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<document id="E410C07508B4FDDD344055E67314536A" ID-CLB-Dataset="55633" ID-DOI="10.11646/palaeoentomology.5.2.3" ID-GBIF-Dataset="6af10238-e40f-4104-9cf6-b00a4aa8ccd5" ID-ISSN="2624-2834" ID-Zenodo-Dep="6530392" ID-ZooBank="1EA06A24-60CF-4EBD-BF9F-BE873E69D613" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652082896052" checkinUser="plazi" docAuthor="HUANG, DI-YING, LIU, QI, LIAN, XIN-NENG, FU, YAN-ZHE &amp; NEL, ANDRÉ" docDate="2022" docId="F736CF0AFFFBF003AE07C9BFFB34FDC7" docLanguage="en" docName="palaeoentomology.5.2.3.pdf" docOrigin="Palaeoentomology 5 (2)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.2.3" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Guangxicalopteryx huashanensis HUANG &amp; LIU &amp; LIAN &amp; FU &amp; NEL 2022, sp. nov." docType="treatment" docUuid="03C70E49-70B3-431B-BE71-DBDB055E85B7" docUuidSource="ZooBank" docVersion="7" lastPageNumber="115" masterDocId="0B0FB772FFFAF000AD2DCA33FFDDFF92" masterDocTitle="A new calopterygid damselfly (Odonata, Zygoptera) from the Oligocene Ningming Basin, Guangxi, South China" masterLastPageNumber="119" masterPageNumber="113" pageNumber="114" updateTime="1699329902818" updateUser="ExternalLinkService">
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<mods:titleInfo id="C8114E72D0A0748204639F98BA8F7690">
<mods:title id="9145644FB09108C9D940421A55A60862">A new calopterygid damselfly (Odonata, Zygoptera) from the Oligocene Ningming Basin, Guangxi, South China</mods:title>
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<mods:namePart id="ED7D1DE463507D27928507971A31F11E">HUANG, DI-YING</mods:namePart>
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<mods:namePart id="4B0E09083287696B90B95A307DE4BAD6">LIU, QI</mods:namePart>
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<mods:namePart id="6DC49368A872DE8CFADF8CD50FF98C9C">LIAN, XIN-NENG</mods:namePart>
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<mods:namePart id="D2EDBD1D17BABF09CA66962086A4D5A3">FU, YAN-ZHE</mods:namePart>
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<mods:namePart id="E9A058C742800D8C058FF80F854A9C8E">NEL, ANDRÉ</mods:namePart>
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<mods:title id="93FD60A9F276D89CD2D10CC063B33BB8">Palaeoentomology</mods:title>
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<mods:date id="6E7F0AF87C94EC197C96161C1B604F43">2022</mods:date>
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<treatment id="F736CF0AFFFBF003AE07C9BFFB34FDC7" ID-DOI="http://doi.org/10.5281/zenodo.6532920" ID-GBIF-Taxon="195277811" ID-Zenodo-Dep="6532920" LSID="urn:lsid:zoobank.org:act:03C70E49-70B3-431B-BE71-DBDB055E85B7" httpUri="http://treatment.plazi.org/id/F736CF0AFFFBF003AE07C9BFFB34FDC7" lastPageId="3" lastPageNumber="115" pageId="1" pageNumber="114">
<subSubSection id="37852D97FFFBF001AE07C9BFFAD0FC34" box="[810,1293,908,935]" pageId="1" pageNumber="114" type="nomenclature">
<paragraph id="7F207E1CFFFBF001AE07C9BFFAD0FC34" blockId="1.[810,1451,908,1046]" box="[810,1293,908,935]" pageId="1" pageNumber="114">
<heading id="2468C970FFFBF001AE07C9BFFAD0FC34" bold="true" box="[810,1293,908,935]" fontSize="10" level="2" pageId="1" pageNumber="114" reason="3">
<emphasis id="4DEBA20EFFFBF001AE07C9BFFAD0FC34" bold="true" box="[810,1293,908,935]" pageId="1" pageNumber="114">
<taxonomicName id="B89F059FFFFBF001AE07C9BFFB70FC35" authority="HUANG &amp; LIU &amp; LIAN &amp; FU &amp; NEL, 2022" authorityName="HUANG &amp; LIU &amp; LIAN &amp; FU &amp; NEL" authorityYear="2022" box="[810,1197,908,935]" class="Insecta" family="Calopterygidae" genus="Guangxicalopteryx" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="114" phylum="Arthropoda" rank="species" species="huashanensis" status="sp. nov.">
<emphasis id="4DEBA20EFFFBF001AE07C9BFFB70FC35" bold="true" box="[810,1197,908,935]" italics="true" pageId="1" pageNumber="114">Guangxicalopteryx huashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="56D81F75FFFBF001A99EC9BFFAD0FC34" box="[1203,1293,908,934]" pageId="1" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="37852D97FFFBF001AE07C982FC32FB84" pageId="1" pageNumber="114" type="description">
<paragraph id="7F207E1CFFFBF001AE07C982FC32FB84" blockId="1.[810,1451,908,1046]" pageId="1" pageNumber="114">
(
<figureCitation id="E7A46299FFFBF001AE1FC982FCA5FC59" box="[818,888,945,971]" captionStart="FIGURE 1" captionStartId="1.[136,235,2040,2064]" captionTargetBox="[154,1431,1331,2012]" captionTargetId="figure-381@1.[151,1436,1327,2016]" captionTargetPageId="1" captionText="FIGURE 1. Fossil locality. A, Map of fossil locality. B, Photograph of fossil collecting." figureDoi="http://doi.org/10.5281/zenodo.6530394" httpUri="https://zenodo.org/record/6530394/files/figure.png" pageId="1" pageNumber="114">Figs 1</figureCitation>
,
<figureCitation id="E7A46299FFFBF001AEABC982FC48FC59" box="[902,917,945,971]" captionStart="FIGURE2" captionStartId="2.[136,235,1690,1714]" captionTargetBox="[215,1372,189,1667]" captionTargetId="figure-122@2.[215,1372,189,1667]" captionTargetPageId="2" captionText="FIGURE2.Guangxicalopteryx huashanensisgen.etsp.nov., holotypeNIGP179898,photographs.A, Counterpart(NIGP179898b). B, Counterpart under alcohol. C, Part (NIGP179898a), covered with a carbonaceous film. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6530396" httpUri="https://zenodo.org/record/6530396/files/figure.png" pageId="1" pageNumber="114">2</figureCitation>
)
<uri id="0B0E721EFFFBF001AE07C9E5FC32FB84" pageId="1" pageNumber="114">
urn:lsid:zoobank.org:act:
<uuid id="0B3944C9FFFBF001A965C9E5FC32FB84" pageId="1" pageNumber="114">03C70E49-70B3-431B-BE71- DBDB055E85B7</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="37852D97FFFBF001AE07CE77FBBCFB11" pageId="1" pageNumber="114" type="materials_examined">
<paragraph id="7F207E1CFFFBF001AE07CE77FBBCFB11" blockId="1.[810,1452,1092,1303]" pageId="1" pageNumber="114">
<emphasis id="4DEBA20EFFFBF001AE07CE77FC44FBCC" bold="true" box="[810,921,1092,1118]" pageId="1" pageNumber="114">Material.</emphasis>
<materialsCitation id="CFF77441FFFBF001AE88CE77FBBCFB11" ID-GBIF-Occurrence="3763880302" collectionCode="NIGP" pageId="1" pageNumber="114" specimenCode="NIGP179898" specimenCount="1" typeStatus="holotype">
<typeStatus id="A024C0BEFFFBF001AE88CE77FBD0FBCC" box="[933,1037,1092,1118]" pageId="1" pageNumber="114" type="holotype">Holotype</typeStatus>
<specimenCode id="2F39D667FFFBF001A934CE77FB6FFBCC" box="[1049,1202,1092,1118]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="1" pageNumber="114">NIGP179898</specimenCode>
a, b, a forewing with basal part (petiole) missing.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="37852D97FFFBF001AE72CEBDFB59FB60" pageId="1" pageNumber="114" type="etymology">
<paragraph id="7F207E1CFFFBF001AE72CEBDFB59FB60" blockId="1.[810,1452,1092,1303]" pageId="1" pageNumber="114">
<emphasis id="4DEBA20EFFFBF001AE72CEBDFC3BFB3A" bold="true" box="[863,998,1166,1192]" pageId="1" pageNumber="114">Etymology.</emphasis>
The specific name huashanensis comes from the famous Huashan rock paintings at the Ningming County, dated 2,000 years ago.
</paragraph>
</subSubSection>
<subSubSection id="37852D97FFFBF001AE72CECFFB43FA85" box="[863,1182,1276,1303]" pageId="1" pageNumber="114" type="diagnosis">
<paragraph id="7F207E1CFFFBF001AE72CECFFB43FA85" blockId="1.[810,1452,1092,1303]" box="[863,1182,1276,1303]" pageId="1" pageNumber="114">
<emphasis id="4DEBA20EFFFBF001AE72CECFFC04FA84" bold="true" box="[863,985,1276,1302]" pageId="1" pageNumber="114">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<caption id="2BE02E94FFFBF001ADA5CDCBFBD9F782" ID-DOI="http://doi.org/10.5281/zenodo.6530394" ID-Zenodo-Dep="6530394" box="[136,1028,2040,2064]" httpUri="https://zenodo.org/record/6530394/files/figure.png" pageId="1" pageNumber="114" startId="1.[136,235,2040,2064]" targetBox="[154,1431,1331,2012]" targetPageId="1">
<paragraph id="7F207E1CFFFBF001ADA5CDCBFBD9F782" blockId="1.[136,1028,2040,2064]" box="[136,1028,2040,2064]" pageId="1" pageNumber="114">
<emphasis id="4DEBA20EFFFBF001ADA5CDCBFED9F782" bold="true" box="[136,260,2040,2064]" pageId="1" pageNumber="114">FIGURE 1.</emphasis>
Fossil locality.
<emphasis id="4DEBA20EFFFBF001AC89CDCBFE6BF782" bold="true" box="[420,438,2040,2064]" pageId="1" pageNumber="114">A</emphasis>
, Map of fossil locality.
<emphasis id="4DEBA20EFFFBF001AF85CDCBFD64F782" bold="true" box="[680,697,2040,2064]" pageId="1" pageNumber="114">B</emphasis>
, Photograph of fossil collecting.
</paragraph>
</caption>
<caption id="2BE02E94FFF8F002ADA5CCA9FB3CF944" ID-DOI="http://doi.org/10.5281/zenodo.6530396" ID-Zenodo-Dep="6530396" httpUri="https://zenodo.org/record/6530396/files/figure.png" pageId="2" pageNumber="115" startId="2.[136,235,1690,1714]" targetBox="[215,1372,189,1667]" targetPageId="2">
<paragraph id="7F207E1CFFF8F002ADA5CCA9FB3CF944" blockId="2.[136,1452,1690,1750]" pageId="2" pageNumber="115">
<emphasis id="4DEBA20EFFF8F002ADA5CCA9FEDCF920" bold="true" box="[136,257,1690,1714]" pageId="2" pageNumber="115">FIGURE 2.</emphasis>
<taxonomicName id="B89F059FFFF8F002AC29CCA9FD8EF920" authority="HUANG &amp; LIU &amp; LIAN &amp; FU &amp; NEL, 2022" authorityName="HUANG &amp; LIU &amp; LIAN &amp; FU &amp; NEL" authorityYear="2022" box="[260,595,1690,1714]" class="Insecta" family="Calopterygidae" genus="Guangxicalopteryx" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="115" phylum="Arthropoda" rank="species" species="huashanensis" status="gen. et sp. nov.">
<emphasis id="4DEBA20EFFF8F002AC29CCA9FD8EF920" box="[260,595,1690,1714]" italics="true" pageId="2" pageNumber="115">Guangxicalopteryx huashanensis</emphasis>
</taxonomicName>
<emphasis id="4DEBA20EFFF8F002AF7BCCA9FD37F920" bold="true" box="[598,746,1690,1714]" pageId="2" pageNumber="115">
<taxonomicNameLabel id="56D81F75FFF8F002AF7BCCA9FD37F920" box="[598,746,1690,1714]" pageId="2" pageNumber="115" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype NIGP179898, photographs.
<emphasis id="4DEBA20EFFF8F002A945CCA9FBA7F920" bold="true" box="[1128,1146,1690,1714]" pageId="2" pageNumber="115">A</emphasis>
, Counterpart (NIGP179898b).
<emphasis id="4DEBA20EFFF8F002ADA5CC8DFF44F944" bold="true" box="[136,153,1726,1750]" pageId="2" pageNumber="115">B</emphasis>
, Counterpart under alcohol.
<emphasis id="4DEBA20EFFF8F002AC91CC8DFE13F944" bold="true" box="[444,462,1726,1750]" pageId="2" pageNumber="115">C</emphasis>
, Part (NIGP179898a), covered with a carbonaceous film. Scale bars = 5 mm.
</paragraph>
</caption>
<subSubSection id="37852D97FFF8F002AD90CD29FE76F8CA" pageId="2" pageNumber="115" type="materials_examined">
<paragraph id="7F207E1CFFF8F002AD90CD29FE76F8CA" blockId="2.[136,778,1818,2061]" pageId="2" pageNumber="115">
<materialsCitation id="CFF77441FFF8F002AD90CD29FE7AF8CA" ID-GBIF-Occurrence="3763880301" county="Ningming County" location="lower Oligocene" pageId="2" pageNumber="115" specimenCount="1" typeStatus="holotype">
<emphasis id="4DEBA20EFFF8F002AD90CD29FE33F8A6" bold="true" box="[189,494,1818,1844]" pageId="2" pageNumber="115">
<typeStatus id="A024C0BEFFF8F002AD90CD29FF25F8A6" box="[189,248,1818,1844]" pageId="2" pageNumber="115">Type</typeStatus>
locality and horizon.
</emphasis>
A locality near
<collectingCounty id="96410690FFF8F002AFB5CD29FF03F8CA" pageId="2" pageNumber="115">Ningming County</collectingCounty>
;
<location id="7A4028C7FFF8F002ADC4CD0DFE7AF8CA" LSID="urn:lsid:plazi:treatment:F736CF0AFFFBF003AE07C9BFFB34FDC7:7A4028C7FFF8F002ADC4CD0DFE7AF8CA" box="[233,423,1854,1880]" county="Ningming County" name="lower Oligocene" pageId="2" pageNumber="115">lower Oligocene</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="37852D97FFF8F003AD90CD51FB34FDC7" lastPageId="3" lastPageNumber="116" pageId="2" pageNumber="115" type="description">
<paragraph id="7F207E1CFFF8F003AD90CD51FB34FDC7" blockId="2.[136,778,1818,2061]" lastBlockId="3.[810,1452,171,597]" lastPageId="3" lastPageNumber="116" pageId="2" pageNumber="115">
<emphasis id="4DEBA20EFFF8F002AD90CD51FE92F8EE" bold="true" box="[189,335,1890,1916]" pageId="2" pageNumber="115">Description.</emphasis>
Wing base destroyed; wing completely darkened, possibly corresponding to a blue metallic original coloration as it frequently occurs among the
<taxonomicName id="B89F059FFFF8F002ADA5CDFDFEEEF87A" authorityName="Selys Longchamps" authorityYear="1850" box="[136,307,1998,2024]" class="Insecta" family="Calopterygidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="115" phylum="Arthropoda" rank="family">Calopterygidae</taxonomicName>
(especially in male specimens); length of preserved part 42.0 mm long, wing
<quantity id="B867D3F9FFF8F002AF75CDC1FD1CF79F" box="[600,705,2034,2061]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="2" pageNumber="115" unit="mm" value="14.2">14.2 mm</quantity>
wide, distance between base and nodus unknown, between arculus and nodus
<emphasis id="4DEBA20EFFF8F002A935CD0CFBE7F8CA" box="[1048,1082,1855,1880]" italics="true" pageId="2" pageNumber="115">ca.</emphasis>
<quantity id="B867D3F9FFF8F002A967CD0CFB71F8CB" box="[1098,1196,1855,1881]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="115" unit="mm" value="7.1">7.1 mm</quantity>
, between nodus and pterostigma
<quantity id="B867D3F9FFF8F002AE90CD51FBF7F8EF" box="[957,1066,1890,1917]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" pageId="2" pageNumber="115" unit="mm" value="29.5">29.5 mm</quantity>
, between pterostigma and wing apex 5.0 mm; pterostigma short but strongly sclerotized,
<quantity id="B867D3F9FFF8F002AE07CD98FC5DF857" box="[810,896,1963,1989]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="115" unit="mm" value="2.1">2.1 mm</quantity>
long,
<quantity id="B867D3F9FFF8F002AEE5CD98FBC2F857" box="[968,1055,1963,1989]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="115" unit="mm" value="0.8">0.8 mm</quantity>
wide, covering five small cells; no pterostigmal brace; discoidal and subdiscoidal spaces not preserved; at least 14 antenodal crossveins preserved but they were probably much more numerous;
<emphasis id="4DEBA20EFFF9F003AF92CCF8FD3CF976" box="[703,737,1739,1764]" italics="true" pageId="3" pageNumber="116">ca.</emphasis>
60 postnodal crossveins between nodus and pterostigma; bases of RP3/4 and IR2 close to arculus but poorly preserved; base of RP2 two cells distad subnodus; RP3/4 apparently forked distally; no oblique vein O; base of IR1 (first longitudinal vein in area between RP1 and RP2) 11 cells distad that of RP2; two long supplementary longitudinal veins between RA and IR1, most basal one being midway between base of RP2 and pterostigma; two shorter intercalaries between these elongate intercalaries; two other such intercalary veins between IR1 and RP2; a strong elongate intercalary longitudinal vein between RP2 and IR2; area between IR2 and RP3/4 greatly broadened distally with several secondary longitudinal veins between them; area between RP3/4 and MA with two such veins in its distal half; postdiscoidal area rather narrow, with a very long intercalary longitudinal vein; 1-3 rows of cells in area between MP and CuAa; 2-3 rows of cells between CuAa and posterior wing margin, with only one row near its base; MP and CuAa weakly curved; CuA not secondarily forked into two strong branches CuAa and CuAb (CuAb very short and weak).
</paragraph>
</subSubSection>
</treatment>
</document>