heredia, Costa Rica); (C) OTU C (GK 2355, road from San Isidro de El General to Cerro de la Muerte, San José, Costa Rica); (D) OTU C (SL 439, Jurutungo, Chiriquí, Panama); (E) OTU C (SL 524, Jurutungo, Chiriquí, Panama); (F) OTU D (SMF 89507, Jurutungo, Chiriquí, Panama); (G) OTU D (SMF 89506, Jurutungo, Chiriquí, Panama); (H) OTU D (SMF 90149, N slope Cerro Pando, Bocas del Toro, Panama); (I) OTU E (SMF 89519, La Nevera, Comarca Ngöbe-Buglé, Panama); (J) OTU E (SL 510, La Nevera, Comarca Ngöbe-Buglé, Panama)." httpUri="https://zenodo.org/record/200704/files/figure.png" pageId="7" pageNumber="8">Figures 1</figureCitation>
FH, 57
</paragraph>
</subSubSection>
<subSubSection id="37CE65337F415337FF3FF8E4FACCF885" box="[151,1284,1800,1823]" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="7F6B36B87F415337FF3FF8E4FACCF885" blockId="7.[151,1284,1800,1823]" box="[151,1284,1800,1823]" pageId="7" pageNumber="8">
<taxonomicName id="B8D44D3B7F415337FF3FF8E4FDE8F885" authority="Poe and Ibanez 2007" authorityName="Poe and Ibanez" authorityYear="2007" box="[151,544,1800,1822]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F415337FF3FF8E4FE8DF885" box="[151,325,1800,1822]" italics="true" pageId="7" pageNumber="8">Anolis pachypus:</emphasis>
Poe and Ibañez 2007
</taxonomicName>
(in part);
<taxonomicName id="B8D44D3B7F415337FD2EF8E6FC0EF885" authority="Savage 2002" authorityName="Savage" authorityYear="2002" box="[646,966,1800,1823]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F415337FD2EF8E6FCF5F885" box="[646,829,1800,1823]" italics="true" pageId="7" pageNumber="8">Norops pachypus:</emphasis>
<bibRefCitation id="1B454B497F415337FCECF8E5FC0EF885" author="Savage" box="[836,966,1801,1822]" pageId="7" pageNumber="8" refString="Savage, J. M. (2002) The amphibians and reptiles of Costa Rica. A herpetofauna between two continents, between two seas. University of Chicago Press, Chicago, xx + 934 pp." type="book" year="2002">Savage 2002</bibRefCitation>
</taxonomicName>
(in part); Köhler 2008 (in part)
</paragraph>
</subSubSection>
<subSubSection id="37CE65337F41533BFF3FF8A0FDACFD3C" lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="7F6B36B87F415337FF3FF8A0FE84F868" blockId="7.[151,1436,1867,2035]" pageId="7" pageNumber="8">
<emphasis id="4DA0EAAA7F415337FF3FF8A0FEC3F8FF" bold="true" box="[151,267,1868,1892]" pageId="7" pageNumber="8">
<typeStatus id="A06F881A7F415337FF3FF8A0FECEF8FF" box="[151,262,1868,1892]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="19C5AE7D7F415337FEBAF8A7FE82F8F8" LSID="urn:lsid:biocol.org:col:34838" box="[274,330,1867,1891]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="8">SMF</collectionCode>
90149 (
<figureCitation id="E7EF2A3D7F415337FE0FF8A7FE39F8F8" box="[423,497,1867,1891]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1851]" captionTargetBox="[305,1272,193,1803]" captionTargetId="figure@3.[305,1272,193,1804]" captionTargetPageId="3" captionText="FIGURE 1. Dewlaps of anoles related to Anolis pachypus from Costa Rica and western Panama, representative of the five OTUs (width of tweezers 3 mm): (A) OTU A (SMF 89729, Sendero La Cascada, Bajo Mono, Chiriquí, Panama); (B) OTU B (GK 2297, road from San José de la Montaña to Birrí, Heredia, Costa Rica); (C) OTU C (GK 2355, road from San Isidro de El General to Cerro de la Muerte, San José, Costa Rica); (D) OTU C (SL 439, Jurutungo, Chiriquí, Panama); (E) OTU C (SL 524, Jurutungo, Chiriquí, Panama); (F) OTU D (SMF 89507, Jurutungo, Chiriquí, Panama); (G) OTU D (SMF 89506, Jurutungo, Chiriquí, Panama); (H) OTU D (SMF 90149, N slope Cerro Pando, Bocas del Toro, Panama); (I) OTU E (SMF 89519, La Nevera, Comarca Ngöbe-Buglé, Panama); (J) OTU E (SL 510, La Nevera, Comarca Ngöbe-Buglé, Panama)." httpUri="https://zenodo.org/record/200704/files/figure.png" pageId="7" pageNumber="8">Figs. 1</figureCitation>
H, 5A and F, 6), adult male, from the north slope of Cerro Pando, leaving the cattle trail to the right after following it to about
<quantity id="B82C9B5D7F415337FDDAF883FD0FF81C" box="[626,711,1903,1927]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="m" value="1000.0">1000 m</quantity>
airline north of the large border monument (
<figureCitation id="E7EF2A3D7F415337FB13F883FB35F81C" box="[1211,1277,1903,1927]" captionStart="FIGURE 8" captionStartId="15.[151,250,1789,1812]" captionTargetBox="[258,1328,193,1767]" captionTargetId="figure@15.[258,1329,193,1768]" captionTargetPageId="15" captionText="FIGURE 8. Habitats of Anolis benedikti: (A C) Pacific slope in Chiriquí province: (A) view over Jurutungo north toward the continental divide (hidden in clouds), the ranges of A. benedikti and A. pachypus meet in the background; (B) forest trail at Jurutungo, one of the lowest collection sites of A. benedikti on the Pacific slope, 2070 m asl; (C) cloud forest near continental divide east of Cerro Pando, 2460 m asl; (D) international border monument on cattle trail along the continental divide, 2450 m asl; (E H) Caribbean slope in Bocas del Toro province: (E) cloud forest at the type locality, 2310 m asl; (F) cloud forest near the type locality, 2390 m asl; (G) clearing for cattle along tributary to Río Changena, 1810 m asl; (H) Río Changena, the lowest collection site on the Caribbean slope, 1640 m asl." httpUri="https://zenodo.org/record/200712/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
D),
<geoCoordinate id="1AE0507F7F415337FA80F883FA5FF81C" box="[1320,1431,1903,1927]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="5" value="8.9333">8.9333°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F415337FF3FF87FFED0F830" box="[151,280,1939,1963]" direction="west" orientation="longitude" pageId="7" pageNumber="8" precision="5" value="-82.7131">82.7131°W</geoCoordinate>
,
<quantity id="B82C9B5D7F415337FE8CF87FFEB2F830" box="[292,378,1939,1963]" metricMagnitude="3" metricUnit="m" metricValue="2.31" pageId="7" pageNumber="8" unit="m" value="2310.0">2310 m</quantity>
asl, Parque Internacional La Amistad (
<collectionCode id="19C5AE7D7F415337FC9DF87FFCB1F830" box="[821,889,1939,1963]" pageId="7" pageNumber="8">PILA</collectionCode>
), Bocas del Toro Province,
<collectingCountry id="07C376287F415337FB1BF878FAC7F830" box="[1203,1295,1940,1963]" name="Panama" pageId="7" pageNumber="8">Panama</collectingCountry>
, close to the border with
<collectingCountry id="07C376287F415337FE8CF85BFE6BF854" box="[292,419,1975,1999]" name="Costa Rica" pageId="7" pageNumber="8">Costa Rica</collectingCountry>
; collected by Andreas Hertz and Sebastian Lotzkat on
<date id="0B6A10787F415337FBB3F85BFB33F854" box="[1051,1275,1975,1999]" pageId="7" pageNumber="8" value="2009-11-19">19 November 2009</date>
; original field number SL 529.
</paragraph>
<caption id="2BAB66307F4E5338FF3FF880FA46F845" httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="8" pageNumber="9" targetBox="[216,1349,193,1860]" targetPageId="8">
<paragraph id="7F6B36B87F4E5338FF3FF880FA46F845" blockId="8.[151,1436,1900,2015]" pageId="8" pageNumber="9">
<emphasis id="4DA0EAAA7F4E5338FF3FF880FEDEF819" bold="true" box="[151,278,1900,1923]" pageId="8" pageNumber="9">FIGURE 5.</emphasis>
Type specimens of
<taxonomicName id="B8D44D3B7F4E5338FE45F880FD5CF819" box="[493,660,1900,1922]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F4E5338FE45F880FD5CF819" box="[493,660,1900,1922]" italics="true" pageId="8" pageNumber="9">Anolis benedikti</emphasis>
</taxonomicName>
in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (CE) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (FO) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female.
</paragraph>
</caption>
<caption id="2BAB66307F4F5339FF3FF847FE4BF87B" httpUri="https://zenodo.org/record/200710/files/figure.png" pageId="9" pageNumber="10" targetBox="[259,1293,206,1930]" targetPageId="9">
<paragraph id="7F6B36B87F4F5339FF3FF847FE4BF87B" blockId="9.[151,1436,1963,2016]" pageId="9" pageNumber="10">
<emphasis id="4DA0EAAA7F4F5339FF3FF847FEDDF85A" bold="true" box="[151,277,1963,1986]" pageId="9" pageNumber="10">FIGURE 6.</emphasis>
Head of Holotype (SMF 90149) of
<emphasis id="4DA0EAAA7F4F5339FD39F840FC88F859" box="[657,832,1964,1986]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="B8D44D3B7F4F5339FD39F840FCF3F859" box="[657,827,1964,1986]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="benedikti">Anolis benedikti</taxonomicName>
:
</emphasis>
(A) lateral (right side), (B) dorsal, and (C) ventral views. Scale bars equal 1 mm.
</paragraph>
</caption>
<caption id="2BAB66307F4C533AFF3FF845FAF7F845" httpUri="https://zenodo.org/record/200711/files/figure.png" pageId="10" pageNumber="11" targetBox="[286,1309,205,1926]" targetPageId="10">
<paragraph id="7F6B36B87F4C533AFF3FF845FAF7F845" blockId="10.[151,1436,1961,2015]" pageId="10" pageNumber="11">
<emphasis id="4DA0EAAA7F4C533AFF3FF845FEDBF824" bold="true" box="[151,275,1961,1984]" pageId="10" pageNumber="11">FIGURE 7.</emphasis>
Designation of nasal scalation types. (A) Type A (SMF 90149); (B) Type B (SMF 92134); (C) Type C (SL 678); (D) Type D (SMF 85264); (E) Type E (SMF 89506); (F) Type F (85272). Scale bars equal 1 mm. See Text for details.
</paragraph>
</caption>
<paragraph id="7F6B36B87F4D533BFF6DFF74FF1EFD1F" blockId="11.[151,1436,151,2012]" pageId="11" pageNumber="12">
<emphasis id="4DA0EAAA7F4D533BFF6DFF74FE8CFF2B" bold="true" box="[197,324,152,176]" pageId="11" pageNumber="12">
<typeStatus id="A06F881A7F4D533BFF6DFF74FEF7FF2B" box="[197,319,152,176]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Collected by Sebastian Lotzkat and Andreas Hertz, if not indicated otherwise. All from the
<collectionCode id="19C5AE7D7F4D533BFAEAFF74FA48FF2B" box="[1346,1408,152,176]" pageId="11" pageNumber="12">PILA</collectionCode>
in
<collectingCountry id="07C376287F4D533BFF3FFF51FF3BFF4F" box="[151,243,189,212]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
. Bocas del Toro: north slope Cerro Pando:
<collectionCode id="19C5AE7D7F4D533BFD4EFF50FCD6FF4F" LSID="urn:lsid:biocol.org:col:34838" box="[742,798,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
89744, male, near
<typeStatus id="A06F881A7F4D533BFC56FF51FBE6FF4E" box="[1022,1070,189,213]" pageId="11" pageNumber="12">type</typeStatus>
locality,
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,
<geoCoordinate id="1AE0507F7F4D533BFABEFF50FA50FF4F" box="[1302,1432,188,212]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="-82.7137">82.7137°W</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFF3FFF33FF25FF6C" box="[151,237,223,247]" metricMagnitude="3" metricUnit="m" metricValue="2.3899999999999997" pageId="11" pageNumber="12" unit="m" value="2390.0">2390 m</quantity>
asl,
<date id="0B6A10787F4D533BFE8CFF33FE0FFF6C" box="[292,455,223,247]" pageId="11" pageNumber="12" value="2009-04-19">19 April 2009</date>
;
<collectionCode id="19C5AE7D7F4D533BFE7CFF33FDC4FF6C" LSID="urn:lsid:biocol.org:col:34838" box="[468,524,223,247]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
897456, male and female, same locality as
<typeStatus id="A06F881A7F4D533BFBB8FF33FBBDFF6C" box="[1040,1141,223,247]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
, 2330 and
<quantity id="B82C9B5D7F4D533BFB5BFF33FA81FF6C" box="[1267,1353,223,247]" metricMagnitude="3" metricUnit="m" metricValue="2.31" pageId="11" pageNumber="12" unit="m" value="2310.0">2310 m</quantity>
asl,
<date id="0B6A10787F4D533BFA28FF33FEDCFE87" pageId="11" pageNumber="12" value="2009-04-20">20 April 2009</date>
;
<collectionCode id="19C5AE7D7F4D533BFE88FEE8FE90FE87" LSID="urn:lsid:biocol.org:col:34838" box="[288,344,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
90148, juvenile male, near
<typeStatus id="A06F881A7F4D533BFD3AFEE9FD0AFE86" box="[658,706,261,285]" pageId="11" pageNumber="12">type</typeStatus>
locality,
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,
<geoCoordinate id="1AE0507F7F4D533BFC0DFEE8FBEEFE87" box="[933,1062,260,284]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="-82.7128">82.7128°W</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFB99FEE8FB4FFE87" box="[1073,1159,260,284]" metricMagnitude="3" metricUnit="m" metricValue="2.28" pageId="11" pageNumber="12" unit="m" value="2280.0">2280 m</quantity>
asl,
<date id="0B6A10787F4D533BFB13FEE8FA50FE87" box="[1211,1432,260,284]" pageId="11" pageNumber="12" value="2009-11-19">19 November 2009</date>
;
<collectionCode id="19C5AE7D7F4D533BFF3FFECBFF07FEA4" LSID="urn:lsid:biocol.org:col:34838" box="[151,207,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
91508, male, SL 6778, subadult male and female, along tributary to Río Changena,
<geoCoordinate id="1AE0507F7F4D533BFB36FECBFAC6FEA4" box="[1182,1294,295,319]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="8.9474">8.9474°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F4D533BFAB2FECBFA50FEA4" box="[1306,1432,295,319]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="82.7098">82.7098°N</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFF3FFEA0FEF9FEFF" box="[151,305,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.0" metricValueMin="1.98" pageId="11" pageNumber="12" unit="m" value="1990.0" valueMax="2000.0" valueMin="1980.0">19802000 m</quantity>
asl,
<date id="0B6A10787F4D533BFECCFEA0FE3FFEFF" box="[356,503,332,356]" pageId="11" pageNumber="12" value="2010-07-13">13 July 2010</date>
;
<collectionCode id="19C5AE7D7F4D533BFDAAFEA0FDF2FEFF" LSID="urn:lsid:biocol.org:col:34838" box="[514,570,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
91507, female, along tributary to Río Changena,
<geoCoordinate id="1AE0507F7F4D533BFBCAFEA0FB19FEFF" box="[1122,1233,332,356]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="8.9524">8.9524°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F4D533BFB74FEA0FA91FEFF" box="[1244,1369,332,356]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="82.7093">82.7093°N</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFACCFEA0FF65FE1C" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="11" pageNumber="12" unit="m" value="1960.0">1960 m</quantity>
asl,
<date id="0B6A10787F4D533BFF48FE83FEBCFE1C" box="[224,372,367,391]" pageId="11" pageNumber="12" value="2010-07-14">14 July 2010</date>
;
<collectionCode id="19C5AE7D7F4D533BFED6FE83FE7EFE1C" LSID="urn:lsid:biocol.org:col:34838" box="[382,438,367,391]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
915056, SL 686, 695, females, Río Changena,
<geoCoordinate id="1AE0507F7F4D533BFC7BFE83FB8AFE1C" box="[979,1090,367,391]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="8.9785">8.9785°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F4D533BFBE5FE83FB02FE1C" box="[1101,1226,367,391]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="82.6901">82.6901°N</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFB7DFE83FAB8FE1C" box="[1237,1392,367,391]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.66" metricValueMin="1.64" pageId="11" pageNumber="12" unit="m" value="1650.0" valueMax="1660.0" valueMin="1640.0">16401660 m</quantity>
asl,
<date id="0B6A10787F4D533BFF3FFE78FE9BFE37" box="[151,339,404,428]" pageId="11" pageNumber="12" value="2010-07-14" valueMax="2010-07-17" valueMin="2010-07-14">1417 July 2010</date>
;
<collectionCode id="19C5AE7D7F4D533BFEF6FE78FE5EFE37" LSID="urn:lsid:biocol.org:col:34838" box="[350,406,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
91509, juvenile, between Río Clarito and Río Changena,
<geoCoordinate id="1AE0507F7F4D533BFBB1FE78FB40FE37" box="[1049,1160,404,428]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="8.9887">8.9887°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F4D533BFB3BFE78FAD9FE37" box="[1171,1297,404,428]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="82.6749">82.6749°N</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFAB3FE78FAB8FE37" box="[1307,1392,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.8199999999999998" pageId="11" pageNumber="12" unit="m" value="1820.0">1820 m</quantity>
asl,
<date id="0B6A10787F4D533BFF3FFE5BFEE6FE54" box="[151,302,439,463]" pageId="11" pageNumber="12" value="2010-07-17">17 July 2010</date>
; Chiriquí: south slope Cerro Pando, Jurutungo:
<collectionCode id="19C5AE7D7F4D533BFCE4FE5BFC4CFE54" LSID="urn:lsid:biocol.org:col:34838" box="[844,900,439,463]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
852767, males,
<geoCoordinate id="1AE0507F7F4D533BFBE5FE5BFB74FE54" box="[1101,1212,439,463]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="8.9083">8.9083°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F4D533BFB61FE5BFA82FE54" box="[1225,1354,439,463]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="-82.7168">82.7168°W</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFAFEFE5BFF3BFE6F" metricMagnitude="3" metricUnit="m" metricValue="2.055" metricValueMax="2.1" metricValueMin="2.01" pageId="11" pageNumber="12" unit="m" value="2055.0" valueMax="2100.0" valueMin="2010.0">2010 2100 m</quantity>
asl, collected by Gunther Köhler on
<date id="0B6A10787F4D533BFD6FFE30FC5AFE6F" box="[711,914,476,501]" pageId="11" pageNumber="12" value="2006-01-11">11 January 2006</date>
;
<collectionCode id="19C5AE7D7F4D533BFC0DFE30FC15FE6F" LSID="urn:lsid:biocol.org:col:34838" box="[933,989,476,500]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
852723, subadult males,
<geoCoordinate id="1AE0507F7F4D533BFA80FE30FA5FFE6F" box="[1320,1431,476,500]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="8.9109">8.9109°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F4D533BFF3FFE13FEDFFD8C" box="[151,279,511,535]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="-82.7144">82.7144°W</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFE8BFE13FEB1FD8C" box="[291,377,511,535]" metricMagnitude="3" metricUnit="m" metricValue="2.06" pageId="11" pageNumber="12" unit="m" value="2060.0">2060 m</quantity>
asl, collected by Gunther Köhler on
<date id="0B6A10787F4D533BFCB1FE13FC1EFD8C" box="[793,982,511,536]" pageId="11" pageNumber="12" value="2006-01-10">10 January 2006</date>
; SL 202, female,
<geoCoordinate id="1AE0507F7F4D533BFB35FE13FAC3FD8C" box="[1181,1291,511,535]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="8.9119">8.9119°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F4D533BFABFFE13FA50FD8C" box="[1303,1432,511,535]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="-82.7095">82.7095°W</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFF3FFDC8FF24FDA7" box="[151,236,548,572]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="11" pageNumber="12" unit="m" value="2250.0">2250 m</quantity>
asl,
<date id="0B6A10787F4D533BFE8AFDC8FE61FDA7" box="[290,425,548,572]" pageId="11" pageNumber="12" value="2008-07-06">6 July 2008</date>
;
<collectionCode id="19C5AE7D7F4D533BFE1EFDC8FE26FDA7" LSID="urn:lsid:biocol.org:col:34838" box="[438,494,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
89505, female,
<geoCoordinate id="1AE0507F7F4D533BFD0DFDC8FCDAFDA7" box="[677,786,548,572]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="8.9111">8.9111°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F4D533BFCB6FDC8FC57FDA7" box="[798,927,548,572]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="-82.7159">82.7159°W</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFC03FDC8FBC9FDA7" box="[939,1025,548,572]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="11" pageNumber="12" unit="m" value="2020.0">2020 m</quantity>
asl,
<date id="0B6A10787F4D533BFB9EFDC8FB77FDA7" box="[1078,1215,548,572]" pageId="11" pageNumber="12" value="2008-07-08">8 July 2008</date>
;
<collectionCode id="19C5AE7D7F4D533BFB62FDC8FACAFDA7" LSID="urn:lsid:biocol.org:col:34838" box="[1226,1282,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
89506, male,
<geoCoordinate id="1AE0507F7F4D533BFF3FFDABFECEFDC4" box="[151,262,583,607]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="8.9121">8.9121°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F4D533BFEBCFDABFE5DFDC4" box="[276,405,583,607]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="-82.7096">82.7096°W</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFE0BFDABFE32FDC4" box="[419,506,583,607]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="12" unit="m" value="2200.0">2200 m</quantity>
asl,
<date id="0B6A10787F4D533BFD9CFDABFD77FDC4" box="[564,703,583,607]" pageId="11" pageNumber="12" value="2008-07-08">8 July 2008</date>
;
<collectionCode id="19C5AE7D7F4D533BFD65FDABFCCDFDC4" LSID="urn:lsid:biocol.org:col:34838" box="[717,773,583,607]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
89507, male,
<geoCoordinate id="1AE0507F7F4D533BFC04FDABFBD2FDC4" box="[940,1050,583,607]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="8.9114">8.9114°N</geoCoordinate>
,
<geoCoordinate id="1AE0507F7F4D533BFB80FDABFB61FDC4" box="[1064,1193,583,607]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="-82.7129">82.7129°W</geoCoordinate>
,
<quantity id="B82C9B5D7F4D533BFB1FFDABFAC6FDC4" box="[1207,1294,583,607]" metricMagnitude="3" metricUnit="m" metricValue="2.07" pageId="11" pageNumber="12" unit="m" value="2070.0">2070 m</quantity>
asl,
<date id="0B6A10787F4D533BFAE0FDABFF1BFD1F" pageId="11" pageNumber="12" value="2008-07-15">15 July 2008</date>
.
</paragraph>
<paragraph id="7F6B36B87F4D533BFF6DFD63FDACFD3C" blockId="11.[151,1436,151,2012]" box="[197,612,655,680]" pageId="11" pageNumber="12">
<emphasis id="4DA0EAAA7F4D533BFF6DFD63FE7FFD33" bold="true" box="[197,439,655,680]" pageId="11" pageNumber="12">Referred specimens.</emphasis>
See Appendix.
</paragraph>
</subSubSection>
<subSubSection id="37CE65337F4D533BFF6DFD5FFDECFA7F" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="7F6B36B87F4D533BFF6DFD5FFDECFA7F" blockId="11.[151,1436,151,2012]" pageId="11" pageNumber="12">
<emphasis id="4DA0EAAA7F4D533BFF6DFD5FFEF7FD57" bold="true" box="[197,319,691,716]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
A medium-sized species (maximum SVL
<quantity id="B82C9B5D7F4D533BFC87FD58FC5EFD57" box="[815,918,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="4.86" pageId="11" pageNumber="12" unit="mm" value="48.6">48.6 mm</quantity>
in males,
<quantity id="B82C9B5D7F4D533BFBBAFD58FBB1FD57" box="[1042,1145,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="4.81" pageId="11" pageNumber="12" unit="mm" value="48.1">48.1 mm</quantity>
in females) of the genus
<taxonomicName id="B8D44D3B7F4D533BFF3FFD34FE74FD74" authority="Poe (2004)" authorityName="Poe" authorityYear="2004" box="[151,444,727,752]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="4DA0EAAA7F4D533BFF3FFD34FF17FD6B" box="[151,223,728,752]" italics="true" pageId="11" pageNumber="12">Anolis</emphasis>
sensu
<bibRefCitation id="1B454B497F4D533BFE90FD35FE74FD74" author="Poe" box="[312,444,727,752]" pageId="11" pageNumber="12" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89." type="journal article" year="2004">Poe (2004)</bibRefCitation>
</taxonomicName>
, that is most similar in external morphology to
<emphasis id="4DA0EAAA7F4D533BFBA0FD34FE9FFC8F" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFBA0FD34FB07FD6B" box="[1032,1231,728,752]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="magnaphallus">A. magnaphallus</taxonomicName>
,
<taxonomicName id="B8D44D3B7F4D533BFB48FD34FABBFD6B" box="[1248,1395,728,752]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pachypus">A. pachypus</taxonomicName>
,
<taxonomicName id="B8D44D3B7F4D533BFA2CFD34FE9BFC8F" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pseudopachypus">A. pseudopachypus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="B8D44D3B7F4D533BFE27FD10FDFEFC8F" box="[399,566,764,788]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="tropidolepis">
<emphasis id="4DA0EAAA7F4D533BFE27FD10FDFEFC8F" box="[399,566,764,788]" italics="true" pageId="11" pageNumber="12">A. tropidolepis</emphasis>
</taxonomicName>
. These four species and
<taxonomicName id="B8D44D3B7F4D533BFCF8FD10FC1FFC8F" box="[848,983,764,788]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F4D533BFCF8FD10FC1FFC8F" box="[848,983,764,788]" italics="true" pageId="11" pageNumber="12">A. benedikti</emphasis>
</taxonomicName>
share narrow toe pads, long legs (tip of fourth toe of adpressed hind limb reaches to a point anterior to eye), a dark interorbital bar, and usually at least the indication of a lyriform marking on the occipital region, keeled dorsal scales on head and body, weakly keeled ventral scales at midbody, and the lack of enlarged postcloacal scales in males.
<taxonomicName id="B8D44D3B7F4D533BFBD6FC84FAF4FC1B" box="[1150,1340,872,896]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F4D533BFBD6FC84FAF4FC1B" box="[1150,1340,872,896]" italics="true" pageId="11" pageNumber="12">Anolis benedikti</emphasis>
</taxonomicName>
can be distinguished readily from all four aforementioned species by the coloration of the male dewlap which is red with a yellow anterior portion comprising just the anterior margin or, at most, the anterior third of the dewlap (versus solid purplish red in
<taxonomicName id="B8D44D3B7F4D533BFEE1FC38FDC2FC77" box="[329,522,980,1004]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="magnaphallus">
<emphasis id="4DA0EAAA7F4D533BFEE1FC38FDC2FC77" box="[329,522,980,1004]" italics="true" pageId="11" pageNumber="12">A. magnaphallus</emphasis>
</taxonomicName>
and
<emphasis id="4DA0EAAA7F4D533BFDEDFC38FD3CFC77" box="[581,756,980,1004]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFDEDFC38FD38FC77" box="[581,752,980,1004]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="tropidolepis">A. tropidolepis</taxonomicName>
,
</emphasis>
red with a central yellow blotch in
<emphasis id="4DA0EAAA7F4D533BFB3FFC38FAE1FC77" box="[1175,1321,980,1004]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFB3FFC38FAEEFC77" box="[1175,1318,980,1004]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pachypus">A. pachypus</taxonomicName>
,
</emphasis>
and solid yellow, sometimes grading into orange-yellow on the posterior portion in
<taxonomicName id="B8D44D3B7F4D533BFC53FC14FB10FB8B" box="[1019,1240,1016,1040]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pseudopachypus">
<emphasis id="4DA0EAAA7F4D533BFC53FC14FB10FB8B" box="[1019,1240,1016,1040]" italics="true" pageId="11" pageNumber="12">A. pseudopachypus</emphasis>
</taxonomicName>
). In addition,
<taxonomicName id="B8D44D3B7F4D533BFA2CFC14FF36FBAF" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F4D533BFA2CFC14FF36FBAF" italics="true" pageId="11" pageNumber="12">A. benedikti</emphasis>
</taxonomicName>
differs from
<emphasis id="4DA0EAAA7F4D533BFE3EFBF0FD3EFBAF" box="[406,758,1052,1076]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFE3EFBF0FD91FBAF" box="[406,601,1052,1076]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="magnaphallus">A. magnaphallus</taxonomicName>
,
<taxonomicName id="B8D44D3B7F4D533BFDCDFBF0FD3AFBAF" box="[613,754,1052,1076]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pachypus">A. pachypus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="B8D44D3B7F4D533BFC98FBF0FC1FFBAF" box="[816,983,1052,1076]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="tropidolepis">
<emphasis id="4DA0EAAA7F4D533BFC98FBF0FC1FFBAF" box="[816,983,1052,1076]" italics="true" pageId="11" pageNumber="12">A. tropidolepis</emphasis>
</taxonomicName>
in having more scale rows between the supraorbital semicircles (usually five or more in
<emphasis id="4DA0EAAA7F4D533BFD4CFBACFCB0FBC3" box="[740,888,1088,1112]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFD4CFBACFCBCFBC3" box="[740,884,1088,1112]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="benedikti">A. benedikti</taxonomicName>
,
</emphasis>
versus four or fewer in
<emphasis id="4DA0EAAA7F4D533BFB03FBACFEC0FBE7" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFB03FBACFABBFBC3" box="[1195,1395,1088,1112]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="magnaphallus">A. magnaphallus</taxonomicName>
,
<taxonomicName id="B8D44D3B7F4D533BFA2CFBACFECCFBE7" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pachypus">A. pachypus</taxonomicName>
,
</emphasis>
and usually four or fewer, rarely five, in
<taxonomicName id="B8D44D3B7F4D533BFCBAFB88FC76FBE7" box="[786,958,1124,1148]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="tropidolepis">
<emphasis id="4DA0EAAA7F4D533BFCBAFB88FC76FBE7" box="[786,958,1124,1148]" italics="true" pageId="11" pageNumber="12">A. tropidolepis</emphasis>
</taxonomicName>
), and more scales between supraorbital semicircles and interparietal plate (usually five or more, rarely four, in
<emphasis id="4DA0EAAA7F4D533BFC7CFB64FBACFB3B" box="[980,1124,1160,1184]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFC7CFB64FBA8FB3B" box="[980,1120,1160,1184]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="benedikti">A. benedikti</taxonomicName>
,
</emphasis>
versus four or fewer in
<emphasis id="4DA0EAAA7F4D533BFA2CFB64FE3CFB5F" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFA2CFB64FEF1FB5F" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="magnaphallus">A. magnaphallus</taxonomicName>
,
<taxonomicName id="B8D44D3B7F4D533BFEEEFB40FE38FB5F" box="[326,496,1196,1220]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="tropidolepis">A. tropidolepis</taxonomicName>
,
</emphasis>
and usually four or fewer, rarely five, in
<taxonomicName id="B8D44D3B7F4D533BFC7CFB40FB97FB5F" box="[980,1119,1196,1220]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F4D533BFC7CFB40FB97FB5F" box="[980,1119,1196,1220]" italics="true" pageId="11" pageNumber="12">A. pachypus</emphasis>
</taxonomicName>
). Furthermore,
<taxonomicName id="B8D44D3B7F4D533BFABCFB40FA54FB5F" box="[1300,1436,1196,1220]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F4D533BFABCFB40FA54FB5F" box="[1300,1436,1196,1220]" italics="true" pageId="11" pageNumber="12">A. benedikti</emphasis>
</taxonomicName>
differs from
<taxonomicName id="B8D44D3B7F4D533BFE8BFB3CFE28FB73" box="[291,480,1232,1256]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="magnaphallus">
<emphasis id="4DA0EAAA7F4D533BFE8BFB3CFE28FB73" box="[291,480,1232,1256]" italics="true" pageId="11" pageNumber="12">A. magnaphallus</emphasis>
</taxonomicName>
,
<emphasis id="4DA0EAAA7F4D533BFE45FB3CFDB5FB73" box="[493,637,1232,1256]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFE45FB3CFDB1FB73" box="[493,633,1232,1256]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pachypus">A. pachypus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="B8D44D3B7F4D533BFD1AFB3CFC90FB73" box="[690,856,1232,1256]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="tropidolepis">
<emphasis id="4DA0EAAA7F4D533BFD1AFB3CFC90FB73" box="[690,856,1232,1256]" italics="true" pageId="11" pageNumber="12">A. tropidolepis</emphasis>
</taxonomicName>
in having a very dark (almost blackish) gray tongue (versus a very light gray tongue in
<taxonomicName id="B8D44D3B7F4D533BFDB3FB18FD10FA97" box="[539,728,1268,1292]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="magnaphallus">
<emphasis id="4DA0EAAA7F4D533BFDB3FB18FD10FA97" box="[539,728,1268,1292]" italics="true" pageId="11" pageNumber="12">A. magnaphallus</emphasis>
</taxonomicName>
,
<emphasis id="4DA0EAAA7F4D533BFD4EFB18FCBDFA97" box="[742,885,1268,1292]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFD4EFB18FCBAFA97" box="[742,882,1268,1292]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pachypus">A. pachypus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="B8D44D3B7F4D533BFC03FB18FB98FA97" box="[939,1104,1268,1292]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="tropidolepis">
<emphasis id="4DA0EAAA7F4D533BFC03FB18FB98FA97" box="[939,1104,1268,1292]" italics="true" pageId="11" pageNumber="12">A. tropidolepis</emphasis>
</taxonomicName>
). At first sight, individuals of
<taxonomicName id="B8D44D3B7F4D533BFF3FFAF4FED6FAAB" box="[151,286,1304,1328]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F4D533BFF3FFAF4FED6FAAB" box="[151,286,1304,1328]" italics="true" pageId="11" pageNumber="12">A. benedikti</emphasis>
</taxonomicName>
might be confused with similarly-colored
<emphasis id="4DA0EAAA7F4D533BFCAAFAF4FCB5FAAB" box="[770,893,1304,1328]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="B8D44D3B7F4D533BFCAAFAF4FCB1FAAB" box="[770,889,1304,1328]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="humilis">A. humilis</taxonomicName>
.
</emphasis>
Yet, regardless of sex or age, individuals of the latter species have at least eight longitudinal rows of greatly enlarged, visibly keeled median dorsal scales that are larger than the ventrals, whereas
<taxonomicName id="B8D44D3B7F4D533BFDAEFA8CFD43FAE3" box="[518,651,1376,1400]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F4D533BFDAEFA8CFD43FAE3" box="[518,651,1376,1400]" italics="true" pageId="11" pageNumber="12">A. benedikti</emphasis>
</taxonomicName>
has a maximum of two slightly enlarged middorsal rows, the scales of which are smaller than the ventrals. Males of
<taxonomicName id="B8D44D3B7F4D533BFD06FA68FCEAFA07" box="[686,802,1412,1436]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="humilis">
<emphasis id="4DA0EAAA7F4D533BFD06FA68FCEAFA07" box="[686,802,1412,1436]" italics="true" pageId="11" pageNumber="12">A. humilis</emphasis>
</taxonomicName>
have a red dewlap with a complete (i.e. bordering the whole dewlap) yellow margin, whereas males of
<taxonomicName id="B8D44D3B7F4D533BFD14FA44FC8AFA5B" box="[700,834,1448,1472]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F4D533BFD14FA44FC8AFA5B" box="[700,834,1448,1472]" italics="true" pageId="11" pageNumber="12">A. benedikti</emphasis>
</taxonomicName>
exhibit yellow coloration only on the anterior portion of their otherwise solid red dewlap.
</paragraph>
</subSubSection>
<subSubSection id="37CE65337F4D5320FF6DFA03FEF5F86A" lastPageId="16" lastPageNumber="17" pageId="11" pageNumber="12" type="description">
<paragraph id="7F6B36B87F4D533CFF6DFA03FD96FD57" blockId="11.[151,1436,151,2012]" lastBlockId="12.[151,1437,151,2033]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="4DA0EAAA7F4D533BFF6DFA03FDDCF993" bold="true" box="[197,532,1519,1544]" pageId="11" pageNumber="12">
Description of the
<typeStatus id="A06F881A7F4D533BFE0EFA1CFDC7F993" box="[422,527,1520,1544]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male as indicated by everted hemipenes and presence of moderate-sized dewlap (
<figureCitation id="E7EF2A3D7F4D533BFF5FF9F8FEFFF9B7" box="[247,311,1556,1580]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1851]" captionTargetBox="[305,1272,193,1803]" captionTargetId="figure@3.[305,1272,193,1804]" captionTargetPageId="3" captionText="FIGURE 1. Dewlaps of anoles related to Anolis pachypus from Costa Rica and western Panama, representative of the five OTUs (width of tweezers 3 mm): (A) OTU A (SMF 89729, Sendero La Cascada, Bajo Mono, Chiriquí, Panama); (B) OTU B (GK 2297, road from San José de la Montaña to Birrí, Heredia, Costa Rica); (C) OTU C (GK 2355, road from San Isidro de El General to Cerro de la Muerte, San José, Costa Rica); (D) OTU C (SL 439, Jurutungo, Chiriquí, Panama); (E) OTU C (SL 524, Jurutungo, Chiriquí, Panama); (F) OTU D (SMF 89507, Jurutungo, Chiriquí, Panama); (G) OTU D (SMF 89506, Jurutungo, Chiriquí, Panama); (H) OTU D (SMF 90149, N slope Cerro Pando, Bocas del Toro, Panama); (I) OTU E (SMF 89519, La Nevera, Comarca Ngöbe-Buglé, Panama); (J) OTU E (SL 510, La Nevera, Comarca Ngöbe-Buglé, Panama)." httpUri="https://zenodo.org/record/200704/files/figure.png" pageId="11" pageNumber="12">Fig. 1</figureCitation>
H); snout-vent length
<quantity id="B82C9B5D7F4D533BFD9BF9F8FD52F9B7" box="[563,666,1556,1580]" metricMagnitude="-2" metricUnit="m" metricValue="4.86" pageId="11" pageNumber="12" unit="mm" value="48.6">48.6 mm</quantity>
; tail length
<quantity id="B82C9B5D7F4D533BFCB7F9F8FC4DF9B7" box="[799,901,1556,1580]" metricMagnitude="-2" metricUnit="m" metricValue="8.17" pageId="11" pageNumber="12" unit="mm" value="81.7">81.7 mm</quantity>
, tail complete; tail slightly compressed in cross section, tail height
<quantity id="B82C9B5D7F4D533BFEC3F9D4FE0CF9CB" box="[363,452,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="12" unit="mm" value="2.2">2.2 mm</quantity>
, tail width
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; axilla to groin distance
<quantity id="B82C9B5D7F4D533BFC01F9D4FBD8F9CB" box="[937,1040,1592,1616]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="11" pageNumber="12" unit="mm" value="18.7">18.7 mm</quantity>
; head length
<quantity id="B82C9B5D7F4D533BFB0BF9D4FAC0F9CB" box="[1187,1288,1592,1616]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="11" pageNumber="12" unit="mm" value="11.9">11.9 mm</quantity>
, head length/ snout-vent length ratio 0.24; snout length
<quantity id="B82C9B5D7F4D533BFDC3F9B0FD0BF9EF" box="[619,707,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="11" pageNumber="12" unit="mm" value="5.3">5.3 mm</quantity>
; head width
<quantity id="B82C9B5D7F4D533BFCF9F9B0FC61F9EF" box="[849,937,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="11" pageNumber="12" unit="mm" value="8.1">8.1 mm</quantity>
; longest toe of adpressed hind limb reaching to snout; shank length
<quantity id="B82C9B5D7F4D533BFE32F993FDCAF90C" box="[410,514,1663,1687]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="11" pageNumber="12" unit="mm" value="12.6">12.6 mm</quantity>
, shank length/head length ratio 1.1; longest finger of extended forelimb reaching about
<quantity id="B82C9B5D7F4D533BFF73F948FEF9F927" box="[219,305,1700,1724]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="11" pageNumber="12" unit="mm" value="4.2">4.2 mm</quantity>
beyond tip of snout; longest finger of adpressed forelimb reaching to anterior insertion of hind limbs; scales on snout keeled; 5 postrostrals; 5 scales between nasals; scales in distinct prefrontal depression distinctly keeled; supraorbital semicircles differentiated, separated by a minimum of 7 scales; supraorbital disc composed of 14 slightly enlarged keeled scales; 3 elongated superciliaries; about 5 rows of small keeled scales extending between enlarged supraorbitals and superciliaries; interparietal plate slightly enlarged, 0.6 x
<quantity id="B82C9B5D7F4D533BFB66F8D8FAE1F8D7" box="[1230,1321,1844,1868]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="12" unit="mm" value="0.3">0.3 mm</quantity>
(length x width), surrounded by small scales; 10 scales present between interparietal plate and supraorbital semicircles; canthal ridge distinct, composed of 5 large (posterior) and 3 small (anterior) canthal scales; 15 scales present between second canthals; 17 scales present between posterior canthals; 93/95 keeled loreal scales in a maximum of 12 horizontal rows; subocular scales granular, subocular row ill-defined; 8 supralabials to level below center of eye; ear opening 0.8 x
<quantity id="B82C9B5D7F4A533CFE0EFF7BFE37FF34" box="[422,511,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="mm" value="1.5">1.5 mm</quantity>
(length x height); mental distinctly wider than long, almost completely divided medially, bordered posteriorly by 7 postmentals; 8 infralabials to level below center of eye; sublabials undifferentiated; slightly keeled granular scales present on chin and throat; dewlap moderate-sized, extending well onto chest, anterior insertion at level of anterior border of orbit, posterior insertion at level of axilla, with about 10 somewhat irregular, partially interrupted gorgetal-sternal rows of 2026 scales per row; dorsum of body with keeled scales, 2 medial rows of slightly enlarged scales, largest dorsal scales subimbricate, about 0.40 x
<quantity id="B82C9B5D7F4A533CFA9FFEA0FA54FEFF" box="[1335,1436,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="13" unit="mm" value="0.35">0.35 mm</quantity>
(length x width); about 66 medial dorsal scales in one head length; about 112 medial dorsal scales between axilla and groin; lateral scales pointed, granular and homogeneous, average size
<quantity id="B82C9B5D7F4A533CFBBAFE78FBB4FE37" box="[1042,1148,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="12" pageNumber="13" unit="mm" value="0.18">0.18 mm</quantity>
in diameter; ventrals at midbody weakly keeled, slightly mucronate, and subimbricate, about 0.30 x
<quantity id="B82C9B5D7F4A533CFBAAFE5BFBA0FE54" box="[1026,1128,439,463]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="13" unit="mm" value="0.4">0.40 mm</quantity>
(length x width); about 42 ventral scales in one head length; about 58 ventral scales between axilla and groin; about 162 scales around midbody; caudal scales strongly keeled, without whorls of enlarged scales; no enlarged postcloacal scales; tubelike axillary pocket not developed; scales on dorsal surface of forelimb strongly keeled, mucronate and imbricate, about 0.30 x
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(length x width); digital pads dilated, dilated pad about 2 times width of non-dilated scales on distal phalanx; distal phalanx narrower than and raised from dilated pad; 23/22 lamellae under phalanges IIIV of fourth toe; 9/10 scales under distal phalanx of fourth toe; 15 lamellae under phalanges IIIV of fourth finger; 9 scales under distal phalanx of fourth toe.
</paragraph>
<paragraph id="7F6B36B87F4A533CFF6DFD3BFC11FCAC" blockId="12.[151,1437,151,2033]" pageId="12" pageNumber="13">The completely everted hemipenis is a stout bilobate organ; sulcus spermaticus bordered by well-developed sulcal lips and opening at base of apex into two broad concave areas, one on each lobe; a large asulcate ridge-like processus present; lobes strongly calyculate, truncus with transverse folds.</paragraph>
<paragraph id="7F6B36B87F4A533CFF6DFCA8FD82FC5C" blockId="12.[151,1437,151,2033]" pageId="12" pageNumber="13">
Coloration in life. The dewlap (
<figureCitation id="E7EF2A3D7F4A533CFD9FFCA8FDB1FCC7" box="[567,633,836,860]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1851]" captionTargetBox="[305,1272,193,1803]" captionTargetId="figure@3.[305,1272,193,1804]" captionTargetPageId="3" captionText="FIGURE 1. Dewlaps of anoles related to Anolis pachypus from Costa Rica and western Panama, representative of the five OTUs (width of tweezers 3 mm): (A) OTU A (SMF 89729, Sendero La Cascada, Bajo Mono, Chiriquí, Panama); (B) OTU B (GK 2297, road from San José de la Montaña to Birrí, Heredia, Costa Rica); (C) OTU C (GK 2355, road from San Isidro de El General to Cerro de la Muerte, San José, Costa Rica); (D) OTU C (SL 439, Jurutungo, Chiriquí, Panama); (E) OTU C (SL 524, Jurutungo, Chiriquí, Panama); (F) OTU D (SMF 89507, Jurutungo, Chiriquí, Panama); (G) OTU D (SMF 89506, Jurutungo, Chiriquí, Panama); (H) OTU D (SMF 90149, N slope Cerro Pando, Bocas del Toro, Panama); (I) OTU E (SMF 89519, La Nevera, Comarca Ngöbe-Buglé, Panama); (J) OTU E (SL 510, La Nevera, Comarca Ngöbe-Buglé, Panama)." httpUri="https://zenodo.org/record/200704/files/figure.png" pageId="12" pageNumber="13">Fig. 1</figureCitation>
H) was recorded as follows: Flame Scarlet (15) with a suggestion of Chrome Orange (16) on posterior three fourths, Orange Yellow (18) on anterior fourth and anterior margin; dewlap scales Sepia (119) and dirty white. Otherwise, no detailed notes of the
<typeStatus id="A06F881A7F4A533CFC1FFC60FBD1FC3F" box="[951,1049,908,932]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
were taken; see
<figureCitation id="E7EF2A3D7F4A533CFB7FFC60FA88FC3F" box="[1239,1344,908,932]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1923]" captionTargetBox="[216,1349,193,1860]" captionTargetId="figure@8.[216,1350,193,1861]" captionTargetPageId="8" captionText="FIGURE 5. Type specimens of Anolis benedikti in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (C E) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (F O) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female." httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="12" pageNumber="13">Figures 5</figureCitation>
A and F for photographs of the
<typeStatus id="A06F881A7F4A533CFE3FFC43FE31FC5C" box="[407,505,943,967]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
in life.
</paragraph>
<paragraph id="7F6B36B87F4A533CFF6DFC38FC3BFB94" blockId="12.[151,1437,151,2033]" pageId="12" pageNumber="13">Coloration after 18 months of preservation in 70% ethanol is similar to that in life, apart from that all reddish shades have faded, especially on the dewlap, which now appears dirty white.</paragraph>
<paragraph id="7F6B36B87F4A533CFF6DFBF7FDB9FA97" blockId="12.[151,1437,151,2033]" pageId="12" pageNumber="13">
<emphasis id="4DA0EAAA7F4A533CFF6DFBF7FEF1FBAF" bold="true" box="[197,313,1051,1076]" pageId="12" pageNumber="13">Variation.</emphasis>
The
<typeStatus id="A06F881A7F4A533CFEDEFBF1FE29FBAE" box="[374,481,1053,1077]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
and referred specimens agree well with the
<typeStatus id="A06F881A7F4A533CFC78FBF0FBFAFBAF" box="[976,1074,1052,1076]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
in terms of general morphology and pholidosis (see
<tableCitation id="325603037F4A533CFEDBFBACFE0DFBC3" box="[371,453,1088,1112]" captionStart="TABLE 1" captionStartId="13.[151,235,396,417]" captionTargetBox="[151,1436,537,1961]" captionTargetPageId="13" captionText="TABLE 1. Selected measurements, proportions and scale characters of Anolis benedikti, A. magnaphallus, A. pseudopachypus, A. pachypus, and A. tropidolepis. Range is followed by mean value and one standard deviation in parentheses. For abbreviations see text. Based on specimens with data on dewlap coloration plus additional specimens (males and females) for which the respective exact geographic origin and morphological key characters allowed for a precise species assignment." httpUri="http://table.plazi.org/id/2BAB66307F4B533DFF3FFE60FAAAFE65" pageId="12" pageNumber="13" tableUuid="2BAB66307F4B533DFF3FFE60FAAAFE65">Table 1</tableCitation>
). Four of the 32 examined specimens (SMF 89507, 91505; SL 686, 695) have only four scales between the supraorbital semicircles. Two specimens (SMF 89507, 91505) have only four scales between the supraorbital semicircles and the interparietal plate. Among the
<specimenCount id="69D2FD317F4A533CFC26FB64FC23FB3B" box="[910,1003,1160,1184]" pageId="12" pageNumber="13" type="adult">17 adult</specimenCount>
males included in statistical analyses, the variability in the nasal region (
<figureCitation id="E7EF2A3D7F4A533CFD80FB40FDA6FB5F" box="[552,622,1196,1220]" captionStart="FIGURE 7" captionStartId="10.[151,250,1961,1984]" captionTargetBox="[286,1309,205,1926]" captionTargetId="figure@10.[247,1339,193,1940]" captionTargetPageId="10" captionText="FIGURE 7. Designation of nasal scalation types. (A) Type A (SMF 90149); (B) Type B (SMF 92134); (C) Type C (SL 678); (D) Type D (SMF 85264); (E) Type E (SMF 89506); (F) Type F (85272). Scale bars equal 1 mm. See Text for details." httpUri="https://zenodo.org/record/200711/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
) was recorded as follows:
<typeStatus id="A06F881A7F4A533CFC00FB41FC17FB5E" box="[936,991,1197,1221]" pageId="12" pageNumber="13">Type</typeStatus>
A one specimen (5.9 %);
<typeStatus id="A06F881A7F4A533CFABCFB41FA84FB5E" box="[1300,1356,1197,1221]" pageId="12" pageNumber="13">Type</typeStatus>
B two specimens (11.8 %);
<typeStatus id="A06F881A7F4A533CFE2FFB3DFE77FB72" box="[391,447,1233,1257]" pageId="12" pageNumber="13">Type</typeStatus>
C four specimens (23.5 %);
<typeStatus id="A06F881A7F4A533CFCA2FB3DFC8AFB72" box="[778,834,1233,1257]" pageId="12" pageNumber="13">Type</typeStatus>
D five specimens (29.4 %);
<typeStatus id="A06F881A7F4A533CFB25FB3DFB0DFB72" box="[1165,1221,1233,1257]" pageId="12" pageNumber="13">Type</typeStatus>
E three specimens (17.6 %);
<typeStatus id="A06F881A7F4A533CFEA0FB19FEF7FA96" box="[264,319,1269,1293]" pageId="12" pageNumber="13">Type</typeStatus>
F two specimens (11.8 %).
</paragraph>
<paragraph id="7F6B36B87F4A533CFF6DFAF4FBD4F9CB" blockId="12.[151,1437,151,2033]" pageId="12" pageNumber="13">
Similar to what we observed in the other four members of the
<taxonomicName id="B8D44D3B7F4A533CFC26FAF4FB80FAAB" box="[910,1096,1304,1328]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F4A533CFC26FAF4FB80FAAB" box="[910,1096,1304,1328]" italics="true" pageId="12" pageNumber="13">Anolis pachypus</emphasis>
</taxonomicName>
complex, the coloration of
<taxonomicName id="B8D44D3B7F4A533CFA2CFAF4FF36FACF" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F4A533CFA2CFAF4FF36FACF" italics="true" pageId="12" pageNumber="13">A. benedikti</emphasis>
</taxonomicName>
is rather variable among individuals, mostly concerning the extent and shape of color patterns, i.e., stripes, blotches, and bands on dorsal and lateral surfaces (compare
<figureCitation id="E7EF2A3D7F4A533CFC25FA8CFC10FAE3" box="[909,984,1376,1400]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1923]" captionTargetBox="[216,1349,193,1860]" captionTargetId="figure@8.[216,1350,193,1861]" captionTargetPageId="8" captionText="FIGURE 5. Type specimens of Anolis benedikti in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (C E) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (F O) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female." httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="12" pageNumber="13">Figs. 5</figureCitation>
F
<collectingCountry id="07C376287F4A533CFC51FA8CFBC6FAE3" box="[1017,1038,1376,1400]" name="Iceland" pageId="12" pageNumber="13">O</collectingCountry>
). Common patterns include a broad middorsal band which is often interspersed with blotches or chevrons, flanked by lateral markings, or disintegrates into diamond-shaped markings posteriorly (
<figureCitation id="E7EF2A3D7F4A533CFD2BFA44FD19FA5B" box="[643,721,1448,1472]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1923]" captionTargetBox="[216,1349,193,1860]" captionTargetId="figure@8.[216,1350,193,1861]" captionTargetPageId="8" captionText="FIGURE 5. Type specimens of Anolis benedikti in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (C E) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (F O) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female." httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="12" pageNumber="13">Figs. 5</figureCitation>
FJ); a narrow light middorsal stripe, often bordered by narrow dark stripes (Figs. KL); and series of triangular markings pointing laterally from middorsum (with or without narrow medial light stripe) which are either offset to form a zig-zag pattern or arranged symmetrically to form a series of diamonds (
<figureCitation id="E7EF2A3D7F4A533CFED6F9F8FE04F9B7" box="[382,460,1556,1580]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1923]" captionTargetBox="[216,1349,193,1860]" captionTargetId="figure@8.[216,1350,193,1861]" captionTargetPageId="8" captionText="FIGURE 5. Type specimens of Anolis benedikti in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (C E) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (F O) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female." httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="12" pageNumber="13">Figs. 5</figureCitation>
M
<collectingCountry id="07C376287F4A533CFE5FF9F8FDC4F9B7" box="[503,524,1556,1580]" name="Iceland" pageId="12" pageNumber="13">O</collectingCountry>
). Most specimens exhibit contrasting markings on dorsal and lateral surfaces of the head, with postocular stripes and an interorbital bar being the most common.
</paragraph>
<paragraph id="7F6B36B87F4A533CFF6DF9B0FBFAF965" blockId="12.[151,1437,151,2033]" pageId="12" pageNumber="13">
Furthermore, these anoles are capable of metachrosis and, therefore, the coloration of a given individual is subject to physiological change according to time or situation. Most individuals encountered at night exhibited light tonalities, but could turn very dark during the day or when handled (
<figureCitation id="E7EF2A3D7F4A533CFC72F94DFBE0F922" box="[986,1064,1697,1721]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1923]" captionTargetBox="[216,1349,193,1860]" captionTargetId="figure@8.[216,1350,193,1861]" captionTargetPageId="8" captionText="FIGURE 5. Type specimens of Anolis benedikti in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (C E) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (F O) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female." httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="12" pageNumber="13">Figs. 5</figureCitation>
KL). Individuals spotted during the day all appeared dark. In the course of metachrosis, the conspicuousness of individual color patterns may vary considerably (as in the postocular stripes between the two females in
<figureCitation id="E7EF2A3D7F4A533CFC08F90AFC23F965" box="[928,1003,1766,1790]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1923]" captionTargetBox="[216,1349,193,1860]" captionTargetId="figure@8.[216,1350,193,1861]" captionTargetPageId="8" captionText="FIGURE 5. Type specimens of Anolis benedikti in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (C E) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (F O) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female." httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="12" pageNumber="13">Figs. 5</figureCitation>
CD).
</paragraph>
<paragraph id="7F6B36B87F4A533DFF6DF8E5FD1DFEC6" blockId="12.[151,1437,151,2033]" lastBlockId="13.[151,1436,151,349]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
The coloration of the male
<typeStatus id="A06F881A7F4A533CFDA9F8E6FDA9F8B9" box="[513,609,1802,1826]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
SMF 89506 (
<figureCitation id="E7EF2A3D7F4A533CFCACF8E5FC98F8BA" box="[772,848,1801,1825]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1851]" captionTargetBox="[305,1272,193,1803]" captionTargetId="figure@3.[305,1272,193,1804]" captionTargetPageId="3" captionText="FIGURE 1. Dewlaps of anoles related to Anolis pachypus from Costa Rica and western Panama, representative of the five OTUs (width of tweezers 3 mm): (A) OTU A (SMF 89729, Sendero La Cascada, Bajo Mono, Chiriquí, Panama); (B) OTU B (GK 2297, road from San José de la Montaña to Birrí, Heredia, Costa Rica); (C) OTU C (GK 2355, road from San Isidro de El General to Cerro de la Muerte, San José, Costa Rica); (D) OTU C (SL 439, Jurutungo, Chiriquí, Panama); (E) OTU C (SL 524, Jurutungo, Chiriquí, Panama); (F) OTU D (SMF 89507, Jurutungo, Chiriquí, Panama); (G) OTU D (SMF 89506, Jurutungo, Chiriquí, Panama); (H) OTU D (SMF 90149, N slope Cerro Pando, Bocas del Toro, Panama); (I) OTU E (SMF 89519, La Nevera, Comarca Ngöbe-Buglé, Panama); (J) OTU E (SL 510, La Nevera, Comarca Ngöbe-Buglé, Panama)." httpUri="https://zenodo.org/record/200704/files/figure.png" pageId="12" pageNumber="13">Figs. 1</figureCitation>
G, 5B and G) was recorded as follows: Dorsal and lateral ground color Antique Brown (37) suffused with Light Drab (119C) and Tawny (38), laterally grading into Olive-Gray (42) mottled with Burnt Umber (22); a Mars Brown (223A) postorbital stripe mottled with Sepia (219) continues to form a dorsolateral band, which disintegrates into a series of blotches around midbody; a few Sepia (119) middorsal blotches present; dorsal and lateral surfaces of head Light Drab (119C); occipital region Mars Brown (223A) mottled with Sepia (219); a Mars Brown (223A) interorbital bar and a preorbital stripe bordered by Sepia (219); a dirty white with a suggestion of Yellow-Green (58) lateral postorbital stripe extending to shoulder; dorsal and lateral surfaces of limbs and tail Antique Brown (37), with irregular Sepia (219), Tawny (38), and Olive- Gray (42) transverse stripes; ventral surface of body and head Pearl Gray (81) suffused with Cinnamon-Rufous (40); ventral surface of tail with a Cinnamon-Rufous (40) midventral stripe; iris Brick Red (132A); tongue Plumbeous (78) with a suggestion of Indigo (73); gular region Flame Scarlet (15); dewlap Flame Scarlet (15) with a small Pearl Gray (81) region at posterior insertion and an Orange Yellow (18) anterior margin; dewlap scales dirty white bordered by Sepia (119) or Sepia (219).
</paragraph>
<caption id="2BAB66307F4B533DFF3FFE60FAAAFE65" ID-Table-UUID="2BAB66307F4B533DFF3FFE60FAAAFE65" httpUri="http://table.plazi.org/id/2BAB66307F4B533DFF3FFE60FAAAFE65" pageId="13" pageNumber="14" targetBox="[151,1436,537,1961]" targetIsTable="true" targetPageId="13">
<paragraph id="7F6B36B87F4B533DFF3FFE60FAAAFE65" blockId="13.[151,1436,395,510]" pageId="13" pageNumber="14">
<emphasis id="4DA0EAAA7F4B533DFF3FFE60FECCFE3A" bold="true" box="[151,260,396,417]" pageId="13" pageNumber="14">TABLE 1.</emphasis>
Selected measurements, proportions and scale characters of
<emphasis id="4DA0EAAA7F4B533DFCC2FE60FEE9FE5B" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="B8D44D3B7F4B533DFCC2FE60FBD8FE39" box="[874,1040,396,418]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="benedikti">Anolis benedikti</taxonomicName>
,
<taxonomicName id="B8D44D3B7F4B533DFBB2FE60FB00FE39" box="[1050,1224,396,418]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="magnaphallus">A. magnaphallus</taxonomicName>
,
<taxonomicName id="B8D44D3B7F4B533DFB7AFE60FA50FE39" box="[1234,1432,396,418]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="pseudopachypus">A. pseudopachypus</taxonomicName>
,
<taxonomicName id="B8D44D3B7F4B533DFF3FFE47FED5FE5B" box="[151,285,426,448]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="pachypus">A. pachypus</taxonomicName>
,
</emphasis>
and
<emphasis id="4DA0EAAA7F4B533DFEC9FE47FDCDFE5B" box="[353,517,426,448]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="B8D44D3B7F4B533DFEC9FE47FDC9FE5B" box="[353,513,426,448]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="tropidolepis">A. tropidolepis</taxonomicName>
.
</emphasis>
Range is followed by mean value and one standard deviation in parentheses. For abbreviations see text. Based on specimens with data on dewlap coloration plus additional specimens (males and females) for which the respective exact geographic origin and morphological key characters allowed for a precise species assignment.
</paragraph>
</caption>
<paragraph id="7F6B36B87F4B533DFE71FDF6FAA9F832" pageId="13" pageNumber="14">
<table id="0DD4C4187F4BACCFFF3FFDF5FA54F832" box="[151,1436,537,1961]" gridcols="6" gridrows="20" pageId="13" pageNumber="14">
<tr id="C1E434FA7F4BACCFFF3FFDF5FA54FDD5" box="[151,1436,537,590]" gridrow="0" pageId="13" pageNumber="14" rowspan-0="1">
<th id="82355D867F4BACCFFE71FDF5FDACFDD5" box="[473,612,537,590]" gridcol="1" gridrow="0" pageId="13" pageNumber="14">
<taxonomicName id="B8D44D3B7F4B533DFE71FDF6FD9DFDB4" box="[473,597,537,559]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F4B533DFE71FDF6FD9DFDB4" box="[473,597,537,559]" italics="true" pageId="13" pageNumber="14">A. pachypus</emphasis>
</taxonomicName>
3 11, Ƥ 5
</th>
<th id="82355D867F4BACCFFD27FDF5FC9BFDD5" box="[655,851,537,590]" gridcol="2" gridrow="0" pageId="13" pageNumber="14">
<taxonomicName id="B8D44D3B7F4B533DFD27FDF6FC9BFDB4" box="[655,851,537,559]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="pseudopachypus">
<emphasis id="4DA0EAAA7F4B533DFD27FDF6FC9BFDB4" box="[655,851,537,559]" italics="true" pageId="13" pageNumber="14">A. pseudopachypus</emphasis>
</taxonomicName>
3 16, Ƥ 5
</th>
<th id="82355D867F4BACCFFCC4FDF5FBDFFDD5" box="[876,1047,537,590]" gridcol="3" gridrow="0" pageId="13" pageNumber="14">
<taxonomicName id="B8D44D3B7F4B533DFCC4FDF6FBDFFDB4" box="[876,1047,537,559]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="magnaphallus">
<emphasis id="4DA0EAAA7F4B533DFCC4FDF6FBDFFDB4" box="[876,1047,537,559]" italics="true" pageId="13" pageNumber="14">A. magnaphallus</emphasis>
</taxonomicName>
3 9, Ƥ 6
</th>
<th id="82355D867F4BACCFFB90FDF5FB06FDD5" box="[1080,1230,537,590]" gridcol="4" gridrow="0" pageId="13" pageNumber="14">
<taxonomicName id="B8D44D3B7F4B533DFB90FDF6FB06FDB4" box="[1080,1230,537,559]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="tropidolepis">
<emphasis id="4DA0EAAA7F4B533DFB90FDF6FB06FDB4" box="[1080,1230,537,559]" italics="true" pageId="13" pageNumber="14">A. tropidolepis</emphasis>
</taxonomicName>
3 10, Ƥ 0
</th>
<th id="82355D867F4BACCFFB5EFDF5FA54FDD5" box="[1270,1436,537,590]" gridcol="5" gridrow="0" pageId="13" pageNumber="14">
<taxonomicName id="B8D44D3B7F4B533DFB5EFDF6FAB8FDB4" box="[1270,1392,537,559]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F4B533DFB5EFDF6FAB8FDB4" box="[1270,1392,537,559]" italics="true" pageId="13" pageNumber="14">A. benedikti</emphasis>
</taxonomicName>
3 20, Ƥ 13
</th>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFD8DFA54FD0D" box="[151,1436,609,662]" gridrow="1" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FFD8DFE0BFD0D" box="[151,451,609,662]" gridcol="0" gridrow="1" pageId="13" pageNumber="14">SVL 3</th>
<td id="82355D867F4BACCFFE71FD8DFDACFD0D" box="[473,612,609,662]" gridcol="1" gridrow="1" pageId="13" pageNumber="14">38.650.7 (45.5 ± 3.9)</td>
<td id="82355D867F4BACCFFD27FD8DFC9BFD0D" box="[655,851,609,662]" gridcol="2" gridrow="1" pageId="13" pageNumber="14">30.045.5 (40.7 ± 4.6)</td>
<td id="82355D867F4BACCFFCC4FD8DFBDFFD0D" box="[876,1047,609,662]" gridcol="3" gridrow="1" pageId="13" pageNumber="14">31.454.1 (47.5 ± 7.1)</td>
<td id="82355D867F4BACCFFB90FD8DFB06FD0D" box="[1080,1230,609,662]" gridcol="4" gridrow="1" pageId="13" pageNumber="14">41.452.3 (47.4 ± 3.5)</td>
<td id="82355D867F4BACCFFB5EFD8DFA54FD0D" box="[1270,1436,609,662]" gridcol="5" gridrow="1" pageId="13" pageNumber="14">34.648.6 (41.6 ± 3.5)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFD45FA54FD45" box="[151,1436,681,734]" gridrow="2" pageId="13" pageNumber="14" rowspan-4="1">
<th id="82355D867F4BACCFFF3FFD45FE0BFD45" box="[151,451,681,734]" gridcol="0" gridrow="2" pageId="13" pageNumber="14">Ƥ</th>
<td id="82355D867F4BACCFFE71FD45FDACFD45" box="[473,612,681,734]" gridcol="1" gridrow="2" pageId="13" pageNumber="14">47.651.1 (49.0 ± 1.3)</td>
<td id="82355D867F4BACCFFD27FD45FC9BFD45" box="[655,851,681,734]" gridcol="2" gridrow="2" pageId="13" pageNumber="14">39.447.3 (45.0 ± 3.7)</td>
<td id="82355D867F4BACCFFCC4FD45FBDFFD45" box="[876,1047,681,734]" gridcol="3" gridrow="2" pageId="13" pageNumber="14">46.558.9 (52.6 ± 5.4)</td>
<td id="82355D867F4BACCFFB5EFD45FA54FD45" box="[1270,1436,681,734]" gridcol="5" gridrow="2" pageId="13" pageNumber="14">34.448.1 (42.5 ± 4.8)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFD1DFA54FCBD" box="[151,1436,753,806]" gridrow="3" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FFD1DFE0BFCBD" box="[151,451,753,806]" gridcol="0" gridrow="3" pageId="13" pageNumber="14">Tail length / SVL 3</th>
<td id="82355D867F4BACCFFE71FD1DFDACFCBD" box="[473,612,753,806]" gridcol="1" gridrow="3" pageId="13" pageNumber="14">1.561.94 (1.81 ± 0.14)</td>
<td id="82355D867F4BACCFFD27FD1DFC9BFCBD" box="[655,851,753,806]" gridcol="2" gridrow="3" pageId="13" pageNumber="14">1.612.06 (1.93 ± 0.12)</td>
<td id="82355D867F4BACCFFCC4FD1DFBDFFCBD" box="[876,1047,753,806]" gridcol="3" gridrow="3" pageId="13" pageNumber="14">1.041.71 (1.58 ± 0.22)</td>
<td id="82355D867F4BACCFFB90FD1DFB06FCBD" box="[1080,1230,753,806]" gridcol="4" gridrow="3" pageId="13" pageNumber="14">1.722.18 (1.99 ± 0.18)</td>
<td id="82355D867F4BACCFFB5EFD1DFA54FCBD" box="[1270,1436,753,806]" gridcol="5" gridrow="3" pageId="13" pageNumber="14">1.222.16 (1.76 ± 0.21)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFCD5FA54FCF5" box="[151,1436,825,878]" gridrow="4" pageId="13" pageNumber="14" rowspan-4="1">
<th id="82355D867F4BACCFFF3FFCD5FE0BFCF5" box="[151,451,825,878]" gridcol="0" gridrow="4" pageId="13" pageNumber="14">Ƥ</th>
<td id="82355D867F4BACCFFE71FCD5FDACFCF5" box="[473,612,825,878]" gridcol="1" gridrow="4" pageId="13" pageNumber="14">1.641.72 (1.69 ± 0.04)</td>
<td id="82355D867F4BACCFFD27FCD5FC9BFCF5" box="[655,851,825,878]" gridcol="2" gridrow="4" pageId="13" pageNumber="14">1.701.78 (1.74 ± 0.04)</td>
<td id="82355D867F4BACCFFCC4FCD5FBDFFCF5" box="[876,1047,825,878]" gridcol="3" gridrow="4" pageId="13" pageNumber="14">1.381.57 (1.48 ± 0.08)</td>
<td id="82355D867F4BACCFFB5EFCD5FA54FCF5" box="[1270,1436,825,878]" gridcol="5" gridrow="4" pageId="13" pageNumber="14">1.421.90 (1.67 ± 0.12)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFC6DFA54FC2D" box="[151,1436,897,950]" gridrow="5" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FFC6DFE0BFC2D" box="[151,451,897,950]" gridcol="0" gridrow="5" pageId="13" pageNumber="14">Tail diameter vertical / 3 horizontal</th>
<td id="82355D867F4BACCFFE71FC6DFDACFC2D" box="[473,612,897,950]" gridcol="1" gridrow="5" pageId="13" pageNumber="14">1.111.31 (1.25 ± 0.07)</td>
<td id="82355D867F4BACCFFD27FC6DFC9BFC2D" box="[655,851,897,950]" gridcol="2" gridrow="5" pageId="13" pageNumber="14">1.061.29 (1.17 ± 0.07)</td>
<td id="82355D867F4BACCFFCC4FC6DFBDFFC2D" box="[876,1047,897,950]" gridcol="3" gridrow="5" pageId="13" pageNumber="14">1.081.45 (1.21 ± 0.10)</td>
<td id="82355D867F4BACCFFB90FC6DFB06FC2D" box="[1080,1230,897,950]" gridcol="4" gridrow="5" pageId="13" pageNumber="14">1.051.31 (1.19 ± 0.08)</td>
<td id="82355D867F4BACCFFB5EFC6DFA54FC2D" box="[1270,1436,897,950]" gridcol="5" gridrow="5" pageId="13" pageNumber="14">1.001.43 (1.19± 0.10)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFC25FA54FC65" box="[151,1436,969,1022]" gridrow="6" pageId="13" pageNumber="14" rowspan-4="1">
<th id="82355D867F4BACCFFF3FFC25FE0BFC65" box="[151,451,969,1022]" gridcol="0" gridrow="6" pageId="13" pageNumber="14">Ƥ</th>
<td id="82355D867F4BACCFFE71FC25FDACFC65" box="[473,612,969,1022]" gridcol="1" gridrow="6" pageId="13" pageNumber="14">1.131.24 (1.18 ± 0.05)</td>
<td id="82355D867F4BACCFFD27FC25FC9BFC65" box="[655,851,969,1022]" gridcol="2" gridrow="6" pageId="13" pageNumber="14">1.131.36 (1.21 ± 0.12)</td>
<td id="82355D867F4BACCFFCC4FC25FBDFFC65" box="[876,1047,969,1022]" gridcol="3" gridrow="6" pageId="13" pageNumber="14">1.101.27 (1.18 ± 0.07)</td>
<td id="82355D867F4BACCFFB5EFC25FA54FC65" box="[1270,1436,969,1022]" gridcol="5" gridrow="6" pageId="13" pageNumber="14">1.121.27 (1.19 ± 0.06)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFBFDFA54FBDD" box="[151,1436,1041,1094]" gridrow="7" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FFBFDFE0BFBDD" box="[151,451,1041,1094]" gridcol="0" gridrow="7" pageId="13" pageNumber="14">HL / SVL 3</th>
<td id="82355D867F4BACCFFE71FBFDFDACFBDD" box="[473,612,1041,1094]" gridcol="1" gridrow="7" pageId="13" pageNumber="14">0.240.28 (0.26 ± 0.01)</td>
<td id="82355D867F4BACCFFD27FBFDFC9BFBDD" box="[655,851,1041,1094]" gridcol="2" gridrow="7" pageId="13" pageNumber="14">0.240.29 (0.26 ± 0.01)</td>
<td id="82355D867F4BACCFFCC4FBFDFBDFFBDD" box="[876,1047,1041,1094]" gridcol="3" gridrow="7" pageId="13" pageNumber="14">0.240.28 (0.26 ± 0.01)</td>
<td id="82355D867F4BACCFFB90FBFDFB06FBDD" box="[1080,1230,1041,1094]" gridcol="4" gridrow="7" pageId="13" pageNumber="14">0.250.26 (0.26 ± 0.01)</td>
<td id="82355D867F4BACCFFB5EFBFDFA54FBDD" box="[1270,1436,1041,1094]" gridcol="5" gridrow="7" pageId="13" pageNumber="14">0.240.31 (0.27 ± 0.02)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFBB5FA54FB15" box="[151,1436,1113,1166]" gridrow="8" pageId="13" pageNumber="14" rowspan-4="1">
<th id="82355D867F4BACCFFF3FFBB5FE0BFB15" box="[151,451,1113,1166]" gridcol="0" gridrow="8" pageId="13" pageNumber="14">Ƥ</th>
<td id="82355D867F4BACCFFE71FBB5FDACFB15" box="[473,612,1113,1166]" gridcol="1" gridrow="8" pageId="13" pageNumber="14">0.250.27 (0.26 ± 0.01)</td>
<td id="82355D867F4BACCFFD27FBB5FC9BFB15" box="[655,851,1113,1166]" gridcol="2" gridrow="8" pageId="13" pageNumber="14">0.240.27 (0.25 ± 0.01)</td>
<td id="82355D867F4BACCFFCC4FBB5FBDFFB15" box="[876,1047,1113,1166]" gridcol="3" gridrow="8" pageId="13" pageNumber="14">0.25 0.26 (0.25 ± 0.01)</td>
<td id="82355D867F4BACCFFB5EFBB5FA54FB15" box="[1270,1436,1113,1166]" gridcol="5" gridrow="8" pageId="13" pageNumber="14">0.25 0.29 (0.27 ± 0.02)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFB4DFA54FB4D" box="[151,1436,1185,1238]" gridrow="9" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FFB4DFE0BFB4D" box="[151,451,1185,1238]" gridcol="0" gridrow="9" pageId="13" pageNumber="14">HL / HW 3</th>
<td id="82355D867F4BACCFFE71FB4DFDACFB4D" box="[473,612,1185,1238]" gridcol="1" gridrow="9" pageId="13" pageNumber="14">1.391.59 (1.50 ± 0.05)</td>
<td id="82355D867F4BACCFFD27FB4DFC9BFB4D" box="[655,851,1185,1238]" gridcol="2" gridrow="9" pageId="13" pageNumber="14">1.431.65 (1.53 ± 0.06)</td>
<td id="82355D867F4BACCFFCC4FB4DFBDFFB4D" box="[876,1047,1185,1238]" gridcol="3" gridrow="9" pageId="13" pageNumber="14">1.421.58 (1.52 ± 0.05)</td>
<td id="82355D867F4BACCFFB90FB4DFB06FB4D" box="[1080,1230,1185,1238]" gridcol="4" gridrow="9" pageId="13" pageNumber="14">1.471.61 (1.54 ± 0.04)</td>
<td id="82355D867F4BACCFFB5EFB4DFA54FB4D" box="[1270,1436,1185,1238]" gridcol="5" gridrow="9" pageId="13" pageNumber="14">1.431.68 (1.52 ± 0.07)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFB05FA54FA85" box="[151,1436,1257,1310]" gridrow="10" pageId="13" pageNumber="14" rowspan-4="1">
<th id="82355D867F4BACCFFF3FFB05FE0BFA85" box="[151,451,1257,1310]" gridcol="0" gridrow="10" pageId="13" pageNumber="14">Ƥ</th>
<td id="82355D867F4BACCFFE71FB05FDACFA85" box="[473,612,1257,1310]" gridcol="1" gridrow="10" pageId="13" pageNumber="14">1.451.56 (1.50 ± 0.05)</td>
<td id="82355D867F4BACCFFD27FB05FC9BFA85" box="[655,851,1257,1310]" gridcol="2" gridrow="10" pageId="13" pageNumber="14">1.511.63 (1.56 ± 0.06)</td>
<td id="82355D867F4BACCFFCC4FB05FBDFFA85" box="[876,1047,1257,1310]" gridcol="3" gridrow="10" pageId="13" pageNumber="14">1.471.58 (1.53 ± 0.04)</td>
<td id="82355D867F4BACCFFB5EFB05FA54FA85" box="[1270,1436,1257,1310]" gridcol="5" gridrow="10" pageId="13" pageNumber="14">1.441.60 (1.52 ± 0.05)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFADDFA54FAFD" box="[151,1436,1329,1382]" gridrow="11" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FFADDFE0BFAFD" box="[151,451,1329,1382]" gridcol="0" gridrow="11" pageId="13" pageNumber="14">Shank length / SVL 3</th>
<td id="82355D867F4BACCFFE71FADDFDACFAFD" box="[473,612,1329,1382]" gridcol="1" gridrow="11" pageId="13" pageNumber="14">0.270.32 (0.28 ± 0.01)</td>
<td id="82355D867F4BACCFFD27FADDFC9BFAFD" box="[655,851,1329,1382]" gridcol="2" gridrow="11" pageId="13" pageNumber="14">0.290.31 (0.30 ± 0.01)</td>
<td id="82355D867F4BACCFFCC4FADDFBDFFAFD" box="[876,1047,1329,1382]" gridcol="3" gridrow="11" pageId="13" pageNumber="14">0.250.27 (0.26 ± 0.01)</td>
<td id="82355D867F4BACCFFB90FADDFB06FAFD" box="[1080,1230,1329,1382]" gridcol="4" gridrow="11" pageId="13" pageNumber="14">0.240.29 (0.27 ± 0.01)</td>
<td id="82355D867F4BACCFFB5EFADDFA54FAFD" box="[1270,1436,1329,1382]" gridcol="5" gridrow="11" pageId="13" pageNumber="14">0.260.33 (0.29 ± 0.02)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFA95FA54FA35" box="[151,1436,1401,1454]" gridrow="12" pageId="13" pageNumber="14" rowspan-4="1">
<th id="82355D867F4BACCFFF3FFA95FE0BFA35" box="[151,451,1401,1454]" gridcol="0" gridrow="12" pageId="13" pageNumber="14">Ƥ</th>
<td id="82355D867F4BACCFFE71FA95FDACFA35" box="[473,612,1401,1454]" gridcol="1" gridrow="12" pageId="13" pageNumber="14">0.260.28 (0.27 ± 0.01)</td>
<td id="82355D867F4BACCFFD27FA95FC9BFA35" box="[655,851,1401,1454]" gridcol="2" gridrow="12" pageId="13" pageNumber="14">0.270.30 (0.28 ± 0.01)</td>
<td id="82355D867F4BACCFFCC4FA95FBDFFA35" box="[876,1047,1401,1454]" gridcol="3" gridrow="12" pageId="13" pageNumber="14">0.220.27 (0.25 ± 0.02)</td>
<td id="82355D867F4BACCFFB5EFA95FA54FA35" box="[1270,1436,1401,1454]" gridcol="5" gridrow="12" pageId="13" pageNumber="14">0.270.33 (0.29 ± 0.02)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FFA2DFA54FA6D" box="[151,1436,1473,1526]" gridrow="13" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FFA2DFE0BFA6D" box="[151,451,1473,1526]" gridcol="0" gridrow="13" pageId="13" pageNumber="14">Axilla-groin distance / 3 SVL</th>
<td id="82355D867F4BACCFFE71FA2DFDACFA6D" box="[473,612,1473,1526]" gridcol="1" gridrow="13" pageId="13" pageNumber="14">0.380.46 (0.41 ± 0.03)</td>
<td id="82355D867F4BACCFFD27FA2DFC9BFA6D" box="[655,851,1473,1526]" gridcol="2" gridrow="13" pageId="13" pageNumber="14">0.340.43 (0.39 ± 0.02)</td>
<td id="82355D867F4BACCFFCC4FA2DFBDFFA6D" box="[876,1047,1473,1526]" gridcol="3" gridrow="13" pageId="13" pageNumber="14">0.370.44 (0.40 ± 0.02)</td>
<td id="82355D867F4BACCFFB90FA2DFB06FA6D" box="[1080,1230,1473,1526]" gridcol="4" gridrow="13" pageId="13" pageNumber="14">0.370.40 (0.38 ± 0.01)</td>
<td id="82355D867F4BACCFFB5EFA2DFA54FA6D" box="[1270,1436,1473,1526]" gridcol="5" gridrow="13" pageId="13" pageNumber="14">0.310.45 (0.39 ± 0.03)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FF9E5FA54F9A5" box="[151,1436,1545,1598]" gridrow="14" pageId="13" pageNumber="14" rowspan-4="1">
<th id="82355D867F4BACCFFF3FF9E5FE0BF9A5" box="[151,451,1545,1598]" gridcol="0" gridrow="14" pageId="13" pageNumber="14">Ƥ</th>
<td id="82355D867F4BACCFFE71F9E5FDACF9A5" box="[473,612,1545,1598]" gridcol="1" gridrow="14" pageId="13" pageNumber="14">0.370.45 (0.41 ± 0.03)</td>
<td id="82355D867F4BACCFFD27F9E5FC9BF9A5" box="[655,851,1545,1598]" gridcol="2" gridrow="14" pageId="13" pageNumber="14">0.400.42 (0.41 ± 0.01)</td>
<td id="82355D867F4BACCFFCC4F9E5FBDFF9A5" box="[876,1047,1545,1598]" gridcol="3" gridrow="14" pageId="13" pageNumber="14">0.370.45 (0.40 ± 0.03)</td>
<td id="82355D867F4BACCFFB5EF9E5FA54F9A5" box="[1270,1436,1545,1598]" gridcol="5" gridrow="14" pageId="13" pageNumber="14">0.340.45 (0.41 ± 0.04)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FF9BDFA54F913" box="[151,1436,1617,1672]" gridrow="15" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FF9BDFE0BF913" box="[151,451,1617,1672]" gridcol="0" gridrow="15" pageId="13" pageNumber="14">Subdigital lamellae under phalanges IIIV of 4th toe</th>
<td id="82355D867F4BACCFFE71F9BDFDACF913" box="[473,612,1617,1672]" gridcol="1" gridrow="15" pageId="13" pageNumber="14">2124 (22.6 ± 1.0)</td>
<td id="82355D867F4BACCFFD27F9BDFC9BF913" box="[655,851,1617,1672]" gridcol="2" gridrow="15" pageId="13" pageNumber="14">2124 (22.3 ± 1.0)</td>
<td id="82355D867F4BACCFFCC4F9BDFBDFF913" box="[876,1047,1617,1672]" gridcol="3" gridrow="15" pageId="13" pageNumber="14">1924 (22.3 ± 1.3)</td>
<td id="82355D867F4BACCFFB90F9BDFB06F913" box="[1080,1230,1617,1672]" gridcol="4" gridrow="15" pageId="13" pageNumber="14">2022 (21.5 ± 0.7)</td>
<td id="82355D867F4BACCFFB5EF9BDFA54F913" box="[1270,1436,1617,1672]" gridcol="5" gridrow="15" pageId="13" pageNumber="14">1925 (22.2 ± 1.4)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FF977FA54F94A" box="[151,1436,1691,1745]" gridrow="16" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FF977FE0BF94A" box="[151,451,1691,1745]" gridcol="0" gridrow="16" pageId="13" pageNumber="14">Number of scales between SS</th>
<td id="82355D867F4BACCFFE71F977FDACF94A" box="[473,612,1691,1745]" gridcol="1" gridrow="16" pageId="13" pageNumber="14">24 (3.2 ± 0.7)</td>
<td id="82355D867F4BACCFFD27F977FC9BF94A" box="[655,851,1691,1745]" gridcol="2" gridrow="16" pageId="13" pageNumber="14">48 (6.2 ± 1.2)</td>
<td id="82355D867F4BACCFFCC4F977FBDFF94A" box="[876,1047,1691,1745]" gridcol="3" gridrow="16" pageId="13" pageNumber="14">34 (3.4 ± 0.5)</td>
<td id="82355D867F4BACCFFB90F977FB06F94A" box="[1080,1230,1691,1745]" gridcol="4" gridrow="16" pageId="13" pageNumber="14">35 (4.1 ± 0.6)</td>
<td id="82355D867F4BACCFFB5EF977FA54F94A" box="[1270,1436,1691,1745]" gridcol="5" gridrow="16" pageId="13" pageNumber="14">48 (5.8 ± 1.1)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FF90FFA54F882" box="[151,1436,1763,1817]" gridrow="17" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FF90FFE0BF882" box="[151,451,1763,1817]" gridcol="0" gridrow="17" pageId="13" pageNumber="14">Number of scales between IP and SS</th>
<td id="82355D867F4BACCFFE71F90FFDACF882" box="[473,612,1763,1817]" gridcol="1" gridrow="17" pageId="13" pageNumber="14">35 (3.4 ± 0.6)</td>
<td id="82355D867F4BACCFFD27F90FFC9BF882" box="[655,851,1763,1817]" gridcol="2" gridrow="17" pageId="13" pageNumber="14">411 (7.6 ± 1.7)</td>
<td id="82355D867F4BACCFFCC4F90FFBDFF882" box="[876,1047,1763,1817]" gridcol="3" gridrow="17" pageId="13" pageNumber="14">24 (3.2 ± 0.5)</td>
<td id="82355D867F4BACCFFB90F90FFB06F882" box="[1080,1230,1763,1817]" gridcol="4" gridrow="17" pageId="13" pageNumber="14">24 (3.5 ± 0.7)</td>
<td id="82355D867F4BACCFFB5EF90FFA54F882" box="[1270,1436,1763,1817]" gridcol="5" gridrow="17" pageId="13" pageNumber="14">410 (7.2 ± 1.5)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FF8C7FA54F8FA" box="[151,1436,1835,1889]" gridrow="18" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FF8C7FE0BF8FA" box="[151,451,1835,1889]" gridcol="0" gridrow="18" pageId="13" pageNumber="14">Number of SPL to level below center of eye</th>
<td id="82355D867F4BACCFFE71F8C7FDACF8FA" box="[473,612,1835,1889]" gridcol="1" gridrow="18" pageId="13" pageNumber="14">67 (6.1 ± 0.4)</td>
<td id="82355D867F4BACCFFD27F8C7FC9BF8FA" box="[655,851,1835,1889]" gridcol="2" gridrow="18" pageId="13" pageNumber="14">68 (6.6 ± 0.7)</td>
<td id="82355D867F4BACCFFCC4F8C7FBDFF8FA" box="[876,1047,1835,1889]" gridcol="3" gridrow="18" pageId="13" pageNumber="14">69 (7.1 ± 0.8)</td>
<td id="82355D867F4BACCFFB90F8C7FB06F8FA" box="[1080,1230,1835,1889]" gridcol="4" gridrow="18" pageId="13" pageNumber="14">58 (6.5 ± 0.9)</td>
<td id="82355D867F4BACCFFB5EF8C7FA54F8FA" box="[1270,1436,1835,1889]" gridcol="5" gridrow="18" pageId="13" pageNumber="14">58 (6.3 ± 0.7)</td>
</tr>
<tr id="C1E434FA7F4BACCFFF3FF89FFA54F832" box="[151,1436,1907,1961]" gridrow="19" pageId="13" pageNumber="14">
<th id="82355D867F4BACCFFF3FF89FFE0BF832" box="[151,451,1907,1961]" gridcol="0" gridrow="19" pageId="13" pageNumber="14">Number of INL to level below center of eye</th>
<td id="82355D867F4BACCFFE71F89FFDACF832" box="[473,612,1907,1961]" gridcol="1" gridrow="19" pageId="13" pageNumber="14">68 (6.9 ± 0.7)</td>
<td id="82355D867F4BACCFFD27F89FFC9BF832" box="[655,851,1907,1961]" gridcol="2" gridrow="19" pageId="13" pageNumber="14">68 (7.2 ± 0.8)</td>
<td id="82355D867F4BACCFFCC4F89FFBDFF832" box="[876,1047,1907,1961]" gridcol="3" gridrow="19" pageId="13" pageNumber="14">69 (7.8 ± 0.7)</td>
<td id="82355D867F4BACCFFB90F89FFB06F832" box="[1080,1230,1907,1961]" gridcol="4" gridrow="19" pageId="13" pageNumber="14">57 (6.7 ± 0.7)</td>
<td id="82355D867F4BACCFFB5EF89FFA54F832" box="[1270,1436,1907,1961]" gridcol="5" gridrow="19" pageId="13" pageNumber="14">58 (6.7 ± 0.8)</td>
</tr>
</table>
</paragraph>
<paragraph id="7F6B36B87F4B533EFB62F857FBCCF99A" blockId="13.[1226,1430,1979,2002]" lastBlockId="14.[151,1437,1201,1988]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
continued next page The dewlap coloration of the juvenile male SMF 90148 from the vicinity of the
<typeStatus id="A06F881A7F48533EFBFBFB5EFB4BFB51" box="[1107,1155,1202,1226]" pageId="14" pageNumber="15">type</typeStatus>
locality was recorded as follows: Flame Scarlet (15), with a suggestion of Chrome Orange (16), grading into Orange Yellow (18) on anterior margin; dewlap scales Sepia (119).
<collectingCountry id="07C376287F48533EFD8FFB1BFD8EFA94" box="[551,582,1271,1295]" name="American Samoa" pageId="14" pageNumber="15">As</collectingCountry>
illustrated in
<figureCitation id="E7EF2A3D7F48533EFD49FB1AFCE1FA95" box="[737,809,1270,1294]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1851]" captionTargetBox="[305,1272,193,1803]" captionTargetId="figure@3.[305,1272,193,1804]" captionTargetPageId="3" captionText="FIGURE 1. Dewlaps of anoles related to Anolis pachypus from Costa Rica and western Panama, representative of the five OTUs (width of tweezers 3 mm): (A) OTU A (SMF 89729, Sendero La Cascada, Bajo Mono, Chiriquí, Panama); (B) OTU B (GK 2297, road from San José de la Montaña to Birrí, Heredia, Costa Rica); (C) OTU C (GK 2355, road from San Isidro de El General to Cerro de la Muerte, San José, Costa Rica); (D) OTU C (SL 439, Jurutungo, Chiriquí, Panama); (E) OTU C (SL 524, Jurutungo, Chiriquí, Panama); (F) OTU D (SMF 89507, Jurutungo, Chiriquí, Panama); (G) OTU D (SMF 89506, Jurutungo, Chiriquí, Panama); (H) OTU D (SMF 90149, N slope Cerro Pando, Bocas del Toro, Panama); (I) OTU E (SMF 89519, La Nevera, Comarca Ngöbe-Buglé, Panama); (J) OTU E (SL 510, La Nevera, Comarca Ngöbe-Buglé, Panama)." httpUri="https://zenodo.org/record/200704/files/figure.png" pageId="14" pageNumber="15">Figs. 1</figureCitation>
FH, the extent of the yellow coloration on the anterior portion of the male dewlap of
<taxonomicName id="B8D44D3B7F48533EFE45FAF5FDBBFAAA" box="[493,627,1305,1329]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F48533EFE45FAF5FDBBFAAA" box="[493,627,1305,1329]" italics="true" pageId="14" pageNumber="15">A. benedikti</emphasis>
</taxonomicName>
varies from covering just the basal part of the anterior margin (
<figureCitation id="E7EF2A3D7F48533EFAE8FAF5FAB7FAAA" box="[1344,1407,1305,1329]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1851]" captionTargetBox="[305,1272,193,1803]" captionTargetId="figure@3.[305,1272,193,1804]" captionTargetPageId="3" captionText="FIGURE 1. Dewlaps of anoles related to Anolis pachypus from Costa Rica and western Panama, representative of the five OTUs (width of tweezers 3 mm): (A) OTU A (SMF 89729, Sendero La Cascada, Bajo Mono, Chiriquí, Panama); (B) OTU B (GK 2297, road from San José de la Montaña to Birrí, Heredia, Costa Rica); (C) OTU C (GK 2355, road from San Isidro de El General to Cerro de la Muerte, San José, Costa Rica); (D) OTU C (SL 439, Jurutungo, Chiriquí, Panama); (E) OTU C (SL 524, Jurutungo, Chiriquí, Panama); (F) OTU D (SMF 89507, Jurutungo, Chiriquí, Panama); (G) OTU D (SMF 89506, Jurutungo, Chiriquí, Panama); (H) OTU D (SMF 90149, N slope Cerro Pando, Bocas del Toro, Panama); (I) OTU E (SMF 89519, La Nevera, Comarca Ngöbe-Buglé, Panama); (J) OTU E (SL 510, La Nevera, Comarca Ngöbe-Buglé, Panama)." httpUri="https://zenodo.org/record/200704/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
F) to covering the anterior third of the dewlap (
<figureCitation id="E7EF2A3D7F48533EFD32FAD0FD13FACF" box="[666,731,1340,1364]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1851]" captionTargetBox="[305,1272,193,1803]" captionTargetId="figure@3.[305,1272,193,1804]" captionTargetPageId="3" captionText="FIGURE 1. Dewlaps of anoles related to Anolis pachypus from Costa Rica and western Panama, representative of the five OTUs (width of tweezers 3 mm): (A) OTU A (SMF 89729, Sendero La Cascada, Bajo Mono, Chiriquí, Panama); (B) OTU B (GK 2297, road from San José de la Montaña to Birrí, Heredia, Costa Rica); (C) OTU C (GK 2355, road from San Isidro de El General to Cerro de la Muerte, San José, Costa Rica); (D) OTU C (SL 439, Jurutungo, Chiriquí, Panama); (E) OTU C (SL 524, Jurutungo, Chiriquí, Panama); (F) OTU D (SMF 89507, Jurutungo, Chiriquí, Panama); (G) OTU D (SMF 89506, Jurutungo, Chiriquí, Panama); (H) OTU D (SMF 90149, N slope Cerro Pando, Bocas del Toro, Panama); (I) OTU E (SMF 89519, La Nevera, Comarca Ngöbe-Buglé, Panama); (J) OTU E (SL 510, La Nevera, Comarca Ngöbe-Buglé, Panama)." httpUri="https://zenodo.org/record/200704/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
H). The female dewlap (
<figureCitation id="E7EF2A3D7F48533EFC51FAD0FB80FACF" box="[1017,1096,1340,1364]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1923]" captionTargetBox="[216,1349,193,1860]" captionTargetId="figure@8.[216,1350,193,1861]" captionTargetPageId="8" captionText="FIGURE 5. Type specimens of Anolis benedikti in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (C E) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (F O) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female." httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="14" pageNumber="15">Figs. 5</figureCitation>
CE) is much smaller than in males, and usually of a whitish color (
<figureCitation id="E7EF2A3D7F48533EFDF2FAB2FD68FAED" box="[602,672,1374,1398]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1923]" captionTargetBox="[216,1349,193,1860]" captionTargetId="figure@8.[216,1350,193,1861]" captionTargetPageId="8" captionText="FIGURE 5. Type specimens of Anolis benedikti in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (C E) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (F O) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female." httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="14" pageNumber="15">Fig. 5</figureCitation>
C). Nevertheless, some female individuals bear reddish coloration around their gular area. These females tend to exhibit a reddish dewlap (
<figureCitation id="E7EF2A3D7F48533EFC62FA6DFBDFFA02" box="[970,1047,1409,1433]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1923]" captionTargetBox="[216,1349,193,1860]" captionTargetId="figure@8.[216,1350,193,1861]" captionTargetPageId="8" captionText="FIGURE 5. Type specimens of Anolis benedikti in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (C E) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (F O) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female." httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="14" pageNumber="15">Figs. 5</figureCitation>
DE), sometimes even resembling that of males, like the female
<typeStatus id="A06F881A7F48533EFE5CFA49FD9CFA26" box="[500,596,1445,1469]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
SMF 89746 (
<figureCitation id="E7EF2A3D7F48533EFD51FA48FCF6FA27" box="[761,830,1444,1468]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1923]" captionTargetBox="[216,1349,193,1860]" captionTargetId="figure@8.[216,1350,193,1861]" captionTargetPageId="8" captionText="FIGURE 5. Type specimens of Anolis benedikti in life: (A) male holotype (SMF 90149); (B) male from Jurutungo, SMF 89506; (C E) female dewlaps: (C) SMF 91506; (D) SL 695; (E) SMF 89746; (F O) variation in dorsal patterns: (F) SMF 90149, male holotype; (G) SMF 89506, male; (H) SMF 89745, male; (I) SMF 91508, male; (J) SL 202, female; (K) SMF 89746, female, at night; (L) SMF 89746 the next day; (M) SMF 91507, female; (N) SL 686, female; (O) SMF 91505, female." httpUri="https://zenodo.org/record/200709/files/figure.png" pageId="14" pageNumber="15">Fig. 5</figureCitation>
E), that was recorded as follows: Flame Scarlet (15) posteriorly, grading into Orange Yellow (18) on anterior half and into dirty white with a suggestion of Chamois (123D) on posterior base; dewlap scales mostly Sepia (119), some dirty white.
</paragraph>
<caption id="2BAB66307F48533EFF37FF4CFE4FFF2D" ID-Table-UUID="2BAB66307F48533EFF37FF4CFE4FFF2D" box="[159,391,160,182]" httpUri="http://table.plazi.org/id/2BAB66307F48533EFF37FF4CFE4FFF2D" pageId="14" pageNumber="15" targetBox="[159,1408,219,1138]" targetIsTable="true" targetPageId="14">
<paragraph id="7F6B36B87F48533EFF37FF4CFE4FFF2D" blockId="14.[159,391,160,182]" box="[159,391,160,182]" pageId="14" pageNumber="15">
<emphasis id="4DA0EAAA7F48533EFF37FF4CFEC4FF2E" bold="true" box="[159,268,160,181]" pageId="14" pageNumber="15">TABLE 1.</emphasis>
(continued)
</paragraph>
</caption>
<paragraph id="7F6B36B87F48533EFE71FF37FAA5FBE9" pageId="14" pageNumber="15">
<table id="0DD4C4187F48ACCFFF37FF37FA48FBE9" box="[159,1408,219,1138]" gridcols="5" gridrows="13" pageId="14" pageNumber="15">
<tr id="C1E434FA7F48ACCFFF37FF37FA48FE94" box="[159,1408,219,271]" gridrow="0" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FF37FD95FE94" box="[159,605,219,271]" gridcol="0" gridrow="0" pageId="14" pageNumber="15">
<taxonomicName id="B8D44D3B7F48533EFE71FF37FD9DFF6A" box="[473,597,219,241]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F48533EFE71FF37FD9DFF6A" box="[473,597,219,241]" italics="true" pageId="14" pageNumber="15">A. pachypus</emphasis>
</taxonomicName>
3 11, Ƥ 5
</th>
<th id="82355D867F48ACCFFD27FF37FC9BFE94" box="[655,851,219,271]" gridcol="1" gridrow="0" pageId="14" pageNumber="15">
<taxonomicName id="B8D44D3B7F48533EFD27FF37FC9BFF6A" box="[655,851,219,241]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pseudopachypus">
<emphasis id="4DA0EAAA7F48533EFD27FF37FC9BFF6A" box="[655,851,219,241]" italics="true" pageId="14" pageNumber="15">A. pseudopachypus</emphasis>
</taxonomicName>
3 16, Ƥ 5
</th>
<th id="82355D867F48ACCFFCC4FF37FBDFFE94" box="[876,1047,219,271]" gridcol="2" gridrow="0" pageId="14" pageNumber="15">
<taxonomicName id="B8D44D3B7F48533EFCC4FF37FBDFFF6A" box="[876,1047,219,241]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="magnaphallus">
<emphasis id="4DA0EAAA7F48533EFCC4FF37FBDFFF6A" box="[876,1047,219,241]" italics="true" pageId="14" pageNumber="15">A. magnaphallus</emphasis>
</taxonomicName>
3 9, Ƥ 6
</th>
<th id="82355D867F48ACCFFB90FF37FB06FE94" box="[1080,1230,219,271]" gridcol="3" gridrow="0" pageId="14" pageNumber="15">
<taxonomicName id="B8D44D3B7F48533EFB90FF37FB06FF6A" box="[1080,1230,219,241]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="tropidolepis">
<emphasis id="4DA0EAAA7F48533EFB90FF37FB06FF6A" box="[1080,1230,219,241]" italics="true" pageId="14" pageNumber="15">A. tropidolepis</emphasis>
</taxonomicName>
3 10, Ƥ 0
</th>
<th id="82355D867F48ACCFFB5EFF37FA48FE94" box="[1270,1408,219,271]" gridcol="4" gridrow="0" pageId="14" pageNumber="15">
<taxonomicName id="B8D44D3B7F48533EFB5EFF37FAB8FF6A" box="[1270,1392,219,241]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F48533EFB5EFF37FAB8FF6A" box="[1270,1392,219,241]" italics="true" pageId="14" pageNumber="15">A. benedikti</emphasis>
</taxonomicName>
3 20, Ƥ 13
</th>
</tr>
<tr id="C1E434FA7F48ACCFFF37FECEFA48FEC3" box="[159,1408,290,344]" gridrow="1" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FECEFD95FEC3" box="[159,605,290,344]" gridcol="0" gridrow="1" pageId="14" pageNumber="15">Total number of loreals 4769 (57.9 ± 6.5)</th>
<td id="82355D867F48ACCFFD27FECEFC9BFEC3" box="[655,851,290,344]" gridcol="1" gridrow="1" pageId="14" pageNumber="15">4396 (68.4 ± 14.5)</td>
<td id="82355D867F48ACCFFCC4FECEFBDFFEC3" box="[876,1047,290,344]" gridcol="2" gridrow="1" pageId="14" pageNumber="15">4897 (65.3 ± 14.1)</td>
<td id="82355D867F48ACCFFB90FECEFB06FEC3" box="[1080,1230,290,344]" gridcol="3" gridrow="1" pageId="14" pageNumber="15">4977 (62.5 ± 9.3)</td>
<td id="82355D867F48ACCFFB5EFECEFA48FEC3" box="[1270,1408,290,344]" gridcol="4" gridrow="1" pageId="14" pageNumber="15">4491 (62.4 ± 11.3)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FE86FA48FE3B" box="[159,1408,362,416]" gridrow="2" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FE86FD95FE3B" box="[159,605,362,416]" gridcol="0" gridrow="2" pageId="14" pageNumber="15">Number of horizontal loreal 711 scale rows (9.6 ± 0.9)</th>
<td id="82355D867F48ACCFFD27FE86FC9BFE3B" box="[655,851,362,416]" gridcol="1" gridrow="2" pageId="14" pageNumber="15">812 (9.9 ± 1.1)</td>
<td id="82355D867F48ACCFFCC4FE86FBDFFE3B" box="[876,1047,362,416]" gridcol="2" gridrow="2" pageId="14" pageNumber="15">712 (9.4 ± 1.3)</td>
<td id="82355D867F48ACCFFB90FE86FB06FE3B" box="[1080,1230,362,416]" gridcol="3" gridrow="2" pageId="14" pageNumber="15">711 (9.1 ± 1.0)</td>
<td id="82355D867F48ACCFFB5EFE86FA48FE3B" box="[1270,1408,362,416]" gridcol="4" gridrow="2" pageId="14" pageNumber="15">712 (9.9 ± 1.2)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FE5EFA48FE73" box="[159,1408,434,488]" gridrow="3" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FE5EFD95FE73" box="[159,605,434,488]" gridcol="0" gridrow="3" pageId="14" pageNumber="15">Number of postrostrals 57 (5.9 ± 0.8)</th>
<td id="82355D867F48ACCFFD27FE5EFC9BFE73" box="[655,851,434,488]" gridcol="1" gridrow="3" pageId="14" pageNumber="15">58 (6.6 ± 0.9)</td>
<td id="82355D867F48ACCFFCC4FE5EFBDFFE73" box="[876,1047,434,488]" gridcol="2" gridrow="3" pageId="14" pageNumber="15">59 (7.0 ± 1.0)</td>
<td id="82355D867F48ACCFFB90FE5EFB06FE73" box="[1080,1230,434,488]" gridcol="3" gridrow="3" pageId="14" pageNumber="15">68 (7.1 ± 0.7)</td>
<td id="82355D867F48ACCFFB5EFE5EFA48FE73" box="[1270,1408,434,488]" gridcol="4" gridrow="3" pageId="14" pageNumber="15">59 (6.6 ± 0.9)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FE16FA48FDAB" box="[159,1408,506,560]" gridrow="4" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FE16FD95FDAB" box="[159,605,506,560]" gridcol="0" gridrow="4" pageId="14" pageNumber="15">Number of postmentals 58 (6.0 ± 0.6)</th>
<td id="82355D867F48ACCFFD27FE16FC9BFDAB" box="[655,851,506,560]" gridcol="1" gridrow="4" pageId="14" pageNumber="15">68 (6.3 ± 0.6)</td>
<td id="82355D867F48ACCFFCC4FE16FBDFFDAB" box="[876,1047,506,560]" gridcol="2" gridrow="4" pageId="14" pageNumber="15">59 (6.6 ± 1.0)</td>
<td id="82355D867F48ACCFFB90FE16FB06FDAB" box="[1080,1230,506,560]" gridcol="3" gridrow="4" pageId="14" pageNumber="15">58 (6.2 ± 0.9)</td>
<td id="82355D867F48ACCFFB5EFE16FA48FDAB" box="[1270,1408,506,560]" gridcol="4" gridrow="4" pageId="14" pageNumber="15">58 (6.3 ± 0.8)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FDAEFA48FDE3" box="[159,1408,578,632]" gridrow="5" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FDAEFD95FDE3" box="[159,605,578,632]" gridcol="0" gridrow="5" pageId="14" pageNumber="15">Number of scales between 69 nasals (7.3 ± 0.9)</th>
<td id="82355D867F48ACCFFD27FDAEFC9BFDE3" box="[655,851,578,632]" gridcol="1" gridrow="5" pageId="14" pageNumber="15">68 (6.6 ± 0.7)</td>
<td id="82355D867F48ACCFFCC4FDAEFBDFFDE3" box="[876,1047,578,632]" gridcol="2" gridrow="5" pageId="14" pageNumber="15">69 (6.9 ± 0.8)</td>
<td id="82355D867F48ACCFFB90FDAEFB06FDE3" box="[1080,1230,578,632]" gridcol="3" gridrow="5" pageId="14" pageNumber="15">68 (7.0 ± 0.5)</td>
<td id="82355D867F48ACCFFB5EFDAEFA48FDE3" box="[1270,1408,578,632]" gridcol="4" gridrow="5" pageId="14" pageNumber="15">510 (6.6 ± 1.1)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FD66FA48FD5A" box="[159,1408,650,705]" gridrow="6" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FD66FD95FD5A" box="[159,605,650,705]" gridcol="0" gridrow="6" pageId="14" pageNumber="15">Number of scales between 2nd 1014 canthals (11.9 ± 1.1)</th>
<td id="82355D867F48ACCFFD27FD66FC9BFD5A" box="[655,851,650,705]" gridcol="1" gridrow="6" pageId="14" pageNumber="15">1321 (16.2 ± 1.8)</td>
<td id="82355D867F48ACCFFCC4FD66FBDFFD5A" box="[876,1047,650,705]" gridcol="2" gridrow="6" pageId="14" pageNumber="15">1014 (11.3 ± 1.1)</td>
<td id="82355D867F48ACCFFB90FD66FB06FD5A" box="[1080,1230,650,705]" gridcol="3" gridrow="6" pageId="14" pageNumber="15">914 (11.3 ± 1.8)</td>
<td id="82355D867F48ACCFFB5EFD66FA48FD5A" box="[1270,1408,650,705]" gridcol="4" gridrow="6" pageId="14" pageNumber="15">1118 (14.0 ± 1.5)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FD39FA48FC91" box="[159,1408,725,778]" gridrow="7" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FD39FD95FC91" box="[159,605,725,778]" gridcol="0" gridrow="7" pageId="14" pageNumber="15">Number of scales between 1116 posterior canthals (13.4 ± 1.2)</th>
<td id="82355D867F48ACCFFD27FD39FC9BFC91" box="[655,851,725,778]" gridcol="1" gridrow="7" pageId="14" pageNumber="15">1524 (18.9 ± 2.1)</td>
<td id="82355D867F48ACCFFCC4FD39FBDFFC91" box="[876,1047,725,778]" gridcol="2" gridrow="7" pageId="14" pageNumber="15">1115 (13.1 ± 1.1)</td>
<td id="82355D867F48ACCFFB90FD39FB06FC91" box="[1080,1230,725,778]" gridcol="3" gridrow="7" pageId="14" pageNumber="15">1015 (12.6 ± 1.7)</td>
<td id="82355D867F48ACCFFB5EFD39FA48FC91" box="[1270,1408,725,778]" gridcol="4" gridrow="7" pageId="14" pageNumber="15">1221 (16.1 ± 1.9)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FCF1FA48FCC9" box="[159,1408,797,850]" gridrow="8" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FCF1FD95FCC9" box="[159,605,797,850]" gridcol="0" gridrow="8" pageId="14" pageNumber="15">Number of medial dorsal 5068 scales in one HL (58.8 ± 5.4)</th>
<td id="82355D867F48ACCFFD27FCF1FC9BFCC9" box="[655,851,797,850]" gridcol="1" gridrow="8" pageId="14" pageNumber="15">5092 (71.4 ± 7.4)</td>
<td id="82355D867F48ACCFFCC4FCF1FBDFFCC9" box="[876,1047,797,850]" gridcol="2" gridrow="8" pageId="14" pageNumber="15">4864 (55.3 ± 3.9)</td>
<td id="82355D867F48ACCFFB90FCF1FB06FCC9" box="[1080,1230,797,850]" gridcol="3" gridrow="8" pageId="14" pageNumber="15">5062 (56.0 ± 4.2)</td>
<td id="82355D867F48ACCFFB5EFCF1FA48FCC9" box="[1270,1408,797,850]" gridcol="4" gridrow="8" pageId="14" pageNumber="15">5276 (63.9 ± 5.6)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FC89FA48FC01" box="[159,1408,869,922]" gridrow="9" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FC89FD95FC01" box="[159,605,869,922]" gridcol="0" gridrow="9" pageId="14" pageNumber="15">Number of medial ventral 3242 scales in one HL (34.8 ± 2.8)</th>
<td id="82355D867F48ACCFFD27FC89FC9BFC01" box="[655,851,869,922]" gridcol="1" gridrow="9" pageId="14" pageNumber="15">3048 (39.3 ± 4.3)</td>
<td id="82355D867F48ACCFFCC4FC89FBDFFC01" box="[876,1047,869,922]" gridcol="2" gridrow="9" pageId="14" pageNumber="15">2638 (30.9 ± 3.7)</td>
<td id="82355D867F48ACCFFB90FC89FB06FC01" box="[1080,1230,869,922]" gridcol="3" gridrow="9" pageId="14" pageNumber="15">3050 (39.2 ± 6.1)</td>
<td id="82355D867F48ACCFFB5EFC89FA48FC01" box="[1270,1408,869,922]" gridcol="4" gridrow="9" pageId="14" pageNumber="15">2644 (37.3 ± 3.7)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FC41FA48FC79" box="[159,1408,941,994]" gridrow="10" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FC41FD95FC79" box="[159,605,941,994]" gridcol="0" gridrow="10" pageId="14" pageNumber="15">Number of scales around 126154 midbody (140.1 ± 8.7)</th>
<td id="82355D867F48ACCFFD27FC41FC9BFC79" box="[655,851,941,994]" gridcol="1" gridrow="10" pageId="14" pageNumber="15">126156 (140.0 ±8.4)</td>
<td id="82355D867F48ACCFFCC4FC41FBDFFC79" box="[876,1047,941,994]" gridcol="2" gridrow="10" pageId="14" pageNumber="15">128148 (137.9 ±6.6)</td>
<td id="82355D867F48ACCFFB90FC41FB06FC79" box="[1080,1230,941,994]" gridcol="3" gridrow="10" pageId="14" pageNumber="15">128152 (137.8 ±6.6)</td>
<td id="82355D867F48ACCFFB5EFC41FA48FC79" box="[1270,1408,941,994]" gridcol="4" gridrow="10" pageId="14" pageNumber="15">128178 (146.2 ±11.7)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FC19FA48FBB1" box="[159,1408,1013,1066]" gridrow="11" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FC19FD95FBB1" box="[159,605,1013,1066]" gridcol="0" gridrow="11" pageId="14" pageNumber="15">Number of dorsal scales 79104 between axilla and groin (90.4 ± 7.0)</th>
<td id="82355D867F48ACCFFD27FC19FC9BFBB1" box="[655,851,1013,1066]" gridcol="1" gridrow="11" pageId="14" pageNumber="15">89119 (105.1 ± 7.6)</td>
<td id="82355D867F48ACCFFCC4FC19FBDFFBB1" box="[876,1047,1013,1066]" gridcol="2" gridrow="11" pageId="14" pageNumber="15">81102 (90.4 ± 6.2)</td>
<td id="82355D867F48ACCFFB90FC19FB06FBB1" box="[1080,1230,1013,1066]" gridcol="3" gridrow="11" pageId="14" pageNumber="15">6692 (78.6 ± 7.7)</td>
<td id="82355D867F48ACCFFB5EFC19FA48FBB1" box="[1270,1408,1013,1066]" gridcol="4" gridrow="11" pageId="14" pageNumber="15">83113 (99.2 ± 8.0)</td>
</tr>
<tr id="C1E434FA7F48ACCFFF37FBD1FA48FBE9" box="[159,1408,1085,1138]" gridrow="12" pageId="14" pageNumber="15">
<th id="82355D867F48ACCFFF37FBD1FD95FBE9" box="[159,605,1085,1138]" gridcol="0" gridrow="12" pageId="14" pageNumber="15">Number of ventral scales 4256 between axilla and groin (50.9 ± 3.7)</th>
<td id="82355D867F48ACCFFD27FBD1FC9BFBE9" box="[655,851,1085,1138]" gridcol="1" gridrow="12" pageId="14" pageNumber="15">4765 (55.8 ± 4.8)</td>
<td id="82355D867F48ACCFFCC4FBD1FBDFFBE9" box="[876,1047,1085,1138]" gridcol="2" gridrow="12" pageId="14" pageNumber="15">4356 (47.9 ± 3.8)</td>
<td id="82355D867F48ACCFFB90FBD1FB06FBE9" box="[1080,1230,1085,1138]" gridcol="3" gridrow="12" pageId="14" pageNumber="15">4866 (53.8 ± 5.4)</td>
<td id="82355D867F48ACCFFB5EFBD1FA48FBE9" box="[1270,1408,1085,1138]" gridcol="4" gridrow="12" pageId="14" pageNumber="15">4163 (52.1 ± 5.2)</td>
</tr>
</table>
</paragraph>
<paragraph id="7F6B36B87F485320FF6DF9E0FAD0FA72" blockId="14.[151,1437,1201,1988]" lastBlockId="16.[151,1437,1247,2033]" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<emphasis id="4DA0EAAA7F48533EFF6DF9E0FE04F9BF" bold="true" box="[197,460,1547,1572]" pageId="14" pageNumber="15">Natural history notes.</emphasis>
By far, most specimens were encountered at night sleeping on vegetation, i.e., on the upper surface of leaves or on twigs, usually
<quantity id="B82C9B5D7F48533EFD2DF9C2FD29F9DD" box="[645,737,1582,1606]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="14" pageNumber="15" unit="m" value="1.25" valueMax="2.0" valueMin="0.5">0.52 m</quantity>
above ground. During daytime, specimens were spotted when fleeing on the ground, mostly seeking shelter between tree buttresses or bushes. The vegetation at the
<typeStatus id="A06F881A7F48533EFAB8F9BEFA88F9F1" box="[1296,1344,1618,1642]" pageId="14" pageNumber="15">type</typeStatus>
locality (
<figureCitation id="E7EF2A3D7F48533EFF37F998FF27F917" box="[159,239,1652,1676]" captionStart="FIGURE 8" captionStartId="15.[151,250,1789,1812]" captionTargetBox="[258,1328,193,1767]" captionTargetId="figure@15.[258,1329,193,1768]" captionTargetPageId="15" captionText="FIGURE 8. Habitats of Anolis benedikti: (A C) Pacific slope in Chiriquí province: (A) view over Jurutungo north toward the continental divide (hidden in clouds), the ranges of A. benedikti and A. pachypus meet in the background; (B) forest trail at Jurutungo, one of the lowest collection sites of A. benedikti on the Pacific slope, 2070 m asl; (C) cloud forest near continental divide east of Cerro Pando, 2460 m asl; (D) international border monument on cattle trail along the continental divide, 2450 m asl; (E H) Caribbean slope in Bocas del Toro province: (E) cloud forest at the type locality, 2310 m asl; (F) cloud forest near the type locality, 2390 m asl; (G) clearing for cattle along tributary to Río Changena, 1810 m asl; (H) Río Changena, the lowest collection site on the Caribbean slope, 1640 m asl." httpUri="https://zenodo.org/record/200712/files/figure.png" pageId="14" pageNumber="15">Figs. 8</figureCitation>
EF) is a lush ridgetop cloud forest rich in palms and with abundant growth of epiphytes that cover virtually every surface. Descending either slope of Cerro Pando, the trees become higher and the canopy cover denser. A detailed description of the forest environments around Cerro Pando was given by
<bibRefCitation id="1B454B497F48533EFB5DF956FA50F94A" author="Myers" box="[1269,1432,1721,1746]" pageId="14" pageNumber="15" refString="Myers, C. W. (1969) The ecological geography of cloud forest in Panama. American Museum Novitates, 2396, 1 - 52." type="journal article" year="1969">Myers (1969)</bibRefCitation>
. Presently, especially below
<quantity id="B82C9B5D7F48533EFE79F930FDE0F96F" box="[465,552,1756,1780]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="m" value="2000.0">2000 m</quantity>
elevation on either slope, major clearings for cattle farming disrupt the forest (
<figureCitation id="E7EF2A3D7F48533EFF37F912FF22F88D" box="[159,234,1790,1814]" captionStart="FIGURE 8" captionStartId="15.[151,250,1789,1812]" captionTargetBox="[258,1328,193,1767]" captionTargetId="figure@15.[258,1329,193,1768]" captionTargetPageId="15" captionText="FIGURE 8. Habitats of Anolis benedikti: (A C) Pacific slope in Chiriquí province: (A) view over Jurutungo north toward the continental divide (hidden in clouds), the ranges of A. benedikti and A. pachypus meet in the background; (B) forest trail at Jurutungo, one of the lowest collection sites of A. benedikti on the Pacific slope, 2070 m asl; (C) cloud forest near continental divide east of Cerro Pando, 2460 m asl; (D) international border monument on cattle trail along the continental divide, 2450 m asl; (E H) Caribbean slope in Bocas del Toro province: (E) cloud forest at the type locality, 2310 m asl; (F) cloud forest near the type locality, 2390 m asl; (G) clearing for cattle along tributary to Río Changena, 1810 m asl; (H) Río Changena, the lowest collection site on the Caribbean slope, 1640 m asl." httpUri="https://zenodo.org/record/200712/files/figure.png" pageId="14" pageNumber="15">Figs. 8</figureCitation>
A and G). In these pastures,
<taxonomicName id="B8D44D3B7F48533EFD82F912FD28F88D" box="[554,736,1790,1814]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F48533EFD82F912FD28F88D" box="[554,736,1790,1814]" italics="true" pageId="14" pageNumber="15">Anolis benedikti</emphasis>
</taxonomicName>
can be found associated with bushes, single trees, or remnants of cut trees. Annual total precipitation at the
<typeStatus id="A06F881A7F48533EFD24F8CEFD74F8A1" box="[652,700,1826,1850]" pageId="14" pageNumber="15">type</typeStatus>
locality is approximately
<quantity id="B82C9B5D7F48533EFC48F8CDFB83F8A2" box="[992,1099,1825,1849]" metricMagnitude="0" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="15" unit="mm" value="2600.0">2600 mm</quantity>
and mean annual temperature approximately 13.6 °C, indicating the presence of the Lower Montane Wet Forest life zone according to the
<bibRefCitation id="1B454B497F48533EFF3FF88AFE95F8E5" author="Holdridge" box="[151,349,1894,1918]" pageId="14" pageNumber="15" refString="Holdridge, L. R. (1967) Life zone ecology. Tropical Science Center, San Jose, Costa Rica, 206 pp." type="book" year="1967">Holdridge (1967)</bibRefCitation>
classification. Our datalogger recordings (N = 205) at the
<typeStatus id="A06F881A7F48533EFBA3F88BFBF3F8E4" box="[1035,1083,1895,1919]" pageId="14" pageNumber="15">type</typeStatus>
locality yielded a temperature range of 11.116.7 °C, with a mean of 12.7 °C and standard deviation ± 1.1 °C. Without exception, our collecting localities of
<taxonomicName id="B8D44D3B7F48533EFEB5F840FE57F85F" box="[285,415,1964,1988]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F48533EFEB5F840FE57F85F" box="[285,415,1964,1988]" italics="true" pageId="14" pageNumber="15">A. benedikti</emphasis>
</taxonomicName>
lie above
<quantity id="B82C9B5D7F48533EFDB8F840FDABF85F" box="[528,611,1964,1988]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="15" unit="m" value="1600.0">1600 m</quantity>
asl on the Caribbean slope and above
<quantity id="B82C9B5D7F48533EFBA0F840FB93F85F" box="[1032,1115,1964,1988]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="15" unit="m" value="1900.0">1900 m</quantity>
asl on the Pacific slope and receive more than
<quantity id="B82C9B5D7F565320FECCFB33FE06FB6C" box="[356,462,1247,1271]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="mm" value="2000.0">2000 mm</quantity>
of annual precipitation, i.e., are assignable to the Lower Montane Wet Forest life zone. In the immediate surroundings of the
<typeStatus id="A06F881A7F565320FD97FAEEFDA7FA81" box="[575,623,1282,1306]" pageId="16" pageNumber="17">type</typeStatus>
locality,
<taxonomicName id="B8D44D3B7F565320FD7FFAEDFC96FA82" box="[727,862,1281,1305]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F565320FD7FFAEDFC96FA82" box="[727,862,1281,1305]" italics="true" pageId="16" pageNumber="17">A. benedikti</emphasis>
</taxonomicName>
was the only reptile species we could find. Reptile species collected on the Caribbean versant along the trail descending the north slope of Cerro Pando (
<figureCitation id="E7EF2A3D7F565320FAA6FAC8FA92FAA7" box="[1294,1370,1316,1340]" captionStart="FIGURE 8" captionStartId="15.[151,250,1789,1812]" captionTargetBox="[258,1328,193,1767]" captionTargetId="figure@15.[258,1329,193,1768]" captionTargetPageId="15" captionText="FIGURE 8. Habitats of Anolis benedikti: (A C) Pacific slope in Chiriquí province: (A) view over Jurutungo north toward the continental divide (hidden in clouds), the ranges of A. benedikti and A. pachypus meet in the background; (B) forest trail at Jurutungo, one of the lowest collection sites of A. benedikti on the Pacific slope, 2070 m asl; (C) cloud forest near continental divide east of Cerro Pando, 2460 m asl; (D) international border monument on cattle trail along the continental divide, 2450 m asl; (E H) Caribbean slope in Bocas del Toro province: (E) cloud forest at the type locality, 2310 m asl; (F) cloud forest near the type locality, 2390 m asl; (G) clearing for cattle along tributary to Río Changena, 1810 m asl; (H) Río Changena, the lowest collection site on the Caribbean slope, 1640 m asl." httpUri="https://zenodo.org/record/200712/files/figure.png" pageId="16" pageNumber="17">Figs. 8</figureCitation>
GH) between
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1640 and
<quantity id="B82C9B5D7F565320FEDCFAABFE64FAC4" box="[372,428,1351,1375]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="m" value="2000.0">2000</quantity>
</date>
m asl were
<taxonomicName id="B8D44D3B7F565320FDE9FAABFCF5FAC4" authority="Peters" authorityName="Peters" box="[577,829,1351,1375]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="capito">
<emphasis id="4DA0EAAA7F565320FDE9FAABFD12FAC4" box="[577,730,1351,1375]" italics="true" pageId="16" pageNumber="17">Anolis capito</emphasis>
(Peters)
</taxonomicName>
,
<taxonomicName id="B8D44D3B7F565320FCE4FAABFBEEFAC4" authority="Peters" authorityName="Peters" box="[844,1062,1351,1375]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="humilis">
<emphasis id="4DA0EAAA7F565320FCE4FAABFC09FAC4" box="[844,961,1351,1375]" italics="true" pageId="16" pageNumber="17">A. humilis</emphasis>
(Peters)
</taxonomicName>
,
<taxonomicName id="B8D44D3B7F565320FB9DFAABFAC2FAC4" authority="Dunn" authorityName="Dunn" box="[1077,1290,1351,1375]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kemptoni">
<emphasis id="4DA0EAAA7F565320FB9DFAABFB76FAC4" box="[1077,1214,1351,1375]" italics="true" pageId="16" pageNumber="17">A. kemptoni</emphasis>
Dunn
</taxonomicName>
,
<taxonomicName id="B8D44D3B7F565320FAB0FAABFF2FFA1A" authority="Cope" authorityName="Cope" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="microtus">
<emphasis id="4DA0EAAA7F565320FAB0FAABFA54FAC4" box="[1304,1436,1351,1375]" italics="true" pageId="16" pageNumber="17">A. microtus</emphasis>
(Cope)
</taxonomicName>
,
<taxonomicName id="B8D44D3B7F565320FF5CFA85FD96FA1A" authority="Taylor" authorityName="Taylor" box="[244,606,1385,1409]" class="Reptilia" family="Gymnophthalmidae" genus="Ptychoglossus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="plicatus">
<emphasis id="4DA0EAAA7F565320FF5CFA85FE3FFA1A" box="[244,503,1385,1409]" italics="true" pageId="16" pageNumber="17">Ptychoglossus plicatus</emphasis>
(Taylor)
</taxonomicName>
, and
<taxonomicName id="B8D44D3B7F565320FD34FA85FC18FA1A" authority="Gunther" authorityName="Gunther" box="[668,976,1385,1409]" class="Reptilia" family="Colubridae" genus="Sibon" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="4DA0EAAA7F565320FD34FA85FC9BFA1A" box="[668,851,1385,1409]" italics="true" pageId="16" pageNumber="17">Sibon annulatus</emphasis>
(Günther)
</taxonomicName>
. Reptile species collected on the Pacific versant between
<date id="0B6A10787F565320FEF2FA60FDCFFA3F" box="[346,519,1420,1444]" pageId="16" pageNumber="17" value="2000" valueMax="2400">
2000 and
<quantity id="B82C9B5D7F565320FE67FA60FDCFFA3F" box="[463,519,1420,1444]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="16" pageNumber="17" unit="m" value="2400.0">2400</quantity>
</date>
m asl, in the high valley known as Jurutungo (
<figureCitation id="E7EF2A3D7F565320FB9CFA60FBB4FA3F" box="[1076,1148,1420,1444]" captionStart="FIGURE 8" captionStartId="15.[151,250,1789,1812]" captionTargetBox="[258,1328,193,1767]" captionTargetId="figure@15.[258,1329,193,1768]" captionTargetPageId="15" captionText="FIGURE 8. Habitats of Anolis benedikti: (A C) Pacific slope in Chiriquí province: (A) view over Jurutungo north toward the continental divide (hidden in clouds), the ranges of A. benedikti and A. pachypus meet in the background; (B) forest trail at Jurutungo, one of the lowest collection sites of A. benedikti on the Pacific slope, 2070 m asl; (C) cloud forest near continental divide east of Cerro Pando, 2460 m asl; (D) international border monument on cattle trail along the continental divide, 2450 m asl; (E H) Caribbean slope in Bocas del Toro province: (E) cloud forest at the type locality, 2310 m asl; (F) cloud forest near the type locality, 2390 m asl; (G) clearing for cattle along tributary to Río Changena, 1810 m asl; (H) Río Changena, the lowest collection site on the Caribbean slope, 1640 m asl." httpUri="https://zenodo.org/record/200712/files/figure.png" pageId="16" pageNumber="17">Figs 8</figureCitation>
AB) and along the trail ascending the south slope of Cerro Pando from Jurutungo, were
<taxonomicName id="B8D44D3B7F565320FCCAFA43FC2EFA5C" box="[866,998,1455,1479]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kemptoni">
<emphasis id="4DA0EAAA7F565320FCCAFA43FC2EFA5C" box="[866,998,1455,1479]" italics="true" pageId="16" pageNumber="17">A. kemptoni</emphasis>
</taxonomicName>
,
<emphasis id="4DA0EAAA7F565320FC5CFA43FA54FA5C" box="[1012,1436,1455,1479]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="B8D44D3B7F565320FC5CFA43FACEFA5C" authority="Sceloporus" authorityName="Sceloporus" box="[1012,1286,1455,1479]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="pachypus">A. pachypus, Sceloporus</taxonomicName>
malachiticus
</emphasis>
Cope,
<taxonomicName id="B8D44D3B7F565320FF48FA3DFDEBFA72" authority="Boulenger" authorityName="Boulenger" box="[224,547,1489,1513]" class="Reptilia" family="Colubridae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="godmani">
<emphasis id="4DA0EAAA7F565320FF48FA3DFE6EFA72" box="[224,422,1489,1513]" italics="true" pageId="16" pageNumber="17">Geophis godmani</emphasis>
Boulenger
</taxonomicName>
,
<taxonomicName id="B8D44D3B7F565320FD87FA3DFCB6FA72" authority="Cope" authorityName="Cope" box="[559,894,1489,1513]" class="Reptilia" family="Colubridae" genus="Rhadinaea" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="calligaster">
<emphasis id="4DA0EAAA7F565320FD87FA3DFCEFFA72" box="[559,807,1489,1513]" italics="true" pageId="16" pageNumber="17">Rhadinaea calligaster</emphasis>
(Cope)
</taxonomicName>
, and
<taxonomicName id="B8D44D3B7F565320FC11FA3DFAD0FA72" authority="Peters." authorityName="Peters." box="[953,1304,1489,1513]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nigroviridis">
<emphasis id="4DA0EAAA7F565320FC11FA3DFB0CFA72" box="[953,1220,1489,1513]" italics="true" pageId="16" pageNumber="17">Bothriechis nigroviridis</emphasis>
Peters.
</taxonomicName>
</paragraph>
<caption id="2BAB66307F49533FFF3FF911FD5EF850" httpUri="https://zenodo.org/record/200712/files/figure.png" pageId="15" pageNumber="16" targetBox="[258,1328,193,1767]" targetPageId="15">
<paragraph id="7F6B36B87F49533FFF3FF911FD5EF850" blockId="15.[151,1437,1789,1995]" pageId="15" pageNumber="16">
<emphasis id="4DA0EAAA7F49533FFF3FF911FEDBF889" bold="true" box="[151,275,1789,1812]" pageId="15" pageNumber="16">FIGURE 8.</emphasis>
Habitats of
<emphasis id="4DA0EAAA7F49533FFE39F911FDF6F888" box="[401,574,1789,1811]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="B8D44D3B7F49533FFE39F911FDF1F888" box="[401,569,1789,1811]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="benedikti">Anolis benedikti</taxonomicName>
:
</emphasis>
(AC) Pacific slope in Chiriquí province: (A) view over Jurutungo north toward the continental divide (hidden in clouds), the ranges of
<taxonomicName id="B8D44D3B7F49533FFD1AF8F1FCE5F8A9" box="[690,813,1820,1842]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F49533FFD1AF8F1FCE5F8A9" box="[690,813,1820,1842]" italics="true" pageId="15" pageNumber="16">A. benedikti</emphasis>
</taxonomicName>
and
<taxonomicName id="B8D44D3B7F49533FFCCAF8F1FC28F8A9" box="[866,992,1820,1842]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F49533FFCCAF8F1FC28F8A9" box="[866,992,1820,1842]" italics="true" pageId="15" pageNumber="16">A. pachypus</emphasis>
</taxonomicName>
meet in the background; (B) forest trail at Jurutungo, one of the lowest collection sites of
<taxonomicName id="B8D44D3B7F49533FFDD3F8D7FD3DF8CA" box="[635,757,1851,1873]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F49533FFDD3F8D7FD3DF8CA" box="[635,757,1851,1873]" italics="true" pageId="15" pageNumber="16">A. benedikti</emphasis>
</taxonomicName>
on the Pacific slope, 2070 m asl; (C) cloud forest near continental divide east of Cerro Pando, 2460 m asl; (D) international border monument on cattle trail along the continental divide, 2450 m asl; (EH) Caribbean slope in Bocas del Toro province: (E) cloud forest at the type locality, 2310 m asl; (F) cloud forest near the type locality, 2390 m asl; (G) clearing for cattle along tributary to Río Changena, 1810 m asl; (H) Río Changena, the lowest collection site on the Caribbean slope, 1640 m asl.
</paragraph>
</caption>
<caption id="2BAB66307F565320FF3FFB8FFD7FFB2C" httpUri="https://zenodo.org/record/200714/files/figure.png" pageId="16" pageNumber="17" targetBox="[193,1407,225,1073]" targetPageId="16">
<paragraph id="7F6B36B87F565320FF3FFB8FFD7FFB2C" blockId="16.[151,1436,1123,1207]" pageId="16" pageNumber="17">
<emphasis id="4DA0EAAA7F565320FF3FFB8FFED3FBE2" bold="true" box="[151,283,1123,1146]" pageId="16" pageNumber="17">FIGURE 9.</emphasis>
Collection localities of
<taxonomicName id="B8D44D3B7F565320FD9AFB88FD16FBE1" box="[562,734,1124,1146]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F565320FD9AFB88FD16FBE1" box="[562,734,1124,1146]" italics="true" pageId="16" pageNumber="17">Anolis benedikti</emphasis>
</taxonomicName>
(circles) and other members of the
<taxonomicName id="B8D44D3B7F565320FB2CFB89FAC1FBE1" box="[1156,1289,1124,1146]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F565320FB2CFB89FAC1FBE1" box="[1156,1289,1124,1146]" italics="true" pageId="16" pageNumber="17">A. pachypus</emphasis>
</taxonomicName>
complex (
<taxonomicName id="B8D44D3B7F565320FA2EFB89FEEEFB02" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="magnaphallus">
<emphasis id="4DA0EAAA7F565320FA2EFB89FEEEFB02" italics="true" pageId="16" pageNumber="17">A. magnaphallus</emphasis>
</taxonomicName>
: squares;
<taxonomicName id="B8D44D3B7F565320FE24FB6FFDC0FB02" box="[396,520,1155,1177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F565320FE24FB6FFDC0FB02" box="[396,520,1155,1177]" italics="true" pageId="16" pageNumber="17">A. pachypus</emphasis>
</taxonomicName>
: upright triangles;
<taxonomicName id="B8D44D3B7F565320FD61FB6FFC44FB02" box="[713,908,1155,1177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="pseudopachypus">
<emphasis id="4DA0EAAA7F565320FD61FB6FFC44FB02" box="[713,908,1155,1177]" italics="true" pageId="16" pageNumber="17">A. pseudopachypus</emphasis>
</taxonomicName>
: inverted triangles) in western Panama and adjacent Costa Rica. Hollow symbols represent type localities.
</paragraph>
</caption>
<paragraph id="7F6B36B87F565320FF6DFA1FFEF5F86A" blockId="16.[151,1437,1247,2033]" pageId="16" pageNumber="17">
<emphasis id="4DA0EAAA7F565320FF6DFA1FFE38F997" bold="true" box="[197,496,1523,1548]" pageId="16" pageNumber="17">Geographic distribution.</emphasis>
<collectingCountry id="07C376287F565320FE53FA18FDD2F997" box="[507,538,1524,1548]" name="American Samoa" pageId="16" pageNumber="17">As</collectingCountry>
presently understood,
<taxonomicName id="B8D44D3B7F565320FC8BFA18FC13F997" box="[803,987,1524,1548]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F565320FC8BFA18FC13F997" box="[803,987,1524,1548]" italics="true" pageId="16" pageNumber="17">Anolis benedikti</emphasis>
</taxonomicName>
inhabits montane elevations along the Talamancan highlands of eastern
<collectingCountry id="07C376287F565320FDBCF9FBFD58F9B4" box="[532,656,1559,1583]" name="Costa Rica" pageId="16" pageNumber="17">Costa Rica</collectingCountry>
and extreme western
<collectingCountry id="07C376287F565320FC25F9F4FC2EF9B4" box="[909,998,1560,1583]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
(
<figureCitation id="E7EF2A3D7F565320FC5FF9FBFBF5F9B4" box="[1015,1085,1559,1583]" captionStart="FIGURE 9" captionStartId="16.[151,250,1123,1146]" captionTargetBox="[193,1407,225,1073]" captionTargetId="figure@16.[151,1436,193,1102]" captionTargetPageId="16" captionText="FIGURE 9. Collection localities of Anolis benedikti (circles) and other members of the A. pachypus complex (A. magnaphallus: squares; A. pachypus: upright triangles; A. pseudopachypus: inverted triangles) in western Panama and adjacent Costa Rica. Hollow symbols represent type localities." httpUri="https://zenodo.org/record/200714/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
). All individuals we examined originated from between
<geoCoordinate id="1AE0507F7F565320FE1DF9D5FDF8F9CA" box="[437,560,1593,1617]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="55" value="-82.728">82.728 °W</geoCoordinate>
and
<geoCoordinate id="1AE0507F7F565320FDC0F9D5FD2CF9CA" box="[616,740,1593,1617]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="55" value="-82.425">82.425 °W</geoCoordinate>
. The species seems to dwell predominantly on the Caribbean versant of the Serranía de Talamanca, where we found it from the continental divide at up to
<quantity id="B82C9B5D7F565320FB63F9B0FAEAF9EF" box="[1227,1314,1628,1652]" metricMagnitude="3" metricUnit="m" metricValue="2.3899999999999997" pageId="16" pageNumber="17" unit="m" value="2390.0">2390 m</quantity>
asl (Cerro Pando). Descending the Caribbean slope, we encountered it down to
<quantity id="B82C9B5D7F565320FC0DF993FC33F90C" box="[933,1019,1663,1687]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="16" pageNumber="17" unit="m" value="1640.0">1640 m</quantity>
asl at the Río Changena,
<quantity id="B82C9B5D7F565320FAB7F993FABDF90C" box="[1311,1397,1663,1687]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="16" pageNumber="17" unit="m" value="1740.0">1740 m</quantity>
asl along the Culebra trail, and
<quantity id="B82C9B5D7F565320FE70F94DFDE7F922" box="[472,559,1697,1721]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="16" pageNumber="17" unit="m" value="1650.0">1650 m</quantity>
asl along the Pianista trail. However, in at least three regions, this species is also found on the Pacific versant, where it occurs next to its close relatives: At Las Tablas, in the Costa Rican Province of Puntarenas next to the Panamanian border, SMF 92134 was found at
<quantity id="B82C9B5D7F565320FB82F90BFBB7F964" box="[1066,1151,1767,1791]" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="16" pageNumber="17" unit="m" value="1960.0">1960 m</quantity>
asl, less than a kilometer from the nearest collected
<taxonomicName id="B8D44D3B7F565320FE6DF8E5FD87F8BA" box="[453,591,1801,1825]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F565320FE6DF8E5FD87F8BA" box="[453,591,1801,1825]" italics="true" pageId="16" pageNumber="17">A. pachypus</emphasis>
</taxonomicName>
at
<quantity id="B82C9B5D7F565320FDDBF8E5FD01F8BA" box="[627,713,1801,1825]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="16" pageNumber="17" unit="m" value="1880.0">1880 m</quantity>
asl. Just a few kilometers east, in Jurutungo (the headwaters of Río Candela in Chiriquí Province,
<collectingCountry id="07C376287F565320FDB5F8C1FDB1F8DF" box="[541,633,1837,1860]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
;
<figureCitation id="E7EF2A3D7F565320FD2CF8C0FD0DF8DF" box="[644,709,1836,1860]" captionStart="FIGURE 8" captionStartId="15.[151,250,1789,1812]" captionTargetBox="[258,1328,193,1767]" captionTargetId="figure@15.[258,1329,193,1768]" captionTargetPageId="15" captionText="FIGURE 8. Habitats of Anolis benedikti: (A C) Pacific slope in Chiriquí province: (A) view over Jurutungo north toward the continental divide (hidden in clouds), the ranges of A. benedikti and A. pachypus meet in the background; (B) forest trail at Jurutungo, one of the lowest collection sites of A. benedikti on the Pacific slope, 2070 m asl; (C) cloud forest near continental divide east of Cerro Pando, 2460 m asl; (D) international border monument on cattle trail along the continental divide, 2450 m asl; (E H) Caribbean slope in Bocas del Toro province: (E) cloud forest at the type locality, 2310 m asl; (F) cloud forest near the type locality, 2390 m asl; (G) clearing for cattle along tributary to Río Changena, 1810 m asl; (H) Río Changena, the lowest collection site on the Caribbean slope, 1640 m asl." httpUri="https://zenodo.org/record/200712/files/figure.png" pageId="16" pageNumber="17">Fig. 8</figureCitation>
A), we found
<taxonomicName id="B8D44D3B7F565320FCCAF8C0FC20F8DF" box="[866,1000,1836,1860]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F565320FCCAF8C0FC20F8DF" box="[866,1000,1836,1860]" italics="true" pageId="16" pageNumber="17">A. benedikti</emphasis>
</taxonomicName>
as low as
<quantity id="B82C9B5D7F565320FBF6F8C0FB7BF8DF" box="[1118,1203,1836,1860]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="16" pageNumber="17" unit="m" value="2020.0">2020 m</quantity>
asl at certain places, less than
<quantity id="B82C9B5D7F565320FEACF8A3FEF7F8FC" box="[260,319,1871,1895]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="m" value="50.0">50 m</quantity>
away from the nearest collection site of
<taxonomicName id="B8D44D3B7F565320FCB2F8A3FC6DF8FC" box="[794,933,1871,1895]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="pachypus">
<emphasis id="4DA0EAAA7F565320FCB2F8A3FC6DF8FC" box="[794,933,1871,1895]" italics="true" pageId="16" pageNumber="17">A. pachypus</emphasis>
</taxonomicName>
. Further east, on the slopes of Volcán Barú north of Boquete, about one kilometer west of the ranger station of Alto Chiquero close to the upper Río Caldera, we collected four males of
<taxonomicName id="B8D44D3B7F565320FE62F878FD99F837" box="[458,593,1940,1964]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="benedikti">
<emphasis id="4DA0EAAA7F565320FE62F878FD99F837" box="[458,593,1940,1964]" italics="true" pageId="16" pageNumber="17">A. benedikti</emphasis>
</taxonomicName>
at
<quantity id="B82C9B5D7F565320FDDCF878FD01F837" box="[628,713,1940,1964]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="16" pageNumber="17" unit="m" value="1920.0">1920 m</quantity>
asl, as well as a single female farther south. These two sites are close to different collection sites (at both lower and higher elevations) as well as to the
<typeStatus id="A06F881A7F565320FB6AF854FB3AF84B" box="[1218,1266,1976,2000]" pageId="16" pageNumber="17">type</typeStatus>
locality of
<emphasis id="4DA0EAAA7F565320FA2CF85BFEF5F86A" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="B8D44D3B7F565320FA2CF85BFEF1F86A" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="magnaphallus">A. magnaphallus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="37CE65337F575321FF6DFF74FBB8FF4F" pageId="17" pageNumber="18" type="etymology">
<paragraph id="7F6B36B87F575321FF6DFF74FBB8FF4F" blockId="17.[151,1436,151,212]" pageId="17" pageNumber="18">
<emphasis id="4DA0EAAA7F575321FF6DFF74FE83FF2B" bold="true" box="[197,331,152,176]" pageId="17" pageNumber="18">Etymology.</emphasis>
The specific name is a patronym for Benedikt Stökl,
<collectingCountry id="07C376287F575321FC60FF7BFBFAFF34" box="[968,1074,151,175]" name="Germany" pageId="17" pageNumber="18">Germany</collectingCountry>
, in recognition of the financial support of taxonomic research provided by Karsten Lutz through the BIOPAT initiative.
</paragraph>
</subSubSection>
</treatment>
</document>