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<document id="2EF8E7DE8BA1148E7CCABC0EEB9A85C5" ID-DOI="10.11646/zootaxa.4759.3.12" ID-GBIF-Dataset="2909060b-18ed-4804-b76a-75be346b2b1c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3741152" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586156510046" checkinUser="plazi" docAuthor="Ahrens, Dirk &amp; Pacheco, Thaynara L." docDate="2020" docId="A22987E44B10FF86F5DCD472FAC5F89A" docLanguage="en" docName="zootaxa.4759.3.12.pdf" docOrigin="Zootaxa 4759 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maladera rugosa Ahrens &amp; Pacheco 2020, new combination" docType="treatment" docVersion="5" lastPageNumber="444" masterDocId="5E10FF9C4B10FF87F54BD275FFA8FFBE" masterDocTitle="Maladera rugosa (Blanchard, 1850) new combination-a valid species and senior synonym of Maladera graeca Petrovitz, 1969 (Coleoptera: Scarabaeidae Melolonthinae: Sericini)" masterLastPageNumber="445" masterPageNumber="443" pageNumber="443" updateTime="1698735700001" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0A7A8AFCBFCFBBEAFD588E6ED6525ACC">Maladera rugosa (Blanchard, 1850) new combination-a valid species and senior synonym of Maladera graeca Petrovitz, 1969 (Coleoptera: Scarabaeidae Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="8C83487D014346A270D560B18A00B7AF">Ahrens, Dirk</mods:namePart>
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<mods:namePart id="682FA39631401E430D90D6BBD0C44D62">Pacheco, Thaynara L.</mods:namePart>
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<mods:title id="051B28100C6AEBC8C2AD8F021B535136">Zootaxa</mods:title>
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<mods:date id="96A48DB6DC65A811C25B4BA17DB2FA24">2020</mods:date>
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<mods:number id="538C5FFC5463DB21001C127A2FFD7C31">2020-04-03</mods:number>
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<mods:detail id="2B82304171EB92209AB6DE3A48791B04" type="volume">
<mods:number id="E98A5C800AB66A9164E6AEE37DA4C7CD">4759</mods:number>
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<mods:classification id="A3C4F098D726FF95B5EA3ABB7B744EC2">journal article</mods:classification>
<mods:identifier id="A3AF7E2807DFED29EB6935C6ADEC943C" type="DOI">10.11646/zootaxa.4759.3.12</mods:identifier>
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<treatment id="A22987E44B10FF86F5DCD472FAC5F89A" ID-DOI="http://doi.org/10.5281/zenodo.3810497" ID-GBIF-Taxon="163437673" ID-Zenodo-Dep="3810497" LSID="urn:lsid:plazi:treatment:A22987E44B10FF86F5DCD472FAC5F89A" httpUri="http://treatment.plazi.org/id/A22987E44B10FF86F5DCD472FAC5F89A" lastPageId="1" lastPageNumber="444" pageId="0" pageNumber="443">
<subSubSection id="629A65794B10FF87F5DCD472FC84F99F" box="[151,812,1543,1570]" pageId="0" pageNumber="443" type="nomenclature">
<paragraph id="2A3F36F24B10FF87F5DCD472FC84F99F" blockId="0.[151,812,1543,1599]" box="[151,812,1543,1570]" pageId="0" pageNumber="443">
<heading id="7177819E4B10FF87F5DCD472FC84F99F" bold="true" box="[151,812,1543,1570]" fontSize="11" level="1" pageId="0" pageNumber="443" reason="1">
<emphasis id="18F4EAE04B10FF87F5DCD472FC84F99F" bold="true" box="[151,812,1543,1570]" pageId="0" pageNumber="443">
<taxonomicName id="ED804D714B10FF87F5DCD472FDE7F99F" authority="(Blanchard, 1850)" authorityName="Ahrens &amp; Pacheco" authorityYear="2020" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[151,591,1543,1570]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosa" status="comb. nov.">
<emphasis id="18F4EAE04B10FF87F5DCD472FECFF99F" bold="true" box="[151,359,1543,1570]" italics="true" pageId="0" pageNumber="443">Maladera rugosa</emphasis>
(
<bibRefCitation id="4E114B034B10FF87F43DD472FDEFF99F" author="Blanchard, M. E." box="[374,583,1543,1569]" pageId="0" pageNumber="443" refId="ref1507" refString="Blanchard, M. E. (1850) Catalogue de la collection Entomologique. Classes des Insectes. Ordre des Coleopteres. Premiere livraison. Gide et Baudry, Paris, 128 pp. https: // doi. org / 10.5962 / bhl. title. 66985" type="book" year="1850">Blanchard, 1850</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="03C7579B4B10FF87F71CD472FC84F99F" box="[599,812,1543,1569]" pageId="0" pageNumber="443" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="629A65794B10FF87F5DCD452FE3CF94D" pageId="0" pageNumber="443" type="reference_group">
<paragraph id="2A3F36F24B10FF87F5DCD452FD84F981" blockId="0.[151,812,1543,1599]" box="[151,556,1574,1599]" pageId="0" pageNumber="443">
<treatmentCitationGroup id="0A9011DC4B10FF87F5DCD452FD84F981" box="[151,556,1574,1599]" pageId="0" pageNumber="443">
<treatmentCitation id="AB2110E34B10FF87F5DCD452FD8FF981" author="Blanchard, M. E." box="[151,551,1574,1599]" page="77" pageId="0" pageNumber="443" year="1850">
<taxonomicName id="ED804D714B10FF87F5DCD452FD8FF981" authority="Blanchard, 1850: 77" authorityName="Blanchard" authorityPageNumber="77" authorityYear="1850" box="[151,551,1574,1599]" class="Insecta" family="Melolonthidae" genus="Omaloplia" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="18F4EAE04B10FF87F5DCD452FEF8F981" box="[151,336,1575,1599]" italics="true" pageId="0" pageNumber="443">Omaloplia rugosa</emphasis>
<bibRefCitation id="4E114B034B10FF87F41CD453FD8FF981" author="Blanchard, M. E." box="[343,551,1574,1599]" pageId="0" pageNumber="443" refId="ref1507" refString="Blanchard, M. E. (1850) Catalogue de la collection Entomologique. Classes des Insectes. Ordre des Coleopteres. Premiere livraison. Gide et Baudry, Paris, 128 pp. https: // doi. org / 10.5962 / bhl. title. 66985" type="book" year="1850">Blanchard, 1850: 77</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2A3F36F24B10FF87F5DCD41AFE3CF94D" blockId="0.[151,1437,1647,2031]" pageId="0" pageNumber="443">
<taxonomicName id="ED804D714B10FF87F5DCD41AFD89F939" authority="(Blanchard, 1850)" authorityName="Ahrens &amp; Pacheco" authorityYear="2020" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[151,545,1647,1671]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="18F4EAE04B10FF87F5DCD41AFEFBF939" box="[151,339,1647,1671]" italics="true" pageId="0" pageNumber="443">Maladera rugosa</emphasis>
(
<bibRefCitation id="4E114B034B10FF87F428D41AFDB1F939" author="Blanchard, M. E." box="[355,537,1647,1671]" pageId="0" pageNumber="443" refId="ref1507" refString="Blanchard, M. E. (1850) Catalogue de la collection Entomologique. Classes des Insectes. Ordre des Coleopteres. Premiere livraison. Gide et Baudry, Paris, 128 pp. https: // doi. org / 10.5962 / bhl. title. 66985" type="book" year="1850">Blanchard, 1850</bibRefCitation>
)
</taxonomicName>
was synonymized by
<bibRefCitation id="4E114B034B10FF87F65DD41AFC15F939" author="Brenske, E." box="[790,957,1647,1671]" pageId="0" pageNumber="443" pagination="345 - 468" refId="ref1556" refString="Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310" type="journal article" year="1897">Brenske (1897)</bibRefCitation>
with
<taxonomicName id="ED804D714B10FF87F6B5D41AFF7EF915" authority="(Faldermann, 1835)" baseAuthorityName="Faldermann" baseAuthorityYear="1835" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="punctatissima">
<emphasis id="18F4EAE04B10FF87F6B5D41AFAACF939" box="[1022,1284,1647,1671]" italics="true" pageId="0" pageNumber="443">Maladera punctatissima</emphasis>
(Faldermann, 1835)
</taxonomicName>
(senior name),
<taxonomicName id="ED804D714B10FF87F432D4E1FD20F915" authority="Kuster, 1849" authorityName="Kuster" authorityYear="1849" box="[377,648,1683,1707]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="elata">
<emphasis id="18F4EAE04B10FF87F432D4E1FE50F915" box="[377,504,1683,1707]" italics="true" pageId="0" pageNumber="443">Serica elata</emphasis>
<bibRefCitation id="4E114B034B10FF87F4B5D4E1FD20F915" author="Kuster, H. C." box="[510,648,1683,1707]" pageId="0" pageNumber="443" refId="ref1821" refString="Kuster, H. C. (1849) Die Kafer Europas nach der Natur beschrieben. Heft 18. Bauer und Raspe, Nurnberg, 2 + 100 pp. https: // doi. org / 10.5962 / bhl. title. 110110" type="book" year="1849">Küster, 1849</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ED804D714B10FF87F7DAD4E6FBABF915" authority="Reitter, 1896" authorityName="Reitter" authorityYear="1896" box="[657,1027,1683,1707]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="armeniaca">
<emphasis id="18F4EAE04B10FF87F7DAD4E6FCDAF915" box="[657,882,1683,1707]" italics="true" pageId="0" pageNumber="443">Maladera armeniaca</emphasis>
Reitter, 1896
</taxonomicName>
, and
<taxonomicName id="ED804D714B10FF87F172D4E6FF71F971" authority="(Fairmaire, 1892)" baseAuthorityName="Fairmaire" baseAuthorityYear="1892" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="fusconitens">
<emphasis id="18F4EAE04B10FF87F172D4E6FA88F915" box="[1081,1312,1683,1707]" italics="true" pageId="0" pageNumber="443">Maladera fusconitens</emphasis>
(
<bibRefCitation id="4E114B034B10FF87F067D4E6FF79F971" author="Fairmaire, M. L." pageId="0" pageNumber="443" pagination="144 - 159" refId="ref1662" refString="Fairmaire, M. L. (1892) Descriptions de Coleopteres des environs d'Akbes (Syrie). Annales de la Societe Entomologique de Belgique, 36, 144 - 159." type="journal article" year="1892">Fairmaire, 1892</bibRefCitation>
)
</taxonomicName>
. These last two were later revalidated by
<bibRefCitation id="4E114B034B10FF87F7C4D4C2FC9CF971" author="Petrovitz, R." box="[655,820,1719,1743]" pageId="0" pageNumber="443" pagination="383 - 400" refId="ref2016" refString="Petrovitz, R. (1969) Ergebnisse zoologischer Sammelreisen in der Tu ¨ rkei: Lamellicornia, Coleoptera. Dritte Folge. Die mediter- ranen Arten der Gattung Maladera Mulsant. Lamellicornia, Coleoptera. Annalen des Naturhistorischen Museums in Wien, 73, 383 - 400." type="journal article" year="1969">Petrovitz (1969</bibRefCitation>
and
<bibRefCitation id="4E114B034B10FF87F62CD4C2FC37F971" author="Petrovitz, R." box="[871,927,1719,1743]" pageId="0" pageNumber="443" pagination="127 - 131" refId="ref1989" refString="Petrovitz, R. (1964) Neue Melolonthidae und Dynastidae aus Europa und Asien. Reichenbachia, 3 (9), 127 - 131." type="journal article" year="1964">1964</bibRefCitation>
, respectively). Since then,
<taxonomicName id="ED804D714B10FF87F1F7D4CDFA81F971" authorityName="Ahrens &amp; Pacheco" authorityYear="2020" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[1212,1321,1720,1743]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="18F4EAE04B10FF87F1F7D4CDFA81F971" box="[1212,1321,1720,1743]" italics="true" pageId="0" pageNumber="443">M. rugosa</emphasis>
</taxonomicName>
status was never questioned again.
</paragraph>
</subSubSection>
<subSubSection id="629A65794B10FF87F58CD48AFF54F851" pageId="0" pageNumber="443" type="materials_examined">
<paragraph id="2A3F36F24B10FF87F58CD48AFCF3F83D" blockId="0.[151,1437,1647,2031]" pageId="0" pageNumber="443">
<emphasis id="18F4EAE04B10FF87F58CD48AFE48F8A9" bold="true" box="[199,480,1791,1815]" pageId="0" pageNumber="443">Type material examined.</emphasis>
<taxonomicName id="ED804D714B10FF87F4ADD48AFD00F8A9" authorityName="Blanchard" authorityYear="1850" box="[486,680,1791,1815]" class="Insecta" family="Melolonthidae" genus="Omaloplia" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="18F4EAE04B10FF87F4ADD48AFD00F8A9" box="[486,680,1791,1815]" italics="true" pageId="0" pageNumber="443">Omaloplia rugosa</emphasis>
</taxonomicName>
,
<materialsCitation id="9AE83CAF4B10FF87F7FED48AFBA4F885" ID-GBIF-Occurrence="2595809699" collectionCode="MNHN" county="Moree" location="Moree" municipality="Brulle Moree" pageId="0" pageNumber="443" specimenCount="1" specimenCount-male="1" stateProvince="Mus" typeStatus="lectotype">
<typeStatus id="F53B88504B10FF87F7FED48AFCB2F8A9" box="[693,794,1791,1815]" pageId="0" pageNumber="443" type="lectotype">lectotype</typeStatus>
,
<specimenCount id="3C86FD7B4B10FF87F668D575FC91F8A9" box="[803,825,1792,1815]" pageId="0" pageNumber="443" type="male">♂</specimenCount>
, present designation: “
<taxonomicName id="ED804D714B10FF87F160D48AFB6DF8A9" authority="Cat." authorityName="Cat." box="[1067,1221,1791,1816]" class="Insecta" family="Melolonthidae" genus="Omaloplia" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosa">O. rugosa Cat.</taxonomicName>
<collectingRegion id="E844F8104B10FF87F181D575FB55F8A9" box="[1226,1277,1792,1815]" country="Turkey" name="Mus" pageId="0" pageNumber="443">Mus</collectingRegion>
.
<collectingCounty id="C35E4E7E4B10FF87F04DD48AFAE3F8A9" box="[1286,1355,1791,1815]" pageId="0" pageNumber="443">Moreé</collectingCounty>
/
<collectingMunicipality id="CA5BAC884B10FF87F011D48AFF74F885" pageId="0" pageNumber="443">Brulle Moreé</collectingMunicipality>
/
<taxonomicName id="ED804D714B10FF87F5A1D556FE2BF885" authority="Cat." authorityName="Cat." box="[234,387,1827,1852]" class="Insecta" family="Melolonthidae" genus="Omaloplia" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosa">O. rugosa Cat.</taxonomicName>
<collectingRegion id="E844F8104B10FF87F4C3D551FE13F885" box="[392,443,1828,1851]" country="Turkey" name="Mus" pageId="0" pageNumber="443">Mus</collectingRegion>
.
<location id="2F5F60294B10FF87F48FD556FDA1F885" LSID="urn:lsid:plazi:treatment:A22987E44B10FF86F5DCD472FAC5F89A:2F5F60294B10FF87F48FD556FDA1F885" box="[452,521,1827,1851]" county="Moree" municipality="Brulle Moree" name="Moree" pageId="0" pageNumber="443" stateProvince="Mus">Moreé</location>
/
<location id="2F5F60294B10FF87F75DD551FC9EF885" LSID="urn:lsid:plazi:treatment:A22987E44B10FF86F5DCD472FAC5F89A:2F5F60294B10FF87F75DD551FC9EF885" box="[534,822,1827,1851]" county="Moree" municipality="Brulle Moree" name="Musem Paris Moree Brulle" pageId="0" pageNumber="443" stateProvince="Mus">Musem Paris Moreé Brullé</location>
4187-33” (
<collectionCode id="4C91AE374B10FF87F6E6D556FBADF885" box="[941,1029,1827,1851]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="443">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="9AE83CAF4B10FF87F15ED556FCFFF83D" ID-GBIF-Occurrence="2595809698" collectionCode="MNHN" county="Paris Moree Marloy" location="Moree" municipality="Marloy Moree" pageId="0" pageNumber="443" specimenCount="1" specimenCount-female="1" stateProvince="Mus" typeStatus="paralectotype">
<typeStatus id="F53B88504B10FF87F15ED556FB01F885" box="[1045,1193,1827,1851]" pageId="0" pageNumber="443" type="paralectotype">Paralectotype</typeStatus>
,
<specimenCount id="3C86FD7B4B10FF87F1F8D551FB61F885" box="[1203,1225,1828,1851]" pageId="0" pageNumber="443" type="female">♀</specimenCount>
: “
<collectingRegion id="E844F8104B10FF87F195D556FABAF885" box="[1246,1298,1827,1851]" country="Turkey" name="Mus" pageId="0" pageNumber="443">Mus</collectingRegion>
.
<collectingCounty id="C35E4E7E4B10FF87F050D556FF4EF8E1" pageId="0" pageNumber="443">Paris Moreé Marloy</collectingCounty>
1833/
<collectingMunicipality id="CA5BAC884B10FF87F465D532FE6FF8E1" box="[302,455,1863,1887]" pageId="0" pageNumber="443">Marloy Moree</collectingMunicipality>
/.
<taxonomicName id="ED804D714B10FF87F4AAD53DFDF2F8E1" authority="Cat." authorityName="Cat." box="[481,602,1863,1888]" class="Insecta" family="Melolonthidae" genus="Omaloplia" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosa">rugosa Cat.</taxonomicName>
<collectingRegion id="E844F8104B10FF87F715D53DFD39F8E1" box="[606,657,1864,1887]" country="Turkey" name="Mus" pageId="0" pageNumber="443">Mus</collectingRegion>
.
<location id="2F5F60294B10FF87F7D1D532FD77F8E1" LSID="urn:lsid:plazi:treatment:A22987E44B10FF86F5DCD472FAC5F89A:2F5F60294B10FF87F7D1D532FD77F8E1" box="[666,735,1863,1887]" county="Paris Moree Marloy" municipality="Marloy Moree" name="Moree" pageId="0" pageNumber="443" stateProvince="Mus">Moreé</location>
/
<taxonomicName id="ED804D714B10FF87F7A6D53DFCF2F8E1" authority="Bl." authorityName="Blanchard" authorityYear="1850" box="[749,858,1863,1888]" class="Insecta" family="Melolonthidae" genus="Omaloplia" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosa">rugosa Bl.</taxonomicName>
<taxonomicName id="ED804D714B10FF87F614D532FB86F8E1" authority="Fald." authorityName="Fald." box="[863,1070,1863,1887]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="443" phylum="Arthropoda" rank="species" species="punctatissima">punctatissima Fald.</taxonomicName>
(hand-written
<location id="2F5F60294B10FF87F182D532FA8CF8E1" LSID="urn:lsid:plazi:treatment:A22987E44B10FF86F5DCD472FAC5F89A:2F5F60294B10FF87F182D532FA8CF8E1" box="[1225,1316,1863,1887]" county="Paris Moree Marloy" municipality="Marloy Moree" name="Brenske" pageId="0" pageNumber="443" stateProvince="Mus">Brenske</location>
)” (
<collectionCode id="4C91AE374B10FF87F009D532FA30F8E1" box="[1346,1432,1863,1887]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="443">MNHN</collectionCode>
, specimen interiorly damaged by dermestids, antenna missing too)
</materialsCitation>
.
</paragraph>
<paragraph id="2A3F36F24B10FF87F58CD5FAFF54F851" blockId="0.[151,1437,1647,2031]" pageId="0" pageNumber="443">
<emphasis id="18F4EAE04B10FF87F58CD5FAFD8BF819" bold="true" box="[199,547,1935,1959]" pageId="0" pageNumber="443">Additional material examined.</emphasis>
<materialsCitation id="9AE83CAF4B10FF87F761D5FAFE15F875" ID-GBIF-Occurrence="2595809696" collectingDate="2005-07-04" collectionCode="ZFMK" collectorName="Ahrens" country="Greece" location="Taygetos Mts." pageId="0" pageNumber="443" specimenCount="1" specimenCount-female="1">
<specimenCount id="3C86FD7B4B10FF87F761D5FAFDFAF819" box="[554,594,1935,1959]" pageId="0" pageNumber="443" type="female">1 ♀</specimenCount>
“678409/
<collectingCountry id="529776624B10FF87F788D5FAFCA6F819" box="[707,782,1935,1959]" name="Greece" pageId="0" pageNumber="443">Greece</collectingCountry>
S-Peleponnes [sic],
<location id="2F5F60294B10FF87F6A1D5E5FB2CF819" LSID="urn:lsid:plazi:treatment:A22987E44B10FF86F5DCD472FAC5F89A:2F5F60294B10FF87F6A1D5E5FB2CF819" box="[1002,1156,1936,1960]" country="Greece" name="Taygetos Mts." pageId="0" pageNumber="443">Taygetos Mts.</location>
, Valley N Krioneri
<date id="5E3E10324B10FF87F01ED5FAFF64F875" pageId="0" pageNumber="443" value="2005-07-04">
<collectingDate id="4E7AE9DA4B10FF87F01ED5FAFF64F875" pageId="0" pageNumber="443" value="2005-07-04">4. July 2005</collectingDate>
</date>
leg.
<collectorName id="877553244B10FF87F44BD5C6FEE7F875" box="[256,335,1971,1995]" pageId="0" pageNumber="443">Ahrens</collectorName>
” (
<collectionCode id="4C91AE374B10FF87F42CD5C6FE1FF875" box="[359,439,1971,1995]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="0" pageNumber="443" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="9AE83CAF4B10FF87F482D5C6FF50F851" ID-GBIF-Occurrence="2595809697" collectionCode="CWAE" collectorName="Apfel" country="Greece" elevation="600" pageId="0" pageNumber="443" specimenCount="1" specimenCount-female="1">
<specimenCount id="3C86FD7B4B10FF87F482D5C6FE59F875" box="[457,497,1971,1995]" pageId="0" pageNumber="443" type="female">1 ♀</specimenCount>
“West-Peloponnes, Eichenwald bei Agia Triada, östl. von Amaliada,
<quantity id="ED789B174B10FF87F19DD5C6FAB4F875" box="[1238,1308,1971,1995]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="0" pageNumber="443" unit="m" value="600.0">
<elevation id="A1ADD1C14B10FF87F19DD5C6FAB4F875" box="[1238,1308,1971,1995]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="0" pageNumber="443" unit="m" value="600.0">600 m</elevation>
</quantity>
, leg.
<collectorName id="877553244B10FF87F01FD5C6FA3BF875" box="[1364,1427,1971,1995]" pageId="0" pageNumber="443">Apfel</collectorName>
” (
<collectionCode id="4C91AE374B10FF87F5D5D5A2FF58F851" box="[158,240,2007,2031]" pageId="0" pageNumber="443">CWAE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="7EFF667A4B11FF86F5DCD1A5FE87FB8F" ID-DOI="http://doi.org/10.5281/zenodo.3741154" ID-Zenodo-Dep="3741154" httpUri="https://zenodo.org/record/3741154/files/figure.png" pageId="1" pageNumber="444" startId="1.[151,250,976,1001]" targetBox="[158,1430,187,945]" targetPageId="1">
<paragraph id="2A3F36F24B11FF86F5DCD1A5FE87FB8F" blockId="1.[151,1436,976,1073]" pageId="1" pageNumber="444">
<emphasis id="18F4EAE04B11FF86F5DCD1A5FEBBFC57" bold="true" box="[151,275,976,1001]" pageId="1" pageNumber="444">FIGURE 1.</emphasis>
Lectotype of
<taxonomicName id="ED804D714B11FF86F4D5D1A4FCA6FC57" authority="(Blanchard, 1850)" authorityName="Ahrens &amp; Pacheco" authorityYear="2020" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[414,782,976,1001]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="rugosa" status="comb. nov.">
<emphasis id="18F4EAE04B11FF86F4D5D1A4FDE6FC57" box="[414,590,977,1001]" italics="true" pageId="1" pageNumber="444">Maladera rugosa</emphasis>
(
<bibRefCitation id="4E114B034B11FF86F710D1A5FCAFFC57" author="Blanchard, M. E." box="[603,775,976,1001]" pageId="1" pageNumber="444" refId="ref1507" refString="Blanchard, M. E. (1850) Catalogue de la collection Entomologique. Classes des Insectes. Ordre des Coleopteres. Premiere livraison. Gide et Baudry, Paris, 128 pp. https: // doi. org / 10.5962 / bhl. title. 66985" type="book" year="1850">Blanchard, 1850</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="03C7579B4B11FF86F653D1A4FC62FC57" box="[792,970,976,1001]" pageId="1" pageNumber="444" rank="species">new combination</taxonomicNameLabel>
. A, aedeagus, left side in lateral view; B, parameres, dorsal view; C, aedeagus, right side in lateral view (scale = 0.5 mm); D, habitus, dorsal view (not to scale); E, associated labels; F, head.
</paragraph>
</caption>
<subSubSection id="629A65794B11FF86F5DCD614FAC5F89A" pageId="1" pageNumber="444" type="discussion">
<paragraph id="2A3F36F24B11FF86F5DCD614FE07FB23" blockId="1.[151,1437,1121,1829]" pageId="1" pageNumber="444">
<emphasis id="18F4EAE04B11FF86F5DCD614FEAEFBC7" bold="true" box="[151,262,1121,1145]" pageId="1" pageNumber="444">Remarks.</emphasis>
The
<typeStatus id="F53B88504B11FF86F476D614FE65FBC7" box="[317,461,1121,1145]" pageId="1" pageNumber="444" type="paralectotype">paralectotype</typeStatus>
(female) specimen is distinctly larger, browner and has a convexly elevated clypeus with a rugosely punctate surface.
</paragraph>
<paragraph id="2A3F36F24B11FF86F58CD6DCFE6EFA27" blockId="1.[151,1437,1121,1829]" pageId="1" pageNumber="444">
<taxonomicName id="ED804D714B11FF86F58CD6DCFDE5FB7F" authority="(Blanchard, 1850)" authorityName="Ahrens &amp; Pacheco" authorityYear="2020" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[199,589,1193,1217]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="rugosa" status="comb. nov.">
<emphasis id="18F4EAE04B11FF86F58CD6DCFE2AFB7F" box="[199,386,1193,1217]" italics="true" pageId="1" pageNumber="444">Maladera rugosa</emphasis>
(
<bibRefCitation id="4E114B034B11FF86F4DAD6DCFDEDFB7F" author="Blanchard, M. E." box="[401,581,1193,1217]" pageId="1" pageNumber="444" refId="ref1507" refString="Blanchard, M. E. (1850) Catalogue de la collection Entomologique. Classes des Insectes. Ordre des Coleopteres. Premiere livraison. Gide et Baudry, Paris, 128 pp. https: // doi. org / 10.5962 / bhl. title. 66985" type="book" year="1850">Blanchard, 1850</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="03C7579B4B11FF86F71FD6DCFCA5FB7F" box="[596,781,1193,1217]" pageId="1" pageNumber="444" rank="species">new combination</taxonomicNameLabel>
was originally described from
<collectingCountry id="529776624B11FF86F115D6DCFB01FB7F" box="[1118,1193,1193,1217]" name="Greece" pageId="1" pageNumber="444">Greece</collectingCountry>
(Moreé,
<emphasis id="18F4EAE04B11FF86F046D6DFFA85FB7F" box="[1293,1325,1194,1217]" italics="true" pageId="1" pageNumber="444">i.e.</emphasis>
,
<collectingRegion id="E844F8104B11FF86F077D6DCFF62FB5B" country="Greece" name="Peloponnisos" pageId="1" pageNumber="444">Peloponnese</collectingRegion>
), as well as
<taxonomicName id="ED804D714B11FF86F403D6BBFDC9FB5B" authority="Petrovitz, 1969" authorityName="Petrovitz" authorityYear="1969" box="[328,609,1229,1253]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="18F4EAE04B11FF86F403D6BBFE1DFB5B" box="[328,437,1230,1253]" italics="true" pageId="1" pageNumber="444">M. graeca</emphasis>
<bibRefCitation id="4E114B034B11FF86F4F0D6B8FDC9FB5B" author="Petrovitz, R." box="[443,609,1229,1253]" pageId="1" pageNumber="444" pagination="383 - 400" refId="ref2016" refString="Petrovitz, R. (1969) Ergebnisse zoologischer Sammelreisen in der Tu ¨ rkei: Lamellicornia, Coleoptera. Dritte Folge. Die mediter- ranen Arten der Gattung Maladera Mulsant. Lamellicornia, Coleoptera. Annalen des Naturhistorischen Museums in Wien, 73, 383 - 400." type="journal article" year="1969">Petrovitz, 1969</bibRefCitation>
</taxonomicName>
. The synonymy of
<taxonomicName id="ED804D714B11FF86F661D6B8FB03FB5B" authority="(Blanchard, 1850)" authorityName="Ahrens &amp; Pacheco" authorityYear="2020" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[810,1195,1229,1253]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="18F4EAE04B11FF86F661D6B8FC4BFB5B" box="[810,995,1229,1253]" italics="true" pageId="1" pageNumber="444">Maladera rugosa</emphasis>
(
<bibRefCitation id="4E114B034B11FF86F6BBD6B8FB0BFB5B" author="Blanchard, M. E." box="[1008,1187,1229,1253]" pageId="1" pageNumber="444" refId="ref1507" refString="Blanchard, M. E. (1850) Catalogue de la collection Entomologique. Classes des Insectes. Ordre des Coleopteres. Premiere livraison. Gide et Baudry, Paris, 128 pp. https: // doi. org / 10.5962 / bhl. title. 66985" type="book" year="1850">Blanchard, 1850</bibRefCitation>
)
</taxonomicName>
with
<taxonomicName id="ED804D714B11FF86F1AED6BBFEC4FAB7" authority="(Faldermann, 1835)" baseAuthorityName="Faldermann" baseAuthorityYear="1835" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="punctatissima">
<emphasis id="18F4EAE04B11FF86F1AED6BBFA34FB5B" box="[1253,1436,1230,1253]" italics="true" pageId="1" pageNumber="444">M. punctatissima</emphasis>
(Faldermann, 1835)
</taxonomicName>
(
<bibRefCitation id="4E114B034B11FF86F430D684FDBCFAB7" author="Brenske, E." box="[379,532,1265,1289]" pageId="1" pageNumber="444" pagination="345 - 468" refId="ref1556" refString="Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310" type="journal article" year="1897">Brenske 1897</bibRefCitation>
;
<bibRefCitation id="4E114B034B11FF86F76BD684FD68FAB7" author="Petrovitz, R." box="[544,704,1265,1289]" pageId="1" pageNumber="444" pagination="383 - 400" refId="ref2016" refString="Petrovitz, R. (1969) Ergebnisse zoologischer Sammelreisen in der Tu ¨ rkei: Lamellicornia, Coleoptera. Dritte Folge. Die mediter- ranen Arten der Gattung Maladera Mulsant. Lamellicornia, Coleoptera. Annalen des Naturhistorischen Museums in Wien, 73, 383 - 400." type="journal article" year="1969">Petrovitz 1969</bibRefCitation>
;
<bibRefCitation id="4E114B034B11FF86F780D684FCFFFAB7" author="Ahrens, D." box="[715,855,1265,1289]" pageId="1" pageNumber="444" pagination="229 - 248" refId="ref1347" refString="Ahrens, D. (2006) Sericinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea- Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248." type="book chapter" year="2006">Ahrens 2006</bibRefCitation>
, 2007;
<bibRefCitation id="4E114B034B11FF86F6EDD684FB0EFAB7" author="Ahrens, D. &amp; Bezdek, A." box="[934,1190,1265,1289]" pageId="1" pageNumber="444" pagination="281 - 317" refId="ref1395" refString="Ahrens, D. &amp; Bezdek, A. (2016) Sericini. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Revised and Updated Edition. Brill, Leiden, Boston, pp. 281 - 317." type="book chapter" year="2016">Ahrens &amp; Bezděk 2016</bibRefCitation>
) was never doubted or ever properly checked while
<bibRefCitation id="4E114B034B11FF86F484D760FDD6FA93" author="Petrovitz, R." box="[463,638,1301,1325]" pageId="1" pageNumber="444" pagination="383 - 400" refId="ref2016" refString="Petrovitz, R. (1969) Ergebnisse zoologischer Sammelreisen in der Tu ¨ rkei: Lamellicornia, Coleoptera. Dritte Folge. Die mediter- ranen Arten der Gattung Maladera Mulsant. Lamellicornia, Coleoptera. Annalen des Naturhistorischen Museums in Wien, 73, 383 - 400." type="journal article" year="1969">Petrovitz (1969)</bibRefCitation>
erected a number of new and closely related species from
<collectingCountry id="529776624B11FF86F1BCD760FAEDFA93" box="[1271,1349,1301,1325]" name="Greece" pageId="1" pageNumber="444">Greece</collectingCountry>
,
<collectingCountry id="529776624B11FF86F01BD760FA34FA93" box="[1360,1436,1301,1325]" name="Turkey" pageId="1" pageNumber="444">Turkey</collectingCountry>
and Near East. Brenske obviously has seen at least one of the
<typeStatus id="F53B88504B11FF86F66CD74CFC2CFAEF" box="[807,900,1337,1361]" pageId="1" pageNumber="444" type="syntype">syntypes</typeStatus>
(the specimen bears a label with his handwriting), but he did not examine the genitalia (his specimen is very likely a female that was strongly damaged by dermestids) of the intact
<typeStatus id="F53B88504B11FF86F44BD7F4FEFBFA27" box="[256,339,1409,1433]" pageId="1" pageNumber="444" type="syntype">syntype</typeStatus>
specimen.
</paragraph>
<paragraph id="2A3F36F24B11FF86F58CD7D0FED7F9F2" blockId="1.[151,1437,1121,1829]" pageId="1" pageNumber="444">
Male genitalia of the
<typeStatus id="F53B88504B11FF86F4E3D7D0FDA2FA03" box="[424,522,1445,1469]" pageId="1" pageNumber="444" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="ED804D714B11FF86F767D7D0FC06FA03" authority="(Blanchard, 1850)" authorityName="Ahrens &amp; Pacheco" authorityYear="2020" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[556,942,1445,1469]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="18F4EAE04B11FF86F767D7D0FD4DFA03" box="[556,741,1445,1469]" italics="true" pageId="1" pageNumber="444">Maladera rugosa</emphasis>
(
<bibRefCitation id="4E114B034B11FF86F7B9D7D0FC0FFA03" author="Blanchard, M. E." box="[754,935,1445,1469]" pageId="1" pageNumber="444" refId="ref1507" refString="Blanchard, M. E. (1850) Catalogue de la collection Entomologique. Classes des Insectes. Ordre des Coleopteres. Premiere livraison. Gide et Baudry, Paris, 128 pp. https: // doi. org / 10.5962 / bhl. title. 66985" type="book" year="1850">Blanchard, 1850</bibRefCitation>
)
</taxonomicName>
, which have been compared with the drawings of the original publication of
<taxonomicName id="ED804D714B11FF86F49AD7BFFD97FA5F" authorityName="Petrovitz" authorityYear="1969" box="[465,575,1482,1505]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="18F4EAE04B11FF86F49AD7BFFD97FA5F" box="[465,575,1482,1505]" italics="true" pageId="1" pageNumber="444">M. graeca</emphasis>
</taxonomicName>
, result to be virtually identical to with the latter, thus,
<taxonomicName id="ED804D714B11FF86F1CFD7BFFA34FA5F" authority="Petrovitz, 1969" authorityName="Petrovitz" authorityYear="1969" box="[1156,1436,1481,1505]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="18F4EAE04B11FF86F1CFD7BFFB5AFA5F" box="[1156,1266,1482,1505]" italics="true" pageId="1" pageNumber="444">M. graeca</emphasis>
<bibRefCitation id="4E114B034B11FF86F1B2D7BCFA34FA5F" author="Petrovitz, R." box="[1273,1436,1481,1505]" pageId="1" pageNumber="444" pagination="383 - 400" refId="ref2016" refString="Petrovitz, R. (1969) Ergebnisse zoologischer Sammelreisen in der Tu ¨ rkei: Lamellicornia, Coleoptera. Dritte Folge. Die mediter- ranen Arten der Gattung Maladera Mulsant. Lamellicornia, Coleoptera. Annalen des Naturhistorischen Museums in Wien, 73, 383 - 400." type="journal article" year="1969">Petrovitz, 1969</bibRefCitation>
</taxonomicName>
has to be considered a junior synonym of
<taxonomicName id="ED804D714B11FF86F705D798FC79F9BB" authority="(Blanchard, 1850)" authorityName="Ahrens &amp; Pacheco" authorityYear="2020" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[590,977,1517,1541]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="18F4EAE04B11FF86F705D798FCAEF9BB" box="[590,774,1517,1541]" italics="true" pageId="1" pageNumber="444">Maladera rugosa</emphasis>
(
<bibRefCitation id="4E114B034B11FF86F65FD798FC61F9BB" author="Blanchard, M. E." box="[788,969,1517,1541]" pageId="1" pageNumber="444" refId="ref1507" refString="Blanchard, M. E. (1850) Catalogue de la collection Entomologique. Classes des Insectes. Ordre des Coleopteres. Premiere livraison. Gide et Baudry, Paris, 128 pp. https: // doi. org / 10.5962 / bhl. title. 66985" type="book" year="1850">Blanchard, 1850</bibRefCitation>
)
</taxonomicName>
. The nearly sympatrical
<taxonomicName id="ED804D714B11FF86F198D798FE96F997" authority="Petrovitz, 1969" authorityName="Petrovitz" authorityYear="1969" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="krueperi">
<emphasis id="18F4EAE04B11FF86F198D798FA34F9BB" box="[1235,1436,1517,1541]" italics="true" pageId="1" pageNumber="444">Maladera krueperi</emphasis>
<bibRefCitation id="4E114B034B11FF86F5DCD464FE96F997" author="Petrovitz, R." box="[151,318,1553,1577]" pageId="1" pageNumber="444" pagination="383 - 400" refId="ref2016" refString="Petrovitz, R. (1969) Ergebnisse zoologischer Sammelreisen in der Tu ¨ rkei: Lamellicornia, Coleoptera. Dritte Folge. Die mediter- ranen Arten der Gattung Maladera Mulsant. Lamellicornia, Coleoptera. Annalen des Naturhistorischen Museums in Wien, 73, 383 - 400." type="journal article" year="1969">Petrovitz, 1969</bibRefCitation>
</taxonomicName>
, also described from
<collectingCountry id="529776624B11FF86F76FD464FDC7F997" box="[548,623,1553,1577]" name="Greece" pageId="1" pageNumber="444">Greece</collectingCountry>
(
<collectingRegion id="E844F8104B11FF86F735D464FCA3F997" box="[638,779,1553,1577]" country="Greece" name="Peloponnisos" pageId="1" pageNumber="444">Peloponnese</collectingRegion>
), seems to be well distinct by the strongly arched dorsal lobe of the right paramere.
</paragraph>
<paragraph id="2A3F36F24B11FF86F58CD42CFAC5F89A" blockId="1.[151,1437,1121,1829]" pageId="1" pageNumber="444">
Given the quite striking morphological similarity of
<taxonomicName id="ED804D714B11FF86F7BCD42CFBD2F9CF" authority="(Blanchard, 1850)" authorityName="Ahrens &amp; Pacheco" authorityYear="2020" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[759,1146,1625,1649]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="18F4EAE04B11FF86F7BCD42CFC19F9CF" box="[759,945,1625,1649]" italics="true" pageId="1" pageNumber="444">Maladera rugosa</emphasis>
(
<bibRefCitation id="4E114B034B11FF86F6F4D42CFBDAF9CF" author="Blanchard, M. E." box="[959,1138,1625,1649]" pageId="1" pageNumber="444" refId="ref1507" refString="Blanchard, M. E. (1850) Catalogue de la collection Entomologique. Classes des Insectes. Ordre des Coleopteres. Premiere livraison. Gide et Baudry, Paris, 128 pp. https: // doi. org / 10.5962 / bhl. title. 66985" type="book" year="1850">Blanchard, 1850</bibRefCitation>
)
</taxonomicName>
with other species, such as
<taxonomicName id="ED804D714B11FF86F5DCD40BFE7AF92B" authority="Petrovitz, 1969" authorityName="Petrovitz" authorityYear="1969" box="[151,466,1661,1685]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="apfelbecki">
<emphasis id="18F4EAE04B11FF86F5DCD40BFE81F92B" box="[151,297,1661,1685]" italics="true" pageId="1" pageNumber="444">M. apfelbecki</emphasis>
<bibRefCitation id="4E114B034B11FF86F464D408FE7AF92B" author="Petrovitz, R." box="[303,466,1661,1685]" pageId="1" pageNumber="444" pagination="383 - 400" refId="ref2016" refString="Petrovitz, R. (1969) Ergebnisse zoologischer Sammelreisen in der Tu ¨ rkei: Lamellicornia, Coleoptera. Dritte Folge. Die mediter- ranen Arten der Gattung Maladera Mulsant. Lamellicornia, Coleoptera. Annalen des Naturhistorischen Museums in Wien, 73, 383 - 400." type="journal article" year="1969">Petrovitz, 1969</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="529776624B11FF86F759D408FDCEF92B" box="[530,614,1661,1685]" name="Albania" pageId="1" pageNumber="444">Albania</collectingCountry>
and
<taxonomicName id="ED804D714B11FF86F7D1D40BFC11F92B" authority="Petrovitz, 1969" authorityName="Petrovitz" authorityYear="1969" box="[666,953,1661,1685]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="villiersi">
<emphasis id="18F4EAE04B11FF86F7D1D40BFCA7F92B" box="[666,783,1661,1685]" italics="true" pageId="1" pageNumber="444">M. villiersi</emphasis>
<bibRefCitation id="4E114B034B11FF86F65DD408FC11F92B" author="Petrovitz, R." box="[790,953,1661,1685]" pageId="1" pageNumber="444" pagination="383 - 400" refId="ref2016" refString="Petrovitz, R. (1969) Ergebnisse zoologischer Sammelreisen in der Tu ¨ rkei: Lamellicornia, Coleoptera. Dritte Folge. Die mediter- ranen Arten der Gattung Maladera Mulsant. Lamellicornia, Coleoptera. Annalen des Naturhistorischen Museums in Wien, 73, 383 - 400." type="journal article" year="1969">Petrovitz, 1969</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="529776624B11FF86F6B0D408FB9EF92B" box="[1019,1078,1661,1685]" name="Syria" pageId="1" pageNumber="444">Syria</collectingCountry>
, the necessity to clarify the status of
<taxonomicName id="ED804D714B11FF86F5FDD4D4FE62F907" authority="Kuster, 1849" authorityName="Kuster" authorityYear="1849" box="[182,458,1697,1721]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="elata">
<emphasis id="18F4EAE04B11FF86F5FDD4D4FE9FF907" box="[182,311,1697,1721]" italics="true" pageId="1" pageNumber="444">Serica elata</emphasis>
<bibRefCitation id="4E114B034B11FF86F40BD4D4FE62F907" author="Kuster, H. C." box="[320,458,1697,1721]" pageId="1" pageNumber="444" refId="ref1821" refString="Kuster, H. C. (1849) Die Kafer Europas nach der Natur beschrieben. Heft 18. Bauer und Raspe, Nurnberg, 2 + 100 pp. https: // doi. org / 10.5962 / bhl. title. 110110" type="book" year="1849">Küster, 1849</bibRefCitation>
</taxonomicName>
(since
<bibRefCitation id="4E114B034B11FF86F750D4D4FD6AF907" author="Brenske, E." box="[539,706,1697,1721]" pageId="1" pageNumber="444" pagination="345 - 468" refId="ref1556" refString="Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310" type="journal article" year="1897">Brenske (1897)</bibRefCitation>
a junior synonym of
<taxonomicName id="ED804D714B11FF86F6FBD4D7FBC1F907" baseAuthorityName="Faldermann" baseAuthorityYear="1835" box="[944,1129,1698,1721]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="punctatissima">
<emphasis id="18F4EAE04B11FF86F6FBD4D7FBC1F907" box="[944,1129,1698,1721]" italics="true" pageId="1" pageNumber="444">M. punctatissima</emphasis>
</taxonomicName>
but unlikely to be synonym of the latter, since it was described from
<collectingCountry id="529776624B11FF86F700D4B0FD7AF963" box="[587,722,1733,1757]" name="Serbia" pageId="1" pageNumber="444">Montenegro</collectingCountry>
), and given a quite well geographically pronounced morphological variation in all species of the subgenus
<taxonomicName id="ED804D714B11FF86F709D49FFC2EF8BF" authority="Medvedev, 1952" authorityName="Medvedev" authorityYear="1952" box="[578,902,1769,1793]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="subGenus" subGenus="Macroserica">
<emphasis id="18F4EAE04B11FF86F709D49FFD61F8BF" box="[578,713,1770,1793]" italics="true" pageId="1" pageNumber="444">Macroserica</emphasis>
Medvedev, 1952
</taxonomicName>
, there is evident a great need of a comprehensive study of the geographical variation of the populations and species in a context of a subsequent taxonomic treatment.
</paragraph>
</subSubSection>
</treatment>
</document>
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