489 lines
57 KiB
XML
489 lines
57 KiB
XML
<document id="604BED0716C3073EECAA4E22908AB4EF" ID-DOI="10.5281/zenodo.156407" ID-GBIF-Dataset="f41d3d44-0e64-4ad1-841f-dea3d4a551e6" ID-ISSN="11755326" ID-Zenodo-Dep="156407" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459595625205" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="E310B73CFF91FFF61E2AFC7C12B3D176" docLanguage="en" docName="zt00164.pdf" docOrigin="Zootaxa 164" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Goniusa caseyi Gusarov, sp. n." docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="1F29CF44FF99FFFB1F22FFEA114FD248" masterDocTitle="A revision of the genus Goniusa Casey, 1906 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="9" updateTime="1698202593239" updateUser="plazi">
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<mods:title id="29A6247E0D45DF33C37DD80CD31B2D07">A revision of the genus Goniusa Casey, 1906 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="4589BEC18B4ADD5DE7F9310EA3FF1D49">Gusarov, Vladimir I.</mods:namePart>
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<mods:title id="3236820E21A9CF554B7E33A56B042A44">Zootaxa</mods:title>
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<mods:date id="EA4034E779477F1BAE8B30B5252F6B95">2003</mods:date>
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<heading id="304EB146FF91FFF31E2AFC7C12CAD1F8" bold="true" box="[264,901,918,944]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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1.
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<taxonomicName id="ACB97DA9FF91FFF31E04FC7C1300D1F8" ID-CoL="3GY2F" authority="Gusarov" authorityName="Gusarov" box="[294,591,918,944]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="caseyi" status="sp. nov.">
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<emphasis id="59CDDA38FF91FFF31E04FC7C1092D1F8" bold="true" box="[294,477,918,944]" italics="true" pageId="8" pageNumber="9">Goniusa caseyi</emphasis>
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Gusarov
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,
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<taxonomicNameLabel id="42FE6743FF91FFF31D7EFC7C13D1D1F8" box="[604,670,918,944]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
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(
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<figureCitation id="F3821AAFFF91FFF31D8CFC7C1267D1F8" box="[686,808,918,944]" captionStart-0="FIGURES 1 5" captionStart-1="FIGURES 6 11" captionStart-2="FIGURES 12 15" captionStartId-0="2.[264,383,1522,1546]" captionStartId-1="4.[264,383,1469,1493]" captionStartId-2="5.[264,383,1787,1811]" captionTargetBox-0="[290,1296,439,1505]" captionTargetBox-1="[298,1287,339,1429]" captionTargetBox-2="[289,1304,581,1774]" captionTargetId-0="figure@2.[280,1306,425,1512]" captionTargetId-1="figure@4.[288,1299,329,1443]" captionTargetId-2="figure@5.[272,1314,569,1776]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1 5. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." captionText-1="FIGURES 6 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 7, 9 11), 0.2 mm (8)." captionText-2="FIGURES 12 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 14)." httpUri-0="https://zenodo.org/record/156408/files/figure.png" httpUri-1="https://zenodo.org/record/156409/files/figure.png" httpUri-2="https://zenodo.org/record/156410/files/figure.png" pageId="8" pageNumber="9">Figs. 115</figureCitation>
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,
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<figureCitation id="F3821AAFFF91FFF31C15FC7C1232D1F8" box="[823,893,918,944]" captionStart-0="FIGURES 20 24" captionStart-1="FIGURES 25 29" captionStart-2="FIGURES 30 36" captionStart-3="FIGURES 37 41" captionStartId-0="9.[264,383,1550,1574]" captionStartId-1="10.[264,383,1768,1792]" captionStartId-2="11.[264,382,1632,1656]" captionStartId-3="12.[264,382,1692,1716]" captionTargetBox-0="[290,1295,579,1513]" captionTargetBox-1="[280,1310,701,1744]" captionTargetBox-2="[288,1303,337,1596]" captionTargetBox-3="[301,1285,783,1653]" captionTargetId-0="figure@9.[282,1306,569,1524]" captionTargetId-1="figure@10.[263,1323,689,1755]" captionTargetId-2="figure@11.[273,1314,326,1608]" captionTargetId-3="figure@12.[288,1299,769,1665]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURES 20 24. Abdominal segment 8 of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (20 21); Aweme, Manitoba (22); and Stonewall, Manitoba (23 24 )). 20 – male tergum 8; 21 – male sternum 8; 22 – apex of male tergum 8; 23 – female tergum 8; 24 – female sternum 8. Scale bar 0.4 mm (20 21, 23 24), 0.2 mm (22)." captionText-1="FIGURES 25 29. Median lobe of aedeagus of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (25 26, 28 29); and Sherborn, Massachusetts (27 )). 25 – median lobe, parameral view; 26 27 – apex of median lobe, parameral view; 28 – median lobe, lateral view; 29 – apex of median lobe, lateral view. Scale bar 0.4 mm (25, 28), 0.2 mm (26 27, 29)." captionText-2="FIGURES 30 36. Genitalia of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (30 31); Texas (32 34); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36 )). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 35 36 – spermatheca. Scale bar 0.1 mm (32 34), 0.2 mm (30 31, 35 36)." captionText-3="FIGURES 37 41. Pronotum of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (37); and Stonewall, Manitoba (38 )) and G. alperti Kistner (holotype (39); female from San Bernardino Mts., California (40), paratype from 10 mi. E of Coville, Washington (41 )). 37, 39 – male pronotum; 38, 40 41 – female pronotum. Scale bar 0.5 mm." httpUri-0="https://zenodo.org/record/156412/files/figure.png" httpUri-1="https://zenodo.org/record/156413/files/figure.png" httpUri-2="https://zenodo.org/record/156414/files/figure.png" httpUri-3="https://zenodo.org/record/156415/files/figure.png" pageId="8" pageNumber="9">2038</figureCitation>
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)
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<subSubSection id="23A355A1FF91FFF31E2AFC0B1213D630" pageId="8" pageNumber="9" type="reference_group">
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<paragraph id="6B06062AFF91FFF31E2AFC0B1214D650" blockId="8.[264,1149,993,1144]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="4BA92104FF91FFF31E2AFC0B1214D650" pageId="8" pageNumber="9">
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<taxonomicName id="ACB97DA9FF91FFF31E2AFC0B13CBD1B0" authority="Casey, 1906: 348" authorityName="Casey" authorityPageNumber="348" authorityYear="1906" box="[264,644,993,1016]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="obtusa">
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<emphasis id="59CDDA38FF91FFF31E2AFC0B10FFD1B0" box="[264,432,993,1016]" italics="true" pageId="8" pageNumber="9">Goniusa obtusa</emphasis>
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:
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<treatmentCitation id="EA18203BFF91FFF31EE7FC0B13CBD1B0" author="Casey" box="[453,644,993,1016]" page="348" pageId="8" pageNumber="9" year="1906">
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<bibRefCitation id="0F287BDBFF91FFF31EE7FC0B1304D1B0" author="Casey" box="[453,587,993,1016]" pageId="8" pageNumber="19" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
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: 348
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</treatmentCitation>
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</taxonomicName>
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(nec
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<treatmentCitation id="EA18203BFF91FFF31DE2FC0B12D2D1B0" author="LeConte" box="[704,925,993,1016]" page="373" pageId="8" pageNumber="9" year="1866">
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<bibRefCitation id="0F287BDBFF91FFF31DE2FC0B122DD1B0" author="LeConte" box="[704,866,993,1016]" pageId="8" pageNumber="19" refString="LeConte, J. L. (1866) Additions to the coleopterous fauna of the United States. No. 1. Proceedings of the Academy of Sciences of Philadelphia, 18, 361 - 394." type="journal article" year="1866">LeConte, 1866</bibRefCitation>
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: 373
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||
</treatmentCitation>
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) (Misidentification).
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<taxonomicName id="ACB97DA9FF91FFF31E2AFBEB1218D650" authority="Casey, 1911: 208 (Misidentification)" authorityName="Casey, 1911: 208 (Misidentification" box="[264,855,1025,1048]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="obtusa">
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||
<emphasis id="59CDDA38FF91FFF31E2AFBEB10FFD650" box="[264,432,1025,1048]" italics="true" pageId="8" pageNumber="9">Goniusa obtusa</emphasis>
|
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:
|
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<treatmentCitation id="EA18203BFF91FFF31EE7FBEB13CCD650" author="Casey" box="[453,643,1025,1048]" page="208" pageId="8" pageNumber="9" year="1911">
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<bibRefCitation id="0F287BDBFF91FFF31EE7FBEB1305D650" author="Casey" box="[453,586,1025,1048]" pageId="8" pageNumber="19" refString="Casey, T. L. (1911) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. The New Era Printing Company, Lancaster, pp. 1 - 245." type="book chapter" year="1911">Casey, 1911</bibRefCitation>
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: 208
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||
</treatmentCitation>
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||
(Misidentification)
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||
</taxonomicName>
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||
.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="6B06062AFF91FFF31E2AFBCB1227D670" blockId="8.[264,1149,993,1144]" box="[264,872,1057,1080]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="4BA92104FF91FFF31E2AFBCB1227D670" box="[264,872,1057,1080]" pageId="8" pageNumber="9">
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<taxonomicName id="ACB97DA9FF91FFF31E2AFBCB122BD670" authority="Fenyes, 1920: 236 (Misidentification)" authorityName="Fenyes, 1920: 236 (Misidentification" box="[264,868,1057,1080]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="obtusa">
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<emphasis id="59CDDA38FF91FFF31E2AFBCB10FFD670" box="[264,432,1057,1080]" italics="true" pageId="8" pageNumber="9">Goniusa obtusa</emphasis>
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:
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<treatmentCitation id="EA18203BFF91FFF31EE7FBCB13DFD670" author="Fenyes" box="[453,656,1057,1080]" page="236" pageId="8" pageNumber="9" year="1920">
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<bibRefCitation id="0F287BDBFF91FFF31EE7FBCB1318D670" author="Fenyes" box="[453,599,1057,1080]" pageId="8" pageNumber="19" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles. Pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
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: 236
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||
</treatmentCitation>
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||
(Misidentification)
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="6B06062AFF91FFF31E2AFBAB1213D630" blockId="8.[264,1149,993,1144]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="4BA92104FF91FFF31E2AFBAB1213D630" pageId="8" pageNumber="9">
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<taxonomicName id="ACB97DA9FF91FFF31E2AFBAB1569D610" authority="Bernhauer & Scheerpeltz, 1926: 597 (Misidentification)" authorityName="Bernhauer & Scheerpeltz, 1926: 597 (Misidentification" box="[264,1062,1089,1112]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="obtusa">
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<emphasis id="59CDDA38FF91FFF31E2AFBAB10FFD610" box="[264,432,1089,1112]" italics="true" pageId="8" pageNumber="9">Goniusa obtusa</emphasis>
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:
|
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<treatmentCitation id="EA18203BFF91FFF31EE7FBAB121DD610" author="Bernhauer" box="[453,850,1089,1112]" page="597" pageId="8" pageNumber="9" year="1926">
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<bibRefCitation id="0F287BDBFF91FFF31EE7FBAB1256D610" author="Bernhauer" box="[453,793,1089,1112]" pageId="8" pageNumber="19" refString="Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer & Scheerpeltz, 1926</bibRefCitation>
|
||
: 597
|
||
</treatmentCitation>
|
||
(Misidentification)
|
||
</taxonomicName>
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||
.
|
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<taxonomicName id="ACB97DA9FF91FFF31E2AFB8B1217D630" authority="Kistner, 1976: 87 (Misidentification)" authorityName="Kistner, 1976: 87 (Misidentification" box="[264,856,1121,1144]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="obtusa">
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<emphasis id="59CDDA38FF91FFF31E2AFB8B10FFD630" box="[264,432,1121,1144]" italics="true" pageId="8" pageNumber="9">Goniusa obtusa</emphasis>
|
||
:
|
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<treatmentCitation id="EA18203BFF91FFF31EE7FB8B13CAD630" author="Kistner" box="[453,645,1121,1144]" page="87" pageId="8" pageNumber="9" year="1976">
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<bibRefCitation id="0F287BDBFF91FFF31EE7FB8B1316D630" author="Kistner" box="[453,601,1121,1144]" pageId="8" pageNumber="19" refString="Kistner, D. H. (1976) Revision and Reclassification of the Genus Goniusa Casey with a Larval Description and Ant Host Records (Coleoptera: Staphylinidae). Sociobiology, 2 (1), 83 - 95." type="journal article" year="1976">Kistner, 1976</bibRefCitation>
|
||
: 87
|
||
</treatmentCitation>
|
||
(Misidentification)
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="23A355A1FF91FFF21E2AFB4312D7D3B6" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="6B06062AFF91FFF31E2AFB43127FD75B" blockId="8.[264,1324,1193,1899]" pageId="8" pageNumber="9">
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<emphasis id="59CDDA38FF91FFF31E2AFB4310F8D68B" bold="true" box="[264,439,1193,1219]" pageId="8" pageNumber="9">
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<typeStatus id="B402B888FF91FFF31E2AFB43100AD68B" box="[264,325,1193,1219]" pageId="8" pageNumber="9">Type</typeStatus>
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material
|
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</emphasis>
|
||
.
|
||
<typeStatus id="B402B888FF91FFF31EEDFB43130ED68B" box="[463,577,1193,1219]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
|
||
:,
|
||
<emphasis id="59CDDA38FF91FFF31D54FB431213D68B" bold="true" box="[630,860,1193,1219]" pageId="8" pageNumber="9">
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<collectingCountry id="13AE46BAFF91FFF31D54FB431213D68B" box="[630,860,1193,1219]" name="United States of America" pageId="8" pageNumber="9">UNITED STATES</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="59CDDA38FF91FFF31C56FB431567D68B" bold="true" box="[884,1064,1193,1219]" pageId="8" pageNumber="9">Massachusetts</emphasis>
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||
: Middlesex Co.:
|
||
<quantity id="AC41ABCFFF91FFF31A24FB431065D6A3" metricMagnitude="3" metricUnit="m" metricValue="7.242048" pageId="8" pageNumber="9" unit="mi" value="4.5">4.5 mi</quantity>
|
||
. W Ashby, from nest of
|
||
<taxonomicName id="ACB97DA9FF91FFF31D47FB381264D6A3" box="[613,811,1234,1259]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="integra">
|
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<emphasis id="59CDDA38FF91FFF31D47FB381264D6A3" box="[613,811,1234,1259]" italics="true" pageId="8" pageNumber="9">Formica integra</emphasis>
|
||
</taxonomicName>
|
||
(mounted with a specimen of
|
||
<taxonomicName id="ACB97DA9FF91FFF31B89FB38146FD6A3" box="[1195,1312,1234,1259]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="integra">
|
||
<emphasis id="59CDDA38FF91FFF31B89FB38146FD6A3" box="[1195,1312,1234,1259]" italics="true" pageId="8" pageNumber="9">F. integra</emphasis>
|
||
</taxonomicName>
|
||
) (A.Newton & M.Thayer),
|
||
<date id="1F0720EAFF91FFF31D1CFB1313F6D75B" box="[574,697,1273,1299]" pageId="8" pageNumber="9" value="1976-04-18">18.iv.1976</date>
|
||
(
|
||
<collectionCode id="0DA89EEFFF91FFF31DEAFB13126BD75B" LSID="urn:lsid:biocol.org:col:34795" box="[712,804,1273,1299]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="9">FMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF91FFF21E1AFACB12D7D3B6" blockId="8.[264,1324,1193,1899]" lastBlockId="9.[264,1325,284,510]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
<typeStatus id="B402B888FF91FFF31E1AFACB10E0D773" box="[312,431,1313,1339]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<emphasis id="59CDDA38FF91FFF31EE9FACB13F9D773" bold="true" box="[459,694,1313,1339]" pageId="8" pageNumber="9">
|
||
<collectingCountry id="13AE46BAFF91FFF31EE9FACB13F9D773" box="[459,694,1313,1339]" name="United States of America" pageId="8" pageNumber="9">UNITED STATES</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="59CDDA38FF91FFF31DF5FACB12C3D773" bold="true" box="[727,908,1313,1339]" pageId="8" pageNumber="9">Massachusetts</emphasis>
|
||
:, (no additional locality data) (
|
||
<collectionCode id="0DA89EEFFF91FFF31E32FAA31027D72B" box="[272,360,1353,1379]" pageId="8" pageNumber="9">KSEM</collectionCode>
|
||
); Middlesex Co.: 4, 4, same data as the
|
||
<typeStatus id="B402B888FF91FFF31C8EFAA3155CD72B" box="[940,1043,1353,1379]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
(
|
||
<specimenCount id="7DBFCDA3FF91FFF31B04FAA315F5D72B" box="[1062,1210,1353,1379]" pageId="8" pageNumber="9" type="generic">6 specimens</specimenCount>
|
||
mounted with specimens of
|
||
<taxonomicName id="ACB97DA9FF91FFF31ECBFA981314D7C3" box="[489,603,1394,1419]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="integra">
|
||
<emphasis id="59CDDA38FF91FFF31ECBFA981314D7C3" box="[489,603,1394,1419]" italics="true" pageId="8" pageNumber="9">F. integra</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<collectionCode id="0DA89EEFFF91FFF31D54FA9B1380D7C3" LSID="urn:lsid:biocol.org:col:34795" box="[630,719,1393,1419]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="9">FMNH</collectionCode>
|
||
,
|
||
<collectionCode id="0DA89EEFFF91FFF31DFEFA9B1268D7C3" box="[732,807,1393,1419]" pageId="8" pageNumber="9">SPSU</collectionCode>
|
||
); 3, ditto but
|
||
<date id="1F0720EAFF91FFF31CD2FA9B153AD7C3" box="[1008,1141,1393,1419]" pageId="8" pageNumber="9" value="1976-07-24">24.vii.1976</date>
|
||
(mounted with specimens of
|
||
<taxonomicName id="ACB97DA9FF91FFF31E8EFA701350D7FB" box="[428,543,1434,1459]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="integra">
|
||
<emphasis id="59CDDA38FF91FFF31E8EFA701350D7FB" box="[428,543,1434,1459]" italics="true" pageId="8" pageNumber="9">F. integra</emphasis>
|
||
</taxonomicName>
|
||
); (mounted with a
|
||
<taxonomicName id="ACB97DA9FF91FFF31C03FA7012C9D7FB" box="[801,902,1434,1459]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF91FFF31C03FA7012C9D7FB" box="[801,902,1434,1459]" italics="true" pageId="8" pageNumber="9">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimen),
|
||
<quantity id="AC41ABCFFF91FFF31B35FA731529D7FB" box="[1047,1126,1433,1459]" metricMagnitude="3" metricUnit="m" metricValue="7.242048" pageId="8" pageNumber="9" unit="mi" value="4.5">4.5 mi</quantity>
|
||
. W Ashby, from ant nest under rocks (A.F.Newton),
|
||
<date id="1F0720EAFF91FFF31D8FFA2B125AD793" box="[685,789,1473,1499]" pageId="8" pageNumber="9" value="1971-05-06">6.v.1971</date>
|
||
;, Westford, from
|
||
<taxonomicName id="ACB97DA9FF91FFF31CDCFA28152CD793" box="[1022,1123,1474,1499]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF91FFF31CDCFA28152CD793" box="[1022,1123,1474,1499]" italics="true" pageId="8" pageNumber="9">Formica</emphasis>
|
||
</taxonomicName>
|
||
nest under rocks (mounted with a
|
||
<taxonomicName id="ACB97DA9FF91FFF31EF1FA001377D44B" box="[467,568,1514,1539]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF91FFF31EF1FA001377D44B" box="[467,568,1514,1539]" italics="true" pageId="8" pageNumber="9">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimen) (A.Newton),
|
||
<date id="1F0720EAFF91FFF31C7DFA03129AD44B" box="[863,981,1513,1539]" pageId="8" pageNumber="9" value="1973-05-27">27.v.1973</date>
|
||
;, Dracut (all
|
||
<collectionCode id="0DA89EEFFF91FFF31B83FA0315B4D44B" LSID="urn:lsid:biocol.org:col:34795" box="[1185,1275,1513,1539]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="9">FMNH</collectionCode>
|
||
);, Framingham (C.A.Frost);, ditto but
|
||
<date id="1F0720EAFF91FFF31DF5F9FB1273D463" box="[727,828,1553,1579]" pageId="8" pageNumber="9" value="1912-05-05">5.v.1912</date>
|
||
(mounted with a
|
||
<taxonomicName id="ACB97DA9FF91FFF31B34F9F81534D463" box="[1046,1147,1554,1579]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF91FFF31B34F9F81534D463" box="[1046,1147,1554,1579]" italics="true" pageId="8" pageNumber="9">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimen);, Sherborn, in nest of
|
||
<taxonomicName id="ACB97DA9FF91FFF31D2CF9D01207D41B" authority="C.A.Frost" authorityName="C.A.Frost" box="[526,840,1593,1619]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rufa">
|
||
<emphasis id="59CDDA38FF91FFF31D2CF9D013FED41B" box="[526,689,1594,1619]" italics="true" pageId="8" pageNumber="9">Formica rufa</emphasis>
|
||
(C.A.Frost)
|
||
</taxonomicName>
|
||
,
|
||
<date id="1F0720EAFF91FFF31C78F9D31296D41B" box="[858,985,1593,1619]" pageId="8" pageNumber="9" value="1916-04-30">30.iv.1916</date>
|
||
;, Sherborn, under stones (mounted with a
|
||
<taxonomicName id="ACB97DA9FF91FFF31EFFF988130DD433" box="[477,578,1634,1659]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF91FFF31EFFF988130DD433" box="[477,578,1634,1659]" italics="true" pageId="8" pageNumber="9">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimen) (C.A.Frost),
|
||
<date id="1F0720EAFF91FFF31C4FF98B129DD433" box="[877,978,1633,1659]" pageId="8" pageNumber="9" value="1915-05-04">4.v.1915</date>
|
||
(all
|
||
<collectionCode id="0DA89EEFFF91FFF31B04F98B1537D433" box="[1062,1144,1633,1659]" pageId="8" pageNumber="9">CASC</collectionCode>
|
||
);
|
||
<collectingCountry id="13AE46BAFF91FFF31BB3F98B15A2D433" box="[1169,1261,1633,1659]" name="Norfolk Island" pageId="8" pageNumber="9">Norfolk</collectingCountry>
|
||
Co.: 2 (mounted with a
|
||
<taxonomicName id="ACB97DA9FF91FFF31D35F9601333D4EB" box="[535,636,1674,1699]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF91FFF31D35F9601333D4EB" box="[535,636,1674,1699]" italics="true" pageId="8" pageNumber="9">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimen), Blue Hills (W.M.
|
||
<collectingCountry id="13AE46BAFF91FFF31CFFF9631568D4EB" box="[989,1063,1673,1699]" name="Isle of Man" pageId="8" pageNumber="9">Mann</collectingCountry>
|
||
),
|
||
<date id="1F0720EAFF91FFF31B1CF96315FDD4EB" box="[1086,1202,1673,1699]" pageId="8" pageNumber="9" value="1912-05-12">12.v.1912</date>
|
||
(
|
||
<collectionCode id="0DA89EEFFF91FFF31BE6F963146FD4EB" LSID="urn:lsid:biocol.org:col:34795" box="[1220,1312,1673,1699]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="9">FMNH</collectionCode>
|
||
);
|
||
<emphasis id="59CDDA38FF91FFF31E2AF95B109BD483" bold="true" box="[264,468,1713,1739]" pageId="8" pageNumber="9">New Hampshire</emphasis>
|
||
: Grafton Co.:, Rumney (mounted with a
|
||
<taxonomicName id="ACB97DA9FF91FFF31CD8F9581510D483" box="[1018,1119,1714,1739]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF91FFF31CD8F9581510D483" box="[1018,1119,1714,1739]" italics="true" pageId="8" pageNumber="9">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimen) (Darlington),
|
||
<date id="1F0720EAFF91FFF31E57F93310BFD4BB" box="[373,496,1753,1779]" pageId="8" pageNumber="9" value="1925-04-24">24.iv.1925</date>
|
||
(
|
||
<collectionCode id="0DA89EEFFF91FFF31D23F9331301D4BB" box="[513,590,1753,1779]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="8" pageNumber="9">CNCI</collectionCode>
|
||
);
|
||
<emphasis id="59CDDA38FF91FFF31D40F93313AFD4BB" bold="true" box="[610,736,1753,1779]" pageId="8" pageNumber="9">New York</emphasis>
|
||
: Tompkins Co.:, Ithaca,
|
||
<date id="1F0720EAFF91FFF31B11F93315FDD4BB" box="[1075,1202,1753,1779]" pageId="8" pageNumber="9" value="1884-08-01">1.viii.1884</date>
|
||
(
|
||
<collectionCode id="0DA89EEFFF91FFF31BE1F9331451D4BB" LSID="urn:lsid:biocol.org:col:34795" box="[1219,1310,1753,1779]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="9">FMNH</collectionCode>
|
||
);
|
||
<emphasis id="59CDDA38FF91FFF31E2AF8EB10D6D553" bold="true" box="[264,409,1793,1819]" pageId="8" pageNumber="9">
|
||
New
|
||
<collectingCountry id="13AE46BAFF91FFF31E6BF8EB10D6D553" box="[329,409,1793,1819]" name="Jersey" pageId="8" pageNumber="9">Jersey</collectingCountry>
|
||
</emphasis>
|
||
: Mercer Co.:, Princeton (mounted with a
|
||
<taxonomicName id="ACB97DA9FF91FFF31CE3F8E81569D553" box="[961,1062,1794,1819]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF91FFF31CE3F8E81569D553" box="[961,1062,1794,1819]" italics="true" pageId="8" pageNumber="9">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimen),
|
||
<date id="1F0720EAFF91FFF31B9BF8EB1464D553" box="[1209,1323,1793,1819]" pageId="8" pageNumber="9" value="1941-05-11">11.v.1941</date>
|
||
(
|
||
<collectionCode id="0DA89EEFFF91FFF31E32F8C31024D50B" LSID="urn:lsid:biocol.org:col:34795" box="[272,363,1833,1859]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="9">FMNH</collectionCode>
|
||
);
|
||
<emphasis id="59CDDA38FF91FFF31E5DF8C31330D50B" bold="true" box="[383,639,1833,1859]" pageId="8" pageNumber="9">
|
||
District of
|
||
<collectingCountry id="13AE46BAFF91FFF31D21F8C31330D50B" box="[515,639,1833,1859]" name="United States of America" pageId="8" pageNumber="9">Columbia</collectingCountry>
|
||
</emphasis>
|
||
: 2 (no additional locality data) (
|
||
<collectionCode id="0DA89EEFFF91FFF31B01F8C3153BD50B" box="[1059,1140,1833,1859]" pageId="8" pageNumber="9">CASC</collectionCode>
|
||
,
|
||
<collectionCode id="0DA89EEFFF91FFF31B5DF8C31599D50B" box="[1151,1238,1833,1859]" pageId="8" pageNumber="9">KSEM</collectionCode>
|
||
); (no additional locality data) (Wickham);, Washington (
|
||
<collectionCode id="0DA89EEFFF91FFF31C83F8BB12BBD523" box="[929,1012,1873,1899]" pageId="8" pageNumber="9">CASC</collectionCode>
|
||
); 4, 3 (3, 2 mounted with
|
||
<taxonomicName id="ACB97DA9FF90FFF21E96FEF61356D37D" box="[436,537,284,309]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF90FFF21E96FEF61356D37D" box="[436,537,284,309]" italics="true" pageId="9" pageNumber="10">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimens), Washington, iv (
|
||
<collectionCode id="0DA89EEFFF90FFF21C5BFEF6129CD37E" LSID="urn:lsid:biocol.org:col:34795" box="[889,979,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="10">FMNH</collectionCode>
|
||
;
|
||
<collectionCode id="0DA89EEFFF90FFF21CC2FEF6157AD37E" box="[992,1077,284,310]" pageId="9" pageNumber="10">KSEM</collectionCode>
|
||
;
|
||
<collectionCode id="0DA89EEFFF90FFF21B60FEF615D3D37E" box="[1090,1180,284,310]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10">NMNH</collectionCode>
|
||
(Casey collection));
|
||
<emphasis id="59CDDA38FF90FFF21E5EFEA9108DD315" bold="true" box="[380,450,323,349]" pageId="9" pageNumber="10">Texas</emphasis>
|
||
: 2, 3, (no locality data) (one female mounted with a
|
||
<taxonomicName id="ACB97DA9FF90FFF21BAFFEAE15BDD315" box="[1165,1266,324,349]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF90FFF21BAFFEAE15BDD315" box="[1165,1266,324,349]" italics="true" pageId="9" pageNumber="10">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimen) (
|
||
<collectionCode id="0DA89EEFFF90FFF21E4BFE86108DD3CE" LSID="urn:lsid:biocol.org:col:34795" box="[361,450,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="10">FMNH</collectionCode>
|
||
,
|
||
<collectionCode id="0DA89EEFFF90FFF21EEDFE861355D3CE" box="[463,538,364,390]" pageId="9" pageNumber="10">SPSU</collectionCode>
|
||
);
|
||
<emphasis id="59CDDA38FF90FFF21D0DFE8113E1D3CD" bold="true" box="[559,686,363,389]" pageId="9" pageNumber="10">
|
||
<collectingCountry id="13AE46BAFF90FFF21D0DFE8113E1D3CD" box="[559,686,363,389]" name="Canada" pageId="9" pageNumber="10">CANADA</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="59CDDA38FF90FFF21DE4FE81120FD3CD" bold="true" box="[710,832,363,389]" pageId="9" pageNumber="10">Manitoba</emphasis>
|
||
: 2,, Aweme (N.Criddle),
|
||
<date id="1F0720EAFF90FFF21B95FE861465D3CE" box="[1207,1322,364,390]" pageId="9" pageNumber="10" value="1919-05-17">17.v.1919</date>
|
||
(
|
||
<collectionCode id="0DA89EEFFF90FFF21E32FE7E102ED3E6" box="[272,353,404,430]" pageId="9" pageNumber="10">CASC</collectionCode>
|
||
,
|
||
<collectionCode id="0DA89EEFFF90FFF21E4FFE7E10F7D3E6" box="[365,440,404,430]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="9" pageNumber="10">CNCI</collectionCode>
|
||
);, Aweme, from
|
||
<taxonomicName id="ACB97DA9FF90FFF21DB7FE7E13B5D3E5" box="[661,762,404,429]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF90FFF21DB7FE7E13B5D3E5" box="[661,762,404,429]" italics="true" pageId="9" pageNumber="10">Formica</emphasis>
|
||
</taxonomicName>
|
||
nest (N.Criddle),
|
||
<date id="1F0720EAFF90FFF21CECFE7E157CD3E6" box="[974,1075,404,430]" pageId="9" pageNumber="10" value="1916-05-02">2.v.1916</date>
|
||
(
|
||
<collectionCode id="0DA89EEFFF90FFF21B60FE7E15DAD3E6" box="[1090,1173,404,430]" pageId="9" pageNumber="10">CASC</collectionCode>
|
||
);, Aweme, from ant nest (N.Criddle),
|
||
<date id="1F0720EAFF90FFF21D65FE5613E3D39E" box="[583,684,444,470]" pageId="9" pageNumber="10" value="1916-05-03">3.v.1916</date>
|
||
(
|
||
<collectionCode id="0DA89EEFFF90FFF21D9FFE561246D39E" box="[701,777,444,470]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="9" pageNumber="10">CNCI</collectionCode>
|
||
);, Aweme (N.Criddle) (
|
||
<collectionCode id="0DA89EEFFF90FFF21B1CFE5615D5D39E" box="[1086,1178,444,470]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10">NMNH</collectionCode>
|
||
(Casey collection));, Stonewall (J.B.
|
||
<collectingCountry id="13AE46BAFF90FFF21D68FE0E13D0D3B6" box="[586,671,484,510]" name="United Kingdom" pageId="9" pageNumber="10">Wallis</collectingCountry>
|
||
),
|
||
<date id="1F0720EAFF90FFF21D96FE0E1260D3B6" box="[692,815,484,510]" pageId="9" pageNumber="10" value="1970-04-23">23.iv.1970</date>
|
||
(
|
||
<collectionCode id="0DA89EEFFF90FFF21C1DFE0E12C4D3B6" box="[831,907,484,510]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="9" pageNumber="10">CNCI</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="3FC656A2FF90FFF21E2AF9E413B2D4C6" httpUri="https://zenodo.org/record/156412/files/figure.png" pageId="9" pageNumber="10" targetBox="[290,1295,579,1513]" targetPageId="9">
|
||
<paragraph id="6B06062AFF90FFF21E2AF9E413B2D4C6" blockId="9.[264,1323,1550,1678]" pageId="9" pageNumber="10">
|
||
<emphasis id="59CDDA38FF90FFF21E2AF9E41085D46E" bold="true" box="[264,458,1550,1574]" pageId="9" pageNumber="10">FIGURES 2024.</emphasis>
|
||
Abdominal segment 8 of
|
||
<taxonomicName id="ACB97DA9FF90FFF21DC7F9E512A5D46E" authority="Gusarov" authorityName="Gusarov" box="[741,1002,1551,1574]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caseyi" status="sp. nov.">
|
||
<emphasis id="59CDDA38FF90FFF21DC7F9E512C6D46E" box="[741,905,1551,1574]" italics="true" pageId="9" pageNumber="10">Goniusa caseyi</emphasis>
|
||
Gusarov
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="59CDDA38FF90FFF21CD7F9E4157ED46E" bold="true" box="[1013,1073,1550,1574]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="42FE6743FF90FFF21CD7F9E4157ED46E" box="[1013,1073,1550,1574]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(paratypes from Washington, D.C. (2021); Aweme, Manitoba (22); and Stonewall, Manitoba (2324)). 20 – male tergum 8; 21 – male sternum 8; 22 – apex of male tergum 8; 23 – female tergum 8; 24 – female sternum 8. Scale bar 0.4 mm (2021, 2324), 0.2 mm (22).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="23A355A1FF90FFF21E1AF95113B1D506" pageId="9" pageNumber="10" type="diagnosis">
|
||
<paragraph id="6B06062AFF90FFF21E1AF95113B1D506" blockId="9.[264,1324,1723,1870]" pageId="9" pageNumber="10">
|
||
<emphasis id="59CDDA38FF90FFF21E1AF95110F7D49D" bold="true" box="[312,440,1723,1749]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
|
||
<taxonomicName id="ACB97DA9FF90FFF21EEAF9561332D49D" box="[456,637,1724,1749]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caseyi">
|
||
<emphasis id="59CDDA38FF90FFF21EEAF9561332D49D" box="[456,637,1724,1749]" italics="true" pageId="9" pageNumber="10">Goniusa caseyi</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="ACB97DA9FF90FFF21CE2F956157CD49D" box="[960,1075,1724,1749]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="alperti">
|
||
<emphasis id="59CDDA38FF90FFF21CE2F956157CD49D" box="[960,1075,1724,1749]" italics="true" pageId="9" pageNumber="10">G. alperti</emphasis>
|
||
</taxonomicName>
|
||
by having pronotum with subparallel lateral sides, reaching maximum width in front of the middle (
|
||
<figureCitation id="F3821AAFFF90FFF21BE2F90E1066D56E" captionStart="FIGURES 37 41" captionStartId="12.[264,382,1692,1716]" captionTargetBox="[301,1285,783,1653]" captionTargetId="figure@12.[288,1299,769,1665]" captionTargetPageId="12" captionText="FIGURES 37 41. Pronotum of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (37); and Stonewall, Manitoba (38 )) and G. alperti Kistner (holotype (39); female from San Bernardino Mts., California (40), paratype from 10 mi. E of Coville, Washington (41 )). 37, 39 – male pronotum; 38, 40 41 – female pronotum. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/156415/files/figure.png" pageId="9" pageNumber="10">Figs. 37 38</figureCitation>
|
||
); antennal segments 510 slightly elongate; larger body and longer apical process of the median lobe of aedeagus (
|
||
<figureCitation id="F3821AAFFF90FFF21D18F8DE13BED506" box="[570,753,1844,1870]" captionStart="FIGURES 25 29" captionStartId="10.[264,383,1768,1792]" captionTargetBox="[280,1310,701,1744]" captionTargetId="figure@10.[263,1323,689,1755]" captionTargetPageId="10" captionText="FIGURES 25 29. Median lobe of aedeagus of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (25 26, 28 29); and Sherborn, Massachusetts (27 )). 25 – median lobe, parameral view; 26 27 – apex of median lobe, parameral view; 28 – median lobe, lateral view; 29 – apex of median lobe, lateral view. Scale bar 0.4 mm (25, 28), 0.2 mm (26 27, 29)." httpUri="https://zenodo.org/record/156413/files/figure.png" pageId="9" pageNumber="10">Figs. 2627, 29</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23A355A1FF93FFF61E1AFEF113EED0D6" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="6B06062AFF93FFF11E1AFEF11399D3E6" blockId="10.[264,1324,283,630]" pageId="10" pageNumber="11">
|
||
<emphasis id="59CDDA38FF93FFF11E1AFEF1109FD37D" bold="true" box="[312,464,283,309]" pageId="10" pageNumber="11">Description.</emphasis>
|
||
Length
|
||
<quantity id="AC41ABCFFF93FFF11D63FEF6139ED37E" box="[577,721,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="4.2" metricValueMin="3.2" pageId="10" pageNumber="11" unit="mm" value="3.7" valueMax="4.2" valueMin="3.2">3.24.2 mm</quantity>
|
||
. Head darker than the rest of the body, brownish black to brown; pronotum, elytra and abdomen light brown to reddish brown; elytra and abdominal segments 35 often lighter than pronotum and abdominal segments 57; legs, antennae and mouthparts brownish red.
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF93FFF11E1AFE56130FD03E" blockId="10.[264,1324,283,630]" pageId="10" pageNumber="11">
|
||
In females head surface glossy, with weak isodiametric microsculpture, with strong punctation, distance between punctures equals 1/21 times their diameter. In males head surface matte, with strong isodiametric microsculpture, with weak, sparse and poorly visible punctation. Temple length to eye length ratio 0.61.0. Antennal articles 410 slightly elongate (
|
||
<figureCitation id="F3821AAFFF93FFF11EF9FDB6137CD03E" box="[475,563,604,630]" captionStart="FIGURES 12 15" captionStartId="5.[264,383,1787,1811]" captionTargetBox="[289,1304,581,1774]" captionTargetId="figure@5.[272,1314,569,1776]" captionTargetPageId="5" captionText="FIGURES 12 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 14)." httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="10" pageNumber="11">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="3FC656A2FF93FFF11E2AF902156DD52F" httpUri="https://zenodo.org/record/156413/files/figure.png" pageId="10" pageNumber="11" targetBox="[280,1310,701,1744]" targetPageId="10">
|
||
<paragraph id="6B06062AFF93FFF11E2AF902156DD52F" blockId="10.[264,1323,1768,1895]" pageId="10" pageNumber="11">
|
||
<emphasis id="59CDDA38FF93FFF11E2AF9021082D548" bold="true" box="[264,461,1768,1792]" pageId="10" pageNumber="11">FIGURES 2529.</emphasis>
|
||
Median lobe of aedeagus of
|
||
<taxonomicName id="ACB97DA9FF93FFF11C3FF9031567D4B7" authority="Gusarov" authorityName="Gusarov" box="[797,1064,1768,1792]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="caseyi" status="sp. nov.">
|
||
<emphasis id="59CDDA38FF93FFF11C3FF903128BD548" box="[797,964,1769,1792]" italics="true" pageId="10" pageNumber="11">Goniusa caseyi</emphasis>
|
||
Gusarov
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="59CDDA38FF93FFF11B14F902153BD548" bold="true" box="[1078,1140,1768,1792]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="42FE6743FF93FFF11B14F902153BD548" box="[1078,1140,1768,1792]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(paratypes from Washington, D.C. (2526, 2829); and Sherborn, Massachusetts (27)). 25 – median lobe, parameral view; 2627 – apex of median lobe, parameral view; 28 – median lobe, lateral view; 29 – apex of median lobe, lateral view. Scale bar 0.4 mm (25, 28), 0.2 mm (2627, 29).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3FC656A2FF92FFF01E2AF98A1592D54A" httpUri="https://zenodo.org/record/156414/files/figure.png" pageId="11" pageNumber="12" targetBox="[288,1303,337,1596]" targetPageId="11">
|
||
<paragraph id="6B06062AFF92FFF01E2AF98A1592D54A" blockId="11.[264,1323,1632,1794]" pageId="11" pageNumber="12">
|
||
<emphasis id="59CDDA38FF92FFF01E2AF98A1084D430" bold="true" box="[264,459,1632,1656]" pageId="11" pageNumber="12">FIGURES 3036.</emphasis>
|
||
Genitalia of
|
||
<taxonomicName id="ACB97DA9FF92FFF01D7DF98B1229D43F" authority="Gusarov" authorityName="Gusarov" box="[607,870,1632,1656]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="caseyi" status="sp. nov.">
|
||
<emphasis id="59CDDA38FF92FFF01D7DF98B124BD430" box="[607,772,1633,1656]" italics="true" pageId="11" pageNumber="12">Goniusa caseyi</emphasis>
|
||
Gusarov
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="59CDDA38FF92FFF01C53F98A12E1D430" bold="true" box="[881,942,1632,1656]" pageId="11" pageNumber="12">
|
||
<taxonomicNameLabel id="42FE6743FF92FFF01C53F98A12E1D430" box="[881,942,1632,1656]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(paratypes from Aweme, Manitoba (3031); Texas (3234); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36)). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 3536 – spermatheca. Scale bar 0.1 mm (3234), 0.2 mm (3031, 3536).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6B06062AFF95FFF71E1AFEF6109FD0D6" blockId="12.[264,1324,284,670]" pageId="12" pageNumber="13">
|
||
Pronotum strongly transverse, with subparallel lateral sides, reaching maximum width in front of the middle (
|
||
<figureCitation id="F3821AAFFF95FFF71D33FEAE13D5D316" box="[529,666,324,350]" captionStart="FIGURES 37 41" captionStartId="12.[264,382,1692,1716]" captionTargetBox="[301,1285,783,1653]" captionTargetId="figure@12.[288,1299,769,1665]" captionTargetPageId="12" captionText="FIGURES 37 41. Pronotum of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (37); and Stonewall, Manitoba (38 )) and G. alperti Kistner (holotype (39); female from San Bernardino Mts., California (40), paratype from 10 mi. E of Coville, Washington (41 )). 37, 39 – male pronotum; 38, 40 41 – female pronotum. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/156415/files/figure.png" pageId="12" pageNumber="13">Figs. 3738</figureCitation>
|
||
), 1.6 times as wide as head, width
|
||
<quantity id="AC41ABCFFF95FFF71B0DFEAE1598D316" box="[1071,1239,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="1.005" metricValueMax="1.07" metricValueMin="0.94" pageId="12" pageNumber="13" unit="mm" value="1.005" valueMax="1.07" valueMin="0.94">0.941.07 mm</quantity>
|
||
, length
|
||
<quantity id="AC41ABCFFF95FFF71E2AFE8610FED3CE" box="[264,433,364,390]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="12" pageNumber="13" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.600.70 mm</quantity>
|
||
, width to length ratio 1.5; in females surface glossy, with weak isodiametric microsculpture; punctation finer than on head, distance between punctures equal to their diameter; in males surface matte, with strong isodiametric microsculpture; and fine and poorly visible punctation. Elytra slightly wider (
|
||
<quantity id="AC41ABCFFF95FFF71C46FE0E155ED3B6" box="[868,1041,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="1.1099999999999999" metricValueMax="1.21" metricValueMin="1.01" pageId="12" pageNumber="13" unit="mm" value="1.1099999999999999" valueMax="1.21" valueMin="1.01">1.011.21 mm</quantity>
|
||
) and longer (
|
||
<quantity id="AC41ABCFFF95FFF71B9BFE0E1076D06E" metricMagnitude="-4" metricUnit="m" metricValue="7.4" metricValueMax="7.9" metricValueMin="6.8999999999999995" pageId="12" pageNumber="13" unit="mm" value="0.74" valueMax="0.79" valueMin="0.69">0.690.79 mm</quantity>
|
||
; measured from humeral angle) than pronotum (pronotal length to elytral length ratio 0.9), 1.5 times wider than long, surface glossy, with fine and weak isodiametric microsculpture, and with fine and slightly asperate punctation, distance between punctures equal to their diameter.
|
||
</paragraph>
|
||
<caption id="3FC656A2FF95FFF71E2AF97612C4D553" httpUri="https://zenodo.org/record/156415/files/figure.png" pageId="12" pageNumber="13" targetBox="[301,1285,783,1653]" targetPageId="12">
|
||
<paragraph id="6B06062AFF95FFF71E2AF97612C4D553" blockId="12.[264,1323,1692,1819]" pageId="12" pageNumber="13">
|
||
<emphasis id="59CDDA38FF95FFF71E2AF9761085D4FC" bold="true" box="[264,458,1692,1716]" pageId="12" pageNumber="13">FIGURES 3741.</emphasis>
|
||
Pronotum of
|
||
<taxonomicName id="ACB97DA9FF95FFF71D46F9771225D4FB" authority="Gusarov" authorityName="Gusarov" box="[612,874,1692,1716]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="caseyi" status="sp. nov.">
|
||
<emphasis id="59CDDA38FF95FFF71D46F9771247D4FC" box="[612,776,1693,1716]" italics="true" pageId="12" pageNumber="13">Goniusa caseyi</emphasis>
|
||
Gusarov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="42FE6743FF95FFF71C56F97612FFD4FC" box="[884,944,1692,1716]" pageId="12" pageNumber="13" rank="species">
|
||
<emphasis id="59CDDA38FF95FFF71C56F97612FFD4FC" bold="true" box="[884,944,1692,1716]" pageId="12" pageNumber="13">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(paratypes from Aweme, Manitoba (37); and Stonewall, Manitoba (38)) and
|
||
<taxonomicName id="ACB97DA9FF95FFF71D98F9551239D49E" authority="Kistner" authorityName="Kistner" box="[698,886,1727,1750]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="alperti">
|
||
<emphasis id="59CDDA38FF95FFF71D98F955126DD49E" box="[698,802,1727,1750]" italics="true" pageId="12" pageNumber="13">G. alperti</emphasis>
|
||
Kistner
|
||
</taxonomicName>
|
||
(holotype (39); female from San Bernardino Mts., California (40), paratype from 10 mi. E of Coville, Washington (41)). 37, 39 – male pronotum; 38, 4041 – female pronotum. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6B06062AFF94FFF61E1AFEF615B6D3CE" blockId="13.[264,1324,284,830]" pageId="13" pageNumber="14">Abdominal terga glossy, with fine and often poorly visible transverse microsculpture, with fine punctation, distance between punctures equals 12 times their diameter on terga 35 and 36 times on terga 67. Apical margin of tergum 7 with white palisade fringe.</paragraph>
|
||
<paragraph id="6B06062AFF94FFF61E1AFE7E10E2D006" blockId="13.[264,1324,284,830]" pageId="13" pageNumber="14">
|
||
In males head with broad medial impression; pronotum with broad medial impression that is half as wide as pronotum, deeper in posterior half, posterolateral portions of the impression with less dense microsculpture, without punctation and pubescence. Sternum 8 with wide apical emargination (
|
||
<figureCitation id="F3821AAFFF94FFF61DBFFDE613B8D06E" box="[669,759,524,550]" captionStart="FIGURES 20 24" captionStartId="9.[264,383,1550,1574]" captionTargetBox="[290,1295,579,1513]" captionTargetId="figure@9.[282,1306,569,1524]" captionTargetPageId="9" captionText="FIGURES 20 24. Abdominal segment 8 of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (20 21); Aweme, Manitoba (22); and Stonewall, Manitoba (23 24 )). 20 – male tergum 8; 21 – male sternum 8; 22 – apex of male tergum 8; 23 – female tergum 8; 24 – female sternum 8. Scale bar 0.4 mm (20 21, 23 24), 0.2 mm (22)." httpUri="https://zenodo.org/record/156412/files/figure.png" pageId="13" pageNumber="14">Fig. 21</figureCitation>
|
||
). Posterior margin of male tergum 8 crenulate (
|
||
<figureCitation id="F3821AAFFF94FFF61E33FDDE10EFD006" box="[273,416,564,590]" captionStart="FIGURES 20 24" captionStartId="9.[264,383,1550,1574]" captionTargetBox="[290,1295,579,1513]" captionTargetId="figure@9.[282,1306,569,1524]" captionTargetPageId="9" captionText="FIGURES 20 24. Abdominal segment 8 of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (20 21); Aweme, Manitoba (22); and Stonewall, Manitoba (23 24 )). 20 – male tergum 8; 21 – male sternum 8; 22 – apex of male tergum 8; 23 – female tergum 8; 24 – female sternum 8. Scale bar 0.4 mm (20 21, 23 24), 0.2 mm (22)." httpUri="https://zenodo.org/record/156412/files/figure.png" pageId="13" pageNumber="14">Figs. 20, 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF94FFF61E1AFDB61332D03E" blockId="13.[264,1324,284,830]" box="[312,637,604,630]" pageId="13" pageNumber="14">
|
||
Aedeagus as in
|
||
<figureCitation id="F3821AAFFF94FFF61ED2FDB61336D03E" box="[496,633,604,630]" captionStart-0="FIGURES 25 29" captionStart-1="FIGURES 30 36" captionStartId-0="10.[264,383,1768,1792]" captionStartId-1="11.[264,382,1632,1656]" captionTargetBox-0="[280,1310,701,1744]" captionTargetBox-1="[288,1303,337,1596]" captionTargetId-0="figure@10.[263,1323,689,1755]" captionTargetId-1="figure@11.[273,1314,326,1608]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 25 29. Median lobe of aedeagus of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (25 26, 28 29); and Sherborn, Massachusetts (27 )). 25 – median lobe, parameral view; 26 27 – apex of median lobe, parameral view; 28 – median lobe, lateral view; 29 – apex of median lobe, lateral view. Scale bar 0.4 mm (25, 28), 0.2 mm (26 27, 29)." captionText-1="FIGURES 30 36. Genitalia of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (30 31); Texas (32 34); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36 )). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 35 36 – spermatheca. Scale bar 0.1 mm (32 34), 0.2 mm (30 31, 35 36)." httpUri-0="https://zenodo.org/record/156413/files/figure.png" httpUri-1="https://zenodo.org/record/156414/files/figure.png" pageId="13" pageNumber="14">Figs. 2534</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF94FFF61E1AFD6E13EED0D6" blockId="13.[264,1324,284,830]" box="[312,673,644,670]" pageId="13" pageNumber="14">
|
||
Spermatheca as in
|
||
<figureCitation id="F3821AAFFF94FFF61D36FD6E13D2D0D6" box="[532,669,644,670]" captionStart="FIGURES 30 36" captionStartId="11.[264,382,1632,1656]" captionTargetBox="[288,1303,337,1596]" captionTargetId="figure@11.[273,1314,326,1608]" captionTargetPageId="11" captionText="FIGURES 30 36. Genitalia of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (30 31); Texas (32 34); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36 )). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 35 36 – spermatheca. Scale bar 0.1 mm (32 34), 0.2 mm (30 31, 35 36)." httpUri="https://zenodo.org/record/156414/files/figure.png" pageId="13" pageNumber="14">Figs. 3536</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23A355A1FF94FFF61E1AFD4112B3D176" pageId="13" pageNumber="14" type="distribution">
|
||
<paragraph id="6B06062AFF94FFF61E1AFD4112CFD0A6" blockId="13.[264,1324,284,830]" pageId="13" pageNumber="14">
|
||
<emphasis id="59CDDA38FF94FFF61E1AFD411097D08D" bold="true" box="[312,472,683,709]" pageId="13" pageNumber="14">Distribution.</emphasis>
|
||
Known from Manitoba, New Hampshire, Massachusetts, New York, New
|
||
<collectingCountry id="13AE46BAFF94FFF61E65FD3E10DED0A6" box="[327,401,724,750]" name="Jersey" pageId="13" pageNumber="14">Jersey</collectingCountry>
|
||
, District of
|
||
<collectingCountry id="13AE46BAFF94FFF61D39FD3E13C1D0A6" box="[539,654,724,750]" name="United States of America" pageId="13" pageNumber="14">Columbia</collectingCountry>
|
||
and Texas (
|
||
<figureCitation id="F3821AAFFF94FFF61C39FD3E123CD0A6" box="[795,883,724,750]" captionStart="FIGURE 57" captionStartId="17.[264,368,1343,1367]" captionTargetBox="[295,1292,323,1314]" captionTargetId="figure@17.[293,1294,320,1316]" captionTargetPageId="17" captionText="FIGURE 57. Geographical distribution of Goniusa caseyi Gusarov, sp. n. and G. alperti Kistner. Black circles and triangles indicate the localities of examined specimens. Open circles and triangles indicate the localities of additional specimens listed by Kistner (1976) but not examined by me. The exact locality of the Texas specimens of Goniusa caseyi is unknown." httpUri="https://zenodo.org/record/156419/files/figure.png" pageId="13" pageNumber="14">Fig. 57</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF94FFF61E1AFD1112B3D176" blockId="13.[264,1324,284,830]" pageId="13" pageNumber="14">
|
||
<emphasis id="59CDDA38FF94FFF61E1AFD111349D15D" bold="true" box="[312,518,763,789]" pageId="13" pageNumber="14">Natural History.</emphasis>
|
||
<taxonomicName id="ACB97DA9FF94FFF61D35FD161382D15D" box="[535,717,764,789]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="caseyi">
|
||
<emphasis id="59CDDA38FF94FFF61D35FD161382D15D" box="[535,717,764,789]" italics="true" pageId="13" pageNumber="14">Goniusa caseyi</emphasis>
|
||
</taxonomicName>
|
||
is associated with ants of the genus
|
||
<taxonomicName id="ACB97DA9FF94FFF61BA0FD1615A8D15D" box="[1154,1255,764,789]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF94FFF61BA0FD1615A8D15D" box="[1154,1255,764,789]" italics="true" pageId="13" pageNumber="14">Formica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="59CDDA38FF94FFF61BDBFD16146CD15D" box="[1273,1315,764,789]" italics="true" pageId="13" pageNumber="14">e.g.</emphasis>
|
||
,
|
||
<taxonomicName id="ACB97DA9FF94FFF61E2AFCCE1087D175" box="[264,456,804,829]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="integra">
|
||
<emphasis id="59CDDA38FF94FFF61E2AFCCE1087D175" box="[264,456,804,829]" italics="true" pageId="13" pageNumber="14">Formica integra</emphasis>
|
||
</taxonomicName>
|
||
(identified by Dr. E.O.Wilson:
|
||
<bibRefCitation id="0F287BDBFF94FFF61C61FCCE12ABD176" author="Kistner" box="[835,996,804,830]" pageId="13" pageNumber="19" refString="Kistner, D. H. (1976) Revision and Reclassification of the Genus Goniusa Casey with a Larval Description and Ant Host Records (Coleoptera: Staphylinidae). Sociobiology, 2 (1), 83 - 95." type="journal article" year="1976">Kistner, 1976</bibRefCitation>
|
||
)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |