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<document id="F79EEC187A399E7686EF29DF5431EAC9" ID-DOI="10.1080/00222930500198403" ID-ISSN="1464-5262" ID-Zenodo-Dep="5220093" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629352853822" checkinUser="felipe" docAuthor="Csuzdi, Csaba" docDate="2005" docId="9A350F00FFECFF84FEDCAC93689FA1D6" docLanguage="en" docName="JNatHist.39.33.3039-3058.pdf" docOrigin="Journal of Natural History 39 (33)" docSource="http://dx.doi.org/10.1080/00222930500198403" docStyle="DocumentStyle:5A9EB2ED5589A523F0236C66DE2A6BA8.4:JNatHist.2005-2007.journal_article.1cover" docStyleId="5A9EB2ED5589A523F0236C66DE2A6BA8" docStyleName="JNatHist.2005-2007.journal_article.1cover" docStyleVersion="4" docTitle="Dichogaster (Dichogaster) thomeana Cognetti 1910" docType="treatment" docVersion="3" lastPageNumber="3042" masterDocId="660C7778FFEFFF80FF9BAF4D6864A530" masterDocTitle="Earthworms (Annelida: Oligochaeta) of Sao Tomé" masterLastPageNumber="3058" masterPageNumber="3039" pageNumber="3041" updateTime="1699069145428" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="690BA3832EF589EA35534B7DC83B679B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E2DD23C57E72963DDCEC2968379EA67A">
<mods:title id="40C61F63A3B26E515B85F9A9961E2B4B">Earthworms (Annelida: Oligochaeta) of Sao Tomé</mods:title>
</mods:titleInfo>
<mods:name id="82CCE590627B4A540DD9A5ED8BF4D11B" type="personal">
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<mods:namePart id="9E9267B4E5CD121A6590E3076DE0B2B6">Csuzdi, Csaba</mods:namePart>
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<mods:typeOfResource id="1F4C38BEA3720E73172E804CDFCBCAAB">text</mods:typeOfResource>
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<mods:title id="42DE3C7F54D2024BD3A3FC4FB34361DD">Journal of Natural History</mods:title>
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<mods:part id="4FED3959BCAAB0A2C5D9FC0A77CA6B00">
<mods:date id="A280679A0C6F4282B28E81BDB9695719">2005</mods:date>
<mods:detail id="F299C33A8DD30BB0B74311EA5962E0E8" type="pubDate">
<mods:number id="B4C34812B1BB9E0C8EA202542FEE91D3">2005-10-28</mods:number>
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<mods:detail id="C2B60A46825A5AA610E5731049A9B4E9" type="volume">
<mods:number id="82F6DCF334CA8DA0306976C4848594E3">39</mods:number>
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<mods:number id="84DF18C75BC7B4B0975CE1B71303B503">33</mods:number>
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<mods:start id="3E7193D529C4698123FCEDE6BA2E839E">3039</mods:start>
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<mods:classification id="4603555B170CA0F20409FFA7B091B0CD">journal article</mods:classification>
<mods:identifier id="CBE8E997AEF6B2188793DDD574A4FF02" type="DOI">10.1080/00222930500198403</mods:identifier>
<mods:identifier id="D58ED6392BBBF4AA303E51E32A2F241A" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="0629BA4F5953BE3545A56CD64BC3C4B0" type="Zenodo-Dep">5220093</mods:identifier>
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<treatment id="9A350F00FFECFF84FEDCAC93689FA1D6" LSID="urn:lsid:plazi:treatment:9A350F00FFECFF84FEDCAC93689FA1D6" httpUri="http://treatment.plazi.org/id/9A350F00FFECFF84FEDCAC93689FA1D6" lastPageId="4" lastPageNumber="3042" pageId="3" pageNumber="3041">
<subSubSection id="5A86ED9DFFECFF83FEDCAC936B80A6C6" box="[327,996,990,1014]" pageId="3" pageNumber="3041" type="nomenclature">
<paragraph id="1223BE16FFECFF83FEDCAC936B80A6C6" blockId="3.[138,996,990,1142]" box="[327,996,990,1014]" pageId="3" pageNumber="3041">
<heading id="496B097AFFECFF83FEDCAC936B80A6C6" box="[327,996,990,1014]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="3041" reason="2">
<taxonomicName id="D59CC595FFECFF83FEDCAC936B80A6C6" authority="Cognetti, 1910" authorityName="Cognetti" authorityYear="1910" box="[327,996,990,1014]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="3041" phylum="Annelida" rank="species" species="thomeana" subGenus="Dichogaster">
<emphasis id="20E86204FFECFF83FEDCAC936B4EA6C6" bold="true" box="[327,810,990,1014]" italics="true" pageId="3" pageNumber="3041">Dichogaster (Dichogaster) thomeana</emphasis>
<bibRefCitation id="760DC3E7FFECFF83FCA8AC936B80A6C6" author="Cognetti de Martiis L" box="[819,996,990,1014]" pageId="3" pageNumber="3041" pagination="79 - 118" refId="ref6590" refString="Cognetti de Martiis L. 1910. Lombrichi raccolti dal Cav. Leonardo Fea nelle Isole del Capo Verde e nel Golfo di Guinea. Annali del Museo Civico di Storia Naturale di Genova 4: 79 - 118." type="journal article" year="1910">Cognetti, 1910</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5A86ED9DFFECFF83FDD2ACB36A86A126" box="[585,738,1022,1046]" pageId="3" pageNumber="3041" type="description">
<paragraph id="1223BE16FFECFF83FDD2ACB36A86A126" blockId="3.[138,996,990,1142]" box="[585,738,1022,1046]" pageId="3" pageNumber="3041">
(
<figureCitation id="8AA7A293FFECFF83FDCBACB36ABEA126" box="[592,730,1022,1046]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStartId-0="4.[382,442,1663,1682]" captionStartId-1="5.[464,524,402,421]" captionStartId-2="5.[138,198,1638,1657]" captionTargetBox-0="[529,770,1288,1622]" captionTargetBox-1="[541,782,135,363]" captionTargetBox-2="[138,1185,882,1602]" captionTargetId-0="figure-145@4.[526,772,1285,1628]" captionTargetId-1="figure-77@5.[539,785,133,366]" captionTargetId-2="figure-85@5.[138,1185,882,1602]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionText-0="Figure 4. D. (D.) thomeana, ventral side of the clitellum." captionText-1="Figure 5. D. (D.) thomeana, spermatheca." captionText-2="Figure 6. D. (D.) thomeana, penial seta: (a) whole seta; (b) middle part of the seta with the characteristic ornamentation; (c) tip of the seta." figureDoi-0="http://doi.org/10.5281/zenodo.5220103" figureDoi-1="http://doi.org/10.5281/zenodo.5220105" figureDoi-2="http://doi.org/10.5281/zenodo.5220107" httpUri-0="https://zenodo.org/record/5220103/files/figure.png" httpUri-1="https://zenodo.org/record/5220105/files/figure.png" httpUri-2="https://zenodo.org/record/5220107/files/figure.png" pageId="3" pageNumber="3041">Figures 4–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5A86ED9DFFECFF83FF11AB536AB9A146" pageId="3" pageNumber="3041" type="reference_group">
<paragraph id="1223BE16FFECFF83FF11AB536AE1A106" blockId="3.[138,996,990,1142]" box="[138,645,1054,1078]" pageId="3" pageNumber="3041">
<treatmentCitationGroup id="328C9938FFECFF83FF11AB536AE1A106" box="[138,645,1054,1078]" pageId="3" pageNumber="3041">
<treatmentCitation id="933D9807FFECFF83FF11AB536AE5A106" author="Cognetti de Martiis L" box="[138,641,1054,1078]" page="100" pageId="3" pageNumber="3041" year="1910">
<taxonomicName id="D59CC595FFECFF83FF11AB536AE5A106" authority="Cognetti 1910, p 100" authorityName="Cognetti" authorityPageNumber="100" authorityYear="1910" box="[138,641,1054,1078]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="3041" phylum="Annelida" rank="species" species="thomeana">
<emphasis id="20E86204FFECFF83FF11AB53691FA106" box="[138,379,1054,1078]" italics="true" pageId="3" pageNumber="3041">Dichogaster thomeana</emphasis>
<bibRefCitation id="760DC3E7FFECFF83FE1FAB536A54A106" author="Cognetti de Martiis L" box="[388,560,1054,1078]" pageId="3" pageNumber="3041" pagination="79 - 118" refId="ref6590" refString="Cognetti de Martiis L. 1910. Lombrichi raccolti dal Cav. Leonardo Fea nelle Isole del Capo Verde e nel Golfo di Guinea. Annali del Museo Civico di Storia Naturale di Genova 4: 79 - 118." type="journal article" year="1910">Cognetti 1910</bibRefCitation>
, p 100
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="1223BE16FFECFF83FF11AB736B4FA166" blockId="3.[138,996,990,1142]" box="[138,811,1086,1110]" pageId="3" pageNumber="3041">
<treatmentCitationGroup id="328C9938FFECFF83FF11AB736B4FA166" box="[138,811,1086,1110]" pageId="3" pageNumber="3041">
<treatmentCitation id="933D9807FFECFF83FF11AB736A83A166" author="Csuzdi Cs" box="[138,743,1086,1110]" page="109" pageId="3" pageNumber="3041" year="1995">
<taxonomicName id="D59CC595FFECFF83FF11AB736A83A166" authority=": Csuzdi 1995, p 109" authorityName=": Csuzdi" authorityPageNumber="109" authorityYear="1995" box="[138,743,1086,1110]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="3041" phylum="Annelida" rank="species" species="greeffi" subGenus="Dichogaster">
<emphasis id="20E86204FFECFF83FF11AB73698BA166" box="[138,495,1086,1110]" italics="true" pageId="3" pageNumber="3041">Dichogaster (Dichogaster) greeffi</emphasis>
:
<bibRefCitation id="760DC3E7FFECFF83FD9BAB736AF3A166" author="Csuzdi Cs" box="[512,663,1086,1110]" pageId="3" pageNumber="3041" pagination="99 - 123" refId="ref6674" refString="Csuzdi Cs. 1995. A catalogue of Benhamiinae species (Oligochaeta, Acanthodrilidae). Annalen des Naturhistorischen Museums in Wien 97 B: 99 - 123." type="journal article" year="1995">Csuzdi 1995</bibRefCitation>
, p 109
</taxonomicName>
</treatmentCitation>
, part.
</treatmentCitationGroup>
</paragraph>
<paragraph id="1223BE16FFECFF83FF11AB136AB9A146" blockId="3.[138,996,990,1142]" box="[138,733,1118,1142]" pageId="3" pageNumber="3041">
<treatmentCitationGroup id="328C9938FFECFF83FF11AB136AB9A146" box="[138,733,1118,1142]" pageId="3" pageNumber="3041">
<treatmentCitation id="933D9807FFECFF83FF11AB136ABDA146" author="Csuzdi Cs" box="[138,729,1118,1142]" page="55" pageId="3" pageNumber="3041" year="2000">
<taxonomicName id="D59CC595FFECFF83FF11AB136ABDA146" authority=": Csuzdi 2000, p 55" authorityName=": Csuzdi" authorityPageNumber="55" authorityYear="2000" box="[138,729,1118,1142]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="3041" phylum="Annelida" rank="species" species="greeffi" subGenus="Dichogaster">
<emphasis id="20E86204FFECFF83FF11AB13698BA146" box="[138,495,1118,1142]" italics="true" pageId="3" pageNumber="3041">Dichogaster (Dichogaster) greeffi</emphasis>
:
<bibRefCitation id="760DC3E7FFECFF83FD9BAB136AF3A146" author="Csuzdi Cs" box="[512,663,1118,1142]" pageId="3" pageNumber="3041" pagination="51 - 80" refId="ref6733" refString="Csuzdi Cs. 2000. A review of Benhamiinae earthworms in the collection of the Natural History Museum, London (Oligochaeta, Acanthodrilidae, Benhamiinae). Opuscula Zoologica Budapest 32: 51 - 80." type="journal article" year="2000">Csuzdi 2000</bibRefCitation>
, p 55
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5A86ED9DFFECFF84FF11ABE3689FA1D6" lastPageId="4" lastPageNumber="3042" pageId="3" pageNumber="3041" type="materials_examined">
<paragraph id="1223BE16FFECFF83FF11ABE3693CA1F6" blockId="3.[138,344,1198,1222]" box="[138,344,1198,1222]" pageId="3" pageNumber="3041">
<emphasis id="20E86204FFECFF83FF11ABE3693CA1F6" box="[138,344,1198,1222]" italics="true" pageId="3" pageNumber="3041">Material examined</emphasis>
</paragraph>
<paragraph id="1223BE16FFECFF83FF11AB936A2BA026" blockId="3.[138,1185,1246,1302]" pageId="3" pageNumber="3041">
AF/5003
<specimenCount id="049A759FFFECFF83FF62AB936937A1C6" box="[249,339,1246,1270]" pageId="3" pageNumber="3041" type="generic">four ex.</specimenCount>
Sao Tome´, on the volcano around Lago Amélia,
<quantity id="D56413F3FFECFF83FC0DAB936B88A1C6" box="[918,1004,1246,1270]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="3041" unit="m" value="1450.0">1450 m</quantity>
asl, from fallen logs, leg. Cs. Csuzdi,
<date id="662298D6FFECFF83FE10ABB36A28A026" box="[395,588,1278,1302]" pageId="3" pageNumber="3041" value="2000-08-30">30 August 2000</date>
.
</paragraph>
<paragraph id="1223BE16FFECFF83FF11AA036897A056" blockId="3.[138,243,1358,1382]" box="[138,243,1358,1382]" pageId="3" pageNumber="3041">
<emphasis id="20E86204FFECFF83FF11AA036897A056" box="[138,243,1358,1382]" italics="true" pageId="3" pageNumber="3041">Diagnosis</emphasis>
</paragraph>
<paragraph id="1223BE16FFECFF83FF11AA336B83A3A5" blockId="3.[138,1185,1406,1685]" pageId="3" pageNumber="3041">
L.
<quantity id="D56413F3FFECFF83FF36AA33694EA0A6" box="[173,298,1406,1430]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="3" pageNumber="3041" unit="mm" value="57.5" valueMax="60.0" valueMin="55.0">55–60 mm</quantity>
, D.
<quantity id="D56413F3FFECFF83FEC4AA3369B1A0A6" box="[351,469,1406,1430]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="3" pageNumber="3041" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
, number of segments 117–126. Colour alive purplish brown, conserved pale. First dorsal pore in 12/13. Clitellum 13–20, prostatic pores in segment 17, 19, spermathecal pores in 7/8, 8/9 (
<figureCitation id="8AA7A293FFECFF83FDB5AAF36AF7A0E6" box="[558,659,1470,1494]" captionStart="Figure 4" captionStartId="4.[382,442,1663,1682]" captionTargetBox="[529,770,1288,1622]" captionTargetId="figure-145@4.[526,772,1285,1628]" captionTargetPageId="4" captionText="Figure 4. D. (D.) thomeana, ventral side of the clitellum." figureDoi="http://doi.org/10.5281/zenodo.5220103" httpUri="https://zenodo.org/record/5220103/files/figure.png" pageId="3" pageNumber="3041">Figure 4</figureCitation>
). Spermatheca ampulla barbell-shaped, duct long muscular, diverticula empty, unilocular with long duct (
<figureCitation id="8AA7A293FFECFF83FCD4AA936BD7A0C6" box="[847,947,1502,1526]" captionStart="Figure 5" captionStartId="5.[464,524,402,421]" captionTargetBox="[541,782,135,363]" captionTargetId="figure-77@5.[539,785,133,366]" captionTargetPageId="5" captionText="Figure 5. D. (D.) thomeana, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5220105" httpUri="https://zenodo.org/record/5220105/files/figure.png" pageId="3" pageNumber="3041">Figure 5</figureCitation>
). Calciferous glands three pairs in 15–17, the first pair is slightly smaller than the others. Meronephridia four to five on each side of the intestine. Penial setae uniform, the fully adult seta
<quantity id="D56413F3FFECFF83FC45A9506C0CA305" box="[990,1128,1565,1589]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="3" pageNumber="3041" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
long and in the middle
<quantity id="D56413F3FFECFF83FEFDA97069A8A365" box="[358,460,1597,1621]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="3041" unit="mm" value="0.04">0.04 mm</quantity>
wide. The seta is slightly curved and the spoon-shaped tip is reclinate. Each seta bears a characteristic ornamentation which are large serrated sculptures on the ental part becoming large thorns towards the ectal part (
<figureCitation id="8AA7A293FFECFF83FCE8A9306BBCA3A5" box="[883,984,1661,1685]" captionStart="Figure 6" captionStartId="5.[138,198,1638,1657]" captionTargetBox="[138,1185,882,1602]" captionTargetId="figure-85@5.[138,1185,882,1602]" captionTargetPageId="5" captionText="Figure 6. D. (D.) thomeana, penial seta: (a) whole seta; (b) middle part of the seta with the characteristic ornamentation; (c) tip of the seta." figureDoi="http://doi.org/10.5281/zenodo.5220107" httpUri="https://zenodo.org/record/5220107/files/figure.png" pageId="3" pageNumber="3041">Figure 6</figureCitation>
).
</paragraph>
<caption id="46E3EE9EFFEBFF84FFE6AC0269D8A64D" ID-DOI="http://doi.org/10.5281/zenodo.5220099" ID-Zenodo-Dep="5220099" httpUri="https://zenodo.org/record/5220099/files/figure.png" pageId="4" pageNumber="3042" startId="4.[125,185,847,866]" targetBox="[125,1174,133,812]" targetPageId="4">
<paragraph id="1223BE16FFEBFF84FFE6AC0269D8A64D" blockId="4.[125,1172,847,893]" pageId="4" pageNumber="3042">
Figure 3.
<taxonomicName id="D59CC595FFEBFF84FF78AC026914A652" authorityName=": Csuzdi" authorityYear="2000" box="[227,368,847,866]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="3042" phylum="Annelida" rank="species" species="greeffi" subGenus="Dichogaster">
<emphasis id="20E86204FFEBFF84FF78AC026914A652" box="[227,368,847,866]" italics="true" pageId="4" pageNumber="3042">D. (D.) greeffi</emphasis>
</taxonomicName>
, penial seta: (a) whole seta; (b) middle part of the seta with the characteristic ornamentation; (c) tip of the seta.
</paragraph>
</caption>
<paragraph id="1223BE16FFEBFF84FFE6ACD368B8A686" blockId="4.[125,220,926,950]" box="[125,220,926,950]" pageId="4" pageNumber="3042">
<emphasis id="20E86204FFEBFF84FFE6ACD368B8A686" box="[125,220,926,950]" italics="true" pageId="4" pageNumber="3042">Remarks</emphasis>
</paragraph>
<paragraph id="1223BE16FFEBFF84FFE6AC83689FA1D6" blockId="4.[125,1172,974,1254]" pageId="4" pageNumber="3042">
This species was synonymized with
<taxonomicName id="D59CC595FFEBFF84FDB1AC826AB5A6D6" authorityName=": Csuzdi" authorityYear="2000" box="[554,721,974,998]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="3042" phylum="Annelida" rank="species" species="greeffi" subGenus="Dichogaster">
<emphasis id="20E86204FFEBFF84FDB1AC826AB5A6D6" box="[554,721,974,998]" italics="true" pageId="4" pageNumber="3042">D. (D.) greeffi</emphasis>
</taxonomicName>
by
<bibRefCitation id="760DC3E7FFEBFF84FC9AAC836BC8A6D6" author="Csuzdi Cs" box="[769,940,974,998]" pageId="4" pageNumber="3042" pagination="99 - 123" refId="ref6674" refString="Csuzdi Cs. 1995. A catalogue of Benhamiinae species (Oligochaeta, Acanthodrilidae). Annalen des Naturhistorischen Museums in Wien 97 B: 99 - 123." type="journal article" year="1995">Csuzdi (1995)</bibRefCitation>
on the basis of the original description of the
<specimenCount id="049A759FFFEBFF84FE41ACA36AE5A136" box="[474,641,1006,1030]" pageId="4" pageNumber="3042" type="juvenile">three juvenile</specimenCount>
specimens of
<taxonomicName id="D59CC595FFEBFF84FCDDACA26C42A136" authorityName="Cognetti" authorityYear="1910" box="[838,1062,1006,1030]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="3042" phylum="Annelida" rank="species" species="thomeana" subGenus="Dichogaster">
<emphasis id="20E86204FFEBFF84FCDDACA26C42A136" box="[838,1062,1006,1030]" italics="true" pageId="4" pageNumber="3042">D. (D.) thomeana</emphasis>
</taxonomicName>
and on examination of the
<typeStatus id="CD2700B4FFEBFF84FEFCAB4369FCA116" box="[359,408,1038,1062]" pageId="4" pageNumber="3042">type</typeStatus>
material of
<taxonomicName id="D59CC595FFEBFF84FDB5AB426AB1A116" authorityName=": Csuzdi" authorityYear="2000" box="[558,725,1038,1062]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="3042" phylum="Annelida" rank="species" species="greeffi" subGenus="Dichogaster">
<emphasis id="20E86204FFEBFF84FDB5AB426AB1A116" box="[558,725,1038,1062]" italics="true" pageId="4" pageNumber="3042">D. (D.) greeffi</emphasis>
</taxonomicName>
housed in the Michaelsen collection, Hamburg. The slight differences in the spermathecae and penial setae were attributed to the juvenile stage of Cognetti’s specimens. SEM study of the penial seta of the new material revealed that its tip is spoon-shaped and reclinate which is markedly different from the sharply hooked seta of
<emphasis id="20E86204FFEBFF84FE18ABC26A4EA196" box="[387,554,1166,1190]" italics="true" pageId="4" pageNumber="3042">
<taxonomicName id="D59CC595FFEBFF84FE18ABC26A42A196" authorityName=": Csuzdi" authorityYear="2000" box="[387,550,1166,1190]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="3042" phylum="Annelida" rank="species" species="greeffi" subGenus="Dichogaster">D. (D.) greeffi</taxonomicName>
.
</emphasis>
Relying upon this observation I conclude that the two names refer to completely different species and
<taxonomicName id="D59CC595FFEBFF84FD3EABE36B14A1F6" authorityName="Cognetti" authorityYear="1910" box="[677,880,1198,1222]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="3042" phylum="Annelida" rank="species" species="thomeana" subGenus="Dichogaster">
<emphasis id="20E86204FFEBFF84FD3EABE36B14A1F6" box="[677,880,1198,1222]" italics="true" pageId="4" pageNumber="3042">D. (D.) thomeana</emphasis>
</taxonomicName>
should be removed from synonymy.
</paragraph>
</subSubSection>
</treatment>
</document>
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