267 lines
33 KiB
XML
267 lines
33 KiB
XML
<document id="90DCF759F16F05F5663A58350872E26C" ID-DOI="10.5281/zenodo.201036" ID-GBIF-Dataset="e32d8614-7fee-4095-921d-fcd6c8dfe476" ID-ISSN="1175-5326" ID-Zenodo-Dep="201036" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460272492680" checkinUser="plazi" docAuthor="Mendoza, Jose Christopher E. & Ng, Peter K. L." docDate="2011" docId="E3508794B30EB92EAEC843E2FE9CFB65" docLanguage="en" docName="zt02809p066.pdf" docOrigin="Zootaxa 2809" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leelumnus radium Mendoza & Ng, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="65" masterDocId="1F69FFECB30CB929AE5F414BFFD4FF8F" masterDocTitle="Leelumnus radium, a new genus and species of pilumnid crab from marine encrusting communities in Singapore (Crustacea: Decapoda: Brachyura: Pilumnidae)" masterLastPageNumber="66" masterPageNumber="58" pageNumber="60" updateTime="1698247077057" updateUser="plazi">
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<mods:mods id="093AF1AEF9607A1252551C4E5E055D15" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0863843ACAB9725A6C9B8A3072FA3DC6">
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<mods:title id="585305D6F52FE8276D6DFE07DBC86C5B">Leelumnus radium, a new genus and species of pilumnid crab from marine encrusting communities in Singapore (Crustacea: Decapoda: Brachyura: Pilumnidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="0B92D0F1512AD738276CD545B08C48E3" type="personal">
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<mods:role id="1C4D7DB83E630F9A8D161110C6E21D6C">
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<mods:roleTerm id="A85062946B7C6698BA562E18C11C6B53">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="104BDF46F24A7808450D7246C2C5909E">Mendoza, Jose Christopher E.</mods:namePart>
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</mods:name>
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<mods:name id="370A9C46A7F0B8C700C5A11032487245" type="personal">
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<mods:role id="5665C81A7B2F1590EF0CAB4B5EBFB6CE">
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<mods:roleTerm id="011604D9DEA7DB8A0B37DBC0F830829A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F1C0280F87D93234FD7835EF07D495C2">Ng, Peter K. L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="38CE27A53F35008D869838B37D112179">text</mods:typeOfResource>
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<mods:relatedItem id="33F78E9A1879B19EA2AEFAAD7359A930" type="host">
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<mods:titleInfo id="7B8B9CA02E2CEEBEF6B5A2C80C14CA0B">
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<mods:title id="9ADDC3EAE60B104819D9931B1738D164">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FA6990C6D797B786B0C33908CC4EA153">
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<mods:date id="DB8D26CBA932BF0D1E66596784ED8093">2011</mods:date>
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<mods:detail id="A6E967B79B6F70E5C2F4DC05C0FB1930" type="volume">
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<mods:number id="17D288480581BC155F088DDABC3817CD">2809</mods:number>
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</mods:detail>
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<mods:extent id="005C4FA3B8FBF57E362C0DB983E39381" unit="page">
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<mods:start id="7AE37E1C1E398A2CF94D1B58CA7FA48E">58</mods:start>
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<mods:end id="5F2BE075DE30EB7256BBCCF1AEFE566C">66</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="BA6E64DC197EE77D4ACBC3E94BE86FFC">journal article</mods:classification>
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<mods:identifier id="DAF11F467FF69337F13B55302A3F62CC" type="DOI">10.5281/zenodo.201036</mods:identifier>
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<mods:identifier id="39008AED16FDE4ACA4E3BE45A32610C6" type="GBIF-Dataset">e32d8614-7fee-4095-921d-fcd6c8dfe476</mods:identifier>
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<mods:identifier id="28657F0F20731F3EB070EC20D2A2FDBE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C6B73D365F0A3D5AB00F0D62FCF5F0A3" type="Zenodo-Dep">201036</mods:identifier>
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</mods:mods>
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<treatment id="E3508794B30EB92EAEC843E2FE9CFB65" ID-DOI="http://doi.org/10.5281/zenodo.6184235" ID-GBIF-Taxon="119395389" ID-Zenodo-Dep="6184235" LSID="urn:lsid:plazi:treatment:E3508794B30EB92EAEC843E2FE9CFB65" httpUri="http://treatment.plazi.org/id/E3508794B30EB92EAEC843E2FE9CFB65" lastPageId="7" lastPageNumber="65" pageId="2" pageNumber="60">
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<subSubSection id="23E36509B30EB92BAEC843E2FE8AFD6B" pageId="2" pageNumber="60" type="nomenclature">
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<paragraph id="6B463682B30EB92BAEC843E2FE36FD4D" blockId="2.[151,482,680,740]" box="[151,482,680,707]" pageId="2" pageNumber="60">
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<heading id="300E81EEB30EB92BAEC843E2FE36FD4D" bold="true" box="[151,482,680,707]" fontSize="11" level="1" pageId="2" pageNumber="60" reason="1">
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<emphasis id="598DEA90B30EB92BAEC843E2FE36FD4D" bold="true" box="[151,482,680,707]" pageId="2" pageNumber="60">
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<taxonomicName id="ACF94D01B30EB92BAEC843E2FEA9FD4C" ID-CoL="3SX5F" box="[151,381,681,707]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="radium" status="sp. nov.">
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<emphasis id="598DEA90B30EB92BAEC843E2FEA9FD4C" bold="true" box="[151,381,681,707]" italics="true" pageId="2" pageNumber="60">Leelumnus radium</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="42BE57EBB30EB92BAFDB43E3FE36FD4D" box="[388,482,680,706]" pageId="2" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="6B463682B30EB92BAEC84387FE8AFD6B" blockId="2.[151,482,680,740]" box="[151,350,716,740]" pageId="2" pageNumber="60">
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(
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<figureCitation id="F3C22A07B30EB92BAEC04387FEDEFD6B" box="[159,266,716,740]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1885,1908]" captionStartId-1="4.[151,250,1978,2001]" captionStartId-2="5.[151,250,1220,1243]" captionTargetBox-0="[151,1436,195,1860]" captionTargetBox-1="[286,1301,193,1956]" captionTargetBox-2="[283,1304,193,1198]" captionTargetId-0="figure@3.[151,1436,195,1861]" captionTargetId-1="figure@4.[286,1301,193,1957]" captionTargetId-2="figure@5.[283,1304,193,1199]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Leelumnus radium gen. et sp. nov. A, holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore; habitus, recently ethanol-preserved coloration; B, paratype male, 14.6 × 9.8 mm (ZRC 2011.0039), colour in life, in-situ on encrusting community." captionText-1="FIGURE 2. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, dorsal view; B, anterior view; C, ventral view." captionText-2="FIGURE 3. Leelumnus radium gen. et sp. nov., holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, major chela, external view; B, minor chela, external view." httpUri-0="https://zenodo.org/record/201037/files/figure.png" httpUri-1="https://zenodo.org/record/201038/files/figure.png" httpUri-2="https://zenodo.org/record/201039/files/figure.png" pageId="2" pageNumber="60">Figs. 1–3</figureCitation>
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,
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<figureCitation id="F3C22A07B30EB92BAF494387FEF1FD6B" box="[278,293,716,740]" captionStart="FIGURE 4" captionStartId="5.[151,250,1938,1961]" captionTargetBox="[151,1436,1354,1917]" captionTargetId="figure@5.[151,1436,1354,1918]" captionTargetPageId="5" captionText="FIGURE 4. Female thoracic sternum and vulvae. A, Leelumnus radium gen. et sp. nov., paratype, 12.7 × 8.0 mm (ZRC 2011.0037), Singapore; B, Aniptumnus quadridentatus (De Man, 1895), 11.0 × 7.1 mm (ZRC 1989.3649 – 3660), Singapore." httpUri="https://zenodo.org/record/201040/files/figure.png" pageId="2" pageNumber="60">4</figureCitation>
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A, 5)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="23E36509B30EB92BAEC84258FB91FBC3" pageId="2" pageNumber="60" type="materials_examined">
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<paragraph id="6B463682B30EB92BAEC84258FB91FBC3" blockId="2.[151,1437,787,2036]" pageId="2" pageNumber="60">
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<emphasis id="598DEA90B30EB92BAEC84258FE54FCA3" bold="true" box="[151,384,787,812]" pageId="2" pageNumber="60">Material examined.</emphasis>
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<typeStatus id="B4428820B30EB92BAFD7425FFE24FCA3" box="[392,496,788,812]" pageId="2" pageNumber="60" type="holotype">Holotype</typeStatus>
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male (18.1 ×
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<quantity id="AC019B67B30EB92BACD2425FFD25FCA3" box="[653,753,788,812]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="2" pageNumber="60" unit="mm" value="11.6">11.6 mm</quantity>
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) (
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<collectionCode id="0DE8AE47B30EB92BAD57425FFC94FCA3" box="[776,832,788,812]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="60">ZRC</collectionCode>
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2011.0036), Paku Buoy,
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<geoCoordinate id="0ECD5045B30EB92BAA02425FFB3BFCA3" box="[1117,1263,788,812]" direction="north" orientation="latitude" pageId="2" pageNumber="60" precision="1" value="1.3923167">01º23.539´N</geoCoordinate>
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,
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<geoCoordinate id="0ECD5045B30EB92BAAA4425FFA4CFCA3" box="[1275,1432,788,812]" direction="east" orientation="longitude" pageId="2" pageNumber="60" precision="1" value="104.002365">104º00.142´E</geoCoordinate>
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, off Changi Point, coll. S.L.M. Teo
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<emphasis id="598DEA90B30EB92BAC754272FDB0FCDF" box="[554,612,824,848]" italics="true" pageId="2" pageNumber="60">et al.</emphasis>
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,
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||
<date id="1F471042B30EB92BAC2A427CFCC4FCC0" box="[629,784,823,848]" pageId="2" pageNumber="60" value="2010-04-15">15 Apr. 2010</date>
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.
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<typeStatus id="B4428820B30EB92BAD434273FC59FCDF" box="[796,909,824,848]" pageId="2" pageNumber="60" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="7DFFFD0BB30EB92BADC5427CFC33FCC0" box="[922,999,823,847]" pageId="2" pageNumber="60" type="male">1 male</specimenCount>
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||
(9.6 ×
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||
<quantity id="AC019B67B30EB92BAA60427CFB4FFCC0" box="[1087,1179,823,847]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="60" unit="mm" value="6.5">6.5 mm</quantity>
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||
),
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||
<specimenCount id="7DFFFD0BB30EB92BAAEF427CFAC7FCC0" box="[1200,1299,823,847]" pageId="2" pageNumber="60" type="female">1 female</specimenCount>
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||
(12.7 × 8.0 mm), 1 ovigerous female, (11.5 ×
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||
<quantity id="AC019B67B30EB92BAC404217FDADFCFB" box="[543,633,860,884]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="60" unit="mm" value="7.5">7.5 mm</quantity>
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) (
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||
<collectionCode id="0DE8AE47B30EB92BACCF4217FD1CFCFB" box="[656,712,860,884]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="60">ZRC</collectionCode>
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||
2011.0037), Lek’s oyster farm, off northeastern Tekong Island, coll. S.L.M. Teo
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||
<emphasis id="598DEA90B30EB92BAF0942CAFE5BFC17" box="[342,399,896,920]" italics="true" pageId="2" pageNumber="60">et al.</emphasis>
|
||
,
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||
<date id="1F471042B30EB92BAFC24234FDEEFC18" box="[413,570,895,920]" pageId="2" pageNumber="60" value="2010-08-26">26 Aug. 2010</date>
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||
;
|
||
<specimenCount id="7DFFFD0BB30EB92BAC184234FD46FC18" box="[583,658,895,919]" pageId="2" pageNumber="60" type="male">1 male</specimenCount>
|
||
(12.6 ×
|
||
<quantity id="AC019B67B30EB92BACAC4234FC9AFC18" box="[755,846,895,919]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="60" unit="mm" value="8.5">8.5 mm</quantity>
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||
),
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||
<specimenCount id="7DFFFD0BB30EB92BAD3E4234FC17FC18" box="[865,963,895,919]" pageId="2" pageNumber="60" type="female">1 female</specimenCount>
|
||
(10.0 ×
|
||
<quantity id="AC019B67B30EB92BAA7C4234FBAEFC18" box="[1059,1146,895,919]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="2" pageNumber="60" unit="mm" value="6.9">6.9 mm</quantity>
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||
) (
|
||
<collectionCode id="0DE8AE47B30EB92BAACE4234FB1DFC18" box="[1169,1225,895,919]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="60">ZRC</collectionCode>
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||
2011.0038), buoy,
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||
<geoCoordinate id="0ECD5045B30EB92BAEC842EFFEFDFC33" box="[151,297,932,956]" direction="north" orientation="latitude" pageId="2" pageNumber="60" precision="1" value="1.24465">01º14.679´N</geoCoordinate>
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||
,
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||
<geoCoordinate id="0ECD5045B30EB92BAF6B42EFFE05FC33" box="[308,465,932,956]" direction="east" orientation="longitude" pageId="2" pageNumber="60" precision="1" value="103.86967">103º52.180´E</geoCoordinate>
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||
, off eastern Sentosa Island, coll. S.L.M. Teo
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||
<emphasis id="598DEA90B30EB92BAD9842EEFC2BFC33" box="[967,1023,932,956]" italics="true" pageId="2" pageNumber="60">et al.</emphasis>
|
||
,
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||
<date id="1F471042B30EB92BAA5242EEFB5BFC33" box="[1037,1167,932,956]" pageId="2" pageNumber="60" value="2011-01-07">7 Jan. 2011</date>
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||
;
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||
<specimenCount id="7DFFFD0BB30EB92BAAC442EEFB26FC33" box="[1179,1266,932,956]" pageId="2" pageNumber="60" type="male">7 males</specimenCount>
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||
(7.1 X 6.0 mm - 16.3 X
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||
<quantity id="AC019B67B30EB92BAEA4428CFEB7FC50" box="[251,355,967,991]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="60" unit="mm" value="10.5">10.5 mm</quantity>
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||
),
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||
<specimenCount id="7DFFFD0BB30EB92BAF2A428CFE35FC50" box="[373,481,967,991]" pageId="2" pageNumber="60" type="female">8 females</specimenCount>
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||
(7 ovigerous, 6.8 X
|
||
<quantity id="AC019B67B30EB92BAC96428CFCCAFC6F" box="[713,798,967,992]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="2" pageNumber="60" unit="mm" value="5.4">5.4 mm</quantity>
|
||
- 12.7 X
|
||
<quantity id="AC019B67B30EB92BADD6428CFC3AFC50" box="[905,1006,967,991]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" pageId="2" pageNumber="60" unit="mm" value="8.75">8.75 mm</quantity>
|
||
) (
|
||
<collectionCode id="0DE8AE47B30EB92BAA5B428CFBE8FC50" box="[1028,1084,967,991]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="60">ZRC</collectionCode>
|
||
2011.0039),
|
||
<specimenCount id="7DFFFD0BB30EB92BAA8E428CFACAFC50" box="[1233,1310,967,991]" pageId="2" pageNumber="60" type="male">1 male</specimenCount>
|
||
, (9.8 X
|
||
<quantity id="AC019B67B30EB92BAB26428CFF10FB8B" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="60" unit="mm" value="6.7">6.7 mm</quantity>
|
||
) 1 ovigerous female (10.1 X
|
||
<quantity id="AC019B67B30EB92BAC4942A7FDBBFB8B" box="[534,623,1004,1028]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="60" unit="mm" value="6.7">6.7 mm</quantity>
|
||
) (NHM),
|
||
<specimenCount id="7DFFFD0BB30EB92BACBC42A7FCE6FB8B" box="[739,818,1004,1028]" pageId="2" pageNumber="60" type="male">1 male</specimenCount>
|
||
, (8.5 X
|
||
<quantity id="AC019B67B30EB92BADCE42A7FC38FB8B" box="[913,1004,1004,1028]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="60" unit="mm" value="5.7">5.7 mm</quantity>
|
||
), 1 ovigerous female, (8.8 X
|
||
<quantity id="AC019B67B30EB92BAB6342A7FA40FB8B" box="[1340,1428,1004,1028]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="60" unit="mm" value="5.9">5.9 mm</quantity>
|
||
) (
|
||
<collectionCode id="0DE8AE47B30EB92BAEC04544FF2DFBA8" LSID="urn:lsid:biocol.org:col:34992" box="[159,249,1039,1063]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="60">RMNH</collectionCode>
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||
), Lek’s oyster farm, off northeastern Tekong Island, amongst encrusting community on submerged nets and ropes, coll. J.C. Mendoza & S.K. Tan,
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||
<date id="1F471042B30EB92BAC27457FFCDAFBC3" box="[632,782,1076,1100]" pageId="2" pageNumber="60" value="2011-02-28">28 Feb. 2011</date>
|
||
. All localities in
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||
<collectingCountry id="13EE7612B30EB92BAD94457FFB94FBC3" box="[971,1088,1076,1100]" name="Singapore" pageId="2" pageNumber="60">Singapore</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23E36509B30EB92EAE9A451CFD12FC48" lastPageId="7" lastPageNumber="65" pageId="2" pageNumber="60" type="description">
|
||
<paragraph id="6B463682B30EB92BAE9A451CFDCAF903" blockId="2.[151,1437,787,2036]" pageId="2" pageNumber="60">
|
||
<emphasis id="598DEA90B30EB92BAE9A451CFE3FFBFF" bold="true" box="[197,491,1111,1136]" pageId="2" pageNumber="60">
|
||
Description of
|
||
<typeStatus id="B4428820B30EB92BAF224513FE32FBFF" box="[381,486,1112,1136]" pageId="2" pageNumber="60" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Carapace transversely subhexagonal, width 1.5–1.6 times length; regions poorly defined; dorsal surface convex, mostly smooth except for patches of minute granules on protogastric and mesobranchial regions, sparsely covered with long and short, plumose setae. Front moderately produced beyond orbits, divided into 2 lobes by V-shaped notch continuing posteriorly as narrow fissure reaching tip of proepistome ventrally; anterior margin of lobes gently concave, granulate, mesial angle more produced than lateral, lateral angle low, not distinctly separated from supraorbital margin; fronto-orbital region relatively narrow transversely,
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||
<emphasis id="598DEA90B30EB92BAB0F4446FAA5FAAB" box="[1360,1393,1293,1316]" italics="true" pageId="2" pageNumber="60">ca.</emphasis>
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||
0.4 times maximum carapace width. Supraorbital margin entire but lined with small granules. Anterolateral margin granulate; exorbital angle low, not prominently projecting; rest of anterolateral margin with 4 distinct teeth; first tooth smallest, connected to exorbital angle by distinct, horizontal, granulate crest; subsequent teeth projecting laterally, curving anteriorly, second broadly triangular, third and fourth relatively more acute. Posterolateral margin almost straight to slightly sinuous, converging towards gently convex posterior carapace margin. Basal antennal article subrectangular, filling orbital hiatus; flagellum long. Antennules folding transversely; proepistome (interantennular septum) narrow. Orbits transverse, relatively short, rounded; eyes filling entire orbital space; distal part of short ocular peduncle covered with minute granules; cornea prominent, rounded. Infraorbital margin distinctly granulate, internal infra-orbital tooth projecting, triangular, visible from dorsal view. Suborbital, subhepatic, pterygostomial regions finely granulate.
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||
</paragraph>
|
||
<paragraph id="6B463682B30EB92BAE9A47D3FEDEF8EB" blockId="2.[151,1437,787,2036]" pageId="2" pageNumber="60">Epistome well-produced; posterior border with acute median projection, with prominent notches laterally. Endostome with well-defined ridges forming channel for efferent water current with endopod of first maxillipeds. Third maxillipeds short, stout; merus quadrate, anterolateral angle rounded, not prominently produced, surface finely granular, especially along margins; palps long; ischium subrectangular, with shallow submedian sulcus; exopod stout, distal edge almost reaching anterior edge of merus, inner subdistal tooth prominent, with well-developed flagellum.</paragraph>
|
||
<paragraph id="6B463682B30EB92BAE9A463BFC4CF87B" blockId="2.[151,1437,787,2036]" pageId="2" pageNumber="60">Thoracic sternum narrow, surface smooth to finely granulate, sparsely setose. Sternites 1, 2 fused; sternites 2, 3 completely separated by suture; sternites 3/4 partially fused, suture demarcated as deep lateral notches; sutures 4/5, 5/6 interrupted medially, 6/7, 7/8 complete; small median, triangular gap between sternites 6, 7. Median line on sternites 4, 6, 7, 8. Sternal press-button on sternite 5, near suture 4/5.</paragraph>
|
||
<caption id="3F86660AB30FB92AAEC84616FEC4F83E" httpUri="https://zenodo.org/record/201037/files/figure.png" pageId="3" pageNumber="61" targetBox="[151,1436,195,1860]" targetPageId="3">
|
||
<paragraph id="6B463682B30FB92AAEC84616FEC4F83E" blockId="3.[151,1436,1885,1969]" pageId="3" pageNumber="61">
|
||
<emphasis id="598DEA90B30FB92AAEC84616FEDBF8FC" bold="true" box="[151,271,1885,1908]" pageId="3" pageNumber="61">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="ACF94D01B30FB92AAF404615FE0BF8FB" box="[287,479,1886,1908]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="61" phylum="Arthropoda" rank="species" species="radium">
|
||
<emphasis id="598DEA90B30FB92AAF404615FE0BF8FB" box="[287,479,1886,1908]" italics="true" pageId="3" pageNumber="61">Leelumnus radium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="598DEA90B30FB92AAFB74615FD5BF8FC" bold="true" box="[488,655,1886,1907]" pageId="3" pageNumber="61">gen. et sp. nov.</emphasis>
|
||
A, holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore; habitus, recently ethanol-preserved coloration; B, paratype male, 14.6 × 9.8 mm (ZRC 2011.0039), colour in life, in-situ on encrusting community.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3F86660AB308B92DAEC846F1FE3DF860" httpUri="https://zenodo.org/record/201038/files/figure.png" pageId="4" pageNumber="62" targetBox="[286,1301,193,1956]" targetPageId="4">
|
||
<paragraph id="6B463682B308B92DAEC846F1FE3DF860" blockId="4.[151,1436,1978,2031]" pageId="4" pageNumber="62">
|
||
<emphasis id="598DEA90B308B92DAEC846F1FED9F85F" bold="true" box="[151,269,1978,2001]" pageId="4" pageNumber="62">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="ACF94D01B308B92DAF4646F1FE03F85F" box="[281,471,1978,2000]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="62" phylum="Arthropoda" rank="species" species="radium">
|
||
<emphasis id="598DEA90B308B92DAF4646F1FE03F85F" box="[281,471,1978,2000]" italics="true" pageId="4" pageNumber="62">Leelumnus radium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="598DEA90B308B92DAF8246F0FDAFF85F" bold="true" box="[477,635,1979,2000]" pageId="4" pageNumber="62">gen. et sp. nov.</emphasis>
|
||
, holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, dorsal view; B, anterior view; C, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3F86660AB309B92CAEC8458FFD8AFB76" httpUri="https://zenodo.org/record/201039/files/figure.png" pageId="5" pageNumber="63" targetBox="[283,1304,193,1198]" targetPageId="5">
|
||
<paragraph id="6B463682B309B92CAEC8458FFD8AFB76" blockId="5.[151,1436,1220,1273]" pageId="5" pageNumber="63">
|
||
<emphasis id="598DEA90B309B92CAEC8458FFED8FB55" bold="true" box="[151,268,1220,1243]" pageId="5" pageNumber="63">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="ACF94D01B309B92CAF46458EFE02FB54" box="[281,470,1221,1243]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="63" phylum="Arthropoda" rank="species" species="radium">
|
||
<emphasis id="598DEA90B309B92CAF46458EFE02FB54" box="[281,470,1221,1243]" italics="true" pageId="5" pageNumber="63">Leelumnus radium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="598DEA90B309B92CAF82458EFDAEFB55" bold="true" box="[477,634,1221,1242]" pageId="5" pageNumber="63">gen. et sp. nov.</emphasis>
|
||
, holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, major chela, external view; B, minor chela, external view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3F86660AB309B92CAEC846D9FAAAF847" httpUri="https://zenodo.org/record/201040/files/figure.png" pageId="5" pageNumber="63" targetBox="[151,1436,1354,1917]" targetPageId="5">
|
||
<paragraph id="6B463682B309B92CAEC846D9FAAAF847" blockId="5.[151,1436,1938,1992]" pageId="5" pageNumber="63">
|
||
<emphasis id="598DEA90B309B92CAEC846D9FEDBF827" bold="true" box="[151,271,1938,1961]" pageId="5" pageNumber="63">FIGURE 4</emphasis>
|
||
. Female thoracic sternum and vulvae. A,
|
||
<taxonomicName id="ACF94D01B309B92CAC9046D8FC44F826" box="[719,912,1939,1961]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="63" phylum="Arthropoda" rank="species" species="radium">
|
||
<emphasis id="598DEA90B309B92CAC9046D8FC44F826" box="[719,912,1939,1961]" italics="true" pageId="5" pageNumber="63">Leelumnus radium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="598DEA90B309B92CADC546D8FB95F827" bold="true" box="[922,1089,1939,1960]" pageId="5" pageNumber="63">gen. et sp. nov.</emphasis>
|
||
, paratype, 12.7 × 8.0 mm (ZRC 2011.0037), Singapore; B,
|
||
<taxonomicName id="ACF94D01B309B92CAFF746F9FCB8F847" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[424,876,1970,1992]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="63" phylum="Arthropoda" rank="species" species="quadridentatus">
|
||
<emphasis id="598DEA90B309B92CAFF746F9FD14F847" box="[424,704,1970,1992]" italics="true" pageId="5" pageNumber="63">Aniptumnus quadridentatus</emphasis>
|
||
(De Man, 1895)
|
||
</taxonomicName>
|
||
, 11.0 × 7.1 mm (ZRC 1989.3649–3660), Singapore.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3F86660AB30AB92FAEC84617FB1BF841" httpUri="https://zenodo.org/record/201041/files/figure.png" pageId="6" pageNumber="64" targetBox="[195,1380,194,1825]" targetPageId="6">
|
||
<paragraph id="6B463682B30AB92FAEC84617FB1BF841" blockId="6.[151,1436,1884,1998]" pageId="6" pageNumber="64">
|
||
<emphasis id="598DEA90B30AB92FAEC84617FEC0F8FD" bold="true" box="[151,276,1884,1907]" pageId="6" pageNumber="64">FIGURE 5.</emphasis>
|
||
<taxonomicName id="ACF94D01B30AB92FAF424617FE08F8FD" box="[285,476,1884,1906]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="64" phylum="Arthropoda" rank="species" species="radium">
|
||
<emphasis id="598DEA90B30AB92FAF424617FE08F8FD" box="[285,476,1884,1906]" italics="true" pageId="6" pageNumber="64">Leelumnus radium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="598DEA90B30AB92FAFBB4616FD5DF8FD" bold="true" box="[484,649,1885,1906]" pageId="6" pageNumber="64">gen. et sp. nov.</emphasis>
|
||
, holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore. A, left third maxilliped, external view (setae denuded); B, anterior thoracic sternum; C, posterior thoracic sternum and penis; D, abdomen with telson; E, coxa, basis-ischium and merus, and, F, propodus and dactylus of right P5, dorsal view; G, left G1, external view; distal tip of left G1, I, external view, J, internal view; H, left G2, external view. Scale bars: A–H = 1.0 mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6B463682B30BB92EAE9A41DCFA9CFEEB" blockId="7.[151,1437,151,1259]" pageId="7" pageNumber="65">Chelipeds distinctly unequal. Inner margins of fused basis-ischium and merus granulate. External surface of carpus rugose, with rounded granules; inner distal angle produced as distinct tooth. Major chela robust; palm inflated, external surface smooth, glabrous; fingers distinctly shorter than palm; fixed finger deflexed, with larger teeth on cutting margin than those of dactylus; dactylus with shallow longitudinal groove on external surface. External surface of minor chela with 2 or 3 longitudinal rows of round granules, setose; fingers shorter than palm; fixed finger deflexed, with 2 longitudinal grooves on external surface, cutting edges with several low teeth.</paragraph>
|
||
<paragraph id="6B463682B30BB92EAE9A4024FEEBFDB3" blockId="7.[151,1437,151,1259]" pageId="7" pageNumber="65">Ambulatory legs moderately long, P3 longest, coxa-to-dactylus length subequal to maximum carapace width; setose, especially on carpus, propodus and dactylus. Articles relatively smooth, devoid of granules. Length of merus about 3.1 times width; distal end with distinct submedian, longitudinal notch, joining up with deep, transverse, subdistal groove. Dactylo-propodal locking mechanism present; distal propodal condyle large, rounded; corresponding tubercle on proximal end of dactylus small, truncatiform. Dactylus nearly straight, ending in a curved, chitinous claw.</paragraph>
|
||
<paragraph id="6B463682B30BB92EAE9A430CFBC9FD28" blockId="7.[151,1437,151,1259]" pageId="7" pageNumber="65">Abdomen with all somites freely articulating, surfaces generally smooth, sparsely setose. Somite 6 subquadrate, lateral margins subparallel. Telson subtriangular, median length subequal to basal width; lateral margins almost straight, tip rounded, reaching level of coxo-sternal condyles of cheliped.</paragraph>
|
||
<paragraph id="6B463682B30BB92EAE9A43FFFD72FC9B" blockId="7.[151,1437,151,1259]" pageId="7" pageNumber="65">G1 sinuous, lateral edge lined with stiff, plumose setae; distal part distinctly flared, flattened, with numerous stiff, spiniform, simple setae. G2 short, sigmoid, about one-fifth length of G1. Penis short, papilliform, emerging from tip of swollen sternal condyle of P5 coxa.</paragraph>
|
||
<paragraph id="6B463682B30BB92EAE9A426BFE1AFCF0" blockId="7.[151,1437,151,1259]" pageId="7" pageNumber="65">
|
||
<emphasis id="598DEA90B30BB92EAE9A426BFE6EFCB7" bold="true" box="[197,442,800,824]" pageId="7" pageNumber="65">Female morphology.</emphasis>
|
||
Similar to that of the male in all non-sexual characters. The vulvae are relatively small and have a small sternal cover originating from the posterior margin. The anterior edge of the vulva does not reach sternal suture 5/6 (
|
||
<figureCitation id="F3C22A07B30BB92EAF38422CFE7EFCF0" box="[359,426,871,895]" captionStart="FIGURE 4" captionStartId="5.[151,250,1938,1961]" captionTargetBox="[151,1436,1354,1917]" captionTargetId="figure@5.[151,1436,1354,1918]" captionTargetPageId="5" captionText="FIGURE 4. Female thoracic sternum and vulvae. A, Leelumnus radium gen. et sp. nov., paratype, 12.7 × 8.0 mm (ZRC 2011.0037), Singapore; B, Aniptumnus quadridentatus (De Man, 1895), 11.0 × 7.1 mm (ZRC 1989.3649 – 3660), Singapore." httpUri="https://zenodo.org/record/201040/files/figure.png" pageId="7" pageNumber="65">Fig. 4</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="6B463682B30BB92EAE9A42C0FD12FC48" blockId="7.[151,1437,151,1259]" pageId="7" pageNumber="65">
|
||
<emphasis id="598DEA90B30BB92EAE9A42C0FEC9FC2B" bold="true" box="[197,285,907,932]" pageId="7" pageNumber="65">Colour.</emphasis>
|
||
The background colour is dirty white with numerous patches of purple on the carapace and pereopods; fngers of chelipeds pink with purple base (
|
||
<figureCitation id="F3C22A07B30BB92EAC2B42E4FD61FC48" box="[628,693,943,967]" captionStart="FIGURE 1" captionStartId="3.[151,250,1885,1908]" captionTargetBox="[151,1436,195,1860]" captionTargetId="figure@3.[151,1436,195,1861]" captionTargetPageId="3" captionText="FIGURE 1. Leelumnus radium gen. et sp. nov. A, holotype male, 18.0 × 11.5 mm (ZRC 2011.0036), Singapore; habitus, recently ethanol-preserved coloration; B, paratype male, 14.6 × 9.8 mm (ZRC 2011.0039), colour in life, in-situ on encrusting community." httpUri="https://zenodo.org/record/201037/files/figure.png" pageId="7" pageNumber="65">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23E36509B30BB92EAE9A429FFF1EFBB8" pageId="7" pageNumber="65" type="etymology">
|
||
<paragraph id="6B463682B30BB92EAE9A429FFF1EFBB8" blockId="7.[151,1437,151,1259]" pageId="7" pageNumber="65">
|
||
<emphasis id="598DEA90B30BB92EAE9A429FFE98FC63" bold="true" box="[197,332,980,1004]" pageId="7" pageNumber="65">Etymology.</emphasis>
|
||
The specific epithet is derived from
|
||
<taxonomicName id="ACF94D01B30BB92EACBA429FFC95FC63" box="[741,833,980,1004]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="65" phylum="Arthropoda" rank="species" species="radium">Radium</taxonomicName>
|
||
, the 88th element on the periodic table of elements, to commemorate the occasion of Dr Lee Seng Tee’s 88th birthday in
|
||
<date id="1F471042B30BB92EAD3142B1FC3EFB9D" box="[878,1002,1018,1042]" pageId="7" pageNumber="65" value="2011-04">April 2011</date>
|
||
. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23E36509B30BB92EAE9A4508FE9CFB65" pageId="7" pageNumber="65" type="discussion">
|
||
<paragraph id="6B463682B30BB92EAE9A4508FE9CFB65" blockId="7.[151,1437,151,1259]" pageId="7" pageNumber="65">
|
||
<emphasis id="598DEA90B30BB92EAE9A4508FEEDFBD4" bold="true" box="[197,313,1091,1115]" pageId="7" pageNumber="65">Remarks.</emphasis>
|
||
<taxonomicName id="ACF94D01B30BB92EAF1C4508FDCCFBD4" box="[323,536,1091,1115]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="65" phylum="Arthropoda" rank="species" species="radium">
|
||
<emphasis id="598DEA90B30BB92EAF1C4508FDCCFBD4" box="[323,536,1091,1115]" italics="true" pageId="7" pageNumber="65">Leelumnus radium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="598DEA90B30BB92EAC7D4508FD0EFBD4" bold="true" box="[546,730,1091,1115]" pageId="7" pageNumber="65">gen. et sp. nov.</emphasis>
|
||
, was collected from encrusting communities on floating and semi-submerged structures around
|
||
<collectingCountry id="13EE7612B30BB92EAC7B452CFD4DFBF0" box="[548,665,1127,1151]" name="Singapore" pageId="7" pageNumber="65">Singapore</collectingCountry>
|
||
. It has not been found in any other benthic habitats in
|
||
<collectingCountry id="13EE7612B30BB92EAB55452CFAAFFBF0" box="[1290,1403,1127,1151]" name="Singapore" pageId="7" pageNumber="65">Singapore</collectingCountry>
|
||
as yet. Due to the volume of shipping that goes through
|
||
<collectingCountry id="13EE7612B30BB92EACAC45C1FCBCFB2D" box="[755,872,1162,1186]" name="Singapore" pageId="7" pageNumber="65">Singapore</collectingCountry>
|
||
, it is possible that this may even be an exotic species that has been transported via fouling communities on the hulls of ships or as larvae in ballast water (see
|
||
<bibRefCitation id="0F684B73B30BB92EAB0A45FBFF22FB65" author="Yeo" pageId="7" pageNumber="65" refString="Yeo, D. C. J., Ahyong, S. T., Lodge, D. M., Ng, P. K. L., Naruse, T. & Lane, D. J. W. (2009) Semisubmersible oil platforms: understudied and potentially major vectors of biofouling-mediated invasions. Biofouling, 26 (2), 179 - 186." type="journal article" year="2009">
|
||
Yeo
|
||
<emphasis id="598DEA90B30BB92EABD745FBFF60FB64" italics="true" pageId="7" pageNumber="65">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="0F684B73B30BB92EAF5E4599FEE9FB65" author="Yeo" box="[257,317,1234,1258]" pageId="7" pageNumber="65" refString="Yeo, D. C. J., Carlton, J. T., Teo, S. L. M. & Ng, P. K. L. (2011) An incoming flood on a cryptic stage: understanding alien crustacean invasions in Southeast Asia. pp. 403 - 417. In: Galil, B. S., Clark, P. F. & Carlton, J. T. (Eds.) In the Wrong Place - Alien Marine Crustaceans: Distribution, Biology and Impacts, Springer Series in Invasion Biology, Volume 6." type="journal article" year="2011">2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |