treatments-xml/data/F7/3D/87/F73D87B3FFF46560FC7AEA75FEE1B373.xml

<document id="1A8FFB668D0384C1197915A0B957CF45" ID-DOI="10.5281/zenodo.158566" ID-GBIF-Dataset="ac5a8900-3ed1-4993-82a2-3de55f724002" ID-ISSN="11755326" ID-Zenodo-Dep="158566" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461038541503" checkinUser="plazi" docAuthor="Jacobus, Luke M. &amp; Sartori, Michel" docDate="2004" docId="F73D87B3FFF46560FC7AEA75FEE1B373" docLanguage="en" docName="zt00785.pdf" docOrigin="Zootaxa 785" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hyrtanella pascalae Jacobus &amp; Sartori, 2004, new species" docType="treatment" docVersion="10" lastPageNumber="10" masterDocId="0B04FFCBFFF66569FD72EC31FF80B43D" masterDocTitle="Review of the genus Hyrtanella (Ephemeroptera: Ephemerellidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1698575045907" updateUser="plazi">
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<mods:title id="3620348C52A730A3AE9A17C29566CCBE">Review of the genus Hyrtanella (Ephemeroptera: Ephemerellidae)</mods:title>
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<paragraph id="7F2B36A5FFF4656BFC7AEA75FD18B260" blockId="2.[264,664,1603,1665]" box="[264,664,1603,1630]" pageId="2" pageNumber="3">
<heading id="246381C9FFF4656BFC7AEA75FD18B260" bold="true" box="[264,664,1603,1630]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="4DE0EAB7FFF4656BFC7AEA75FD18B260" bold="true" box="[264,664,1603,1630]" pageId="2" pageNumber="3">
<taxonomicName id="B8944D26FFF4656BFC7AEA75FE79B263" ID-CoL="3P5K2" box="[264,505,1604,1630]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="pascalae" status="sp. nov.">
<emphasis id="4DE0EAB7FFF4656BFC7AEA75FE79B263" bold="true" box="[264,505,1604,1630]" italics="true" pageId="2" pageNumber="3">Hyrtanella pascalae</emphasis>
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,
<taxonomicNameLabel id="56D357CCFFF4656BFF7AEA72FD18B260" box="[520,664,1603,1629]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
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<paragraph id="7F2B36A5FFF4656BFC7AEA56FE4DB2BC" blockId="2.[264,664,1603,1665]" box="[264,461,1639,1665]" pageId="2" pageNumber="3">
<figureCitation id="E7AF2A20FFF4656BFC7AEA56FEF7B2BC" box="[264,375,1639,1665]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="2" pageNumber="3">Figs. 1–8</figureCitation>
,
<figureCitation id="E7AF2A20FFF4656BFCF6EA56FE4DB2BC" box="[388,461,1639,1665]" captionStart-0="FIGURES 10 – 14" captionStart-1="FIGURES 15 – 18" captionStart-2="FIGURES 19 – 20" captionStartId-0="5.[264,382,1620,1644]" captionStartId-1="6.[264,382,1784,1808]" captionStartId-2="8.[264,382,1002,1026]" captionTargetBox-0="[283,1286,320,1607]" captionTargetBox-1="[269,1316,317,1772]" captionTargetBox-2="[264,1322,283,989]" captionTargetId-0="figure@5.[640,1159,796,1112]" captionTargetId-1="figure@6.[269,1317,317,1047]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionText-0="FIGURES 10 – 14. Hyrtanella pascalae, n. sp., larvae: Fig 10. Foreleg. Fig. 11. Detail of apex of tibia. Fig. 12. Detail of foreclaw. Fig. 13. Middle femur. Fig. 14. Hind femur." captionText-1="FIGURES 15 – 18. Hyrtanella pascalae, n. sp., male subimago: 15 Photograph of specimen in alcohol. 16. Forewing. 17. Hindwing. 18. Genitalia, ventral view." captionText-2="FIGURES 19 – 20. Eggs (scale bars = 10 m): 19. Hyrtanella pascalae, n. sp. 20. Undetermined Ephemerellinae, West Malaysia." httpUri-0="https://zenodo.org/record/158568/files/figure.png" httpUri-1="https://zenodo.org/record/169630/files/figure.png" httpUri-2="https://zenodo.org/record/169631/files/figure.png" pageId="2" pageNumber="3">10–19</figureCitation>
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<subSubSection id="378E652EFFF4656EFC7AEA87FB44B0C6" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3" type="description">
<paragraph id="7F2B36A5FFF4656BFC7AEA87FD3EB2C4" blockId="2.[264,1324,1718,1905]" pageId="2" pageNumber="3">
<emphasis id="4DE0EAB7FFF4656BFC7AEA87FE2CB2ED" bold="true" box="[264,428,1718,1744]" pageId="2" pageNumber="3">Descriptions.</emphasis>
<emphasis id="4DE0EAB7FFF4656BFCCBEA86FE7FB2ED" box="[441,511,1719,1744]" italics="true" pageId="2" pageNumber="3">Larva</emphasis>
(
<figureCitation id="E7AF2A20FFF4656BFF67EA86FD0AB2EC" box="[533,650,1719,1745]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="2" pageNumber="3">Figs. 1–8</figureCitation>
,
<figureCitation id="E7AF2A20FFF4656BFFEFEA86FD65B2EC" box="[669,741,1719,1745]" captionStart="FIGURES 10 – 14" captionStartId="5.[264,382,1620,1644]" captionTargetBox="[283,1286,320,1607]" captionTargetId="figure@5.[640,1159,796,1112]" captionTargetPageId="5" captionText="FIGURES 10 – 14. Hyrtanella pascalae, n. sp., larvae: Fig 10. Foreleg. Fig. 11. Detail of apex of tibia. Fig. 12. Detail of foreclaw. Fig. 13. Middle femur. Fig. 14. Hind femur." httpUri="https://zenodo.org/record/158568/files/figure.png" pageId="2" pageNumber="3">10–14</figureCitation>
)—Length: body 4.0–
<quantity id="B86C9B40FFF4656BFE83EA86FBD4B2EC" box="[1009,1108,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="3" unit="mm" value="4.4">4.4 mm</quantity>
; caudal filaments
<quantity id="B86C9B40FFF4656BFC7AEAEEFE18B2C4" box="[264,408,1759,1785]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.1" metricValueMin="1.4" pageId="2" pageNumber="3" unit="mm" value="1.75" valueMax="2.1" valueMin="1.4">1.4–2.1 mm</quantity>
. Body color light brown.
</paragraph>
<paragraph id="7F2B36A5FFF4656AFC4AEB36FE5DB6D3" blockId="2.[264,1324,1718,1905]" lastBlockId="3.[264,1324,284,1870]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Head: vertex with two pairs of short suboccipital spines. No ocellar bumps. Inner margin of antennal base with short projections. Frontal shelf not expanded. Genae slightly produced. Long hairlike setae present along anterior margin of frontal shelf and sparsely present below compound eye. Clypeus with anterior row of long hairlike setae. Labrum (
<figureCitation id="E7AF2A20FFF5656AFC63ED75FED4B563" box="[273,340,324,350]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
) width nearly
<date id="0B2A1065FFF5656AFF73ED75FD9EB563" box="[513,542,324,350]" pageId="3" pageNumber="4">2x</date>
length, with shallow, very broad notch anteriorly; dorsal face with scattered long hairlike setae on lateral margin and with several rows of brushlike setae anteriorly (
<figureCitation id="E7AF2A20FFF5656AFCF8EDA5FE51B593" box="[394,465,404,430]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
); ventral face with simple or fimbriate setae on lateral margins and dense transverse row of 7–9 long, stout setae (
<figureCitation id="E7AF2A20FFF5656AFE71ED8DFCD0B5EB" box="[771,848,444,470]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
). Hypopharynx with superlinguae size nearly subequal to linguae size (
<figureCitation id="E7AF2A20FFF5656AFFF1EDD5FD4CB5C3" box="[643,716,484,510]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
); lingua with row of short setae on anterior margin; superlingua with row of long, thin setae on margin, with setae decreasing in length anteriorly. Mandibles (
<figureCitation id="E7AF2A20FFF5656AFCA7EE05FDC9B673" box="[469,585,564,590]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="3" pageNumber="4">Figs. 5, 6</figureCitation>
) with lateral setae relatively long. Maxilla (
<figureCitation id="E7AF2A20FFF5656AF92CEE05FB27B673" box="[1118,1191,564,590]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
) with palp reduced; apex of galea relatively sharp with 3 incisors, and with no setae distally on outer margin; lacinia with 2 stout spinous processes (dentisetae) and with double row of 4 stout setae. Labium (
<figureCitation id="E7AF2A20FFF5656AFCB4EE9DFD90B6FB" box="[454,528,684,710]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
) reduced; palp segments short; terminal segment length about 3 times basal width.
</paragraph>
<paragraph id="7F2B36A5FFF5656AFC4AEECDFB62B123" blockId="3.[264,1324,284,1870]" pageId="3" pageNumber="4">
Thorax: Prothorax width subequal to head width, with no anterolateral projections and no dorsal spines; anterior margin upturned and not receded; ventral spines absent. Mesonotum with forewingpads base darkly pigmented and no spine between forewingpads. Coxae with slight dorsolateral projections; projections with hairlike setae. Legs with marginal long hairlike setae; femora generally flattened. Forefemur (
<figureCitation id="E7AF2A20FFF5656AF94AEFADFB0DB78B" box="[1080,1165,924,950]" captionStart="FIGURES 10 – 14" captionStartId="5.[264,382,1620,1644]" captionTargetBox="[283,1286,320,1607]" captionTargetId="figure@5.[640,1159,796,1112]" captionTargetPageId="5" captionText="FIGURES 10 – 14. Hyrtanella pascalae, n. sp., larvae: Fig 10. Foreleg. Fig. 11. Detail of apex of tibia. Fig. 12. Detail of foreclaw. Fig. 13. Middle femur. Fig. 14. Hind femur." httpUri="https://zenodo.org/record/158568/files/figure.png" pageId="3" pageNumber="4">Fig. 10</figureCitation>
) with incomplete narrow transverse band of dorsally projecting bristlelike setae; dorsal edge with single stout bristlelike seta medially; ventral edge with few chalazae medially; anterior face with no dorsal projections; foretibial projection short (
<figureCitation id="E7AF2A20FFF5656AFEF2E825FC56B013" box="[896,982,1044,1070]" captionStart="FIGURES 10 – 14" captionStartId="5.[264,382,1620,1644]" captionTargetBox="[283,1286,320,1607]" captionTargetId="figure@5.[640,1159,796,1112]" captionTargetPageId="5" captionText="FIGURES 10 – 14. Hyrtanella pascalae, n. sp., larvae: Fig 10. Foreleg. Fig. 11. Detail of apex of tibia. Fig. 12. Detail of foreclaw. Fig. 13. Middle femur. Fig. 14. Hind femur." httpUri="https://zenodo.org/record/158568/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
). Claws bent, with short subdistal setae; palisade of elongate denticles on inner distolateral margin and few denticles present proximally (
<figureCitation id="E7AF2A20FFF5656AFC8AE855FDD1B043" box="[504,593,1124,1150]" captionStart="FIGURES 10 – 14" captionStartId="5.[264,382,1620,1644]" captionTargetBox="[283,1286,320,1607]" captionTargetId="figure@5.[640,1159,796,1112]" captionTargetPageId="5" captionText="FIGURES 10 – 14. Hyrtanella pascalae, n. sp., larvae: Fig 10. Foreleg. Fig. 11. Detail of apex of tibia. Fig. 12. Detail of foreclaw. Fig. 13. Middle femur. Fig. 14. Hind femur." httpUri="https://zenodo.org/record/158568/files/figure.png" pageId="3" pageNumber="4">Fig. 12</figureCitation>
). Dorsal edge of midfemur (
<figureCitation id="E7AF2A20FFF5656AFED6E855FC7BB043" box="[932,1019,1124,1150]" captionStart="FIGURES 10 – 14" captionStartId="5.[264,382,1620,1644]" captionTargetBox="[283,1286,320,1607]" captionTargetId="figure@5.[640,1159,796,1112]" captionTargetPageId="5" captionText="FIGURES 10 – 14. Hyrtanella pascalae, n. sp., larvae: Fig 10. Foreleg. Fig. 11. Detail of apex of tibia. Fig. 12. Detail of foreclaw. Fig. 13. Middle femur. Fig. 14. Hind femur." httpUri="https://zenodo.org/record/158568/files/figure.png" pageId="3" pageNumber="4">Fig. 13</figureCitation>
) with single stout bristlelike seta, but seta situated more distally than similar seta of forefemora; ventral edge with numerous chalazae, with proximal chalazae bearing long, bristlelike setae and distal chalazae with blunt, stout setae. Dorsal edge of hindfemur (
<figureCitation id="E7AF2A20FFF5656AFEE7E8EDFC69B0CB" box="[917,1001,1244,1270]" captionStart="FIGURES 10 – 14" captionStartId="5.[264,382,1620,1644]" captionTargetBox="[283,1286,320,1607]" captionTargetId="figure@5.[640,1159,796,1112]" captionTargetPageId="5" captionText="FIGURES 10 – 14. Hyrtanella pascalae, n. sp., larvae: Fig 10. Foreleg. Fig. 11. Detail of apex of tibia. Fig. 12. Detail of foreclaw. Fig. 13. Middle femur. Fig. 14. Hind femur." httpUri="https://zenodo.org/record/158568/files/figure.png" pageId="3" pageNumber="4">Fig. 14</figureCitation>
) with 6–8 chalazae bearing long, bristlelike setae; ventral edge with only blunt and stout chalazae on distal half.
</paragraph>
<paragraph id="7F2B36A5FFF5656AFC4AE91DFC18B373" blockId="3.[264,1324,284,1870]" pageId="3" pageNumber="4">
Abdomen: Gills similar to those of
<taxonomicName id="B8944D26FFF5656AFFA4E91DFCFCB178" box="[726,892,1324,1349]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="christineae">
<emphasis id="4DE0EAB7FFF5656AFFA4E91DFCFCB178" box="[726,892,1324,1349]" italics="true" pageId="3" pageNumber="4">H. christineae</emphasis>
</taxonomicName>
(see
<bibRefCitation id="1B054B54FFF5656AFEC8E91DFBC7B17B" author="Kluge" box="[954,1095,1324,1350]" pageId="3" pageNumber="12" refString="Kluge, N. (2004) The Phylogenetic System of Ephemeroptera, Kluwer, Dordrecht, The Netherlands, 442 pp." type="book" year="2004">Kluge 2004</bibRefCitation>
: Figs. 94a–e). Gills 3 brown with no apparent distal pigmentation; gills 4–7 translucent. Gills 3 operculate, with medial transverse weak band; gills 4 operculate. Dorsal lamellae of gills 4–6 with strong projection on inner proximal margin; projection folded dorsally; gills with slight projection on outer proximal margin. Dorsal lamellae truncate apically. Gills 3–6 ventral lamellae bifid, with dorsal and ventral lobules; lobules tips acute. Gills 7 narrow; point of attachment medial. Terga 1 and 2 with long hairlike setae on posterior margins. Terga 3–7 or 4–7 with pair of blunt oblique ridges and with short, blunt, paired spines on posterior margins; spines and ridges form inner portion of prominent gill chamber. Tergum 8 sculpted posterolaterally, and with single, short, blunt, median spine and blunt posterolateral projections. Tergum 9 with single, short, blunt, median spine and weak oblique ridges. Segments 6–8 with lateral margins strongly upturned, forming outer portion of prominent gill chamber. Abdominal sterna flattened, with no maculation and no long setae. Caudal filaments pale, with dense whorls of long hairlike setae.
</paragraph>
<caption id="2BEB662DFFF2656DFC7AEADDFB31B356" httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="4" pageNumber="5" targetBox="[265,1321,327,1759]" targetPageId="4">
<paragraph id="7F2B36A5FFF2656DFC7AEADDFB31B356" blockId="4.[264,1323,1772,1900]" pageId="4" pageNumber="5">
<emphasis id="4DE0EAB7FFF2656DFC7AEADDFDA4B339" bold="true" box="[264,548,1772,1796]" pageId="4" pageNumber="5">FIGURES 1–9. Figs 1–8.</emphasis>
<taxonomicName id="B8944D26FFF2656DFF5EEADCFC87B339" box="[556,775,1773,1796]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pascalae" status="sp. nov.">
<emphasis id="4DE0EAB7FFF2656DFF5EEADCFC87B339" box="[556,775,1773,1796]" italics="true" pageId="4" pageNumber="5">Hyrtanella pascalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="56D357CCFFF2656DFE64EADDFCD2B33E" box="[790,850,1772,1795]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
, larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium.
<emphasis id="4DE0EAB7FFF2656DFF35EB65FD0EB351" bold="true" box="[583,654,1876,1900]" pageId="4" pageNumber="5">Fig. 9.</emphasis>
<taxonomicName id="B8944D26FFF2656DFFE6EB64FC04B351" box="[660,900,1877,1900]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="christineae">
<emphasis id="4DE0EAB7FFF2656DFFE6EB64FC04B351" box="[660,900,1877,1900]" italics="true" pageId="4" pageNumber="5">Hyrtanella christineae</emphasis>
</taxonomicName>
, larva: Detail of labial palp.
</paragraph>
</caption>
<caption id="2BEB662DFFF3656CFC7AEA65FBBCB2B3" httpUri="https://zenodo.org/record/158568/files/figure.png" pageId="5" pageNumber="6" targetBox="[283,1286,320,1607]" targetPageId="5">
<paragraph id="7F2B36A5FFF3656CFC7AEA65FBBCB2B3" blockId="5.[264,1322,1620,1678]" pageId="5" pageNumber="6">
<emphasis id="4DE0EAB7FFF3656CFC7AEA65FE50B251" bold="true" box="[264,464,1620,1644]" pageId="5" pageNumber="6">FIGURES 10–14.</emphasis>
<taxonomicName id="B8944D26FFF3656CFCABEA65FD34B256" box="[473,692,1620,1643]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pascalae" status="sp. nov.">
<emphasis id="4DE0EAB7FFF3656CFCABEA65FD34B256" box="[473,692,1620,1643]" italics="true" pageId="5" pageNumber="6">Hyrtanella pascalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="56D357CCFFF3656CFFB1EA65FC80B256" box="[707,768,1620,1643]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
, larvae: Fig 10. Foreleg. Fig. 11. Detail of apex of tibia. Fig. 12. Detail of foreclaw. Fig. 13. Middle femur. Fig. 14. Hind femur.
</paragraph>
</caption>
<paragraph id="7F2B36A5FFF3656CFC4AEADBFD39B311" blockId="5.[264,1322,1770,1836]" pageId="5" pageNumber="6">
<emphasis id="4DE0EAB7FFF3656CFC4AEADBFE73B33E" box="[312,499,1770,1795]" italics="true" pageId="5" pageNumber="6">Male Subimago</emphasis>
(
<figureCitation id="E7AF2A20FFF3656CFF74EADBFD11B339" box="[518,657,1770,1796]" captionStart="FIGURES 15 – 18" captionStartId="6.[264,382,1784,1808]" captionTargetBox="[269,1316,317,1772]" captionTargetId="figure@6.[269,1317,317,1047]" captionTargetPageId="6" captionText="FIGURES 15 – 18. Hyrtanella pascalae, n. sp., male subimago: 15 Photograph of specimen in alcohol. 16. Forewing. 17. Hindwing. 18. Genitalia, ventral view." httpUri="https://zenodo.org/record/169630/files/figure.png" pageId="5" pageNumber="6">Figs. 15–18</figureCitation>
)—Length: body (excluding caudal filaments)
<quantity id="B86C9B40FFF3656CF9B7EADBFAA6B339" box="[1221,1318,1770,1796]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="6" unit="mm" value="5.5">5.5 mm</quantity>
; forewing 5.0 mm; hindwing
<quantity id="B86C9B40FFF3656CFF2AEB23FD36B311" box="[600,694,1810,1836]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="6" unit="mm" value="0.9">0.9 mm</quantity>
.
</paragraph>
<caption id="2BEB662DFFF0656FFC7AEAC9FC1BB30F" httpUri="https://zenodo.org/record/169630/files/figure.png" pageId="6" pageNumber="7" targetBox="[269,1316,317,1772]" targetPageId="6">
<paragraph id="7F2B36A5FFF0656FFC7AEAC9FC1BB30F" blockId="6.[264,1323,1784,1842]" pageId="6" pageNumber="7">
<emphasis id="4DE0EAB7FFF0656FFC7AEAC9FE4EB32D" bold="true" box="[264,462,1784,1808]" pageId="6" pageNumber="7">FIGURES 15–18.</emphasis>
<emphasis id="4DE0EAB7FFF0656FFCA7EAC8FD34B32D" box="[469,692,1785,1808]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="B8944D26FFF0656FFCA7EAC8FD30B32D" box="[469,688,1785,1808]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pascalae" status="sp. nov.">Hyrtanella pascalae</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="56D357CCFFF0656FFFC9EAC9FD75B332" box="[699,757,1784,1807]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
, male subimago: 15 Photograph of specimen in alcohol. 16. Forewing. 17. Hindwing. 18. Genitalia, ventral view.
</paragraph>
</caption>
<paragraph id="7F2B36A5FFF1656EFC4AED2DFDABB593" blockId="7.[264,1325,284,1838]" pageId="7" pageNumber="8">Head: Vertex yellowbrown; antennae with scape and pedicel light yellow, flagellar segments light brown. Ocelli light yellow; median ocellus tinted with grey on posterior margin and lateral ocelli tinted with grey on inner magin. Compound eye with upper part yellow, lower part black.</paragraph>
<paragraph id="7F2B36A5FFF1656EFC4AED8DFD1AB77E" blockId="7.[264,1325,284,1838]" pageId="7" pageNumber="8">
Thorax: Prothorax brown with purple band on proximal margin. Mesoscutellum yellowbrown with light yellow maculae. Mesosterna light yellow with purple narrow longitudinal band medially. Legs yellowbrown, with tarsi pale; each leg with claws dissimilar (one blunt and paddlelike and one hooklike). Forewing (
<figureCitation id="E7AF2A20FFF1656EFEC7EE05FBC7B673" box="[949,1095,564,590]" captionStart="FIGURES 15 – 18" captionStartId="6.[264,382,1784,1808]" captionTargetBox="[269,1316,317,1772]" captionTargetId="figure@6.[269,1317,317,1047]" captionTargetPageId="6" captionText="FIGURES 15 – 18. Hyrtanella pascalae, n. sp., male subimago: 15 Photograph of specimen in alcohol. 16. Forewing. 17. Hindwing. 18. Genitalia, ventral view." httpUri="https://zenodo.org/record/169630/files/figure.png" pageId="7" pageNumber="8">Figs. 15, 16</figureCitation>
) brown, with dark brown basally in subcostal field; few crossveins, mainly in radial and median fields; posterior margin with some irregular intercalaries; one intercalary between IMP and MP2, and one between MP2 and CuA;
<specimenCount id="6992FD2CFFF1656EFF14EE9EFC8EB6F4" box="[614,782,687,713]" pageId="7" pageNumber="8" type="intercalary">3 intercalaries</specimenCount>
between CuA and CuP. Axillary cords (plumidia) long. Hindwing (
<figureCitation id="E7AF2A20FFF1656EFF43EEE8FD46B6CE" box="[561,710,729,755]" captionStart="FIGURES 15 – 18" captionStartId="6.[264,382,1784,1808]" captionTargetBox="[269,1316,317,1772]" captionTargetId="figure@6.[269,1317,317,1047]" captionTargetPageId="6" captionText="FIGURES 15 – 18. Hyrtanella pascalae, n. sp., male subimago: 15 Photograph of specimen in alcohol. 16. Forewing. 17. Hindwing. 18. Genitalia, ventral view." httpUri="https://zenodo.org/record/169630/files/figure.png" pageId="7" pageNumber="8">Figs. 15, 17</figureCitation>
) brown with dark brown basally in subcostal and radial fields; crossveins absent; costa with welldeveloped process at middle; tip of process angled away from wing base.
</paragraph>
<paragraph id="7F2B36A5FFF1656EFC4AEF60FDEAB066" blockId="7.[264,1325,284,1838]" pageId="7" pageNumber="8">
Abdomen: Terga
<date id="0B2A1065FFF1656EFF78EF60FD30B756" box="[522,688,849,875]" pageId="7" pageNumber="8" value="1910-03-01" valueMax="1910-08-01" valueMin="1910-03-01">1–3 and 8–10</date>
yellow; terga 4–7 light brown; each with thin black band on anterior and posterior margins. Tergum 3 with pair of brown spots. Sterna brown, but with sterna 8 and 9 yellow. Gill socket vestiges on segments 3–7, decreasing in size posteriorly. Genitalia (
<figureCitation id="E7AF2A20FFF1656EFF61EFF8FDEAB7DE" box="[531,618,969,995]" captionStart="FIGURES 15 – 18" captionStartId="6.[264,382,1784,1808]" captionTargetBox="[269,1316,317,1772]" captionTargetId="figure@6.[269,1317,317,1047]" captionTargetPageId="6" captionText="FIGURES 15 – 18. Hyrtanella pascalae, n. sp., male subimago: 15 Photograph of specimen in alcohol. 16. Forewing. 17. Hindwing. 18. Genitalia, ventral view." httpUri="https://zenodo.org/record/169630/files/figure.png" pageId="7" pageNumber="8">Fig. 18</figureCitation>
) with posterior margin of subgenital plate convex; forceps threesegmented; forceps segment 3 elongate. Penes fused and compact, with small median incision; outer, lateral margins subparallel distally and concave proximally. Caudal filaments broken and missing.
</paragraph>
<paragraph id="7F2B36A5FFF1656EFC4AE858FD22B0BE" blockId="7.[264,1325,284,1838]" box="[312,674,1129,1155]" pageId="7" pageNumber="8">
<emphasis id="4DE0EAB7FFF1656EFC4AE858FDA6B0BF" box="[312,550,1129,1154]" italics="true" pageId="7" pageNumber="8">Female Subimago—</emphasis>
Unknown.
</paragraph>
<paragraph id="7F2B36A5FFF1656EFC4AE8A0FD98B096" blockId="7.[264,1325,284,1838]" box="[312,536,1169,1195]" pageId="7" pageNumber="8">
<emphasis id="4DE0EAB7FFF1656EFC4AE8A0FE1DB097" box="[312,413,1169,1194]" italics="true" pageId="7" pageNumber="8">Adults—</emphasis>
Unknown.
</paragraph>
<paragraph id="7F2B36A5FFF1656EFC4AE888FB44B0C6" blockId="7.[264,1325,284,1838]" pageId="7" pageNumber="8">
<emphasis id="4DE0EAB7FFF1656EFC4AE888FEE7B0EF" box="[312,359,1209,1234]" italics="true" pageId="7" pageNumber="8">Egg</emphasis>
(
<figureCitation id="E7AF2A20FFF1656EFC05E888FE4AB0EE" box="[375,458,1209,1235]" captionStart="FIGURES 19 – 20" captionStartId="8.[264,382,1002,1026]" captionTargetBox="[264,1322,283,989]" captionTargetPageId="8" captionText="FIGURES 19 – 20. Eggs (scale bars = 10 m): 19. Hyrtanella pascalae, n. sp. 20. Undetermined Ephemerellinae, West Malaysia." httpUri="https://zenodo.org/record/169631/files/figure.png" pageId="7" pageNumber="8">Fig. 19</figureCitation>
)—shape ovoid; one polar cap; chorion generally smooth, without reticulations; micropyles in equatorial position; large KCTs between equator and poles.
</paragraph>
</subSubSection>
<subSubSection id="378E652EFFF1656EFC4AE938FD1BB1FE" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="7F2B36A5FFF1656EFC4AE938FD1BB1FE" blockId="7.[264,1325,284,1838]" pageId="7" pageNumber="8">
<emphasis id="4DE0EAB7FFF1656EFC4AE938FE38B11E" bold="true" box="[312,440,1289,1315]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
<emphasis id="4DE0EAB7FFF1656EFCB2E938FD93B11F" box="[448,531,1289,1314]" italics="true" pageId="7" pageNumber="8">Larvae</emphasis>
of
<taxonomicName id="B8944D26FFF1656EFF4CE938FD4BB11F" box="[574,715,1289,1314]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pascalae">
<emphasis id="4DE0EAB7FFF1656EFF4CE938FD4BB11F" box="[574,715,1289,1314]" italics="true" pageId="7" pageNumber="8">H. pascalae</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="B8944D26FFF1656EFEBAE938FBF0B11F" box="[968,1136,1289,1314]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="christineae">
<emphasis id="4DE0EAB7FFF1656EFEBAE938FBF0B11F" box="[968,1136,1289,1314]" italics="true" pageId="7" pageNumber="8">H. christineae</emphasis>
</taxonomicName>
by being much smaller and by having bluntly expanded genae, no basal projections on the mandibles, the third segment of the labial palp (
<figureCitation id="E7AF2A20FFF1656EFFFFE968FD53B14E" box="[653,723,1369,1395]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
) more slender than that of
<taxonomicName id="B8944D26FFF1656EF968E968FB41B14F" box="[1050,1217,1369,1394]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="christineae">
<emphasis id="4DE0EAB7FFF1656EF968E968FB41B14F" box="[1050,1217,1369,1394]" italics="true" pageId="7" pageNumber="8">H. christineae</emphasis>
</taxonomicName>
(
<figureCitation id="E7AF2A20FFF1656EF9A1E968FA9BB14E" box="[1235,1307,1369,1395]" captionStart="FIGURES 1 – 9" captionStartId="4.[264,382,1772,1796]" captionTargetBox="[265,1321,327,1759]" captionTargetId="figure@4.[661,882,805,1154]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Figs 1 – 8. Hyrtanella pascalae, n. sp., larvae: 1. Labrum (left, dorsal view; right, ventral view). 2. Detail of brushlike setae on dorsal surface of labrum. 3. Detail of transverse row of setae on ventral surface of labrum. 4. Hypopharynx (right half). 5. Right mandible. 6. Left mandible. 7. Maxilla. 8. Labium. Fig. 9. Hyrtanella christineae, larva: Detail of labial palp." httpUri="https://zenodo.org/record/158567/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
), no dorsal projections on the upper surface of the forefemur, and no ventral projections on the thoracic and abdominal sterna.
</paragraph>
</subSubSection>
<subSubSection id="378E652EFFF1656EFC4AE9E0FD25B2E3" pageId="7" pageNumber="8" type="description">
<paragraph id="7F2B36A5FFF1656EFC4AE9E0FD3EB25B" blockId="7.[264,1325,284,1838]" pageId="7" pageNumber="8">
<emphasis id="4DE0EAB7FFF1656EFC4AE9E0FE78B1D7" box="[312,504,1489,1514]" italics="true" pageId="7" pageNumber="8">Male subimagos</emphasis>
(
<figureCitation id="E7AF2A20FFF1656EFF7BE9E0FDDDB1D6" box="[521,605,1489,1515]" captionStart="FIGURES 15 – 18" captionStartId="6.[264,382,1784,1808]" captionTargetBox="[269,1316,317,1772]" captionTargetId="figure@6.[269,1317,317,1047]" captionTargetPageId="6" captionText="FIGURES 15 – 18. Hyrtanella pascalae, n. sp., male subimago: 15 Photograph of specimen in alcohol. 16. Forewing. 17. Hindwing. 18. Genitalia, ventral view." httpUri="https://zenodo.org/record/169630/files/figure.png" pageId="7" pageNumber="8">Fig. 15</figureCitation>
) of
<taxonomicName id="B8944D26FFF1656EFFFCE9E0FC9AB1D7" box="[654,794,1489,1514]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pascalae">
<emphasis id="4DE0EAB7FFF1656EFFFCE9E0FC9AB1D7" box="[654,794,1489,1514]" italics="true" pageId="7" pageNumber="8">H. pascalae</emphasis>
</taxonomicName>
differ from
<taxonomicName id="B8944D26FFF1656EFEDBE9E0FBA8B1D7" box="[937,1064,1489,1514]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="4DE0EAB7FFF1656EFEDBE9E0FBA8B1D7" box="[937,1064,1489,1514]" italics="true" pageId="7" pageNumber="8">Hyrtanella</emphasis>
</taxonomicName>
female adults beyond the usual sexual differences by having an abdomen that is not constricted, one small intercalary between MP2 and CuA (
<figureCitation id="E7AF2A20FFF1656EFF01EA10FD4AB206" box="[627,714,1569,1595]" captionStart="FIGURES 15 – 18" captionStartId="6.[264,382,1784,1808]" captionTargetBox="[269,1316,317,1772]" captionTargetId="figure@6.[269,1317,317,1047]" captionTargetPageId="6" captionText="FIGURES 15 – 18. Hyrtanella pascalae, n. sp., male subimago: 15 Photograph of specimen in alcohol. 16. Forewing. 17. Hindwing. 18. Genitalia, ventral view." httpUri="https://zenodo.org/record/169630/files/figure.png" pageId="7" pageNumber="8">Fig. 16</figureCitation>
), and the costal process of the hindwing with the tip angled away from the body (
<figureCitation id="E7AF2A20FFF1656EFF2BEA7DFD30B25B" box="[601,688,1612,1638]" captionStart="FIGURES 15 – 18" captionStartId="6.[264,382,1784,1808]" captionTargetBox="[269,1316,317,1772]" captionTargetId="figure@6.[269,1317,317,1047]" captionTargetPageId="6" captionText="FIGURES 15 – 18. Hyrtanella pascalae, n. sp., male subimago: 15 Photograph of specimen in alcohol. 16. Forewing. 17. Hindwing. 18. Genitalia, ventral view." httpUri="https://zenodo.org/record/169630/files/figure.png" pageId="7" pageNumber="8">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="7F2B36A5FFF1656EFC4AEA45FD25B2E3" blockId="7.[264,1325,284,1838]" pageId="7" pageNumber="8">
<emphasis id="4DE0EAB7FFF1656EFC4AEA45FE47B2B0" box="[312,455,1652,1677]" italics="true" pageId="7" pageNumber="8">Male adults</emphasis>
very likely have genital forceps segment 3 much longer than wide and penes lobes that are compact and nearly fused. This probable diagnosis is based on our examination of the male subimago.
</paragraph>
</subSubSection>
<subSubSection id="378E652EFFF16561FC4AEADAFC3FB125" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="etymology">
<paragraph id="7F2B36A5FFF1656EFC4AEADAFB10B313" blockId="7.[264,1325,284,1838]" pageId="7" pageNumber="8">
<emphasis id="4DE0EAB7FFF1656EFC4AEADAFE44B338" bold="true" box="[312,452,1771,1797]" pageId="7" pageNumber="8">Etymology.</emphasis>
This species is named in honor of Dr. Pascale Derleth (
<collectingCountry id="07837635FFF1656EF9F3EADDFB47B33B" box="[1153,1223,1772,1798]" name="Switzerland" pageId="7" pageNumber="8">Swiss</collectingCountry>
Federal Institute of Technology, Lausanne), who collected much of the
<typeStatus id="A02F8807FFF1656EFE9CEB25FBA0B313" box="[1006,1056,1812,1838]" pageId="7" pageNumber="8">type</typeStatus>
material.
</paragraph>
<caption id="2BEB662DFFFE6561FC7AEFDBFDE1B01A" httpUri="https://zenodo.org/record/169631/files/figure.png" pageId="8" pageNumber="9" targetBox="[264,1322,283,989]" targetPageId="8">
<paragraph id="7F2B36A5FFFE6561FC7AEFDBFDE1B01A" blockId="8.[264,1323,1002,1063]" pageId="8" pageNumber="9">
<emphasis id="4DE0EAB7FFFE6561FC7AEFDBFE50B03F" bold="true" box="[264,464,1002,1026]" pageId="8" pageNumber="9">FIGURES 19–20.</emphasis>
Eggs (scale bars = 10 m): 19.
<taxonomicName id="B8944D26FFFE6561FE49EFDBFB96B03C" box="[827,1046,1002,1025]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pascalae" status="sp. nov.">
<emphasis id="4DE0EAB7FFFE6561FE49EFDBFB96B03C" box="[827,1046,1002,1025]" italics="true" pageId="8" pageNumber="9">Hyrtanella pascalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="56D357CCFFFE6561F957EFDBFBE0B03C" box="[1061,1120,1002,1025]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
20. Undetermined Ephemerellinae, West Malaysia.
</paragraph>
</caption>
<paragraph id="7F2B36A5FFFE6561FC4AE8B4FC3FB125" blockId="8.[264,1325,1157,1904]" pageId="8" pageNumber="9">
<emphasis id="4DE0EAB7FFFE6561FC4AE8B4FD9EB0A2" bold="true" box="[312,542,1157,1183]" pageId="8" pageNumber="9">Larval variability.</emphasis>
General color patterns vary from the typical light brown to dark brown. This variation appears to be between populations, and it is not dependent on the instar number. The number, relative development, and placement of chalazae on the ventral edge of the femora varies slightly between populations.
</paragraph>
</subSubSection>
<subSubSection id="378E652EFFFE6561FC4AE914FB0CB34D" pageId="8" pageNumber="9" type="description">
<paragraph id="7F2B36A5FFFE6561FC4AE914FD9CB2C5" blockId="8.[264,1325,1157,1904]" pageId="8" pageNumber="9">
<emphasis id="4DE0EAB7FFFE6561FC4AE914FE21B102" bold="true" box="[312,417,1317,1343]" pageId="8" pageNumber="9">Ecology.</emphasis>
<taxonomicName id="B8944D26FFFE6561FCD8E917FD1AB102" box="[426,666,1318,1343]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pascalae">
<emphasis id="4DE0EAB7FFFE6561FCD8E917FD1AB102" box="[426,666,1318,1343]" italics="true" pageId="8" pageNumber="9">Hyrtanella pascalae</emphasis>
</taxonomicName>
was collected commonly in the study areas of
<collectingCountry id="07837635FFFE6561F9A2E917FEABB155" name="Indonesia" pageId="8" pageNumber="9">Indonesia</collectingCountry>
, and sometimes it was locally abundant. It was sampled from stream riffles and runs with different substrate compositions, including silt, gravel, cobble, and boulders. Flow velocity at these sites ranged from 0.2–2.0 m/s; water depth was between 0.3–3.0 m; and at least some sites had a moderate silt load. Water temperature during the sampling period varied from 24.1°–26.4°C (Derleth 2003; Danuta Zaranko, pers. comm.). Multivariate analyses have shown that
<taxonomicName id="B8944D26FFFE6561FF48EA27FD46B212" box="[570,710,1558,1583]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pascalae">
<emphasis id="4DE0EAB7FFFE6561FF48EA27FD46B212" box="[570,710,1558,1583]" italics="true" pageId="8" pageNumber="9">H. pascalae</emphasis>
</taxonomicName>
is more abundant in larger streams, and its distribution is positively correlated with the opening of the canopy, stream width, and water temperature (Derleth 2003). With respect to the influence of logging activities in the area,
<taxonomicName id="B8944D26FFFE6561F87CEA57FEF0B29A" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pascalae">
<emphasis id="4DE0EAB7FFFE6561F87CEA57FEF0B29A" italics="true" pageId="8" pageNumber="9">H. pascalae</emphasis>
</taxonomicName>
is characterised by population density increasing during, and until six months after, logging. However, the population density decreases one to three years after logging activity (Derleth 2003).
</paragraph>
<paragraph id="7F2B36A5FFFE6561FC4AEB37FB0CB34D" blockId="8.[264,1325,1157,1904]" pageId="8" pageNumber="9">
All localities where
<taxonomicName id="B8944D26FFFE6561FF55EB37FD34B322" box="[551,692,1798,1823]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pascalae">
<emphasis id="4DE0EAB7FFFE6561FF55EB37FD34B322" box="[551,692,1798,1823]" italics="true" pageId="8" pageNumber="9">H. pascalae</emphasis>
</taxonomicName>
has been collected are situated between 100 and 400 meters above sea level, whereas collecting locales of
<taxonomicName id="B8944D26FFFE6561FEFAEB1FFBB0B37A" box="[904,1072,1838,1863]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="christineae">
<emphasis id="4DE0EAB7FFFE6561FEFAEB1FFBB0B37A" box="[904,1072,1838,1863]" italics="true" pageId="8" pageNumber="9">H. christineae</emphasis>
</taxonomicName>
in Sabah are located between 900 and
<quantity id="B86C9B40FFFE6561FCA4EB67FDB2B34D" box="[470,562,1878,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="9" unit="m" value="1600.0">1600 m</quantity>
. These species may represent altitudinal vicariants.
</paragraph>
</subSubSection>
<subSubSection id="378E652EFFFF6560FC4AED2AFEE1B373" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="7F2B36A5FFFF6560FC4AED2AFDFFB593" blockId="9.[264,1325,283,1870]" pageId="9" pageNumber="10">
<emphasis id="4DE0EAB7FFFF6560FC4AED2AFDAEB508" bold="true" box="[312,558,283,309]" pageId="9" pageNumber="10">Material examined.</emphasis>
<typeStatus id="A02F8807FFFF6560FF45ED2DFD29B50B" box="[567,681,284,310]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="07837635FFFF6560FFC7ED2DFD7DB50B" box="[693,765,284,310]" name="Iceland" pageId="9" pageNumber="10">Island</collectingCountry>
of Borneo—INDONESIA,
<emphasis id="4DE0EAB7FFFF6560F93EED2AFAA2B508" bold="true" box="[1100,1314,283,309]" pageId="9" pageNumber="10">East Kalimantan</emphasis>
: Malinau Basin—
<emphasis id="4DE0EAB7FFFF6560FCAAED75FD77B560" box="[472,759,324,349]" italics="true" pageId="9" pageNumber="10">Seturan River tributary</emphasis>
: Temalat (Sungai Guang) trib., Loc. Seturan (2000petak 43),
<geoCoordinate id="1AA05062FFFF6560FC95ED5DFDF7B5BB" box="[487,631,364,390]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="2.9913888">2°59'29&quot; N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FFFFED5DFCB5B5BB" box="[653,821,364,390]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.55806">116°33'29&quot; E</geoCoordinate>
,
<date id="0B2A1065FFFF6560FE39ED5DFC56B5BB" box="[843,982,364,390]" pageId="9" pageNumber="10" value="2000-06-18">18/VI/2000</date>
, P Derleth, JL Gattolliat, (B0811),
<specimenCount id="6992FD2CFFFF6560FC05EDA5FE5DB593" box="[375,477,404,430]" pageId="9" pageNumber="10" type="female">1 female</specimenCount>
larva [
<collectionCode id="1985AE60FFFF6560FF5FEDA5FDF3B593" box="[557,627,404,430]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
].
</paragraph>
<paragraph id="7F2B36A5FFFF6560FC4AED8DFE11B1AB" blockId="9.[264,1325,283,1870]" pageId="9" pageNumber="10">
<typeStatus id="A02F8807FFFF6560FC4AED8DFE2EB5EB" box="[312,430,444,470]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
:
<emphasis id="4DE0EAB7FFFF6560FCC9ED8DFD39B5E8" box="[443,697,444,469]" italics="true" pageId="9" pageNumber="10">same locale and date</emphasis>
,
<specimenCount id="6992FD2CFFFF6560FFB8ED8DFCB7B5EB" box="[714,823,444,470]" pageId="9" pageNumber="10" type="larva">11 larvae</specimenCount>
[
<collectionCode id="1985AE60FFFF6560FE3AED8DFC0EB5EB" box="[840,910,444,470]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; same locale, but
<date id="0B2A1065FFFF6560F91DED8DFA8DB5EB" box="[1135,1293,444,470]" pageId="9" pageNumber="10" value="2000-08-16">16/VIII/2000</date>
, P Derleth, R Schlaepfer, (B0812), two larvae [
<collectionCode id="1985AE60FFFF6560FE6EEDD5FCE3B5C3" box="[796,867,484,510]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; same locale, but
<date id="0B2A1065FFFF6560F936EDD5FB4FB5C3" box="[1092,1231,484,510]" pageId="9" pageNumber="10" value="2001-04-16">16/IV/2001</date>
, P Derleth, (B0813) one larva [
<collectionCode id="1985AE60FFFF6560FF57EE3DFDECB61B" box="[549,620,524,550]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
];
<emphasis id="4DE0EAB7FFFF6560FF0DEE3DFC69B618" box="[639,1001,524,549]" italics="true" pageId="9" pageNumber="10">other Seturan River tributaries</emphasis>
: Tamalang trib., Loc. Seturan (2001petak 57),
<geoCoordinate id="1AA05062FFFF6560FF6EEE05FD1AB673" box="[540,666,564,590]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="2.9894445">2°59'22&quot;N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FFDDEE05FCD7B673" box="[687,855,564,590]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.50806">116°30'29&quot; E</geoCoordinate>
,
<date id="0B2A1065FFFF6560FE1EEE05FB80B673" box="[876,1024,564,590]" pageId="9" pageNumber="10" value="2000-07-18">18/VII/2000</date>
, P Derleth, F Béboux, (B0511), two larvae [
<collectionCode id="1985AE60FFFF6560FF7CEE6DFDD4B64B" box="[526,596,604,630]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; Tamalang trib., Loc. Seturan (2001petak 57),
<geoCoordinate id="1AA05062FFFF6560F9D5EE6DFAA5B64B" box="[1191,1317,604,630]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="2.9894445">2°59'22&quot;N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FC7AEEB5FE25B6A3" box="[264,421,644,670]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.51278">116°30'46&quot; E</geoCoordinate>
,
<date id="0B2A1065FFFF6560FCC3EEB5FDBBB6A3" box="[433,571,644,670]" pageId="9" pageNumber="10" value="2001-04-10">10/IV/2001</date>
, P Derleth, (B0513),
<specimenCount id="6992FD2CFFFF6560FE46EEB5FC20B6A3" box="[820,928,644,670]" pageId="9" pageNumber="10" type="larva">16 larvae</specimenCount>
[
<collectionCode id="1985AE60FFFF6560FEDCEEB5FC75B6A3" box="[942,1013,644,670]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; Tamalang trib., Loc. Seturan (2001petak 57),
<geoCoordinate id="1AA05062FFFF6560FF54EE9DFD24B6FB" box="[550,676,684,710]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="2.9972222">2°59'50&quot;N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FFCAEE9DFCDDB6FB" box="[696,861,684,710]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.50806">116°30'29&quot; E</geoCoordinate>
,
<date id="0B2A1065FFFF6560FE00EE9DFB86B6FB" box="[882,1030,684,710]" pageId="9" pageNumber="10" value="2000-07-19">19/VII/2000</date>
, P Derleth, F Béboux, (B0521),
<specimenCount id="6992FD2CFFFF6560FC0FEEE5FE6EB6D3" box="[381,494,724,750]" pageId="9" pageNumber="10" type="larva">14 larvae</specimenCount>
[
<collectionCode id="1985AE60FFFF6560FF73EEE5FDC8B6D3" box="[513,584,724,750]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; same locale, but
<date id="0B2A1065FFFF6560FE40EEE5FC40B6D3" box="[818,960,724,750]" pageId="9" pageNumber="10" value="2000-08-05">5/VIII/2000</date>
, P Derleth, (B0521PLS), one male subimago [
<collectionCode id="1985AE60FFFF6560FCACEECDFDA4B72B" box="[478,548,764,790]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; Bengahau trib., Loc. Seturan (2001petak 57),
<geoCoordinate id="1AA05062FFFF6560F9D5EECDFAA5B72B" box="[1191,1317,764,790]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="2.9894445">2°59'22&quot;N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FC7AEF15FE1FB703" box="[264,415,804,830]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.51278">116°30'46&quot;E</geoCoordinate>
,
<date id="0B2A1065FFFF6560FCD9EF15FDBAB703" box="[427,570,804,830]" pageId="9" pageNumber="10" value="2000-08-08">8/VIII/2000</date>
, P Derleth, (B0531), five larvae [
<collectionCode id="1985AE60FFFF6560FEB2EF15FB86B703" box="[960,1030,804,830]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; same locale, but
<date id="0B2A1065FFFF6560F9ADEF15FEC5B75B" pageId="9" pageNumber="10" value="2001-04-11">11/IV/ 2001</date>
, P Derleth, B Feldmeyer, (B0533), seven larvae [
<collectionCode id="1985AE60FFFF6560FEF3EF7DFC48B75B" box="[897,968,844,870]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; Bengahau trib., Loc. Seturan (2001petak 57),
<geoCoordinate id="1AA05062FFFF6560FC9AEF45FDE6B7B3" box="[488,614,884,910]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="2.9894445">2°59'22&quot;N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FF0FEF45FCA6B7B3" box="[637,806,884,910]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.51278">116°30'46&quot; E</geoCoordinate>
,
<date id="0B2A1065FFFF6560FE4FEF45FC5BB7B3" box="[829,987,884,910]" pageId="9" pageNumber="10" value="2000-08-19">19/VIII/2000</date>
, P Derleth, R Schlaepfer (B0541), ten larvae [
<collectionCode id="1985AE60FFFF6560FF70EFADFDE0B78B" box="[514,608,924,950]" pageId="9" pageNumber="10">BMNH</collectionCode>
,
<collectionCode id="1985AE60FFFF6560FF1DEFADFD45B78B" box="[623,709,924,950]" pageId="9" pageNumber="10">FAMU</collectionCode>
,
<collectionCode id="1985AE60FFFF6560FFA1EFADFC97B78B" box="[723,791,924,950]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; Kipah trib., Loc. Seturan (1998petak 28),
<geoCoordinate id="1AA05062FFFF6560FC7AEFF5FE0DB7E3" box="[264,397,964,990]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="3.03">3°01'48&quot; N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FCE8EFF5FDB7B7E3" box="[410,567,964,990]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.496666">116°29'48&quot; E</geoCoordinate>
,
<date id="0B2A1065FFFF6560FF36EFF5FD4DB7E3" box="[580,717,964,990]" pageId="9" pageNumber="10" value="2001-03-29">29/III/2001</date>
, P Derleth, (B0913), one larva [
<collectionCode id="1985AE60FFFF6560F930EFF5FB09B7E3" box="[1090,1161,964,990]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; Seturan riv., Loc. Seturan (unlogged),
<geoCoordinate id="1AA05062FFFF6560FF44EFDDFD3CB03B" box="[566,700,1004,1030]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="3.0013888">3°00'05&quot; N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FFBBEFDDFCE7B03B" box="[713,871,1004,1030]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.513336">116°30'48&quot; E</geoCoordinate>
,
<date id="0B2A1065FFFF6560FE07EFDDFC7DB03B" box="[885,1021,1004,1030]" pageId="9" pageNumber="10" value="2001-03-28">28/III/2001</date>
, P Derleth, B Feldmeyer, (B1313),
<specimenCount id="6992FD2CFFFF6560FC09E825FE69B013" box="[379,489,1044,1070]" pageId="9" pageNumber="10" type="larva">50 larvae</specimenCount>
[
<collectionCode id="1985AE60FFFF6560FC89E825FDC1B013" box="[507,577,1044,1070]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; Riv. Seturan, Loc. Seturan, Aff. cours principal,
<geoCoordinate id="1AA05062FFFF6560F9D2E825FAA4B013" box="[1184,1316,1044,1070]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="3.0013888">3°00’05”N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FC7AE80DFE24B06B" box="[264,420,1084,1110]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.513336">116°30’48”E</geoCoordinate>
,
<date id="0B2A1065FFFF6560FCC1E80DFDBBB06B" box="[435,571,1084,1110]" pageId="9" pageNumber="10" value="2001-03-28">28/III/2001</date>
, P Derleth, B Feldmeyer, three larvae [
<collectionCode id="1985AE60FFFF6560F967E80DFBE7B06B" box="[1045,1127,1084,1110]" pageId="9" pageNumber="10">PERC</collectionCode>
]; unnamed trib., Loc. Seturan (unlogged),
<geoCoordinate id="1AA05062FFFF6560FF4BE855FD40B043" box="[569,704,1124,1150]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="2.9969444">2°59'49&quot; N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FFBFE855FCECB043" box="[717,876,1124,1150]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.52278">116°31'22&quot; E</geoCoordinate>
,
<date id="0B2A1065FFFF6560FE09E855FB85B043" box="[891,1029,1124,1150]" pageId="9" pageNumber="10" value="2001-04-17">17/IV/2001</date>
, P Derleth, M Sartori, B Feldmeyer, (B1513), one larva [
<collectionCode id="1985AE60FFFF6560FFF2E8BDFD47B09B" box="[640,711,1164,1190]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
].
<emphasis id="4DE0EAB7FFFF6560FFA9E8BDFC5DB098" box="[731,989,1164,1189]" italics="true" pageId="9" pageNumber="10">Rian River tributaries</emphasis>
: Belakau trib., Loc. Langap
<bibRefCitation id="1B054B54FFFF6560FC7AE885FE6DB0F3" author="Sud" box="[264,493,1204,1230]" pageId="9" pageNumber="12" refString="Studemann, D. &amp; Landolt, P. (1997) Eggs of Ephemerellidae (Ephemeroptera). In: Landolt, P. &amp; Sartori, M. (Ed) Ephemeroptera &amp; Plecoptera: Biology-Ecology-Systematics, MTL, Fribourg, Switzerland, pp. 362 - 371." type="book chapter" year="1997">Sud (1997petak 6)</bibRefCitation>
,
<geoCoordinate id="1AA05062FFFF6560FC8BE885FDFFB0F3" box="[505,639,1204,1230]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="3.0677776">3°04'04&quot; N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FFF9E885FCA9B0F3" box="[651,809,1204,1230]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.507225">116°30'26&quot; E</geoCoordinate>
, Alt.
<quantity id="B86C9B40FFFF6560FE18E885FC38B0F3" box="[874,952,1204,1230]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="m" value="100.0">100 m</quantity>
,
<date id="0B2A1065FFFF6560FEB1E885FBCDB0F3" box="[963,1101,1204,1230]" pageId="9" pageNumber="10" value="2001-04-20">20/IV/2001</date>
, P Derleth, M Sartori, (B0113), one larva [
<collectionCode id="1985AE60FFFF6560FF3AE8EDFD0FB0CB" box="[584,655,1244,1270]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; Ngayo trib., Loc. Langap Sud (1995),
<geoCoordinate id="1AA05062FFFF6560F9EBE8EDFAA6B0CB" box="[1177,1318,1244,1270]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="3.03">3°01'48&quot; N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FC7AE935FE27B123" box="[264,423,1284,1310]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.496666">116°29'48&quot; E</geoCoordinate>
, Alt.
<quantity id="B86C9B40FFFF6560FC9EE935FDBBB123" box="[492,571,1284,1310]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="10" unit="m" value="160.0">160 m</quantity>
,
<date id="0B2A1065FFFF6560FF3AE935FD52B123" box="[584,722,1284,1310]" pageId="9" pageNumber="10" value="2000-06-14">14/VI/2000</date>
, (B0410), seven larvae [
<collectionCode id="1985AE60FFFF6560FE87E935FBB0B123" box="[1013,1072,1284,1310]" pageId="9" pageNumber="10">LIPI</collectionCode>
,
<collectionCode id="1985AE60FFFF6560F94CE935FB02B123" box="[1086,1154,1284,1310]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; same locale, but
<date id="0B2A1065FFFF6560FC4BE91DFE3FB17B" box="[313,447,1324,1350]" pageId="9" pageNumber="10" value="2000-07-08">8/VII/2000</date>
, P Derleth, (B0411), two larvae [
<collectionCode id="1985AE60FFFF6560FE2EE91DFC23B17B" box="[860,931,1324,1350]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
]; Ngayo trib., Loc. Langap Sud (1995),
<geoCoordinate id="1AA05062FFFF6560FC14E965FE6CB153" box="[358,492,1364,1390]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="3.0822222">3°04'56&quot; N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560FC8BE965FD18B153" box="[505,664,1364,1390]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="116.51611">116°30'58&quot; E</geoCoordinate>
, Alt.
<quantity id="B86C9B40FFFF6560FFA9E965FCA9B153" box="[731,809,1364,1390]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="10" unit="m" value="160.0">160 m</quantity>
,
<date id="0B2A1065FFFF6560FE44E965FC49B153" box="[822,969,1364,1390]" pageId="9" pageNumber="10" value="2000-07-13">13/VII/2000</date>
, P Derleth, (B0431), four larvae [
<collectionCode id="1985AE60FFFF6560FC4DE94DFE06B1AB" box="[319,390,1404,1430]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
].
</paragraph>
<paragraph id="7F2B36A5FFFF6560FC4AE995FD26B183" blockId="9.[264,1325,283,1870]" box="[312,678,1444,1470]" pageId="9" pageNumber="10">
Other Material (not
<typeStatus id="A02F8807FFFF6560FF51E995FD1AB183" box="[547,666,1444,1470]" pageId="9" pageNumber="10" type="paratype">paratypes</typeStatus>
):
</paragraph>
<paragraph id="7F2B36A5FFFF6560FC4AE9FDFEE1B373" blockId="9.[264,1325,283,1870]" pageId="9" pageNumber="10">
<collectingCountry id="07837635FFFF6560FC4AE9FDFE00B1DB" box="[312,384,1484,1510]" name="Iceland" pageId="9" pageNumber="10">Island</collectingCountry>
of Borneo—INDONESIA,
<emphasis id="4DE0EAB7FFFF6560FF99E9FDFC4BB1DB" bold="true" box="[747,971,1484,1510]" pageId="9" pageNumber="10">East Kalimantan</emphasis>
: Kelinjau R., about
<quantity id="B86C9B40FFFF6560F99AE9FDFAABB1DB" box="[1256,1323,1484,1510]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="km" value="10.0">10km</quantity>
upstream of Atan R., at logging staging area above rapids,
<geoCoordinate id="1AA05062FFFF6560FEBEE9C5FBA5B233" box="[972,1061,1524,1550]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="0.75">0°45’N</geoCoordinate>
,
<geoCoordinate id="1AA05062FFFF6560F941E9C5FB25B233" box="[1075,1189,1524,1550]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="116.25">116°15’E</geoCoordinate>
,
<date id="0B2A1065FFFF6560F9C1E9C5FAA8B233" box="[1203,1320,1524,1550]" pageId="9" pageNumber="10" value="1997-01-23">23/I/1997</date>
, one larva [
<collectionCode id="1985AE60FFFF6560FCF8EA2DFE5DB20B" box="[394,477,1564,1590]" pageId="9" pageNumber="10">PERC</collectionCode>
].
<collectingCountry id="07837635FFFF6560FC81EA2DFDBBB20B" box="[499,571,1564,1590]" name="Iceland" pageId="9" pageNumber="10">Island</collectingCountry>
of Borneo—MALAYSIA (east),
<emphasis id="4DE0EAB7FFFF6560FEA1EA2DFBA1B20B" bold="true" box="[979,1057,1564,1590]" pageId="9" pageNumber="10">Sabah</emphasis>
: Liwagu R at bridge, Ranau,
<quantity id="B86C9B40FFFF6560FC1AEA75FE2BB263" box="[360,427,1604,1630]" metricMagnitude="2" metricUnit="m" metricValue="3.35" pageId="9" pageNumber="10" unit="m" value="335.0">335m</quantity>
elev.,
<date id="0B2A1065FFFF6560FF76EA75FD4DB263" box="[516,717,1604,1630]" pageId="9" pageNumber="10" value="1972-08-11" valueMax="1972-08-16" valueMin="1972-08-11">11–16/VIII/1972</date>
, GF&amp;CG Edmunds, WL&amp;JG Peters, four larvae [
<collectionCode id="1985AE60FFFF6560FC7DEA5DFEE1B2BB" box="[271,353,1644,1670]" pageId="9" pageNumber="10">PERC</collectionCode>
]; Mantukungan R, S of Poring Hot Springs Station,
<quantity id="B86C9B40FFFF6560FE83EA5DFBB4B2BB" box="[1009,1076,1644,1670]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="9" pageNumber="10" unit="m" value="365.0">365m</quantity>
elev.,
<date id="0B2A1065FFFF6560F9F8EA5DFAA8B2BB" box="[1162,1320,1644,1670]" pageId="9" pageNumber="10" value="1972-08-12">12/VIII/1972</date>
, WL&amp;JG Peters, five larvae [
<collectionCode id="1985AE60FFFF6560FF27EAA5FD27B293" box="[597,679,1684,1710]" pageId="9" pageNumber="10">PERC</collectionCode>
]; Sungai Moyog (26°C),
<quantity id="B86C9B40FFFF6560FEBCEAA5FB83B293" box="[974,1027,1684,1710]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="9" pageNumber="10" unit="mi" value="3.0">3 mi</quantity>
E of Penampung,
<date id="0B2A1065FFFF6560F9A9EAA5FEEDB2EB" pageId="9" pageNumber="10" value="1978-09-27" valueMax="1978-09-29" valueMin="1978-09-27">27–29/ IX/1978</date>
, GF&amp;
<collectingCountry id="07837635FFFF6560FCC4EA8DFE5FB2EB" box="[438,479,1724,1750]" name="Switzerland" pageId="9" pageNumber="10">CH</collectingCountry>
Edmunds, seven larvae [
<collectionCode id="1985AE60FFFF6560FE7EEA8DFCDFB2EB" box="[780,863,1724,1750]" pageId="9" pageNumber="10">PERC</collectionCode>
]; Sungai Moyog at tributary
<quantity id="B86C9B40FFFF6560F9CBEA8DFB70B2EB" box="[1209,1264,1724,1750]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="9" pageNumber="10" unit="mi" value="8.0">8 mi</quantity>
E of Penampung,
<date id="0B2A1065FFFF6560FCDBEAD5FD9BB2C3" box="[425,539,1764,1790]" pageId="9" pageNumber="10" value="1978-10-02">2/X/1978</date>
, GF&amp;
<collectingCountry id="07837635FFFF6560FF19EAD5FD14B2C3" box="[619,660,1764,1790]" name="Switzerland" pageId="9" pageNumber="10">CH</collectingCountry>
Edmunds, one larva (mouthparts on slide) [
<collectionCode id="1985AE60FFFF6560F9BCEAD5FAA1B2C3" box="[1230,1313,1764,1790]" pageId="9" pageNumber="10">PERC</collectionCode>
]; Penompange River, Crocher National Park,
<date id="0B2A1065FFFF6560FE6CEB3DFC2DB31B" box="[798,941,1804,1830]" pageId="9" pageNumber="10" value="2003-08-08">8/VIII/2003</date>
, M Whiting team, seven larvae [
<collectionCode id="1985AE60FFFF6560FC7DEB05FED6B373" box="[271,342,1844,1870]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="9" pageNumber="10">MZL</collectionCode>
].
</paragraph>
</subSubSection>
</treatment>
</document>