206 lines
30 KiB
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206 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/dez.65.21000" ID-GBIF-Dataset="03cce405-5887-4474-b040-35de18e8d020" ID-GBIF-Taxon="182223665" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="77D7AEC9294554E5BBDCABA37901022A" ID-Zenodo-Dep="1166397" ID-ZooBank="73DEE0F32BB04A21B4455E168FE50F54" LSID="urn:lsid:zoobank.org:act:572EF2F8-5BF7-4663-8F53-74100E27DF5E" ModsDocID="1860-1324-65-1-1" checkinTime="1621372489409" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="CB0F8B6C344341BF29928F5D4AD9E56B" docLanguage="en" docName="DeutEntomolZeit 65(1): 1-63" docOrigin="Deutsche Entomologische Zeitschrift 65 (1)" docPubDate="2018-01-18" docSource="http://dx.doi.org/10.3897/dez.65.21000" docTitle="Mecyclothorax mouensis Moore & Liebherr 2018, sp. n." docType="treatment" docUuid="572EF2F8-5BF7-4663-8F53-74100E27DF5E" docUuidSource="ZooBank" docVersion="3" id="77D7AEC9294554E5BBDCABA37901022A" lastPageNumber="37" masterDocId="77D7AEC9294554E5BBDCABA37901022A" masterDocTitle="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel" masterLastPageNumber="63" masterPageNumber="1" pageId="36" pageNumber="37" updateTime="1643465225333" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Liebherr, James K.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9831-884X</mods:nameIdentifier>
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<mods:affiliation>Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:nameIdentifier type="email">jkl5@cornell.edu</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-01-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>65</mods:number>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>63</mods:end>
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<mods:url>http://dx.doi.org/10.3897/dez.65.21000</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.65.21000</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-1324-1-1</mods:identifier>
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<mods:identifier type="ZooBank">73DEE0F32BB04A21B4455E168FE50F54</mods:identifier>
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<mods:identifier type="Pensoft-UUID">77D7AEC9294554E5BBDCABA37901022A</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1166397</mods:identifier>
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<treatment ID-GBIF-Taxon="182223665" LSID="urn:lsid:zoobank.org:act:572EF2F8-5BF7-4663-8F53-74100E27DF5E" httpUri="http://treatment.plazi.org/id/CB0F8B6C344341BF29928F5D4AD9E56B" lastPageNumber="37" pageId="36" pageNumber="37">
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<subSubSection pageId="36" pageNumber="37" type="nomenclature">
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<paragraph pageId="36" pageNumber="37">
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<pageBreakToken pageId="36" pageNumber="37" start="start">13</pageBreakToken>
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.
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<taxonomicName LSID="http://zoobank.org/572EF2F8-5BF7-4663-8F53-74100E27DF5E" authority="Moore & Liebherr" authorityName="Moore & Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax mouensis" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="mouensis" status="sp. n.">Mecyclothorax mouensis Moore & Liebherr</taxonomicName>
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<taxonomicNameLabel pageId="36" pageNumber="37">sp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Thoracic structures of Mecyclothorax and Amblytelus spp. A-B. Pronotum, dorsal view: A, M. laterorectus; B, M. mouensis. C-F. Prosternum, ventral view: C, M. fleutiauxi; D, M. curtus; E, M. blackburni; F, M. moorei. G-J. Elytral apices, dorsal view: G, M. fleutiauxi; H, M. laterorectus; I, M. lissus; J, A. curtus. K-L. Elytral lateral margin showing eighth stria, lateral view: K, M. ambiguus; L, M. blackburni. Abbreviations include: aag, anteapical groove; acc, accessory elytral dorsal setae in interval 5; as, apical elytral seta; ca, carina laterad stria 7; ls, lateral elytral setae; ri, ridge mesad stria 8; sa, subapical elytral seta; sas, subapical sinuation of elytron; scs, supracarinal elytral setae; 8, medial portion of stria 8." figureDoi="10.3897/dez.65.21000.figure2" httpUri="https://binary.pensoft.net/fig/179958" pageId="36" pageNumber="37">Figures 2B</figureCitation>
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<figureCitation captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="36" pageNumber="37">, 17M-N</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F19" captionText="Figure 18. Paired left (above) and right (below) parameres of Mecyclothorax (Phacothorax) spp. (ectal view): A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. manautei; F, M. paniensis; G, M. mouensis." figureDoi="10.3897/dez.65.21000.figure18" httpUri="https://binary.pensoft.net/fig/179974" pageId="36" pageNumber="37">, 18G</figureCitation>
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<figureCitation captionStart="Figure 20" captionStartId="F21" captionText="Figure 20. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Humboldt, 580 m); E, M. kanak (Mt. Humboldt, 630 m); F, M. kanak (Mt. Dzumac); G, M. kanak (Col de Yate); H, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="36" pageNumber="37">, 20C</figureCitation>
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<figureCitation captionStart="Figure 23" captionStartId="F24" captionText="Figure 23. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Dzumac; common oviduct artificially darkened to show configuration); E, M. kanak (Pic du Grand Kaori); F, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure23" httpUri="https://binary.pensoft.net/fig/179980" pageId="36" pageNumber="37">, 23C</figureCitation>
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<figureCitation captionStart="Figure 24" captionStartId="F25" captionText="Figure 24. Left female gonocoxa of Mecyclothorax (Phacothorax) spp., ventral view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak; E, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure24" httpUri="https://binary.pensoft.net/fig/179981" pageId="36" pageNumber="37">, 24C</figureCitation>
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<figureCitation captionStart="Figure 25" captionStartId="F26" captionText="Figure 25. Geographical distributions of Mecyclothorax (Phacothorax) spp. (asterisked locality attributed to M. mouensis based on non-type female specimen)." figureDoi="10.3897/dez.65.21000.figure25" httpUri="https://binary.pensoft.net/fig/179982" pageId="36" pageNumber="37">, 25</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="diagnosis">
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<paragraph pageId="36" pageNumber="37">Diagnosis.</paragraph>
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<paragraph pageId="36" pageNumber="37">
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This species, along with
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<taxonomicName genus="M." lsidName="M. kanak" pageId="36" pageNumber="37" rank="species" species="kanak">
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<emphasis italics="true" pageId="36" pageNumber="37">M. kanak</emphasis>
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</taxonomicName>
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and
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<taxonomicName genus="M." lsidName="M. picdupinsensis" pageId="36" pageNumber="37" rank="species" species="picdupinsensis">
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<emphasis italics="true" pageId="36" pageNumber="37">M. picdupinsensis</emphasis>
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</taxonomicName>
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, comprises a triplet of cryptic sibling species best determined using the male aedeagal median lobe. To that end, this species is characterized by a median lobe with the apex dorsally and ventrally subparallel beyond the ostial opening, and with the
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<normalizedToken originalValue="lobe’s">lobe's</normalizedToken>
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apical face narrowly flattened (Fig.
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<figureCitation captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="36" pageNumber="37">17M</figureCitation>
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). All three species share broadly hemiovoid elytra, with this species exhibiting a broadly rounded basal groove across the humerus. Individuals of this species can be best diagnosed externally from those of the following two species by the broader elytral lateral marginal depression outside the anterior series of lateral elytral setae (Fig.
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<figureCitation captionStart="Figure 20" captionStartId="F21" captionText="Figure 20. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Humboldt, 580 m); E, M. kanak (Mt. Humboldt, 630 m); F, M. kanak (Mt. Dzumac); G, M. kanak (Col de Yate); H, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="36" pageNumber="37">20C</figureCitation>
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). The anterior lateral elytral setae are situated in the lateral depression in beetles of this species, whereas they lie on the upraised lateral reaches of the elytral disc in
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<taxonomicName genus="M." lsidName="M. kanak" pageId="36" pageNumber="37" rank="species" species="kanak">
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<emphasis italics="true" pageId="36" pageNumber="37">M. kanak</emphasis>
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</taxonomicName>
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and
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<taxonomicName genus="M." lsidName="M. picdupinsensis" pageId="36" pageNumber="37" rank="species" species="picdupinsensis">
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<emphasis italics="true" pageId="36" pageNumber="37">M. picdupinsensis</emphasis>
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</taxonomicName>
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. Also the sutural stria is deep and smooth on the disc and striae 2-3 are evident and traceable to nearly obsolete in
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<taxonomicName genus="M." lsidName="M. mouensis" pageId="36" pageNumber="37" rank="species" species="mouensis">
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<emphasis italics="true" pageId="36" pageNumber="37">M. mouensis</emphasis>
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</taxonomicName>
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. This differs from the sutural stria in beetles of the following two species where the sutural stria is crenulate to indistinctly punctate on the disc. Standardized body length 2.8-3.4 mm. Chaetotaxy +/+//+/-//+/2/+/+.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="description">
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<paragraph pageId="36" pageNumber="37">Description</paragraph>
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<paragraph pageId="36" pageNumber="37">
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(n = 5). Head capsule elongate, eyes small, very convex, ocular lobe-genal juncture evenly curved, a very shallow groove indicating limit of ocular lobe; 11 ommatidia along horizontal diameter of eye; ocular ratio 1.43-1.48, ocular lobe ratio 0.73-0.77, EyL/EyD = 2.17-2.38; frontal grooves narrowly incised, sinuously convergent to pit at frontoclypeal suture, briefly extended onto clypeus; mandibles moderately elongate, mandibular ratio 1.73; ligula narrowed to a moderately broad, slightly convex anterior margin, the two ligular setae separated by one setal diameter; paraglossae thin, extended twice as far beyond ligular margin as their basal length to margin, apex visibly spiculate (100
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<normalizedToken originalValue="×">x</normalizedToken>
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); antennae moderately elongate, antennomere 9 length 2.0
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<normalizedToken originalValue="×">x</normalizedToken>
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maximal breadth; antennomere 3 glabrous except for apical ring of setae. Pronotum very transverse, distinctly cordate, lateral margins sinuately concave anterad obtuse-rounded hind angles (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Thoracic structures of Mecyclothorax and Amblytelus spp. A-B. Pronotum, dorsal view: A, M. laterorectus; B, M. mouensis. C-F. Prosternum, ventral view: C, M. fleutiauxi; D, M. curtus; E, M. blackburni; F, M. moorei. G-J. Elytral apices, dorsal view: G, M. fleutiauxi; H, M. laterorectus; I, M. lissus; J, A. curtus. K-L. Elytral lateral margin showing eighth stria, lateral view: K, M. ambiguus; L, M. blackburni. Abbreviations include: aag, anteapical groove; acc, accessory elytral dorsal setae in interval 5; as, apical elytral seta; ca, carina laterad stria 7; ls, lateral elytral setae; ri, ridge mesad stria 8; sa, subapical elytral seta; sas, subapical sinuation of elytron; scs, supracarinal elytral setae; 8, medial portion of stria 8." figureDoi="10.3897/dez.65.21000.figure2" httpUri="https://binary.pensoft.net/fig/179958" pageId="36" pageNumber="37">2B</figureCitation>
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,
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<figureCitation captionStart="Figure 20" captionStartId="F21" captionText="Figure 20. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Humboldt, 580 m); E, M. kanak (Mt. Humboldt, 630 m); F, M. kanak (Mt. Dzumac); G, M. kanak (Col de Yate); H, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="36" pageNumber="37">20C</figureCitation>
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), MPW/BPW = 2.43-2.82, MPW/PL = 1.35-1.39; front angles slightly protruded, obtuse, APW/BPW = 1.42-1.50; median base unmargined basally, trapezoidally depressed relative to disc, the median longitudinal impression divided basally into two transverse impressions that isolate disc from base, each transverse basal impression terminated laterally in a short laterally arcuate laterobasal depression; median longitudinal impression finely incised on disc, terminated anteriorly at anterior transverse impression, a narrow fovea at posterior juncture with basal transverse impressions; anterior transverse impression shallow, broad, traceable to front angles; proepisternum separated from prosternum by a shallow groove anteriorly, and deep, distinct groove ventrally; smooth anteapical impression well developed laterally, continuous though shallower ventrally; prosternal process broadly, shallowly depressed between procoxae, the shallow depression extended 1/2 distance toward anterior prothoracic margin. Elytra broadly hemiovoid, humeri well-extended laterally before lateral margins curve posteriorly; basal elytral groove posteriorly curved laterad obtuse humeral angle; MEW/EL = 0.85-0.91; sutural stria well developed in basal 2/3, striae 2-3 traceable on disc near positions of dorsal elytral setae, stria 4 indicated by an series of intermittent longitudinal depressions; only stria 8 evident on elytral apex; elytra appressed and conjoined apically, the sutural margin upraised at apex. Pterothoracic mesepisternal anterior furrow with five to six depressions in one to two irregular rows; mesosternal-mesepisternal suture complete (as in Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Thoracic structures of Mecyclothorax spp. A-B. Lateral view of elytral epipleuron and meso-and metapleurites: A, M. laterorectus; B, M. fleutiauxi. C-D. Metathoracic flight wing: C, M. punctipennis, showing fully developed flight wing folded under elytron; D, M. lissus, stenopterous wing rudiment that extends beyond metanotum, rudimentary wing venation present at base of strap. Abbreviations include: mms, mesepisternal-mesosternal suture; msp, mesepimeron; mst, mesepisternum; mtp, metepimeron; mtt, metepisternum." figureDoi="10.3897/dez.65.21000.figure3" httpUri="https://binary.pensoft.net/fig/179959" pageId="36" pageNumber="37">3a</figureCitation>
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); metepisternum length and breadth subequal; metepisternal-metepimeral suture complete though fine laterally. Abdomen with elongate crescent-shaped depression along suture between first and second ventrite, second ventrite only slightly depressed within crescent; suture between second and third ventrites reduced though traceable laterally; ventrites 3-6 with broad, shallow, linear plaques near lateral margin. Microsculpture of frons reduced, surface glossy, indistinct transverse mesh-sculpticell breadth 2
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<normalizedToken originalValue="×">x</normalizedToken>
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length-visible over portions of vertex; pronotal disc glossy but with transverse sculpticells visible outside areas of reflection, trapezoidal median base glossy with indistinct transverse mesh anterad laterobasal depression; elytra opalescent, disc covered with very finely separated transverse lines, elytral apex covered with elongate transverse sculpticells and lines.
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</paragraph>
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<paragraph pageId="36" pageNumber="37">
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<emphasis italics="true" pageId="36" pageNumber="37">Male genitalia</emphasis>
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(n = 8). Antecostal margin of mediotergite IX angulate, not extended (Fig.
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<figureCitation captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="36" pageNumber="37">17N</figureCitation>
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); right paramere narrow, with basal cuff articulating with elongate, whiplike apical extension (Fig.
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<figureCitation captionStart="Figure 18" captionStartId="F19" captionText="Figure 18. Paired left (above) and right (below) parameres of Mecyclothorax (Phacothorax) spp. (ectal view): A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. manautei; F, M. paniensis; G, M. mouensis." figureDoi="10.3897/dez.65.21000.figure18" httpUri="https://binary.pensoft.net/fig/179974" pageId="36" pageNumber="37">18G</figureCitation>
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), six setae along ventral margin to complement to longer apical setae; left paramere narrow basally, evenly narrowed to whiplike apex; aedeagal median lobe robust, broadest medially basad apex of ostial opening, apex subparallel dorsoventrally beyond ostium, with apical face somewhat flattened (Fig.
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<figureCitation captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="36" pageNumber="37">17M</figureCitation>
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); aedeagal median lobe internal sac with broad, basal dorsal plate and a large field of macrospicules apicad the plate, but flagellum and flagellar sheath not present.
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</paragraph>
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<paragraph pageId="36" pageNumber="37">
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<emphasis italics="true" pageId="36" pageNumber="37">Female reproductive tract</emphasis>
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(n = 1). Bursa copulatrix length subequal to circumference, its surface thin, translucent, slightly wrinkled (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F24" captionText="Figure 23. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Dzumac; common oviduct artificially darkened to show configuration); E, M. kanak (Pic du Grand Kaori); F, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure23" httpUri="https://binary.pensoft.net/fig/179980" pageId="36" pageNumber="37">23C</figureCitation>
|
||
); spermathecal duct entering near bursa-common oviduct juncture with duct oriented toward right side of bursa; elongate laminar helminthoid sclerite present at base of spermathecal duct; spermatheca of only slightly greater circumference than spermathecal duct, spermathecal gland duct entering at base of spermathecal reservoir; ligular apophysis present on common oviduct; basal gonocoxite 1 with apical fringe of two setae laterally, a very small setae may be present immediately mesad, and several smaller setae along medial margin (Fig.
|
||
<figureCitation captionStart="Figure 24" captionStartId="F25" captionText="Figure 24. Left female gonocoxa of Mecyclothorax (Phacothorax) spp., ventral view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak; E, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure24" httpUri="https://binary.pensoft.net/fig/179981" pageId="36" pageNumber="37">24C</figureCitation>
|
||
); medial margin bearing a dense field of microtrichia basally, this field about 3-4 microtrichia broad and appearing like the hooklike surface of a Velcro® closure; gonocoxite 2 moderately broad basally, basal width 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
medial length; two gracile lateral ensiform setae of moderate length present.
|
||
</paragraph>
|
||
<paragraph pageId="36" pageNumber="37">
|
||
<emphasis italics="true" pageId="36" pageNumber="37">Types</emphasis>
|
||
- Holotype male (MNHN): NEW CALEDONIA / Mt Mou summit / 24 May 1984 / G. Monteith & D. Cook // Q.M. Berlesate 659 / 22.04S X 166.21E / Rainforest, 1200 n / Litter // QUEENSLAND / MUSEUM LOAN / DATE: Nov. 2003 / No. LEN-1686 (green label) // HOLOTYPE /
|
||
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
|
||
/
|
||
<taxonomicName authorityName="Moore & Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax mouensis" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="mouensis">Mecyclothorax mouensis</taxonomicName>
|
||
/ B.P. Moore & J.K. / Liebherr 2017 (black-bordered red label).
|
||
</paragraph>
|
||
<paragraph pageId="36" pageNumber="37">
|
||
Paratypes (33 specimens). NEW CALEDONIA: Mts. Koghis, [code PA 58],
|
||
<geoCoordinate degrees="22" direction="south" minutes="11" orientation="latitude" precision="925" value="-22.183332">22°11'S</geoCoordinate>
|
||
<geoCoordinate degrees="166" direction="east" minutes="31" orientation="longitude" precision="925" value="166.51666">166°31'E</geoCoordinate>
|
||
, 30-viii-1970, Franz (NHMW, 3), ~5 km N Noumea, 550 m el.,
|
||
<geoCoordinate degrees="22" direction="south" minutes="10.5" orientation="latitude" precision="92" value="-22.175">22 10.5'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="166" direction="east" minutes="30.3" orientation="longitude" precision="92" value="166.505">166 30.3'E</geoCoordinate>
|
||
, forest litter, 25-xi-2009, Schuh (NHMW, 1), forest, 520 m el.,
|
||
<geoCoordinate degrees="22" direction="south" minutes="10.7" orientation="latitude" precision="92" value="-22.178333">22°10.7'S</geoCoordinate>
|
||
<geoCoordinate degrees="166" direction="east" minutes="30.3" orientation="longitude" precision="92" value="166.505">166°30.3'E</geoCoordinate>
|
||
, sifted litter, 25-x-2008, Wanat (MNHW, 9), La Roussette, 600 m el.,
|
||
<geoCoordinate degrees="22" direction="south" minutes="10.8" orientation="latitude" precision="92" value="-22.18">22°10.8'S</geoCoordinate>
|
||
<geoCoordinate degrees="166" direction="east" minutes="30.7" orientation="longitude" precision="92" value="166.51167">166°30.7'E</geoCoordinate>
|
||
, sifted litter, 27-x-2008, Wanat (MNHW, 4), track entrance, 500 m el.,
|
||
<geoCoordinate degrees="22" direction="south" minutes="11" orientation="latitude" precision="925" value="-22.183332">22°11'S</geoCoordinate>
|
||
<geoCoordinate degrees="166" direction="east" minutes="31" orientation="longitude" precision="925" value="166.51666">166°31'E</geoCoordinate>
|
||
, berlesate, 06-v-2005, lot 12264, Monteith (QMB, 4), 500 m el.,
|
||
<geoCoordinate degrees="22" direction="south" minutes="10" orientation="latitude" precision="925" value="-22.166666">22°10'S</geoCoordinate>
|
||
<geoCoordinate degrees="166" direction="east" minutes="31" orientation="longitude" precision="925" value="166.51666">166°31'E</geoCoordinate>
|
||
, 26-viii-1978, S. & J. Peck (CNC, 1), 400-500 m el.,
|
||
<geoCoordinate degrees="22" direction="south" minutes="11" orientation="latitude" precision="925" value="-22.183332">22°11'S</geoCoordinate>
|
||
<geoCoordinate degrees="166" direction="east" minutes="31" orientation="longitude" precision="925" value="166.51666">166°31'E</geoCoordinate>
|
||
, primary forest, 18-19-x-1998,
|
||
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
|
||
(MBC, 1; MHNG, 6); Mt. Mou, summit, 1200 m el.,
|
||
<geoCoordinate degrees="22" direction="south" minutes="04" orientation="latitude" precision="925" value="-22.066668">22°04'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="166" direction="east" minutes="21" orientation="longitude" precision="925" value="166.35">166°21'E</geoCoordinate>
|
||
, rainforest litter, 24-v-1984, Monteith & Cook (QMB, 2), top camp, 1150 m el.,
|
||
<geoCoordinate degrees="22" direction="south" minutes="04" orientation="latitude" precision="925" value="-22.066668">22°04'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="166" direction="east" minutes="21" orientation="longitude" precision="925" value="166.35">166°21'E</geoCoordinate>
|
||
, berlesate, sieved litter, 27-xii-2004, lot 12013, Monteith (QMB, 2). Non-type female specimen:
|
||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken originalValue="Cachée">Cachee</normalizedToken>
|
||
, 250 m el.,
|
||
<geoCoordinate degrees="22" direction="south" minutes="11.5" orientation="latitude" precision="92" value="-22.191668">22°11.5'S</geoCoordinate>
|
||
<geoCoordinate degrees="166" direction="east" minutes="47.2" orientation="longitude" precision="92" value="166.78667">166°47.2'E</geoCoordinate>
|
||
, sifted litter, 26-x-2008, Wanat (MNHW, 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="36" pageNumber="37" type="etymology">
|
||
<paragraph pageId="36" pageNumber="37">Etymology.</paragraph>
|
||
<paragraph pageId="36" pageNumber="37">
|
||
As in the species above, this species epithet elides the type locality, Mt. Mou, with the genitive, adjectival ending -ensis. As Dr. Barry P. Moore had both diagnosed this species from the following
|
||
<taxonomicName genus="M." lsidName="M. kanak" pageId="36" pageNumber="37" rank="species" species="kanak">
|
||
<emphasis italics="true" pageId="36" pageNumber="37">M. kanak</emphasis>
|
||
</taxonomicName>
|
||
through the use of male genitalic characters, and had chosen a holotype specimen, he is given senior author status for this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="36" pageNumber="37" type="distribution">
|
||
<paragraph pageId="36" pageNumber="37">Distribution and habitat.</paragraph>
|
||
<paragraph pageId="36" pageNumber="37">
|
||
This species is restricted to the southern portion of Grande Terre, from Mt. Mou on the west, Mts. Koghis north of Noumea, and with a easternmost record from
|
||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken originalValue="Cachée">Cachee</normalizedToken>
|
||
based on a single non-type female specimen (Fig.
|
||
<figureCitation captionStart="Figure 25" captionStartId="F26" captionText="Figure 25. Geographical distributions of Mecyclothorax (Phacothorax) spp. (asterisked locality attributed to M. mouensis based on non-type female specimen)." figureDoi="10.3897/dez.65.21000.figure25" httpUri="https://binary.pensoft.net/fig/179982" pageId="36" pageNumber="37">25</figureCitation>
|
||
). Collection localities range 250-1200 m elevation. Given that 24 of the 35 specimens are labeled from Berlese samples or sieved litter, and another 7 specimens (1, S. & J. Peck, CNC; 6,
|
||
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
|
||
, MHNG) were collected by coleopterists studying the microcoleopteran litter fauna, it can be concluded that this species occupies the ground-level litter layer.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |