219 lines
27 KiB
XML
219 lines
27 KiB
XML
<document id="55D1A15780BCC3736AB836032F708F2C" ID-DOI="10.1080/14772019.2016.1245680" ID-ISSN="1478-0941" ID-Zenodo-Dep="10883052" ID-ZooBank="3EBCAEF3-27C2-4216-9F18-89F195FA534F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1711483395715" checkinUser="felipe" docAuthor="Foster, William J., Danise, Silvia & Twitchett, Richard J." docDate="2017" docId="C53B0B4D8057E8236C4DFCB68F54278D" docLanguage="en" docName="JSystPalaeontol.15.10.851-877.pdf" docOrigin="Journal of Systematic Palaeontology 15 (10)" docSource="http://dx.doi.org/10.1080/14772019.2016.1245680" docStyle="DocumentStyle:602474738E3D100F0F13788B53F61545.2:JSystPalaeontol.2016-.journal_article.open" docStyleId="602474738E3D100F0F13788B53F61545" docStyleName="JSystPalaeontol.2016-.journal_article.open" docStyleVersion="2" docTitle="Nucinella nakremi Foster & Danise & Twitchett 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="858" masterDocId="390273358050E82B6D0FFFE98D67263C" masterDocTitle="A silicified Early Triassic marine assemblage from Svalbard" masterLastPageNumber="877" masterPageNumber="851" pageNumber="857" updateTime="1711483788606" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="97567A2D8ABD51FDF3F37D4CAF1BC3FA" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="D9D6DD3415D6E365EB9DFC8E5558531E">
|
||
<mods:title id="12122529FEFBF2CBFDA68DAB33F343E7">A silicified Early Triassic marine assemblage from Svalbard</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="E6391FA2634A95990970336D09CE6BED" type="personal">
|
||
<mods:role id="17C439A0089B0EAC29C9A1192200DE3A">
|
||
<mods:roleTerm id="8E56D4D6B4C6CA5F5F534049D5E2B585">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2A782B8A469B673F7D1F6D707317010D">Foster, William J.</mods:namePart>
|
||
<mods:affiliation id="EB03580901C150915F9886B1890D50C2">Jackson School of Geosciences, University of Texas, Austin, TX 78712, USA; & Earth Sciences, Plymouth University, Plymouth, PL 4 8 AA, UK; & Department of Earth Sciences, Natural History Museum, London, SW 7 5 BD, UK;</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="9C77EC02A50F3905A60873FEEB7A7ABC" type="personal">
|
||
<mods:role id="3DB503499114751E1F4C829A09D4B93A">
|
||
<mods:roleTerm id="81C3E4D37ADECE40DA8CC96E95CE2D9B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5A9B13F8E52DE8715502CEDFD54E2A1E">Danise, Silvia</mods:namePart>
|
||
<mods:affiliation id="45222B36E0AFBE218E28D41066736144">Earth Sciences, Plymouth University, Plymouth, PL 4 8 AA, UK; & Department of Geology, University of Georgia, Athens, GA 30602, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="22CFF90D61EB06AAE3E2FCAAAE913F33" type="personal">
|
||
<mods:role id="C9A51EC0D1F39547786385ABA7443E74">
|
||
<mods:roleTerm id="1779DFF65D17DFAD8C0160D53F00FED9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="41B5E991E915E49CA523D5B10CB54281">Twitchett, Richard J.</mods:namePart>
|
||
<mods:affiliation id="4F0595428A8F9BFE98F93C44A87CBA2B">Department of Earth Sciences, Natural History Museum, London, SW 7 5 BD, UK;</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="24A896A5C7CEF37887C580F29558B762">text</mods:typeOfResource>
|
||
<mods:relatedItem id="55520DEEF5041B79A7C2E4284A80702E" type="host">
|
||
<mods:titleInfo id="EBD3B36A7CA092294A0253BF94C97D4A">
|
||
<mods:title id="BFEE2E71F468EAD9DCD6CF423B17A11A">Journal of Systematic Palaeontology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="8025B17B766BC9BECF8C837F8AF7B2D8">
|
||
<mods:date id="7F4DA478D2F9ED192396AD1034BEBEB2">2017</mods:date>
|
||
<mods:detail id="81437D9ED73666312776DCBDA8D908D9" type="pubDate">
|
||
<mods:number id="360CE85511F955A04FEFEBF4FA27A6B1">2016-11-10</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CF2F00E9370C97E46460CAA62382CEA4" type="volume">
|
||
<mods:number id="1F31BB267EFCD0473B4B534E70DED44E">15</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="FF6481202808296C14657CFBB6970401" type="issue">
|
||
<mods:number id="4C78B6AA8AEAF43335BD9689F3D635D1">10</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="A68A154D0C8006B4D1CBB17CF730FDAE" unit="page">
|
||
<mods:start id="4599C3CE93975E81647A061C9ED8D33B">851</mods:start>
|
||
<mods:end id="9F9BA37E10541277A57EE5C15817696C">877</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="519B8CEF110811C8E68EC37E93D5B308">
|
||
<mods:url id="B99B1CC8566387083446F688006EBEAE">http://dx.doi.org/10.1080/14772019.2016.1245680</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="F32640CED758F76BD8DA1D127B1EA9FB">journal article</mods:classification>
|
||
<mods:identifier id="74C9EC6DAB245F74EFCA717AD5B768D2" type="DOI">10.1080/14772019.2016.1245680</mods:identifier>
|
||
<mods:identifier id="60DC3B38DE459FC79255F702BF7D0309" type="ISSN">1478-0941</mods:identifier>
|
||
<mods:identifier id="B85ACF464D6085EEC44F1E645A4337ED" type="Zenodo-Dep">10883052</mods:identifier>
|
||
<mods:identifier id="02B168603F579E28B191661147382C51" type="ZooBank">3EBCAEF3-27C2-4216-9F18-89F195FA534F</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="C53B0B4D8057E8236C4DFCB68F54278D" ID-DOI="http://doi.org/10.5281/zenodo.10883096" ID-Zenodo-Dep="10883096" LSID="urn:lsid:plazi:treatment:C53B0B4D8057E8236C4DFCB68F54278D" httpUri="http://treatment.plazi.org/id/C53B0B4D8057E8236C4DFCB68F54278D" lastPageId="8" lastPageNumber="858" pageId="7" pageNumber="857">
|
||
<subSubSection id="0588E9D08057E82C6C4DFCB68F0C254A" box="[322,619,863,887]" pageId="7" pageNumber="857" type="nomenclature">
|
||
<paragraph id="4D2DBA5B8057E82C6C4DFCB68F0C254A" blockId="7.[322,619,863,918]" box="[322,619,863,887]" pageId="7" pageNumber="857">
|
||
<heading id="16650D378057E82C6C4DFCB68F0C254A" box="[322,619,863,887]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="857" reason="8">
|
||
<taxonomicName id="8A92C1D88057E82C6C4DFCB68F76254B" authority="Foster & Danise & Twitchett, 2017" authorityName="Foster & Danise & Twitchett" authorityYear="2017" box="[322,529,863,887]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="species" species="nakremi" status="sp. nov.">
|
||
<emphasis id="7FE666498057E82C6C4DFCB68F76254B" bold="true" box="[322,529,863,887]" italics="true" pageId="7" pageNumber="857">Nucinella nakremi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="64D5DB328057E82C6F18FCB68F0C254A" box="[535,619,863,886]" pageId="7" pageNumber="857" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0588E9D08057E82C6CA1FC968C9925AA" box="[430,510,895,918]" pageId="7" pageNumber="857" type="description">
|
||
<paragraph id="4D2DBA5B8057E82C6CA1FC968C9925AA" blockId="7.[322,619,863,918]" box="[430,510,895,918]" pageId="7" pageNumber="857">
|
||
(
|
||
<figureCitation id="D5A9A6DE8057E82C6CB8FC968C9225AA" box="[439,501,895,918]" captionStart="Figure 4" captionStartId="7.[159,227,566,587]" captionTargetBox="[175,1452,183,534]" captionTargetId="figure-571@7.[175,1452,183,534]" captionTargetPageId="7" captionText="Figure 4. Nucinella nakremi sp. nov. A, holotype, NHMUK PI MB 1219, external view, left valve; B, paratype, NHMUK PI MB 1220, external view, left valve; C, holotype, NHMUK PI MB 1219, interior view, left valve; D, paratype, NHMUK PI MB 1220, interior view (transposed hinge), left valve. Scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10883060" httpUri="https://zenodo.org/record/10883060/files/figure.png" pageId="7" pageNumber="857">Fig. 4</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0588E9D08057E82C6D90FC598C96221B" pageId="7" pageNumber="857" type="diagnosis">
|
||
<paragraph id="4D2DBA5B8057E82C6D90FC598C96221B" blockId="7.[159,781,944,1063]" pageId="7" pageNumber="857">
|
||
<emphasis id="7FE666498057E82C6D90FC598C7325FB" bold="true" box="[159,276,944,967]" pageId="7" pageNumber="857">Diagnosis.</emphasis>
|
||
A small
|
||
<taxonomicName id="8A92C1D88057E82C6C8AFC598C8A25FB" authorityName="Wood" authorityYear="1851" box="[389,493,944,967]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7FE666498057E82C6C8AFC598C8A25FB" box="[389,493,944,967]" italics="true" pageId="7" pageNumber="857">Nucinella</emphasis>
|
||
</taxonomicName>
|
||
having a nuculoid shape, smooth shell except for growth lines. Prosogyrate beak, one to three subumbonal teeth. Ligament amphidetic and does not invade the hinge plate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0588E9D08057E82C6D90FBAB8C272390" pageId="7" pageNumber="857" type="materials_examined">
|
||
<paragraph id="4D2DBA5B8057E82C6D90FBAB8FD12244" blockId="7.[159,781,1090,1144]" pageId="7" pageNumber="857">
|
||
<materialsCitation id="FDFAB0068057E82C6D90FBAB8FD12244" collectionCode="NHMUK, PI" pageId="7" pageNumber="857" specimenCode="MB 1219" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="7FE666498057E82C6D90FBAB8C692265" bold="true" box="[159,270,1090,1113]" pageId="7" pageNumber="857">
|
||
<typeStatus id="922904F98057E82C6D90FBAB8C6E2265" box="[159,265,1090,1113]" pageId="7" pageNumber="857" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Disarticulated left valve,
|
||
<collectionCode id="2B83229E8057E82C6F47FBAB8FCB2265" box="[584,684,1090,1113]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="857" type="Museum">NHMUK</collectionCode>
|
||
<collectionCode id="2B83229E8057E82C6FB3FBAB8FB32265" box="[700,724,1090,1113]" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="857">PI</collectionCode>
|
||
<specimenCode id="1D3412208057E82C6FEBFBAB8DB02244" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="7" pageNumber="857">MB 1219</specimenCode>
|
||
, LD-04; length = 1.1 mm, height = 1.1 mm.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="4D2DBA5B8057E82C6D90FB7A8C4822D6" blockId="7.[159,781,1171,1258]" pageId="7" pageNumber="857">
|
||
<materialsCitation id="FDFAB0068057E82C6D90FB7A8C4B22D6" collectionCode="NHMUK, PI" pageId="7" pageNumber="857" specimenCode="MB 1220" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="7FE666498057E82C6D90FB7A8C692296" bold="true" box="[159,270,1171,1194]" pageId="7" pageNumber="857">
|
||
<typeStatus id="922904F98057E82C6D90FB7A8C6E2296" box="[159,265,1171,1194]" pageId="7" pageNumber="857" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Disarticulated left valve,
|
||
<collectionCode id="2B83229E8057E82C6F47FB7A8FCB2296" box="[584,684,1171,1194]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="857" type="Museum">NHMUK</collectionCode>
|
||
<collectionCode id="2B83229E8057E82C6FB3FB7A8FB32296" box="[700,724,1171,1194]" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="857">PI</collectionCode>
|
||
<specimenCode id="1D3412208057E82C6FEBFB7A8DB022F6" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="7" pageNumber="857">MB 1220</specimenCode>
|
||
, LD-04; length = 0.9 mm, height = 0.8 mm (transposed hinge)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4D2DBA5B8057E82C6D90FAED8C122307" blockId="7.[159,781,1284,1339]" pageId="7" pageNumber="857">
|
||
<emphasis id="7FE666498057E82C6D90FAED8C332327" bold="true" box="[159,340,1284,1307]" pageId="7" pageNumber="857">Other material.</emphasis>
|
||
<materialsCitation id="FDFAB0068057E82C6C6DFAED8C162307" collectionCode="NHMUK, PI" pageId="7" pageNumber="857" specimenCode="MB 1221-1222" specimenCount="2">
|
||
<specimenCount id="5B9471D28057E82C6C6DFAED8F6C2327" box="[354,523,1284,1307]" count="2" pageId="7" pageNumber="857" type="generic">Two specimens</specimenCount>
|
||
from LD-04 (
|
||
<collectionCode id="2B83229E8057E82C6FA8FAED8E6A2327" box="[679,781,1284,1307]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="857" type="Museum">NHMUK</collectionCode>
|
||
<collectionCode id="2B83229E8057E82C6D90FACD8DD02307" box="[159,183,1316,1339]" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="857">PI</collectionCode>
|
||
<specimenCode id="1D3412208057E82C6DB2FACD8C0E2307" box="[189,361,1316,1339]" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="7" pageNumber="857">MB 1221–1222</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4D2DBA5B8057E82C6D90FABF8C272390" blockId="7.[159,781,1366,1452]" pageId="7" pageNumber="857">
|
||
<emphasis id="7FE666498057E82C6D90FABF8CE62351" bold="true" box="[159,385,1366,1389]" pageId="7" pageNumber="857">Derivation of name.</emphasis>
|
||
Named after Dr Hans Arne Nakrem in recognition of his work on Permian and Triassic fossils from
|
||
<collectingCountry id="3585FACB8057E82C6DD5FA7C8C5B2390" box="[218,316,1429,1452]" name="Svalbard and Jan Mayen" pageId="7" pageNumber="857">Svalbard</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0588E9D08057E82C6D90FA2E8F2920E1" pageId="7" pageNumber="857" type="description">
|
||
<paragraph id="4D2DBA5B8057E82C6D90FA2E8F2920E1" blockId="7.[159,781,1479,1757]" pageId="7" pageNumber="857">
|
||
<emphasis id="7FE666498057E82C6D90FA2E8C4D23E2" bold="true" box="[159,298,1479,1502]" pageId="7" pageNumber="857">Description.</emphasis>
|
||
Shell small, nuculoid and ovate. Posterior dorsal margin distinct, slightly incurved; posterior margin rounded. Inequilateral, prosogyrate, with beaks close to anterior margin. Umbo prominent. Smooth, thin shell with very weak growth lines. Monomyarian: posterior adductor muscle scar absent; anterior adductor large, oval. One to two subumbonal, pointed blade-like teeth plus one anterior tooth. Ligament amphidetic, prominent, external does not invade the hinge plate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0588E9D08057E8236D90F91E8F54278D" lastPageId="8" lastPageNumber="858" pageId="7" pageNumber="857" type="discussion">
|
||
<paragraph id="4D2DBA5B8057E82C6D90F91E89D323EC" blockId="7.[159,781,1783,1934]" lastBlockId="7.[845,1467,763,1934]" pageId="7" pageNumber="857">
|
||
<emphasis id="7FE666498057E82C6D90F91E8C692132" bold="true" box="[159,270,1783,1806]" pageId="7" pageNumber="857">Remarks.</emphasis>
|
||
These specimens differ from other described nucinellid species in having fewer hinge teeth, a more elliptical shape and a prosogyrate beak. Such differences may occur during the ontogeny of
|
||
<taxonomicName id="8A92C1D88057E82C6F1DF8BE8F1D2152" authorityName="Wood" authorityYear="1851" box="[530,634,1879,1902]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7FE666498057E82C6F1DF8BE8F1D2152" box="[530,634,1879,1902]" italics="true" pageId="7" pageNumber="857">Nucinella</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<bibRefCitation id="2903C7AA8057E82C6FB8F8BE8DB321B2" author="Bernard, F." pageId="7" pageNumber="857" pagination="1 - 208" refId="ref16190" refString="Bernard, F. 1898. Recherches ontog´enetiques et morphologiques sur la coquille des lamellibranches. Premi`ere partie. Taxodontes et Anisomyaires. Annales des Sciences Naturelles. Zoologie et Paleontologie´, 8, 1 - 208." type="journal article" year="1898">Bernard 1898</bibRefCitation>
|
||
) and so are not sufficient for assignment to a separate genus. These specimens are, however, considered to represent a separate species rather than an intermediate ontogenetic stage between the protoconch and adult stage of
|
||
<taxonomicName id="8A92C1D88057E82C6E42FCB28960254E" authorityName="Foster & Danise & Twitchett" authorityYear="2017" box="[845,1031,859,882]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="species" species="taylori">
|
||
<emphasis id="7FE666498057E82C6E42FCB28960254E" box="[845,1031,859,882]" italics="true" pageId="7" pageNumber="857">Nucinella taylori</emphasis>
|
||
</taxonomicName>
|
||
. During ontogeny, the shape, size and position of nucinellid subombonal teeth also vary: in earlier stages of development they are more rounded and later they develop a chevron-blade shape with the older teeth making space below the beak for thinner, newer ones (
|
||
<bibRefCitation id="2903C7AA8057E82C6E83FC138946222D" author="Bernard, F." box="[908,1057,1018,1041]" pageId="7" pageNumber="857" pagination="1 - 208" refId="ref16190" refString="Bernard, F. 1898. Recherches ontog´enetiques et morphologiques sur la coquille des lamellibranches. Premi`ere partie. Taxodontes et Anisomyaires. Annales des Sciences Naturelles. Zoologie et Paleontologie´, 8, 1 - 208." type="journal article" year="1898">Bernard 1898</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2903C7AA8057E82C693FFC1389AA222D" author="La Perna, R." box="[1072,1229,1018,1041]" pageId="7" pageNumber="857" pagination="571 - 577" refId="ref19380" refString="La Perna, R. 2004. Nucinella alibrandi (Conti, 1864) and N. seguenzae (Dall, 1898), the last European nucinellids (Bivalvia, Protobranchia). Rivista Italiana di Paleontologica e Stratigrafia, 110, 571 - 577." type="journal article" year="2004">
|
||
<taxonomicName id="8A92C1D88057E82C693FFC1389F6222D" box="[1072,1169,1018,1041]" class="Lingulata" family="Lingulidae" genus="Lingularia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lingulida" pageId="7" pageNumber="857" phylum="Brachiopoda" rank="species" species="perna">La Perna</taxonomicName>
|
||
2004
|
||
</bibRefCitation>
|
||
). In contrast, the subumbonal teeth of specimens assigned to
|
||
<taxonomicName id="8A92C1D88057E82C681FFBF388ED220D" authorityName="Foster & Danise & Twitchett" authorityYear="2017" box="[1296,1418,1050,1073]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="species" species="nakremi">
|
||
<emphasis id="7FE666498057E82C681FFBF388ED220D" box="[1296,1418,1050,1073]" italics="true" pageId="7" pageNumber="857">N. nakremi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8A92C1D88057E82C6E42FBD38ED4226D" authorityName="Foster & Danise & Twitchett" authorityYear="2017" box="[845,947,1082,1105]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="species" species="taylori">
|
||
<emphasis id="7FE666498057E82C6E42FBD38ED4226D" box="[845,947,1082,1105]" italics="true" pageId="7" pageNumber="857">N. taylori</emphasis>
|
||
</taxonomicName>
|
||
have comparable shapes, and so indicate a similar stage of development. Furthermore, because the position of the ligament in nucinellids is fixed after metamorphosis (
|
||
<bibRefCitation id="2903C7AA8057E82C690DFB7389FE228D" author="Bernard, F." box="[1026,1177,1178,1201]" pageId="7" pageNumber="857" pagination="1 - 208" refId="ref16190" refString="Bernard, F. 1898. Recherches ontog´enetiques et morphologiques sur la coquille des lamellibranches. Premi`ere partie. Taxodontes et Anisomyaires. Annales des Sciences Naturelles. Zoologie et Paleontologie´, 8, 1 - 208." type="journal article" year="1898">Bernard 1898</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2903C7AA8057E82C69A4FB738829228D" author="La Perna, R." box="[1195,1358,1178,1201]" pageId="7" pageNumber="857" pagination="571 - 577" refId="ref19380" refString="La Perna, R. 2004. Nucinella alibrandi (Conti, 1864) and N. seguenzae (Dall, 1898), the last European nucinellids (Bivalvia, Protobranchia). Rivista Italiana di Paleontologica e Stratigrafia, 110, 571 - 577." type="journal article" year="2004">
|
||
<taxonomicName id="8A92C1D88057E82C69A4FB738868228D" box="[1195,1295,1178,1201]" class="Lingulata" family="Lingulidae" genus="Lingularia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lingulida" pageId="7" pageNumber="857" phylum="Brachiopoda" rank="species" species="perna">La Perna</taxonomicName>
|
||
2004
|
||
</bibRefCitation>
|
||
), and the only subsequent ontogenetic change is an increase in ligament size with age, the differences in ligament position between
|
||
<taxonomicName id="8A92C1D88057E82C6EA2FB108943232C" authorityName="Foster & Danise & Twitchett" authorityYear="2017" box="[941,1060,1273,1296]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="species" species="nakremi">
|
||
<emphasis id="7FE666498057E82C6EA2FB108943232C" box="[941,1060,1273,1296]" italics="true" pageId="7" pageNumber="857">N. nakremi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8A92C1D88057E82C6956FB1089A7232C" authorityName="Foster & Danise & Twitchett" authorityYear="2017" box="[1113,1216,1273,1296]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="species" species="taylori">
|
||
<emphasis id="7FE666498057E82C6956FB1089A7232C" box="[1113,1216,1273,1296]" italics="true" pageId="7" pageNumber="857">N. taylori</emphasis>
|
||
</taxonomicName>
|
||
cannot be ontogenetic. If the specimens assigned to
|
||
<taxonomicName id="8A92C1D88057E82C6989FAF0899A230C" authorityName="Foster & Danise & Twitchett" authorityYear="2017" box="[1158,1277,1305,1328]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="species" species="nakremi">
|
||
<emphasis id="7FE666498057E82C6989FAF0899A230C" box="[1158,1277,1305,1328]" italics="true" pageId="7" pageNumber="857">N. nakremi</emphasis>
|
||
</taxonomicName>
|
||
were included as an intermediate ontogenetic stage of
|
||
<taxonomicName id="8A92C1D88057E82C69EEFAD0882E236C" authorityName="Foster & Danise & Twitchett" authorityYear="2017" box="[1249,1353,1337,1360]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="species" species="taylori">
|
||
<emphasis id="7FE666498057E82C69EEFAD0882E236C" box="[1249,1353,1337,1360]" italics="true" pageId="7" pageNumber="857">N. taylori</emphasis>
|
||
</taxonomicName>
|
||
, the ontogenetic pattern would not match any known nucinellid (cf.
|
||
<bibRefCitation id="2903C7AA8057E82C6E77FA90896D23AC" author="Bernard, F." box="[888,1034,1401,1424]" pageId="7" pageNumber="857" pagination="1 - 208" refId="ref16190" refString="Bernard, F. 1898. Recherches ontog´enetiques et morphologiques sur la coquille des lamellibranches. Premi`ere partie. Taxodontes et Anisomyaires. Annales des Sciences Naturelles. Zoologie et Paleontologie´, 8, 1 - 208." type="journal article" year="1898">Bernard 1898</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2903C7AA8057E82C6917FA9089D323AC" author="La Perna, R." box="[1048,1204,1401,1424]" pageId="7" pageNumber="857" pagination="571 - 577" refId="ref19380" refString="La Perna, R. 2004. Nucinella alibrandi (Conti, 1864) and N. seguenzae (Dall, 1898), the last European nucinellids (Bivalvia, Protobranchia). Rivista Italiana di Paleontologica e Stratigrafia, 110, 571 - 577." type="journal article" year="2004">
|
||
<taxonomicName id="8A92C1D88057E82C6917FA90891F23AC" box="[1048,1144,1401,1424]" class="Lingulata" family="Lingulidae" genus="Lingularia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lingulida" pageId="7" pageNumber="857" phylum="Brachiopoda" rank="species" species="perna">La Perna</taxonomicName>
|
||
2004
|
||
</bibRefCitation>
|
||
), and, therefore, the differences in the subumbonal teeth and the position of the ligament support their separation.
|
||
</paragraph>
|
||
<paragraph id="4D2DBA5B8057E8236E68FA318FBF2768" blockId="7.[845,1467,763,1934]" lastBlockId="8.[159,781,189,340]" lastPageId="8" lastPageNumber="858" pageId="7" pageNumber="857">
|
||
A specimen of
|
||
<taxonomicName id="8A92C1D88057E82C691FFA3189BF23D3" authority="Foster & Danise & Twitchett, 2017" authorityName="Foster & Danise & Twitchett" authorityYear="2017" box="[1040,1240,1496,1519]" class="Bivalvia" family="Nucinellidae" genus="Nucinella" kingdom="Animalia" order="Solemyida" pageId="7" pageNumber="857" phylum="Mollusca" rank="species" species="nakremi" status="sp. nov.">
|
||
<emphasis id="7FE666498057E82C691FFA3189BF23D3" box="[1040,1240,1496,1519]" italics="true" pageId="7" pageNumber="857">Nucinella nakremi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="64D5DB328057E82C69EEFA31885023D3" box="[1249,1335,1496,1519]" pageId="7" pageNumber="857" rank="species">sp. nov.</taxonomicNameLabel>
|
||
includes an example of a transposed hinge on a left valve (
|
||
<figureCitation id="D5A9A6DE8057E82C6858FA1188C92033" box="[1367,1454,1528,1551]" captionStart="Figure 4" captionStartId="7.[159,227,566,587]" captionTargetBox="[175,1452,183,534]" captionTargetId="figure-571@7.[175,1452,183,534]" captionTargetPageId="7" captionText="Figure 4. Nucinella nakremi sp. nov. A, holotype, NHMUK PI MB 1219, external view, left valve; B, paratype, NHMUK PI MB 1220, external view, left valve; C, holotype, NHMUK PI MB 1219, interior view, left valve; D, paratype, NHMUK PI MB 1220, interior view (transposed hinge), left valve. Scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10883060" httpUri="https://zenodo.org/record/10883060/files/figure.png" pageId="7" pageNumber="857">Fig. 4D</figureCitation>
|
||
). Instead of the normal left valve arrangement of three hinge teeth and a lateral secondary ridge creating a secondary socket, this specimen has two hinge teeth and a lateral tooth (i.e. the normal right valve arrangement). An alternative interpretation is that this specimen represents an earlier ontogenetic stage, but this is rejected because even though the specimen is slightly smaller, its lateral tooth is more prominent than in a typical left valve arrangement and both the subumbonal and lateral teeth appear to fit with the expected corresponding valve arrangement. In addition, it has been reported that in the early stages of nucinellid development the second lateral tooth is very small and closer to the subumbonal teeth than later in ontogeny (
|
||
<bibRefCitation id="2903C7AA8058E8236CA9FF348F5926C8" author="Bernard, F." box="[422,574,221,244]" pageId="8" pageNumber="858" pagination="1 - 208" refId="ref16190" refString="Bernard, F. 1898. Recherches ontog´enetiques et morphologiques sur la coquille des lamellibranches. Premi`ere partie. Taxodontes et Anisomyaires. Annales des Sciences Naturelles. Zoologie et Paleontologie´, 8, 1 - 208." type="journal article" year="1898">Bernard 1898</bibRefCitation>
|
||
), which is not the case with this specimen. Transposed hinges have been reported in a number of bivalve families, but this is the first reported occurrence in a species of
|
||
<taxonomicName id="8A92C1D88058E8236F47FED48FB32768" baseAuthorityName="Oliver & Taylor" baseAuthorityYear="2012" box="[584,724,317,340]" class="Bivalvia" family="Nucinellidae" kingdom="Animalia" order="Solemyida" pageId="8" pageNumber="858" phylum="Mollusca" rank="family">Nucinellidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4D2DBA5B8058E8236D90FE938F54278D" blockId="8.[159,781,378,433]" pageId="8" pageNumber="858">
|
||
<emphasis id="7FE666498058E8236D90FE938C5227AD" bold="true" box="[159,309,378,401]" pageId="8" pageNumber="858">Mode of life.</emphasis>
|
||
Shallow infaunal, fully motile, slow, chemosymbiotic (Oliver & Taylor 2012).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |