365 lines
39 KiB
XML
365 lines
39 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e83035" ID-Pensoft-Pub="1864-8312-80-439" ID-Pensoft-UUID="3F4E21B524465471831D09738C2304A6" ID-ZooBank="62FFB94CEBF441639F22881435EFC37C" ModsDocID="1864-8312-80-439" checkinTime="1663021027595" checkinUser="pensoft" docAuthor="Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha" docDate="2022" docId="9AA3C8A3056751F0A754D36347DE9153" docLanguage="en" docName="ArthropodSystPhyl 80: 439-494" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-09-12" docSource="http://dx.doi.org/10.3897/asp.80.e83035" docTitle="Amphineurus Skuse 1890" docType="treatment" docVersion="1" id="3F4E21B524465471831D09738C2304A6" lastPageNumber="439" masterDocId="3F4E21B524465471831D09738C2304A6" masterDocTitle="Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)" masterLastPageNumber="494" masterPageNumber="439" pageNumber="439" updateTime="1663021027595" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Santos, Daubian</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1220-1267</mods:nameIdentifier>
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<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">daubians@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Santos, Rodrigo dos Reis</mods:namePart>
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<mods:affiliation>Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ribeiro, Guilherme Cunha</mods:namePart>
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<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>439</mods:start>
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<mods:end>494</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e83035</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e83035</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-439</mods:identifier>
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<mods:identifier type="ZooBank">62FFB94CEBF441639F22881435EFC37C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">3F4E21B524465471831D09738C2304A6</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:9AA3C8A3056751F0A754D36347DE9153" httpUri="http://treatment.plazi.org/id/9AA3C8A3056751F0A754D36347DE9153" lastPageNumber="439" pageId="0" pageNumber="439">
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<subSubSection pageId="0" pageNumber="439" type="nomenclature">
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<paragraph pageId="0" pageNumber="439">
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3.1. Genus
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<taxonomicName LSID="9AA3C8A3-0567-51F0-A754-D36347DE9153" authority="Skuse, 1890" authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">Amphineurus Skuse, 1890</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="439" type="reference_group">
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<paragraph pageId="0" pageNumber="439">
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<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">Amphineurus</taxonomicName>
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Skuse, 1890: 800 (as a subgenus of
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<taxonomicName class="Insecta" family="Limoniidae" genus="Rhypholophus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhypholophus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">Rhypholophus</taxonomicName>
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). Type-species:
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<taxonomicName baseAuthorityName="Skuse" baseAuthorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (A.) umbraticus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="umbraticus" subGenus="A.">Amphineurus (A.) umbraticus</taxonomicName>
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Skuse, 1890 (as
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<taxonomicName class="Insecta" family="Limoniidae" genus="Rhypholophus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhypholophus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">Rhypholophus</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="439" type="diagnosis">
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<paragraph pageId="0" pageNumber="439">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="439">
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<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
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</taxonomicName>
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can be diagnosed by the following combination of characters: gonocoxite bifid; wing with R2+3+4 aligned with R2+3. When there is an element R4+5 (and therefore, no R2+3), members of the genus are distinguished from other
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<taxonomicName lsidName="" pageId="0" pageNumber="439" rank="subFamily" subFamily="Chioneinae">Chioneinae</taxonomicName>
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by the closed discal cell.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="439" type="remarks">
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<paragraph pageId="0" pageNumber="439">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="439">
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There is great diversity in the male terminalia of
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<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
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</taxonomicName>
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(Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Schematic illustration of the male terminalia of representatives of Amphineurus (dorsal view). A A. (A.) Amphineurus patya; B A. (A.) superbus; C A. (A.) pita; D A. (A.) umbraticus. - Abbreviations: ad spur, anterodorsal spur; aed, aedeagus; aed sh, sheath of aedeagus; b spur, basal spur; cgonst, clasper of gonostylus; db clasper, dorsal branch of clasper of gonostylus; db lobe, dorsal branch of lobe of gonostylus; f, foramen of male tergite X; goncx, gonocoxite; la spur, latero-apical spur; lb sh, lateral branch of sheath of aedeagus; lgonst, lobe of gonostylus; lh, lateral humps; mh, medial humps; pb sh, posterior branches of sheath of aedeagus; t 9, male tergite IX; t 10, male tergite X; vb clasper, ventral branch of clasper of gonostylus; vb lobe, ventral branch of lobe of gonostylus." figureDoi="10.3897/asp.80.e83035.figure1" httpUri="https://binary.pensoft.net/fig/740554" pageId="0" pageNumber="439">1</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Schematic illustration of the male terminalia of representatives of Amphineurus (dorsal view). A A. (A.) maculosus; B A. (Nothormosia) edentulus; C A. (A.) perdecorus; D A. (Nesormosia) fatuus. - Abbreviations: aed, aedeagus; aed sh, sheath of aedeagus; cgonst, clasper of gonostylus; db clasper, dorsal branch of clasper of gonostylus; goncx, gonocoxite; gonst, gonostylus; lb sh, lateral branch of sheath of aedeagus; lgonst, lobe of gonostylus; ms lobe, mesal lobe of gonocoxite; t 9, male tergite IX; vb clasper, ventral branch of clasper of gonostylus." figureDoi="10.3897/asp.80.e83035.figure2" httpUri="https://binary.pensoft.net/fig/740555" pageId="0" pageNumber="439">2</figureCitation>
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). There are many variations from the simplified pattern of
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<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (A.) patya" order="Diptera" pageId="0" pageNumber="439" rank="species" species="patya" subGenus="A.">A. (A.) patya</taxonomicName>
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Theischinger (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Schematic illustration of the male terminalia of representatives of Amphineurus (dorsal view). A A. (A.) Amphineurus patya; B A. (A.) superbus; C A. (A.) pita; D A. (A.) umbraticus. - Abbreviations: ad spur, anterodorsal spur; aed, aedeagus; aed sh, sheath of aedeagus; b spur, basal spur; cgonst, clasper of gonostylus; db clasper, dorsal branch of clasper of gonostylus; db lobe, dorsal branch of lobe of gonostylus; f, foramen of male tergite X; goncx, gonocoxite; la spur, latero-apical spur; lb sh, lateral branch of sheath of aedeagus; lgonst, lobe of gonostylus; lh, lateral humps; mh, medial humps; pb sh, posterior branches of sheath of aedeagus; t 9, male tergite IX; t 10, male tergite X; vb clasper, ventral branch of clasper of gonostylus; vb lobe, ventral branch of lobe of gonostylus." figureDoi="10.3897/asp.80.e83035.figure1" httpUri="https://binary.pensoft.net/fig/740554" pageId="0" pageNumber="439">1A</figureCitation>
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) to the asymmetric disposition of
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<taxonomicName authority="" family="Limoniidae" kingdom="Animalia" lsidName="A. (Nesormosia)" order="Diptera" pageId="0" pageNumber="439" rank="subGenus" subGenus="Nesormosia">A. (Nesormosia)</taxonomicName>
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(Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Schematic illustration of the male terminalia of representatives of Amphineurus (dorsal view). A A. (A.) maculosus; B A. (Nothormosia) edentulus; C A. (A.) perdecorus; D A. (Nesormosia) fatuus. - Abbreviations: aed, aedeagus; aed sh, sheath of aedeagus; cgonst, clasper of gonostylus; db clasper, dorsal branch of clasper of gonostylus; goncx, gonocoxite; gonst, gonostylus; lb sh, lateral branch of sheath of aedeagus; lgonst, lobe of gonostylus; ms lobe, mesal lobe of gonocoxite; t 9, male tergite IX; vb clasper, ventral branch of clasper of gonostylus." figureDoi="10.3897/asp.80.e83035.figure2" httpUri="https://binary.pensoft.net/fig/740555" pageId="0" pageNumber="439">2D</figureCitation>
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). Therefore, defining a broad diagnosis for
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<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
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</taxonomicName>
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is not so easy.
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.15067" author="Skuse, FAA" journalOrPublisher="International Journal of Dipterological Research" pageId="0" pageNumber="439" refId="B41" refString="Skuse, FAA, 1890. Diptera of Australia, Part 7. The Tipulidae brevipalpi. Proceedings of the Linnaean Society of New South Wales (2)4: 757-892. https://doi.org/10.5962/bhl.part.15067" title="Diptera of Australia, Part 7. The Tipulidae brevipalpi. Proceedings of the Linnaean Society of New South Wales (2) 4: 757 - 892." url="https://doi.org/10.5962/bhl.part.15067" year="1890">Skuse (1890)</bibRefCitation>
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defined
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<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
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</taxonomicName>
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as a subgenus of
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<taxonomicName class="Insecta" family="Limoniidae" genus="Rhypholophus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhypholophus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="439">Rhypholophus</emphasis>
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</taxonomicName>
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.
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<normalizedToken originalValue="Skuse’s">Skuse's</normalizedToken>
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definition fits better to the current subgenus
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<taxonomicName authority="" family="Limoniidae" kingdom="Animalia" lsidName="A. (Amphineurus)" order="Diptera" pageId="0" pageNumber="439" rank="subGenus" subGenus="Amphineurus">A. (Amphineurus)</taxonomicName>
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but does not portray the reality of other species of
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<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
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</taxonomicName>
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. His diagnosis included the absence of the discal cell, the radial sector with four branches, "second longitudinal vein angulated", and hind femora one third longer than the intermediate pair. These features are non-exclusive or variable in the genus.
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</paragraph>
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<caption doi="10.3897/asp.80.e83035.figure1" httpUri="https://binary.pensoft.net/fig/740554" pageId="0" pageNumber="439" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="439">
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<emphasis bold="true" pageId="0" pageNumber="439">Figure 1.</emphasis>
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Schematic illustration of the male terminalia of representatives of
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<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
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</taxonomicName>
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(dorsal view).
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<emphasis bold="true" pageId="0" pageNumber="439">A</emphasis>
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<taxonomicName family="Limoniidae" genus="A." kingdom="Animalia" lsidName="A." order="Diptera" pageId="0" pageNumber="439" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="439">A.</emphasis>
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</taxonomicName>
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(
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<taxonomicName family="Limoniidae" genus="A." kingdom="Animalia" lsidName="A." order="Diptera" pageId="0" pageNumber="439" rank="genus">A.</taxonomicName>
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)
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<taxonomicName authorityName="Theischinger" authorityYear="1994" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus patya" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="patya">
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<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus patya</emphasis>
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</taxonomicName>
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;
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<emphasis bold="true" pageId="0" pageNumber="439">B</emphasis>
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<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (A.) superbus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="superbus" subGenus="A.">A. (A.) superbus</taxonomicName>
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;
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<emphasis bold="true" pageId="0" pageNumber="439">C</emphasis>
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<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (A.) pita" order="Diptera" pageId="0" pageNumber="439" rank="species" species="pita" subGenus="A.">A. (A.) pita</taxonomicName>
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;
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<emphasis bold="true" pageId="0" pageNumber="439">D</emphasis>
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<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (A.) umbraticus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="umbraticus" subGenus="A.">A. (A.) umbraticus</taxonomicName>
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. - Abbreviations:
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<emphasis bold="true" pageId="0" pageNumber="439">ad spur</emphasis>
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, anterodorsal spur;
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<emphasis bold="true" pageId="0" pageNumber="439">aed</emphasis>
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, aedeagus;
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<emphasis bold="true" pageId="0" pageNumber="439">aed sh</emphasis>
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, sheath of aedeagus;
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<emphasis bold="true" pageId="0" pageNumber="439">b spur</emphasis>
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, basal spur;
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<emphasis bold="true" pageId="0" pageNumber="439">cgonst</emphasis>
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, clasper of gonostylus;
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<emphasis bold="true" pageId="0" pageNumber="439">db clasper</emphasis>
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, dorsal branch of clasper of gonostylus;
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<emphasis bold="true" pageId="0" pageNumber="439">db lobe</emphasis>
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, dorsal branch of lobe of gonostylus;
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<emphasis bold="true" pageId="0" pageNumber="439">f</emphasis>
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, foramen of male tergite X;
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<emphasis bold="true" pageId="0" pageNumber="439">goncx</emphasis>
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, gonocoxite;
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<emphasis bold="true" pageId="0" pageNumber="439">la spur</emphasis>
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, latero-apical spur;
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<emphasis bold="true" pageId="0" pageNumber="439">lb sh</emphasis>
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, lateral branch of sheath of aedeagus;
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<emphasis bold="true" pageId="0" pageNumber="439">lgonst</emphasis>
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, lobe of gonostylus;
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<emphasis bold="true" pageId="0" pageNumber="439">lh</emphasis>
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, lateral humps;
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<emphasis bold="true" pageId="0" pageNumber="439">mh</emphasis>
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, medial humps;
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<emphasis bold="true" pageId="0" pageNumber="439">pb sh</emphasis>
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, posterior branches of sheath of aedeagus;
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<emphasis bold="true" pageId="0" pageNumber="439">t9</emphasis>
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, male tergite IX;
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<emphasis bold="true" pageId="0" pageNumber="439">t10</emphasis>
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, male tergite X;
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<emphasis bold="true" pageId="0" pageNumber="439">vb clasper</emphasis>
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, ventral branch of clasper of gonostylus;
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<emphasis bold="true" pageId="0" pageNumber="439">vb lobe</emphasis>
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, ventral branch of lobe of gonostylus.
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</paragraph>
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</caption>
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<caption doi="10.3897/asp.80.e83035.figure2" httpUri="https://binary.pensoft.net/fig/740555" pageId="0" pageNumber="439" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="439">
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<emphasis bold="true" pageId="0" pageNumber="439">Figure 2.</emphasis>
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Schematic illustration of the male terminalia of representatives of
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<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
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</taxonomicName>
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(dorsal view).
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<emphasis bold="true" pageId="0" pageNumber="439">A</emphasis>
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<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (A.) maculosus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="maculosus" subGenus="A.">A. (A.) maculosus</taxonomicName>
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;
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<emphasis bold="true" pageId="0" pageNumber="439">B</emphasis>
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<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (Nothormosia) edentulus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="edentulus" subGenus="Nothormosia">A. (Nothormosia) edentulus</taxonomicName>
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;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">C</emphasis>
|
||
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (A.) perdecorus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="perdecorus" subGenus="A.">A. (A.) perdecorus</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">D</emphasis>
|
||
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (Nesormosia) fatuus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="fatuus" subGenus="Nesormosia">A. (Nesormosia) fatuus</taxonomicName>
|
||
. - Abbreviations:
|
||
<emphasis bold="true" pageId="0" pageNumber="439">aed</emphasis>
|
||
, aedeagus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">aed sh</emphasis>
|
||
, sheath of aedeagus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">cgonst</emphasis>
|
||
, clasper of gonostylus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">db clasper</emphasis>
|
||
, dorsal branch of clasper of gonostylus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">goncx</emphasis>
|
||
, gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">gonst</emphasis>
|
||
, gonostylus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">lb sh</emphasis>
|
||
, lateral branch of sheath of aedeagus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">lgonst</emphasis>
|
||
, lobe of gonostylus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">ms lobe</emphasis>
|
||
, mesal lobe of gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">t9</emphasis>
|
||
, male tergite IX;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">vb clasper</emphasis>
|
||
, ventral branch of clasper of gonostylus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="439">
|
||
<bibRefCitation DOI="https://doi.org/10.4039/Ent52141-6" author="Alexander, CP" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="439" pagination="52 - 63" refId="B1" refString="Alexander, CP, 1920. New or little-known crane-flies in the Queensland museum (Tipulidae, Diptera). Memoirs of the Queensland Museum 7: 52 - 63, DOI: https://doi.org/10.4039/Ent52141-6" title="New or little-known crane-flies in the Queensland museum (Tipulidae, Diptera)." url="https://doi.org/10.4039/Ent52141-6" volume="7" year="1920">Alexander (1920)</bibRefCitation>
|
||
allocated
|
||
<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
|
||
</taxonomicName>
|
||
as a subgenus of
|
||
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Ormosia" kingdom="Plantae" lsidName="Ormosia" order="Fabales" pageId="0" pageNumber="439" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Ormosia</emphasis>
|
||
</taxonomicName>
|
||
after highlighting the abundant and conspicuous wing macrotrichia (Fig.
|
||
<figureCitation captionStart="Figure 5. A" captionStartId="F5" captionText="Figure 5. A Wing of Amphineurus (Rhamphoneurus) caleuchus sp. nov. (male); B detail of wing of Amphineurus (Rhamphoneurus) immaculatus sp. nov. - Abbreviation: arc, arculus." figureDoi="10.3897/asp.80.e83035.figure5a" httpUri="https://binary.pensoft.net/fig/740558" pageId="0" pageNumber="439">5B</figureCitation>
|
||
).
|
||
<bibRefCitation author="Edwards, FW" journalOrPublisher="Transactions of the New Zealand Institute" pageId="0" pageNumber="439" pagination="265 - 352" refId="B19" refString="Edwards, FW, 1923. A preliminary revision of the crane-flies of New Zealand (Anisopodidae, Tanyderidae, Tipulidae). Transactions of the New Zealand Institute 54: 265 - 352" title="A preliminary revision of the crane-flies of New Zealand (Anisopodidae, Tanyderidae, Tipulidae)." volume="54" year="1923">Edwards (1923)</bibRefCitation>
|
||
redefined the genus to support two New Zealand species.
|
||
<normalizedToken originalValue="Edwards’">Edwards'</normalizedToken>
|
||
diagnosis includes the short mid-leg, short and hairy wing, and radius four-branched. Nevertheless, these features occur in other
|
||
<taxonomicName lsidName="" pageId="0" pageNumber="439" rank="subFamily" subFamily="Chioneinae">Chioneinae</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Edwards, FW" journalOrPublisher="Transactions of the New Zealand Institute" pageId="0" pageNumber="439" pagination="265 - 352" refId="B19" refString="Edwards, FW, 1923. A preliminary revision of the crane-flies of New Zealand (Anisopodidae, Tanyderidae, Tipulidae). Transactions of the New Zealand Institute 54: 265 - 352" title="A preliminary revision of the crane-flies of New Zealand (Anisopodidae, Tanyderidae, Tipulidae)." volume="54" year="1923">Edwards (1923)</bibRefCitation>
|
||
also referred to the fact that the medial vein forks in M3+4 and M1+2 and that the anal vein is sinuous. These features are plastic in the genus.
|
||
<bibRefCitation author="Alexander, CP" journalOrPublisher="Diptera of Patagonia and South Chile" pageId="0" pageNumber="439" pagination="1 - 240" refId="B7" refString="Alexander, CP, 1929a. Diptera of Patagonia and South Chile. Part I. Crane-flies (Tipulidae, Trichoceridae, Tanyderidae). Diptera of Patagonia and South Chile 1: 1 - 240" title="Diptera of Patagonia and South Chile. Part I. Crane-flies (Tipulidae, Trichoceridae, Tanyderidae)." volume="1" year="1929 a">Alexander (1929a)</bibRefCitation>
|
||
used the definition of
|
||
<bibRefCitation author="Edwards, FW" journalOrPublisher="Transactions of the New Zealand Institute" pageId="0" pageNumber="439" pagination="265 - 352" refId="B19" refString="Edwards, FW, 1923. A preliminary revision of the crane-flies of New Zealand (Anisopodidae, Tanyderidae, Tipulidae). Transactions of the New Zealand Institute 54: 265 - 352" title="A preliminary revision of the crane-flies of New Zealand (Anisopodidae, Tanyderidae, Tipulidae)." volume="54" year="1923">Edwards (1923)</bibRefCitation>
|
||
and emphasized the characteristic "abundant trichiation" of the genus. However, in the same article,
|
||
<bibRefCitation author="Alexander, CP" journalOrPublisher="Diptera of Patagonia and South Chile" pageId="0" pageNumber="439" pagination="1 - 240" refId="B7" refString="Alexander, CP, 1929a. Diptera of Patagonia and South Chile. Part I. Crane-flies (Tipulidae, Trichoceridae, Tanyderidae). Diptera of Patagonia and South Chile 1: 1 - 240" title="Diptera of Patagonia and South Chile. Part I. Crane-flies (Tipulidae, Trichoceridae, Tanyderidae)." volume="1" year="1929 a">Alexander (1929a)</bibRefCitation>
|
||
described the genus
|
||
<taxonomicName authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Maietta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maietta" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Maietta</emphasis>
|
||
</taxonomicName>
|
||
, which has more wing vestiture than
|
||
<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
|
||
</taxonomicName>
|
||
. Some species of the Neotropical subgenus
|
||
<taxonomicName authority="" family="Limoniidae" kingdom="Animalia" lsidName="A. (Rhamphoneurus)" order="Diptera" pageId="0" pageNumber="439" rank="subGenus" subGenus="Rhamphoneurus">A. (Rhamphoneurus)</taxonomicName>
|
||
have less macrotrichia than other subgenera. No more inclusive diagnosis was proposed and recent revisions (e.g.,
|
||
<bibRefCitation author="Theischinger, G" journalOrPublisher="Stapfia" pageId="0" pageNumber="439" pagination="37 - 276" refId="B42" refString="Theischinger, G, 1994. The Limoniinae (Diptera: Tipulidae) of Australia V. The genera Helius Le Peletier and Serville, Toxorhina Loew, Limonia Meigen (part), Tonnoiromyia Alexander and Collessophila gen. nov. (all tribe Limoniini) and Atarba Osten-Sacken, Amphineurus Skuse, Erioptera Meigen, Cheilotrichia Rossi, Gonomyia Meigen and Idiocera Dale (all tribe Eriopterini). Stapfia 36: 37 - 276" title="The Limoniinae (Diptera: Tipulidae) of Australia V. The genera Helius Le Peletier and Serville, Toxorhina Loew, Limonia Meigen (part), Tonnoiromyia Alexander and Collessophila gen. nov. (all tribe Limoniini) and Atarba Osten-Sacken, Amphineurus Skuse, Erioptera Meigen, Cheilotrichia Rossi, Gonomyia Meigen and Idiocera Dale (all tribe Eriopterini)." volume="36" year="1994">Theischinger 1994</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Theischinger, G" journalOrPublisher="Stapfia" pageId="0" pageNumber="439" pagination="1 - 18" refId="B43" refString="Theischinger, G, 1996. The Limoniinae (Diptera: Tipulidae) of Australia VI. New and insufficiently known species of Toxorhina Loew, Limonia Meigen, Atarba Osten Sacken, Amphineurus Skuse, Gonomyia Meigen and Molophilus Curtis. Stapfia 44: 1 - 18" title="The Limoniinae (Diptera: Tipulidae) of Australia VI. New and insufficiently known species of Toxorhina Loew, Limonia Meigen, Atarba Osten Sacken, Amphineurus Skuse, Gonomyia Meigen and Molophilus Curtis." volume="44" year="1996">1996</bibRefCitation>
|
||
) redefined only the diagnosis of the subgenus
|
||
<taxonomicName authority="" family="Limoniidae" kingdom="Animalia" lsidName="A. (Amphineurus)" order="Diptera" pageId="0" pageNumber="439" rank="subGenus" subGenus="Amphineurus">A. (Amphineurus)</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="439">
|
||
<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
|
||
</taxonomicName>
|
||
has an 180
|
||
<normalizedToken originalValue="º">°</normalizedToken>
|
||
rotation of male terminalia (in the words of Edwards: "undergone permanent torsion").
|
||
<bibRefCitation author="Edwards, FW" journalOrPublisher="Transactions of the New Zealand Institute" pageId="0" pageNumber="439" pagination="265 - 352" refId="B19" refString="Edwards, FW, 1923. A preliminary revision of the crane-flies of New Zealand (Anisopodidae, Tanyderidae, Tipulidae). Transactions of the New Zealand Institute 54: 265 - 352" title="A preliminary revision of the crane-flies of New Zealand (Anisopodidae, Tanyderidae, Tipulidae)." volume="54" year="1923">Edwards (1923)</bibRefCitation>
|
||
also shows that
|
||
<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
|
||
</taxonomicName>
|
||
has a long, finger-like gonocoxite process (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Schematic illustration of the male terminalia of representatives of Amphineurus (dorsal view). A A. (A.) Amphineurus patya; B A. (A.) superbus; C A. (A.) pita; D A. (A.) umbraticus. - Abbreviations: ad spur, anterodorsal spur; aed, aedeagus; aed sh, sheath of aedeagus; b spur, basal spur; cgonst, clasper of gonostylus; db clasper, dorsal branch of clasper of gonostylus; db lobe, dorsal branch of lobe of gonostylus; f, foramen of male tergite X; goncx, gonocoxite; la spur, latero-apical spur; lb sh, lateral branch of sheath of aedeagus; lgonst, lobe of gonostylus; lh, lateral humps; mh, medial humps; pb sh, posterior branches of sheath of aedeagus; t 9, male tergite IX; t 10, male tergite X; vb clasper, ventral branch of clasper of gonostylus; vb lobe, ventral branch of lobe of gonostylus." figureDoi="10.3897/asp.80.e83035.figure1" httpUri="https://binary.pensoft.net/fig/740554" pageId="0" pageNumber="439">1A</figureCitation>
|
||
). This is evidence of the bifid gonocoxite, a condition present (Fig.
|
||
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Amphineurus (Rhamphoneurus) stigamticus sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure29" httpUri="https://binary.pensoft.net/fig/740582" pageId="0" pageNumber="439">29G</figureCitation>
|
||
) in some
|
||
<taxonomicName lsidName="" pageId="0" pageNumber="439" rank="subFamily" subFamily="Chioneinae">Chioneinae</taxonomicName>
|
||
with inverted male terminalia. This gonocoxal process forms a protective hood (
|
||
<bibRefCitation DOI="https://doi.org/10.1007/s12312-020-00750-x" author="Kramer, J" journalOrPublisher="Dipterists Forum Cranefly Recording Scheme Newsletter" pageId="0" pageNumber="439" pagination="6 - 7" refId="B24" refString="Kramer, J, 2020. The rotation of the hypopygium in Molophilus and other Chioneinae. Bulletin of the Dipterists Forum 89; Cranefly News. Dipterists Forum Cranefly Recording Scheme Newsletter 35: 6 - 7, DOI: https://doi.org/10.1007/s12312-020-00750-x" title="The rotation of the hypopygium in Molophilus and other Chioneinae. Bulletin of the Dipterists Forum 89; Cranefly News." url="https://doi.org/10.1007/s12312-020-00750-x" volume="35" year="2020">Kramer 2020</bibRefCitation>
|
||
) for the appendices of the inverted terminalia.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="439">
|
||
The
|
||
<taxonomicName lsidName="" pageId="0" pageNumber="439" rank="subFamily" subFamily="Chioneinae">Chioneinae</taxonomicName>
|
||
genera with inverted terminalia bear great diversity and frequently they have hairy wings similar to
|
||
<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
|
||
</taxonomicName>
|
||
. The great amount of wing vestiture resembles
|
||
<taxonomicName authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Maietta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maietta" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Maietta</emphasis>
|
||
</taxonomicName>
|
||
Alexander. The wings of some
|
||
<taxonomicName authority="" family="Limoniidae" kingdom="Animalia" lsidName="A. (Amphineurus)" order="Diptera" pageId="0" pageNumber="439" rank="subGenus" subGenus="Amphineurus">A. (Amphineurus)</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Most parsimonious tree (part) yielded by parsimony analysis with implied weighting (K = 3), depicting recovered relationships among outgroup taxa. Numbers at tree branches designate nodes (see 4.2.). - Abbreviations: Aus, Australia; NC, New Caledonia; NG, New Guinea; NZ, New Zealand, SAm, South America; Tas, Tasmania." figureDoi="10.3897/asp.80.e83035.figure32" httpUri="https://binary.pensoft.net/fig/740585" pageId="0" pageNumber="439">32</figureCitation>
|
||
: Node 18) have a distinctive radial sector such as seen in
|
||
<taxonomicName class="Insecta" family="Limoniidae" genus="Molophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Molophilus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Molophilus</emphasis>
|
||
</taxonomicName>
|
||
Curtis. Some
|
||
<taxonomicName authority="" family="Limoniidae" kingdom="Animalia" lsidName="A. (Amphineurus)" order="Diptera" pageId="0" pageNumber="439" rank="subGenus" subGenus="Amphineurus">A. (Amphineurus)</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Most parsimonious tree (part) yielded by parsimony analysis with implied weighting (K = 3), depicting recovered relationships among outgroup taxa. Numbers at tree branches designate nodes (see 4.2.). - Abbreviations: Aus, Australia; NC, New Caledonia; NG, New Guinea; NZ, New Zealand, SAm, South America; Tas, Tasmania." figureDoi="10.3897/asp.80.e83035.figure32" httpUri="https://binary.pensoft.net/fig/740585" pageId="0" pageNumber="439">32</figureCitation>
|
||
: Node 23) resembles
|
||
<taxonomicName class="Insecta" family="Limoniidae" genus="Hoplolabis" kingdom="Animalia" lsidName="Hoplolabis" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Hoplolabis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Savchenko" authorityYear="1976" class="Insecta" family="Limoniidae" genus="Parilisia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parilisia" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Parilisia</emphasis>
|
||
</taxonomicName>
|
||
Savchenko). The trifid claspers of
|
||
<taxonomicName authority="" family="Limoniidae" kingdom="Animalia" lsidName="A. (Rhamphoneurus)" order="Diptera" pageId="0" pageNumber="439" rank="subGenus" subGenus="Rhamphoneurus">A. (Rhamphoneurus)</taxonomicName>
|
||
are observed in genera such as
|
||
<taxonomicName class="Insecta" family="Limoniidae" genus="Hoplolabis" kingdom="Animalia" lsidName="Hoplolabis" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Hoplolabis</emphasis>
|
||
</taxonomicName>
|
||
Osten Sacken and
|
||
<taxonomicName authorityName="Rossi" authorityYear="1848" class="Insecta" family="Limoniidae" genus="Cheilotrichia" kingdom="Animalia" lsidName="Cheilotrichia" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Cheilotrichia</emphasis>
|
||
</taxonomicName>
|
||
Rossi. Despite that, in the group of genera with a bifid gonocoxite and inverted terminalia, vein R2+3 is normally aligned with R2+3+4. The exception is a clade of
|
||
<taxonomicName authority="" family="Limoniidae" kingdom="Animalia" lsidName="A. (Amphineurus)" order="Diptera" pageId="0" pageNumber="439" rank="subGenus" subGenus="Amphineurus">A. (Amphineurus)</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Most parsimonious tree (part) yielded by parsimony analysis with implied weighting (K = 3), depicting recovered relationships among outgroup taxa. Numbers at tree branches designate nodes (see 4.2.). - Abbreviations: Aus, Australia; NC, New Caledonia; NG, New Guinea; NZ, New Zealand, SAm, South America; Tas, Tasmania." figureDoi="10.3897/asp.80.e83035.figure32" httpUri="https://binary.pensoft.net/fig/740585" pageId="0" pageNumber="439">32</figureCitation>
|
||
: Node 18) that shows resemblances in the radial sector with
|
||
<taxonomicName class="Insecta" family="Limoniidae" genus="Molophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Molophilus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Molophilus</emphasis>
|
||
</taxonomicName>
|
||
(where R2+3+4 is lacking), but the former has a closed discal cell among other wing features which places it within
|
||
<taxonomicName authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |