treatments-xml/data/9A/2E/14/9A2E144DDC53560E8C1AD97BF089175A.xml

<document ID-DOI="http://dx.doi.org/10.3897/dez.69.89864" ID-Pensoft-Pub="1860-1324-2-219" ID-Pensoft-UUID="9EE9E317E8195E3A94756A0DBBB1C35B" ID-ZooBank="BFE2BE4759E9450A807079B702A38625" ModsDocID="1860-1324-69-2-219" checkinTime="1671200981815" checkinUser="pensoft" docAuthor="Souma, Jun" docDate="2022" docId="9A2E144DDC53560E8C1AD97BF089175A" docLanguage="en" docName="DeutEntomolZeit 69(2): 219-281" docOrigin="Deutsche Entomologische Zeitschrift 69 (2)" docPubDate="2022-12-15" docSource="http://dx.doi.org/10.3897/dez.69.89864" docTitle="Stephanitis (Norba) aperta Horvath 1912" docType="treatment" docVersion="1" id="9EE9E317E8195E3A94756A0DBBB1C35B" lastPageNumber="219" masterDocId="9EE9E317E8195E3A94756A0DBBB1C35B" masterDocTitle="Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan" masterLastPageNumber="281" masterPageNumber="219" pageNumber="219" updateTime="1671200981815" updateUser="pensoft">
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<mods:title>Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Souma, Jun</mods:namePart>
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<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, Japan &amp; Research Fellowship for Young Scientists (DC 1), Japan Society for the Promotion of Science, Tokyo, Japan</mods:affiliation>
<mods:nameIdentifier type="email">kodokusignal@gmail.com</mods:nameIdentifier>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-12-15</mods:number>
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<mods:number>69</mods:number>
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<taxonomicName LSID="9A2E144D-DC53-560E-8C1A-D97BF089175A" authority="Horvath, 1912" authorityName="Horvath" authorityYear="1912" class="Insecta" family="Tingidae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">
Stephanitis (Norba) aperta 
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
, 1912
</taxonomicName>
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<subSubSection pageId="0" pageNumber="219" type="description">
<paragraph pageId="0" pageNumber="219">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male habitus of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua from Honshu; B. S. (Norba) aperta from Honshu; C. S. (N.) exigua from Minamidaito Island, Daito Islands; D. S. (N.) hayashii sp. nov. from Senaga Island, central part of Ryukyu Islands; E, F. S. (N.) hiurai from Kakeroma (E) and Takara (F) islands, central part of Ryukyu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure2" httpUri="https://binary.pensoft.net/fig/782629" pageId="0" pageNumber="219">[Japanese name: Tabu-gunbai] Figs 2B</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female habitus of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua from Honshu; B. S. (Norba) aperta from Honshu; C. S. (N.) exigua from Okinawa Island, central part of Ryukyu Islands; D. S. (N.) hayashii sp. nov. from Aguni Island, central part of Ryukyu Islands; E, F. S. (N.) hiurai from Okinawa (E) and Takara (F) Islands, central part of Ryukyu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure4" httpUri="https://binary.pensoft.net/fig/782631" pageId="0" pageNumber="219">, 4B</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Teratological forms of two Stephanitis species from Japan, dorsal view: A. S. (Norba) aperta from Izu-Oshima Island, northern part of Izu Islands; B. S. (Stephanitis) takeyai from Kyushu. Arrows indicate antennae with segmental oligomery and malformed paranotum. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure6" httpUri="https://binary.pensoft.net/fig/782633" pageId="0" pageNumber="219">, 6A</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure7" httpUri="https://binary.pensoft.net/fig/782634" pageId="0" pageNumber="219">, 7B</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure9" httpUri="https://binary.pensoft.net/fig/782636" pageId="0" pageNumber="219">, 9B</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure11" httpUri="https://binary.pensoft.net/fig/782638" pageId="0" pageNumber="219">, 11B</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure13" httpUri="https://binary.pensoft.net/fig/782640" pageId="0" pageNumber="219">, 13B</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Male hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure15" httpUri="https://binary.pensoft.net/fig/782642" pageId="0" pageNumber="219">, 15B</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Female hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure17" httpUri="https://binary.pensoft.net/fig/782644" pageId="0" pageNumber="219">, 17B</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Rostra of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure19" httpUri="https://binary.pensoft.net/fig/782646" pageId="0" pageNumber="219">, 19B</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure21" httpUri="https://binary.pensoft.net/fig/782648" pageId="0" pageNumber="219">, 21B</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure23" httpUri="https://binary.pensoft.net/fig/782650" pageId="0" pageNumber="219">, 23B</figureCitation>
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Line drawings of the pronota of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Abbreviations: ca, calli; co, collar; ho, hood; lc, lateral carina; mc, median carina; pa, paranotum; pd, pronotal disc; pp, posterior process. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure25" httpUri="https://binary.pensoft.net/fig/782652" pageId="0" pageNumber="219">, 25B</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Line drawings of the pronota of four Stephanitis species from Japan, lateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov. Abbreviations: bu, buccula; ca, costal area; da, discoidal area; hl, hypocostal lamina; ho, hood; lc, lateral carina; mc, median carina; pa, paranotum. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure27" httpUri="https://binary.pensoft.net/fig/782654" pageId="0" pageNumber="219">, 27B</figureCitation>
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Line drawings of the pygophore of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D, E. S. (N.) hayashii sp. nov. from Kakeroma (D) and Aguni (E) Islands, central part of Ryukyu Islands; F. S. (N.) hiurai. Scale bars: 0.1 mm." figureDoi="10.3897/dez.69.89864.figure29" httpUri="https://binary.pensoft.net/fig/782656" pageId="0" pageNumber="219">, 29B</figureCitation>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Line drawings of the paramere of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D, E. S. (N.) hayashii sp. nov. from Kakeroma (D) and Aguni (E) Islands, central part of Ryukyu Islands; F. S. (N.) hiurai. Scale bars: 0.1 mm. Arrows indicate important morphological differences." figureDoi="10.3897/dez.69.89864.figure31" httpUri="https://binary.pensoft.net/fig/782658" pageId="0" pageNumber="219">, 31B</figureCitation>
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Three specimens of Stephanitis species from Japan. A. Holotype of S. (Norba) vitrea [= S. (N.) aperta] deposited in ELKU and its labels; B. Suspected syntype of S. (Stephanitis) propinqua [= S. (N.) aperta] deposited in Matsumura's collection of ELHU and its labels; C. Syntype of S. (N.) exigua [= S. (S.) pyrioides] deposited in Matsumura's collection of ELHU and its labels." figureDoi="10.3897/dez.69.89864.figure34" httpUri="https://binary.pensoft.net/fig/782661" pageId="0" pageNumber="219">, 34A, B</figureCitation>
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Living individuals of three species of Stephanitis from Japan: A-C. S. (Stephanitis) ambigua from Honshu, male (A) and fifth (B) and fourth (C) instar nymphs; D-G. S. (Norba) aperta from Honshu, male (D), female (E) and fifth (F) and fourth (G) instar nymphs; H, I. S. (N.) exigua from Okinawa Island, central part of Ryukyu Islands, male (H) and fifth instar nymph (I)." figureDoi="10.3897/dez.69.89864.figure40" httpUri="https://binary.pensoft.net/fig/782667" pageId="0" pageNumber="219">, 40D-G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="reference_group">
<paragraph pageId="0" pageNumber="219">
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Tingidae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">Stephanitis (Norba) aperta</taxonomicName>
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
, 1912: 335. Syntype(s): Japan: Sakuna [= Honshu, Chiba-ken, Sakuna of former Toyofusa-mura in early 20th Century (current Tateyama-shi, Sakuna)]; HNHM. 
<normalizedToken originalValue="“Sakuna”">&quot;Sakuna&quot;</normalizedToken>
was considered to be a misspelling of 
<normalizedToken originalValue="“Satsuma”">&quot;Satsuma&quot;</normalizedToken>
(
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="65 - 84" refId="B86" refString="Takeya, C, 1931. Some Tingitidae of the Japanese Empire. Mushi 4: 65 - 84" title="Some Tingitidae of the Japanese Empire." volume="4" year="1931">Takeya 1931</bibRefCitation>
: 77).
</paragraph>
<paragraph pageId="0" pageNumber="219">
<taxonomicName class="Insecta" family="Tingidae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) vitrea" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="vitrea" subGenus="Norba">Stephanitis (Norba) vitrea</taxonomicName>
Takeya, 1931: 74. Holotype (Fig. 
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Three specimens of Stephanitis species from Japan. A. Holotype of S. (Norba) vitrea [= S. (N.) aperta] deposited in ELKU and its labels; B. Suspected syntype of S. (Stephanitis) propinqua [= S. (N.) aperta] deposited in Matsumura's collection of ELHU and its labels; C. Syntype of S. (N.) exigua [= S. (S.) pyrioides] deposited in Matsumura's collection of ELHU and its labels." figureDoi="10.3897/dez.69.89864.figure34" httpUri="https://binary.pensoft.net/fig/782661" pageId="0" pageNumber="219">34A</figureCitation>
): Japan: Yakushima Is., Onoaida-Ambo [= Ryukyu Islands, Yakushima Island, between Onoaida and Anbo]; ELKU. Synonymised with 
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Tingidae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="Norba">Stephanitis (Norba) exigua</taxonomicName>
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
, 1912 by 
<bibRefCitation author="Takeya, C" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="219" pagination="167 - 176" refId="B90" refString="Takeya, C, 1953. Notes on the Tingidae of Shikoku, Japan (Hemiptera). Transactions of the Shikoku Entomological Society 3 (7): 167 - 176" title="Notes on the Tingidae of Shikoku, Japan (Hemiptera)." volume="3" year="1953">Takeya (1953</bibRefCitation>
: 168), with 
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Tingidae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">Stephanitis (Norba) aperta</taxonomicName>
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
, 1912 by 
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">Takeya (1963</bibRefCitation>
: 38).
</paragraph>
<paragraph pageId="0" pageNumber="219">
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Tingidae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua">Stephanitis exigua</taxonomicName>
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
, 1912: 
<bibRefCitation author="Takeya, C" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="219" pagination="167 - 176" refId="B90" refString="Takeya, C, 1953. Notes on the Tingidae of Shikoku, Japan (Hemiptera). Transactions of the Shikoku Entomological Society 3 (7): 167 - 176" title="Notes on the Tingidae of Shikoku, Japan (Hemiptera)." volume="3" year="1953">Takeya (1953</bibRefCitation>
: 168) (distribution: part). Misidentification (
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">Takeya 1963</bibRefCitation>
: 38).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="references">
<paragraph pageId="0" pageNumber="219">References.</paragraph>
<paragraph pageId="0" pageNumber="219">
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="65 - 84" refId="B86" refString="Takeya, C, 1931. Some Tingitidae of the Japanese Empire. Mushi 4: 65 - 84" title="Some Tingitidae of the Japanese Empire." volume="4" year="1931">Takeya (1931</bibRefCitation>
: 75) (distribution: part); 
<bibRefCitation author="Drake, CJ" journalOrPublisher="Notes d'Entomologie Chinoise" pageId="0" pageNumber="219" pagination="45 - 56" refId="B12" refString="Drake, CJ, 1948. New species of Stephanitis Stal including a list of species of the World. Notes d'Entomologie Chinoise 12 (6): 45 - 56" title="New species of Stephanitis Stal including a list of species of the World." volume="12" year="1948">Drake (1948</bibRefCitation>
: 54) (checklist: 
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Stephanitis</taxonomicName>
); 
<bibRefCitation author="Takeya, C" journalOrPublisher="Kurume University Journal" pageId="0" pageNumber="219" pagination="5 - 28" refId="B89" refString="Takeya, C, 1951b. A tentative list of the Tingidae of Japan and her adjacent territories (Hemiptera). Kurume University Journal 4 (1): 5 - 28" title="A tentative list of the Tingidae of Japan and her adjacent territories (Hemiptera)." volume="4" year="1951 b">Takeya (1951b</bibRefCitation>
: 13) (checklist: Japan); 
<bibRefCitation author="Drake, CJ" journalOrPublisher="Quarternary Journal of Taiwan Museum" pageId="0" pageNumber="219" pagination="87 - 101" refId="B14" refString="Drake, CJ, Maa, TC, 1953. Chinese and other Oriental Tingoidea (Hemiptera). Quarternary Journal of Taiwan Museum 6 (2): 87 - 101" title="Chinese and other Oriental Tingoidea (Hemiptera)." volume="6" year="1953">Drake and Maa (1953</bibRefCitation>
: 100) (checklist: 
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
); 
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">Takeya (1963</bibRefCitation>
: 38) (distribution: part); 
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="271 - 280" refId="B44" refString="Miyamoto, S, 1964a. Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera). Kontyu 32 (2): 271 - 280" title="Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 a">Miyamoto (1964a</bibRefCitation>
: 274) (distribution: part); 
<bibRefCitation DOI="https://doi.org/10.5479/si.03629236.243.1" author="Drake, CJ" journalOrPublisher="Iowa State College Journal of Science" pageId="0" pageNumber="219" refId="B18" refString="Drake, CJ, Ruhoff, FA, 1965. Lacebugs of the world, a catalog (Hemiptera: Tingidae). Bulletin of the United States National Museum 243: [i-viii +] 1-634[, pls. frontis piece + 1-56]. https://doi.org/10.5479/si.03629236.243.1" title="Lacebugs of the world, a catalog (Hemiptera: Tingidae). Bulletin of the United States National Museum 243: [i-viii +] 1 - 634 [, pls. frontis piece + 1 - 56]." url="https://doi.org/10.5479/si.03629236.243.1" year="1965">Drake and Ruhoff (1965</bibRefCitation>
: 366) (catalog); 
<bibRefCitation DOI="https://doi.org/10.5109/22784" author="Lee, CE" journalOrPublisher="Mushi" pageId="0" pageNumber="219" refId="B37" refString="Lee, CE, 1969. Morphological and phylogenic studies on the larvae and male genitalia of the East Asiatic Tingidae (Heteroptera). Journal of the Faculty of Agriculture, Kyushu University 15(2): 137-256[, pl. 1-16]. https://doi.org/10.5109/22784" title="Morphological and phylogenic studies on the larvae and male genitalia of the East Asiatic Tingidae (Heteroptera). Journal of the Faculty of Agriculture, Kyushu University 15 (2): 137 - 256 [, pl. 1 - 16]." url="https://doi.org/10.5109/22784" year="1969">Lee (1969</bibRefCitation>
: 246) (male genitalia); 
<bibRefCitation author="Jing, HL" journalOrPublisher="Niho-no-Mushi" pageId="0" pageNumber="219" refId="B31" refString="Jing, HL, 1981. Tingidae. In: Hsiao TY, Ren SZ, Zheng LY, Jing HL, Zou HG, Liu SL: A handbook for the determination of the Chinese Hemiptera-Heteroptera (Volume II). Science Press, Beijing, China, 271-368[, pls. 42-50]. [in Chinese with English summary]." title="Tingidae. In: Hsiao TY, Ren SZ, Zheng LY, Jing HL, Zou HG, Liu SL: A handbook for the determination of the Chinese Hemiptera-Heteroptera (Volume II). Science Press, Beijing, China, 271 - 368 [, pls. 42 - 50]. [in Chinese with English summary]." year="1981">Jing (1981</bibRefCitation>
: 349) (monograph); 
<bibRefCitation author="Miyamoto, S" editor="Hirashima, Y" journalOrPublisher="I. Laboratory of Entomology, Faculty of Agriculture, Kyushu University, Fukuoka, Japan" pageId="0" pageNumber="219" pagination="151 - 188" refId="B50" refString="Miyamoto, S, Yasunaga, T, 1989. Heteroptera. In: Hirashima, Y, Ed., A Check List of Japanese Insects. I. Laboratory of Entomology, Faculty of Agriculture, Kyushu University, Fukuoka, Japan: 151 - 188" title="Heteroptera." volumeTitle="A Check List of Japanese Insects." year="1989">Miyamoto and Yasunaga (1989</bibRefCitation>
: 168) (checklist: Japan); 
<bibRefCitation author="Yasunaga, T" journalOrPublisher=", Tokyo, Japan" pageId="0" pageNumber="219" refId="B114" refString="Yasunaga, T, Takai, M, Yamashita, I, Kawamura, M, Kawasawa, T, 1993. A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" title="A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd." year="1993">Yasunaga et al. (1993</bibRefCitation>
: 178) (monograph: part); 
<bibRefCitation author="Pericart, J" editor="Aukema, B" journalOrPublisher="Vol. 2, Cimicomorpha I. Netherlands Entomological Society, Amsterdam, Netherlands" pageId="0" pageNumber="219" pagination="3 - 78" refId="B57" refString="Pericart, J, Golub, VB, 1996. Superfamily Tingoidea Laporte, 1832. In: Aukema, B, Rieger, C, Eds., Catalogue of the Heteroptera of the Palaearctic Region. Vol. 2, Cimicomorpha I. Netherlands Entomological Society, Amsterdam, Netherlands: 3 - 78" title="Superfamily Tingoidea Laporte, 1832." volumeTitle="Catalogue of the Heteroptera of the Palaearctic Region." year="1996">
<normalizedToken originalValue="Péricart">Pericart</normalizedToken>
and Golub (1996
</bibRefCitation>
: 58) (catalogue: Palaearctic); 
<bibRefCitation author="Tomokuni, M" journalOrPublisher="Memoirs of the National Science Museum" pageId="0" pageNumber="219" pagination="35 - 55" refId="B104" refString="Tomokuni, M, Hayashi, M, Usui, T, 2000. Hemiptera excluding Sternorrhyncha from the Garden of the Imperial Palace, Tokyo, Japan. Memoirs of the National Science Museum 36: 35 - 55" title="Hemiptera excluding Sternorrhyncha from the Garden of the Imperial Palace, Tokyo, Japan." volume="36" year="2000">Tomokuni et al. (2000</bibRefCitation>
: 38) (distribution); 
<bibRefCitation author="Tomokuni, M" journalOrPublisher="Memoirs of the National Science Museum" pageId="0" pageNumber="219" pagination="397 - 408" refId="B98" refString="Tomokuni, M, 2005. Heteroptera (Insecta) from the Akasaka Imperial Gardens and the Tokiwamatsu Imperial Villa, Tokyo, Japan. Memoirs of the National Science Museum 39: 397 - 408" title="Heteroptera (Insecta) from the Akasaka Imperial Gardens and the Tokiwamatsu Imperial Villa, Tokyo, Japan." volume="39" year="2005">Tomokuni (2005</bibRefCitation>
: 400) (distribution); 
<bibRefCitation author="Tomokuni, M" journalOrPublisher="Memoirs of the National Science Museum" pageId="0" pageNumber="219" pagination="349 - 354" refId="B99" refString="Tomokuni, M, 2006a. Additional material of Heteroptera (Insecta) collected at the Imperial Palace, the Akasaka Imperial Gardens, and the Tokiwamatsu Imperial Villa, Tokyo Japan. Memoirs of the National Science Museum 43: 349 - 354" title="Additional material of Heteroptera (Insecta) collected at the Imperial Palace, the Akasaka Imperial Gardens, and the Tokiwamatsu Imperial Villa, Tokyo Japan." volume="43" year="2006 a">Tomokuni (2006a</bibRefCitation>
: 350) (distribution); 
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni (2012</bibRefCitation>
: 205) (monograph: part); 
<bibRefCitation author="Yiu, V" journalOrPublisher="Hong Kong Entomological Society, Hong Kong, China" pageId="0" pageNumber="219" refId="B115" refString="Yiu, V, Yip, CH, 2012. A photographic guide to Hong Kong true bugs. Hong Kong Entomological Society, Hong Kong, China" title="A photographic guide to Hong Kong true bugs." year="2012">Yiu and Yip (2012</bibRefCitation>
: 83) (distribution); 
<bibRefCitation author="Yano, S" journalOrPublisher="Bulletin of Omogo mountain museum" pageId="0" pageNumber="219" pagination="23 - 52" refId="B113" refString="Yano, S, Takechi, L, Kikuhara, Y, 2013. Heteroptera of Ehime Prefecture, Shikoku, Japan (Insecta, Hemiptera) I. Bulletin of Omogo mountain museum 5: 23 - 52" title="Heteroptera of Ehime Prefecture, Shikoku, Japan (Insecta, Hemiptera) I." volume="5" year="2013">Yano et al. (2013</bibRefCitation>
: 25) (distribution); 
<bibRefCitation author="Tomokuni, M" journalOrPublisher="Memoirs of the National Science Museum" pageId="0" pageNumber="219" pagination="359 - 369" refId="B101" refString="Tomokuni, M, 2014. Heteroptera from the Garden of the Imperial Palace, Tokyo, Japan. Memoirs of the National Science Museum 50: 359 - 369" title="Heteroptera from the Garden of the Imperial Palace, Tokyo, Japan." volume="50" year="2014">Tomokuni (2014</bibRefCitation>
: 362) (distribution); 
<bibRefCitation author="Nozaki, T" journalOrPublisher="Rostria" pageId="0" pageNumber="219" pagination="1 - 40" refId="B53" refString="Nozaki, T, Nozaki, Y, Uki, K, Tsukada, T, 2015. The heteropteran fauna of Shimokoshiki Island, Kagoshima Prefecture, Japan. Rostria 58: 1 - 40" title="The heteropteran fauna of Shimokoshiki Island, Kagoshima Prefecture, Japan." volume="58" year="2015">Nozaki et al. (2015</bibRefCitation>
: 9) (distribution); 
<bibRefCitation author="Yamada, K" editor="Hayashi, M" journalOrPublisher="Touka-shobo, Fukuoka, Japan" pageId="0" pageNumber="219" pagination="429 - 435" refId="B111" refString="Yamada, K, Ishikawa, T, 2016. Family Tingidae. In: Hayashi, M, Tomokuni, M, Yoshizawa, K, Ishikawa, T, Eds., Catalogue of the insects of Japan Volume 4 Paraneoptera. Touka-shobo, Fukuoka, Japan: 429 - 435" title="Family Tingidae." volumeTitle="Catalogue of the insects of Japan Volume 4 Paraneoptera." year="2016">Yamada and Ishikawa (2016</bibRefCitation>
: 433) (checklist: Japan); 
<bibRefCitation author="Maehara, S" journalOrPublisher="Insekuto" pageId="0" pageNumber="219" pagination="144 - 149" refId="B41" refString="Maehara, S, 2017. 栃木県で採集されたカメムシ類について [True bugs collected from Tochigi-ken, Honshu, Japan]. Insekuto 68 (2): 144 - 149" title="栃木県で採集されたカメムシ類について [True bugs collected from Tochigi-ken, Honshu, Japan]." volume="68" year="2017">Maehara (2017</bibRefCitation>
: 146) (distribution); 
<bibRefCitation author="Okochi, A" journalOrPublisher="Sukashiba" pageId="0" pageNumber="219" pagination="1 - 4" refId="B54" refString="Okochi, A, 2019. 山陰地方で採集したグンバイムシ [Tingidae collected from Sanin district, Honshu, Japan]. Sukashiba 66: 1 - 4" title="山陰地方で採集したグンバイムシ [Tingidae collected from Sanin district, Honshu, Japan]." volume="66" year="2019">Okochi (2019</bibRefCitation>
: 2) (distribution); 
<bibRefCitation author="Souma, J" journalOrPublisher="Fauna Ryukyuana" pageId="0" pageNumber="219" pagination="29 - 39" refId="B73" refString="Souma, J, 2021b. Lace bugs from Yakushima Island, Ryukyu Archipelago, Japan (Hemiptera: Heteroptera: Tingidae). Fauna Ryukyuana 63: 29 - 39" title="Lace bugs from Yakushima Island, Ryukyu Archipelago, Japan (Hemiptera: Heteroptera: Tingidae)." volume="63" year="2021 b">Souma (2021b</bibRefCitation>
: 31) (distribution).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="materials_examined">
<paragraph pageId="0" pageNumber="219">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="219">
<materialsCitation collectingDate="1929-08-03" collectingDateMax="2022-09-16" collectingDateMin="1929-08-03" collectorName="H. Hori. The, T. Mita, J. Souma, S. Maehara, A. Saito, M. Tomokuni, M. Takakuwa, K. Nakano, G. Mashima, Y. Tamadera, G. Yamamoto, T. Esaki, A. Yamasaki, S. Miyamoto, K. Goto, R. Ito, Yoshida, I. Hiura, N. Tsuji, Hirashima, Y. Miyatake, Y. Uehara, Hori, Y. Obae, N. Kaneko, K. Morimoto, Y. Watanabe, C. Takeya, Y. Hirashima, T. Saeki, H. Yoshitake, M. Sakai" country="JAPAN" county="Takeya" elevation="450" latitude="34.969086" location="Therefore" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" specimenCount="2631" specimenCount-female="1557" specimenCount-male="1074" stateProvince="Tochigi Pref." typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="219">
<typeStatus>Holotype</typeStatus>
of 
<taxonomicName class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) vitrea" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="vitrea" subGenus="Norba">Stephanitis (Norba) vitrea</taxonomicName>
<collectingCounty>Takeya</collectingCounty>
, 1931
</emphasis>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU) (Fig. 
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Three specimens of Stephanitis species from Japan. A. Holotype of S. (Norba) vitrea [= S. (N.) aperta] deposited in ELKU and its labels; B. Suspected syntype of S. (Stephanitis) propinqua [= S. (N.) aperta] deposited in Matsumura's collection of ELHU and its labels; C. Syntype of S. (N.) exigua [= S. (S.) pyrioides] deposited in Matsumura's collection of ELHU and its labels." figureDoi="10.3897/dez.69.89864.figure34" httpUri="https://binary.pensoft.net/fig/782661" pageId="0" pageNumber="219">34A</figureCitation>
), 
<collectingCountry name="Japan">JAPAN</collectingCountry>
: 
<collectingMunicipality>Ryukyu Islands</collectingMunicipality>
(northern part): &quot;Yakushima 
<normalizedToken originalValue="Onoaida-Ambô”">Onoaida-Ambo&quot;</normalizedToken>
[= Yakushima Island, 
<locationDeviation firstLocation="Onoaida" secondLocation="Anbo">between Onoaida and Anbo</locationDeviation>
(approximate coordinates: 
<geoCoordinate degrees="30" direction="north" minutes="16" orientation="latitude" precision="1" seconds="08.3" value="30.268972">30°16'08.3&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="130" direction="east" minutes="36" orientation="longitude" precision="1" seconds="44.2" value="130.61229">130°36'44.2&quot;E</geoCoordinate>
)], 
<collectingDate value="1929-08-03">3.viii.1929</collectingDate>
, leg. 
<collectorName>H. Hori. The</collectorName>
labels shown in Fig. 
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Three specimens of Stephanitis species from Japan. A. Holotype of S. (Norba) vitrea [= S. (N.) aperta] deposited in ELKU and its labels; B. Suspected syntype of S. (Stephanitis) propinqua [= S. (N.) aperta] deposited in Matsumura's collection of ELHU and its labels; C. Syntype of S. (N.) exigua [= S. (S.) pyrioides] deposited in Matsumura's collection of ELHU and its labels." figureDoi="10.3897/dez.69.89864.figure34" httpUri="https://binary.pensoft.net/fig/782661" pageId="0" pageNumber="219">34A</figureCitation>
were created after (
<collectorName>T. Mita</collectorName>
, pers. comm. 2021), but the condition of the specimen matched the photograph of the original description (
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="65 - 84" refId="B86" refString="Takeya, C, 1931. Some Tingitidae of the Japanese Empire. Mushi 4: 65 - 84" title="Some Tingitidae of the Japanese Empire." volume="4" year="1931">Takeya 1931</bibRefCitation>
). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:17B2C8D518696A55762089A3D14FF5A6" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Therefore" stateProvince="Tochigi Pref.">Therefore</location>
, the female individual seems to correspond to the 
<typeStatus>holotype</typeStatus>
of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) vitrea" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="vitrea" subGenus="N.">S. (N.) vitrea</taxonomicName>
. 
<emphasis bold="true" pageId="0" pageNumber="219">
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:F795DFBEFBBE85C6E28FAC3D722B4C31" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Suspected" stateProvince="Tochigi Pref.">Suspected</location>
<typeStatus>syntype</typeStatus>
of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) propinqua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="propinqua" subGenus="Stephanitis">S. (Stephanitis) propinqua</taxonomicName>
<normalizedToken originalValue="Horváth">
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:04CA6BD62ED1E1334559AA22EFEF2DEF" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Horvath" stateProvince="Tochigi Pref.">Horvath</location>
</normalizedToken>
, 1912
</emphasis>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELHU) (Fig. 
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Three specimens of Stephanitis species from Japan. A. Holotype of S. (Norba) vitrea [= S. (N.) aperta] deposited in ELKU and its labels; B. Suspected syntype of S. (Stephanitis) propinqua [= S. (N.) aperta] deposited in Matsumura's collection of ELHU and its labels; C. Syntype of S. (N.) exigua [= S. (S.) pyrioides] deposited in Matsumura's collection of ELHU and its labels." figureDoi="10.3897/dez.69.89864.figure34" httpUri="https://binary.pensoft.net/fig/782661" pageId="0" pageNumber="219">34B</figureCitation>
), 
<collectingCountry name="Japan">JAPAN</collectingCountry>
: 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:FB335F5E3862249076C3AF78C0AA5486" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Kyushu" stateProvince="Tochigi Pref.">Kyushu</location>
: 
<normalizedToken originalValue="“カゴシマ”">&quot;カゴシマ&quot;</normalizedToken>
[= Kagoshima-ken (approximate coordinates: 
<geoCoordinate degrees="31" direction="north" minutes="35" orientation="latitude" precision="1" seconds="59.5" value="31.599861">31°35'59.5&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="130" direction="east" minutes="32" orientation="longitude" precision="1" seconds="59.6" value="130.5499">130°32'59.6&quot;E</geoCoordinate>
)], 
<normalizedToken originalValue="“7/10”">&quot;7/10&quot;</normalizedToken>
[= 10.vii]. 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:71AD0A8D6790CA4481D6CB7D80AE535B" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="This" stateProvince="Tochigi Pref.">This</location>
specimen was labelled as &quot; 
<taxonomicName class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis gratiosa" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis gratiosa</emphasis>
</taxonomicName>
Horv.&quot; (apparently an unpublished name) and 
<normalizedToken originalValue="“カゴシマグンバイ”">&quot;カゴシマグンバイ&quot;</normalizedToken>
(unknown 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:E6849035D84F820637E3AFB7E8C431BF" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Japanese" stateProvince="Tochigi Pref.">Japanese</location>
name). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:8440233EDA7FCD0880126A1CBE36716D" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="According" stateProvince="Tochigi Pref.">According</location>
to the original description (
<bibRefCitation author="Horvath, G" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="219" pagination="319 - 339" refId="B26" refString="Horvath, G, 1912. Species generis Tingitidarum Stephanitis. Annales Historico-Naturales Musei Nationalis Hungarici 10: 319 - 339" title="Species generis Tingitidarum Stephanitis." volume="10" year="1912">
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
1912
</bibRefCitation>
), the 
<typeStatus>syntype</typeStatus>
(s) of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) propinqua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="propinqua" subGenus="S.">S. (S.) propinqua</taxonomicName>
was (were) deposited in ELHU and this female individual is the only specimen in 
<normalizedToken originalValue="Matsumura’s">
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:065C851D6DA09D3CFCF24512D81068FB" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Matsumura's" stateProvince="Tochigi Pref.">Matsumura's</location>
</normalizedToken>
collection of ELHU matching the label data of 
<typeStatus>syntype</typeStatus>
(s) of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) propinqua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="propinqua" subGenus="S.">S. (S.) propinqua</taxonomicName>
. 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:92A74183D24CFFB5A3250081582DBC15" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="However" stateProvince="Tochigi Pref.">However</location>
, the morphological characteristics of the specimen match the original description of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
(
<bibRefCitation author="Horvath, G" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="219" pagination="319 - 339" refId="B26" refString="Horvath, G, 1912. Species generis Tingitidarum Stephanitis. Annales Historico-Naturales Musei Nationalis Hungarici 10: 319 - 339" title="Species generis Tingitidarum Stephanitis." volume="10" year="1912">
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
1912
</bibRefCitation>
), but not that of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) propinqua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="propinqua" subGenus="S.">S. (S.) propinqua</taxonomicName>
. 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:615BDFD5D9F4904DF59CD76811E295A6" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Conversely" stateProvince="Tochigi Pref.">Conversely</location>
, the general habitus of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) propinqua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="propinqua" subGenus="S.">S. (S.) propinqua</taxonomicName>
recorded from Korea by previous studies (
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" refId="B87" refString="Takeya, C, 1932. Some Corean lace-bugs. Mushi 5: 8-13[, tab. 1]." title="Some Corean lace-bugs. Mushi 5: 8 - 13 [, tab. 1]." year="1932">Takeya 1932</bibRefCitation>
, 
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">1963</bibRefCitation>
) differs from that of all the 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:AB29B7489C9CA328CCF83D94CB5C1A49" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="East Asian" stateProvince="Tochigi Pref.">East Asian</location>
species of 
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
(present study). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:89C53E9C937FB6BBD902F176921153A4" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Therefore" stateProvince="Tochigi Pref.">Therefore</location>
, if there is no 
<typeStatus>syntype</typeStatus>
of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) propinqua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="propinqua" subGenus="S.">S. (S.) propinqua</taxonomicName>
in other institutions, then 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) propinqua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="propinqua" subGenus="S.">S. (S.) propinqua</taxonomicName>
should be synonymised with 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
and a new name should be given to &quot; 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) propinqua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="propinqua" subGenus="S.">S. (S.) propinqua</taxonomicName>
&quot; distributed in Korea. 
<emphasis bold="true" pageId="0" pageNumber="219">Non-types</emphasis>
(
<specimenCount type="male">534 ♂♂</specimenCount>
<specimenCount type="female">777 ♀♀</specimenCount>
16 nymphs), 
<collectingCountry name="Japan">JAPAN</collectingCountry>
: Honshu: Ibaraki-ken, Higashiibaraki-gun, Oarai-machi, Isohama-cho, 
<collectingDate value="2020-09-12">12.ix.2020</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">7 ♂♂</specimenCount>
<specimenCount type="female">13 ♀♀</specimenCount>
3 nymphs, TUA); 
<collectingRegion country="Japan" name="Tochigi">Tochigi Pref.</collectingRegion>
, Uchino, Watarase-yusuichi, 
<collectingDate value="2016-11-26">26.xi.2016</collectingDate>
, leg. 
<collectorName>S. Maehara</collectorName>
(
<specimenCount type="male">4 ♂♂</specimenCount>
<specimenCount type="female">2 ♀♀</specimenCount>
, TUA); as above but 
<collectingDate value="2020-10-02">2.x.2020</collectingDate>
(
<specimenCount type="male">4 ♂♂</specimenCount>
<specimenCount type="female">3 ♀♀</specimenCount>
, TUA); Chiba-ken, Tateyama-shi, Ohto (approximate coordinates: 
<geoCoordinate degrees="34" direction="north" minutes="58" orientation="latitude" precision="1" seconds="19.8" value="34.972168">34°58'19.8&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="139" direction="east" minutes="52" orientation="longitude" precision="1" seconds="33.9" value="139.87608">139°52'33.9&quot;E</geoCoordinate>
), 
<collectingDate value="2021-05-22">22.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, TUA); Chiba-ken, Tateyama-shi, Shimosanagura, 
<collectingDate value="2021-05-22">22.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">2 ♀♀</specimenCount>
, TUA); Chiba-ken, Tateyama-shi, Tateyama, 
<collectingDate value="2021-05-22">22.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">2 ♂♂</specimenCount>
<specimenCount type="female">6 ♀♀</specimenCount>
, TUA); as above but 
<collectingDate value="2021-05-23">23.v.2021</collectingDate>
(
<specimenCount type="female">4 ♀♀</specimenCount>
, ELKU; 
<specimenCount type="male">2 ♂♂</specimenCount>
<specimenCount type="female">2 ♀♀</specimenCount>
1 nymph, TUA); Chiba-ken, Tateyama-shi, Sunozaki, 
<collectingDate value="2021-05-23">23.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">11 ♂♂</specimenCount>
<specimenCount type="female">32 ♀♀</specimenCount>
, TUA); Chiba-ken, Tateyama-shi, Sunozaki Shrine, 
<collectingDate value="2021-05-23">23.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">2 ♀♀</specimenCount>
, TUA); Chiba-ken, Tateyama-shi, Masaki, 
<collectingDate value="2021-05-23">23.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">7 ♂♂</specimenCount>
<specimenCount type="female">14 ♀♀</specimenCount>
, TUA); Chiba-ken, Tateyama-shi, Higashinagata, 
<collectingDate value="2021-05-24">24.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, TUA); Chiba-ken, Tateyama-shi, Yamogi (approximate coordinates: 
<geoCoordinate degrees="34" direction="north" minutes="58" orientation="latitude" precision="1" seconds="08.7" value="34.969086">34°58'08.7&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="139" direction="east" minutes="53" orientation="longitude" precision="1" seconds="34.4" value="139.89288">139°53'34.4&quot;E</geoCoordinate>
), 
<collectingDate value="2021-05-24">24.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">2 ♂♂</specimenCount>
<specimenCount type="female">4 ♀♀</specimenCount>
, TUA); Chiba-ken, Minamiboso-shi, Chikura-cho, Minamiasai, 
<collectingDate value="2021-05-24">24.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, TUA); Chiba-ken, Kamogawa-shi, Yomogi, 
<collectingDate value="2021-05-25">25.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, TUA); 
<collectingRegion country="Japan" name="Tokyo">Tokyo</collectingRegion>
Met., Imperial Palace, 
<collectingDate value="1996-06-05">5.vi.1996</collectingDate>
, leg. 
<collectorName>A. Saito</collectorName>
(
<specimenCount type="male">7 ♂♂</specimenCount>
<specimenCount type="female">7 ♀♀</specimenCount>
, NSMT); as above but 
<collectingDate value="1996-10-16">16.x.1996</collectingDate>
, leg. 
<collectorName>M. Tomokuni</collectorName>
(
<specimenCount type="male">8 ♂♂</specimenCount>
<specimenCount type="female">15 ♀♀</specimenCount>
, NSMT); 
<collectingDate value="1997-11-19">19.xi.1997</collectingDate>
, leg. 
<collectorName>M. Tomokuni</collectorName>
(
<specimenCount type="male">9 ♂♂</specimenCount>
<specimenCount type="female">6 ♀♀</specimenCount>
, NSMT); as above but 
<collectingDate value="1998-05-25">25.v.1998</collectingDate>
(
<specimenCount type="female">2 ♀♀</specimenCount>
, NSMT); as above but 
<collectingDate value="1998-05-28">28.v.1998</collectingDate>
, leg. 
<collectorName>M. Tomokuni</collectorName>
(
<specimenCount type="male">4 ♂♂</specimenCount>
<specimenCount type="female">3 ♀♀</specimenCount>
, NSMT); as above but 
<collectingDate value="2009-07-16">16.vii.2009</collectingDate>
, leg. 
<collectorName>M. Tomokuni</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, NSMT); as above but 
<collectingDate value="2012-05-28">28.v.2012</collectingDate>
, leg. 
<collectorName>M. Tomokuni</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, NSMT); as above but 
<collectingDate value="2012-07-23">23.vii.2012</collectingDate>
, leg. 
<collectorName>M. Tomokuni</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">2 ♀♀</specimenCount>
, NSMT); as above but 
<collectingDate value="2012-09-10">10.ix.2012</collectingDate>
, leg. 
<collectorName>M. Tomokuni</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, NSMT); Tokyo-to, Minato-ku, Shiba-koen, 
<collectingDate value="2022-07-08">8.vii.2022</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">4 ♂♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, TUA); Kanagawa-ken, Sagamihara-shi, Minami-ku, Shimomizo, 
<collectingDate value="2021-11-15">15.xi.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">9 ♂♂</specimenCount>
<specimenCount type="female">8 ♀♀</specimenCount>
3 nymphs, TUA); Kanagawa-ken, Sagamihara-shi, Chuo-ku, Tanashioda, 
<collectingDate value="2019-05-19">19.v.2019</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">2 ♀♀</specimenCount>
, TUA); Kanagawa-ken, Sagamihara-shi, Chuo-ku, Tana, 
<collectingDate value="2019-05-19">19.v.2019</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">22 ♂♂</specimenCount>
<specimenCount type="female">16 ♀♀</specimenCount>
, TUA); as above but 
<collectingDate value="2021-11-17">17.xi.2021</collectingDate>
(
<specimenCount type="male">9 ♂♂</specimenCount>
<specimenCount type="female">11 ♀♀</specimenCount>
, TUA); Kanagawa-ken, Sagamihara-shi, Midori-ku, Oshima, 
<collectingDate value="2021-11-17">17.xi.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">3 ♀♀</specimenCount>
1 nymph, TUA); Kanagawa-ken, Zama-shi, Iriyanishi, 
<collectingDate value="2022-01-01">1.i.2022</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">14 ♀♀</specimenCount>
, TUA); Kanagawa-ken, Ebina-shi, Kamiimaizumi, 
<collectingDate value="2022-01-01">1.i.2022</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, TUA); Kanagawa-ken, Atsugi-shi, Funako, 
<collectingDate value="2017-05-29">29.v.2017</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">37 ♂♂</specimenCount>
<specimenCount type="female">41 ♀♀</specimenCount>
, TUA); as above but 
<collectingDate value="2017-05-30">30.v.2017</collectingDate>
(
<specimenCount type="male">32 ♂♂</specimenCount>
<specimenCount type="female">30 ♀♀</specimenCount>
, TUA); as above but 
<collectingDate value="2017-05-31">31.v.2017</collectingDate>
(
<specimenCount type="female">4 ♀♀</specimenCount>
, ELKU; 
<specimenCount type="male">52 ♂♂</specimenCount>
<specimenCount type="female">55 ♀♀</specimenCount>
, TUA); as above but 
<collectingDate value="2017-06-02">2.vi.2017</collectingDate>
(
<specimenCount type="male">6 ♂♂</specimenCount>
<specimenCount type="female">16 ♀♀</specimenCount>
, TUA); as above but 
<collectingDate value="2017-06-05">5.vi.2017</collectingDate>
(
<specimenCount type="male">30 ♂♂</specimenCount>
<specimenCount type="female">17 ♀♀</specimenCount>
, TUA); Kanagawa-ken, Atsugi-shi, Nanasawa, 
<collectingDate value="2017-06-06">6.vi.2017</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, TUA); Kanagawa-ken, Aiko-gun, Aikawa-machi, Nakatsu, 
<collectingDate value="2021-05-26">26.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">3 ♂♂</specimenCount>
<specimenCount type="female">11 ♀♀</specimenCount>
, TUA); as above but 
<collectingDate value="2021-11-15">15.xi.2021</collectingDate>
(
<specimenCount type="female">3 ♀♀</specimenCount>
, TUA); Kanagawa-ken, Yokohama-shi, Isogo-ku, Negishihachiman Shrine, 
<collectingDate value="1999-05-31">31.v.1999</collectingDate>
, leg. 
<collectorName>M. Takakuwa</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, KPMNH); Kanagawa-ken, Yokohama-shi, Kanazawa-ku, Noukendaimori, 
<collectingDate value="2017-06-15">15.vi.2017</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">3 ♂♂</specimenCount>
, TUA); Kanagawa-ken, Yokosuka-shi, Kamoi, 
<collectingDate value="2017-06-27">27.vi.2017</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">4 ♀♀</specimenCount>
, TUA); Kanagawa-ken, Chigasaki-shi, Yanagishima, 
<collectingDate value="2019-06-01">1.vi.2019</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">10 ♂♂</specimenCount>
<specimenCount type="female">7 ♀♀</specimenCount>
, TUA); Kanagawa-ken, Hadano-shi, Tsurumaki, 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:44B494CFC6DADF6C6C5BE63EBDCB9785" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Mt. Azuma" stateProvince="Tochigi Pref.">Mt. Azuma</location>
, 
<collectingDate value="2021-11-19">19.xi.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">2 ♂♂</specimenCount>
<specimenCount type="female">5 ♀♀</specimenCount>
, TUA); Kanagawa-ken, Hadano-shi, Soya, 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:E9689C57C45573ED9BCFAAAE792E27E4" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Mt. Kobo" stateProvince="Tochigi Pref.">Mt. Kobo</location>
, 
<collectingDate value="2021-11-19">19.xi.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">4 ♀♀</specimenCount>
, TUA); Kanagawa-ken, Ashigarashimo-gun, Manazuru-machi, Manazuru, 
<collectingDate value="2021-05-23">23.v.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">11 ♂♂</specimenCount>
<specimenCount type="female">30 ♀♀</specimenCount>
, TUA); Niigata-ken, Niigata-shi, Nishi-ku, Igarashi-2no-cho, 
<collectingDate value="2015-07-19">19.vii.2015</collectingDate>
, leg. 
<collectorName>K. Nakano</collectorName>
(
<specimenCount type="female">2 ♀♀</specimenCount>
, TUA); as above but 
<collectingDate value="2021-06-16">16.vi.2021</collectingDate>
, leg. 
<collectorName>G. Mashima</collectorName>
(
<specimenCount type="male">12 ♂♂</specimenCount>
<specimenCount type="female">14 ♀♀</specimenCount>
, TUA); Niigata-ken, Niigata-shi, Chuo-ku, Sekiya, Nishikaigan Park, 
<collectingDate value="2016-10-22">22.x.2016</collectingDate>
, leg. 
<collectorName>K. Nakano</collectorName>
(
<specimenCount type="male">4 ♂♂</specimenCount>
<specimenCount type="female">10 ♀♀</specimenCount>
, TUA); Niigata-ken, Niigata-shi, Chuo-ku, Seigoro, Toyanogata Park, 
<collectingDate value="2016-10-30">30.x.2016</collectingDate>
, leg. 
<collectorName>K. Nakano</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">4 ♀♀</specimenCount>
, TUA); Niigata-ken, Niigata-shi, Higashi-ku, Matsuzono, 
<collectingDate value="2019-07-21">21.vii.2019</collectingDate>
, leg. 
<collectorName>K. Nakano</collectorName>
(
<specimenCount type="male">2 ♂♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, TUA); Niigata-ken, Niigata-shi, Nishi-ku, Akatsuka, Sakata, 
<collectingDate value="2019-10-09">9.x.2019</collectingDate>
, leg. 
<collectorName>K. Nakano</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">3 ♀♀</specimenCount>
, TUA); Niigata-ken, Niigata-shi, Kita-ku, Nigorikawa, 
<collectingDate value="2020-06-07">7.vi.2020</collectingDate>
, leg. 
<collectorName>K. Nakano</collectorName>
(
<specimenCount type="female">5 ♀♀</specimenCount>
, TUA); Niigata-ken, Niigata-shi, Chuo-ku, Nishifunami-cho, 
<collectingDate value="2021-06-12">12.vi.2021</collectingDate>
, leg. 
<collectorName>G. Mashima</collectorName>
(
<specimenCount type="male">12 ♂♂</specimenCount>
<specimenCount type="female">17 ♀♀</specimenCount>
, TUA); Niigata-ken, Niigata-shi, Chuo-ku, Bandaijima, 
<collectingDate value="2021-08-30">30.viii.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">18 ♂♂</specimenCount>
<specimenCount type="female">10 ♀♀</specimenCount>
, TUA); Shizuoka-ken, Shimoda-shi, Suzaki, Tsumekizaki, 
<collectingDate value="2020-07-21">21.vii.2020</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
, TUA); Shizuoka-ken, Hamamatsu-shi, Nishi-ku, Kamigaya-cho, 
<collectingDate value="2017-06-16">16.vi.2017</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">15 ♂♂</specimenCount>
<specimenCount type="female">33 ♀♀</specimenCount>
, TUA); Shizuoka-ken, Numazu-shi, Kamikanuki, Higashihongo-cho, 
<collectingDate value="2021-12-27">27.xii.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, TUA); Yamaguchi-ken, Shimonoseki-shi, Mimosusogawa-cho, 
<collectingDate value="2022-09-16">16.ix.2022</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">3 ♀♀</specimenCount>
, ELKU). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:0AE856AE72B74FF04BE54947B9CD7B29" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Izu Islands" stateProvince="Tochigi Pref.">Izu Islands</location>
(northern part): Izu-Oshima Island: Okada, Minatono-mieru-oka, 
<collectingDate value="2019-06-04">4.vi.2019</collectingDate>
, leg. 
<collectorName>Y. Tamadera</collectorName>
(
<specimenCount type="male">19 ♂♂</specimenCount>
<specimenCount type="female">37 ♀♀</specimenCount>
, TUA). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:1A923A388EE788D470A62ED57D24464E" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Jogashima Island" stateProvince="Tochigi Pref.">Jogashima Island</location>
: 
<collectingDate value="2019-06-04">4.vi.2019</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">11 ♂♂</specimenCount>
<specimenCount type="female">9 ♀♀</specimenCount>
, TUA). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:F8BA00DEAD646E5E84A96C6EE30504C0" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Ebisu Island" stateProvince="Tochigi Pref.">Ebisu Island</location>
: 
<collectingDate value="2020-07-21">21.vii.2020</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">3 ♂♂</specimenCount>
, TUA). Shikoku: Ehime Pref., Kashima, 
<collectingDate value="1971-06-02">2.vi.1971</collectingDate>
, leg. 
<collectorName>M. Tomokuni</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, NSMT); Tosa, Nakagawa, 
<collectingDate value="1953-08-19">19.viii.1953</collectingDate>
, leg. 
<collectorName>G. Yamamoto</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
, ELKU); Kochi Pref., Cape Ashizuri, 
<collectingDate value="1971-06-07">7.vi.1971</collectingDate>
, leg. 
<collectorName>M. Tomokuni</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, NSMT); Kochi-ken, Kochi-shi, Hitsuzan-cho, 
<collectingDate value="2020-06-30">30.vi.2020</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, TUA). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:2124E9E302B627B215E3AA66595D541C" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Okinoshima Island" stateProvince="Tochigi Pref.">Okinoshima Island</location>
(Kochi Prefecture): 
<collectingDate value="1951-08-11">11.viii.1951</collectingDate>
, leg. 
<collectorName>T. Esaki</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU; 
<specimenCount type="female">1 ♀</specimenCount>
, KUM). Kyushu: Prov. Buzen, Kokura, 
<collectingDate value="1951-11-20">20.xi.1951</collectingDate>
, leg. 
<collectorName>A. Yamasaki</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
, KUM); Fukuoka, Tachibanayama, 
<collectingDate value="1961-07-23">23.vii.1961</collectingDate>
, leg. 
<collectorName>S. Miyamoto</collectorName>
(
<specimenCount type="male">6 ♂♂</specimenCount>
<specimenCount type="female">8 ♀♀</specimenCount>
, KUM); as above but 
<collectingDate value="1961-09-08">8.ix.1961</collectingDate>
(
<specimenCount type="female">1 ♀</specimenCount>
, KUM); Fukuoka-ken, Fukuoka-shi, Nishi-ku, Motooka, Kyushu University, 
<collectingDate value="2020-05-23">23.v.2020</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); Fukuoka-ken, Itoshima-shi, Tomari, 
<collectingDate value="2022-05-16">16.v.2022</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); as above but 
<collectingDate value="2022-07-10">10.vii.2022</collectingDate>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); Kumamoto-Pref., Kumamoto-City, Kuwamizuhonmachi, 
<collectingDate value="2021-01-04">4.i.2021</collectingDate>
, leg. 
<collectorName>K. Goto</collectorName>
(
<specimenCount type="female">2 ♀♀</specimenCount>
, ELKU); 
<normalizedToken originalValue="Ôita">Oita</normalizedToken>
Pref., Saiki-shi, Yonouzu, Tsurumisaki, 
<collectingDate value="2017-07-02">2.vii.2017</collectingDate>
, leg. 
<collectorName>R. Ito</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); 
<normalizedToken originalValue="Ôita">Oita</normalizedToken>
Pref., Saiki-shi, Kamiura, Niinameura, 
<collectingDate value="2020-07-19">19.vii.2020</collectingDate>
, leg. 
<collectorName>R. Ito</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">3 ♀♀</specimenCount>
, ELKU); Miyazaki Pref., Nichinan-shi, Miyaura, 
<collectingDate value="2018-05-12">12.v.2018</collectingDate>
, leg. 
<collectorName>R. Ito</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); Miyazaki Pref., 
<normalizedToken originalValue="Hyûga-shi">Hyuga-shi</normalizedToken>
, Okuragahama, 
<collectingDate value="2019-06-01">1.vi.2019</collectingDate>
, leg. 
<collectorName>R. Ito</collectorName>
(
<specimenCount type="female">2 ♀♀</specimenCount>
, ELKU); Kagoshima, 
<collectingDate value="1953-05-21">21.v.1953</collectingDate>
, leg. 
<collectorName>S. Miyamoto</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, KUM); Oosumi, Izashiki~Ootomari, 
<collectingDate value="1953-05-25">25.v.1953</collectingDate>
, leg. 
<collectorName>Yoshida</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
, ELKU); Osumi, Sata, 
<collectingDate value="1953-05-29">29.v.1953</collectingDate>
, leg. 
<collectorName>Yoshida</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); Osumi, Sata Cape, 
<collectingDate value="1953-05-30">30.v.1953</collectingDate>
, leg. 
<collectorName>I. Hiura</collectorName>
(
<specimenCount type="female">4 ♀♀</specimenCount>
, KUM); Kagoshima-ken, Kagoshima-shi, Shiroyama-cho, 
<collectingDate value="2017-07-04">4.vii.2017</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
, ELKU); Kagoshima, Minamiosumi-T., 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:34B19DC2F47F2B1037C659D5805B0C27" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Sugiyama-dani Valley" stateProvince="Tochigi Pref.">Sugiyama-dani Valley</location>
, 
<collectingDate value="2017-07-31">31.vii.2017</collectingDate>
, leg. 
<collectorName>N. Tsuji</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); Kagoshima Pref., 
<normalizedToken originalValue="Minamiôsumi-chô">Minamiosumi-cho</normalizedToken>
, Sata, Hetsuka, 
<collectingDate value="2020-05-30">30.v.2020</collectingDate>
, leg. 
<collectorName>R. Ito</collectorName>
(
<specimenCount type="male">9 ♂♂</specimenCount>
<specimenCount type="female">6 ♀♀</specimenCount>
, ELKU). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:96B0CF248C363507B3B222B4B9508598" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Okinoshima Island" stateProvince="Tochigi Pref.">Okinoshima Island</location>
(Fukuoka Prefecture): 
<collectingDate value="1958-07-25" valueMax="1958-07-28" valueMin="1958-07-25">25-28.vii.1958</collectingDate>
, leg. 
<collectorName>Hirashima</collectorName>
, Murakami &amp; 
<collectorName>Y. Miyatake</collectorName>
(
<specimenCount type="male">35 ♂♂</specimenCount>
<specimenCount type="female">70 ♀♀</specimenCount>
, ELKU; 
<specimenCount type="female">1 ♀</specimenCount>
, KUM). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:61AE583C53AF4231F1C334A175E8049D" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Tsushima Island" stateProvince="Tochigi Pref.">Tsushima Island</location>
: Izuhara-machi, Kitazato, Kamisaka, 
<collectingDate value="2022-07-27">27.vii.2022</collectingDate>
, leg. 
<collectorName>Y. Uehara</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:1CB4F01AEAEE649FCD7600D75884B633" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Amakusa Islands" stateProvince="Tochigi Pref.">Amakusa Islands</location>
: 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:5750A7BDC5FD1C81DAC1FBEC09D521AF" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Shimoshima Island" stateProvince="Tochigi Pref.">Shimoshima Island</location>
: Tomioka, 
<collectingDate value="1931-09-12">12.ix.1931</collectingDate>
, leg. 
<collectorName>Hori</collectorName>
&amp; 
<normalizedToken originalValue="Chô">Cho</normalizedToken>
(
<specimenCount type="male">2 ♂♂</specimenCount>
<specimenCount type="female">4 ♀♀</specimenCount>
, ELKU; 
<specimenCount type="female">1 ♀</specimenCount>
, KUM). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:A7DF2AF4F8BD21C809875658530A5A11" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Goto Islands" stateProvince="Tochigi Pref.">Goto Islands</location>
: 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:74E44561C5D74AB7168822EE8F2B9D11" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Fukue Island" stateProvince="Tochigi Pref.">Fukue Island</location>
: 
<collectingDate value="1962-09-01">1. ix.1962</collectingDate>
, leg. 
<collectorName>S. Miyamoto</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, KUM); as above but leg. 
<collectorName>S. Miyamoto</collectorName>
&amp; Kawarabata (
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); Inuyamaze, 
<collectingDate value="1962-09-02">2.ix.1962</collectingDate>
, leg. 
<collectorName>S. Miyamoto</collectorName>
(
<specimenCount type="male">2 ♂♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, KUM); as above but (
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, KUM); Arakawa, 
<collectingDate value="1962-09-03">3.ix.1962</collectingDate>
, leg. 
<collectorName>S. Miyamoto</collectorName>
(
<specimenCount type="female">2 ♀♀</specimenCount>
, KUM). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:77726827CD6171ABD24322A3E16B67EB" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Sakura Island" stateProvince="Tochigi Pref.">Sakura Island</location>
: Kurokami-cho, 
<collectingDate value="2021-07-27">27.vii.2021</collectingDate>
, leg. 
<collectorName>Y. Obae</collectorName>
(
<specimenCount type="male">18 ♂♂</specimenCount>
<specimenCount type="female">13 ♀♀</specimenCount>
, TUA). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:01EFE10C36CCEA8EF75A6576D19C5C58" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Koshiki Islands" stateProvince="Tochigi Pref.">Koshiki Islands</location>
: 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:CBB3202E092D3EC21374203BFB059A58" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Kamikoshiki Island" stateProvince="Tochigi Pref.">Kamikoshiki Island</location>
: Nakano, 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:4F4DCD851400846FE6AA7F9F5D3F7592" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Mt. Tomeki" stateProvince="Tochigi Pref.">Mt. Tomeki</location>
, 
<collectingDate value="2019-05-02">2.v.2019</collectingDate>
, leg. 
<collectorName>N. Kaneko</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">4 ♀♀</specimenCount>
1 nymph, TUA). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:AE718BCC51F0BC809708E0B499E5A783" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Shimokoshiki Island" stateProvince="Tochigi Pref.">Shimokoshiki Island</location>
: Teuchi, 
<collectingDate value="1960-08-27" valueMax="1960-08-29" valueMin="1960-08-27">27-29.viii.1960</collectingDate>
, leg. 
<collectorName>K. Morimoto</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">2 ♀♀</specimenCount>
, ELKU); 
<collectingDate value="1975-05-25">25.v.1975</collectingDate>
, leg. 
<collectorName>Y. Watanabe</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, TUA). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:8BDB1AA08E4930C9269AAFCAFD3A51CA" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Ryukyu Islands" stateProvince="Tochigi Pref.">Ryukyu Islands</location>
(northern part): 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:60A015D54D15D20FC41F3E729B0B4E37" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Tanegashima Island" stateProvince="Tochigi Pref.">Tanegashima Island</location>
: Nishinoomote, 
<collectingDate value="1952-08-31">31.viii.1952</collectingDate>
, leg. 
<collectorName>C. Takeya</collectorName>
&amp; 
<collectorName>Y. Hirashima</collectorName>
(
<specimenCount type="male">2 ♂♂</specimenCount>
, ELKU; 
<specimenCount type="male">2 ♂♂</specimenCount>
2 nymphs, KUM); Nakatane-cho, Masuda, 
<collectingDate value="2021-07-10">10.vii.2021</collectingDate>
, leg. 
<collectorName>T. Saeki</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, TUA). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:7FC2D464850F1400CF5429E10C6C6C6E" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Yakushima Island" stateProvince="Tochigi Pref.">Yakushima Island</location>
: 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:4005E572A1F4BB3BC36C6B2776295758" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Onoaida" stateProvince="Tochigi Pref.">Onoaida</location>
, 
<collectingDate value="1952-08-26">26.viii.1952</collectingDate>
, leg. 
<collectorName>C. Takeya</collectorName>
&amp; 
<collectorName>Y. Hirashima</collectorName>
(
<specimenCount type="male">3 ♂♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); as above but 
<collectingDate value="1952-08-27">27.viii.1952</collectingDate>
(
<specimenCount type="male">8 ♂♂</specimenCount>
<specimenCount type="female">11 ♀♀</specimenCount>
1 nymph, ELKU); Miyanoura, 
<collectingDate value="1952-08-28">28.viii.1952</collectingDate>
, leg. 
<collectorName>C. Takeya</collectorName>
&amp; 
<collectorName>Y. Hirashima</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">4 ♀♀</specimenCount>
, ELKU); as above but 
<collectingDate value="2022-05-18">18.v.2022</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, TUA); 
<normalizedToken originalValue="Shiratani-unsuikyô">Shiratani-unsuikyo</normalizedToken>
, alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" unit="m" value="450.0" valueMax="600.0" valueMin="300.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" unit="m" value="450.0" valueMax="600.0" valueMin="300.0">300-600 m</elevation>
</quantity>
, 
<collectingDate value="2017-07-14">14.vii.2017</collectingDate>
, leg. 
<collectorName>R. Ito</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
, ELKU); Kurio, 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:A14A5E3A84D0800CAB35E2D772F9EE96" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Koyojigawa For. Rd." stateProvince="Tochigi Pref.">Koyojigawa For. Rd.</location>
, 
<collectingDate value="2021-07-07">7.vii.2021</collectingDate>
, leg. 
<collectorName>T. Saeki</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
, ELKU); Kurio, 
<collectingDate value="2021-08-15">15.viii.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">3 ♂♂</specimenCount>
, ELKU); Koseda, 
<collectingDate value="2021-08-17">17.viii.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
, ELKU); as above but 
<collectingDate value="2022-05-19">19.v.2022</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">10 ♂♂</specimenCount>
<specimenCount type="female">14 ♀♀</specimenCount>
, TUA); Funayuki, 
<collectingDate value="2021-08-17">17.viii.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); Hirauchi, 
<collectingDate value="2021-08-19">19.viii.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">4 ♀♀</specimenCount>
, ELKU); 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:37FB97F8CF554E7B4BEA221E47D2EAC0" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Anbo" stateProvince="Tochigi Pref.">Anbo</location>
, 
<collectingDate value="2021-08-20">20.viii.2021</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">9 ♂♂</specimenCount>
<specimenCount type="female">14 ♀♀</specimenCount>
, ELKU); Tabugawa, 
<collectingDate value="2022-05-18">18.v.2022</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">3 ♀♀</specimenCount>
, TUA). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:36492117B6624AB7D4CC5B716BE2A52F" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Nakanoshima Island" stateProvince="Tochigi Pref.">Nakanoshima Island</location>
: 
<collectingDate value="1953-06-03" valueMax="1953-06-13" valueMin="1953-06-03">3-13.vi.1953</collectingDate>
, leg. 
<collectorName>S. Miyamoto</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); Okizaki, 
<collectingDate value="2017-07-05">5.vii.2017</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="female">2 ♀♀</specimenCount>
, TUA); as above but 
<collectingDate value="2017-07-06">6.vii.2017</collectingDate>
(
<specimenCount type="female">1 ♀</specimenCount>
, TUA); Kusuki, 
<collectingDate value="2017-07-07">7.vii.2017</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">4 ♂♂</specimenCount>
<specimenCount type="female">7 ♀♀</specimenCount>
, TUA). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:10478133DF3097259AE6C53FE921D67D" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Taira Island" stateProvince="Tochigi Pref.">Taira Island</location>
: 
<normalizedToken originalValue="Shûraku">Shuraku</normalizedToken>
, 
<collectingDate value="2016-10-08" valueMax="2016-10-10" valueMin="2016-10-08">8-10.x.2016</collectingDate>
, leg. 
<collectorName>H. Yoshitake</collectorName>
(
<specimenCount type="female">3 ♀♀</specimenCount>
, TUA); Higashinohama, 
<collectingDate value="2016-10-08" valueMax="2016-10-10" valueMin="2016-10-08">8-10.x.2016</collectingDate>
, leg. 
<collectorName>H. Yoshitake</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, TUA). 
<location LSID="urn:lsid:plazi:treatment:9A2E144DDC53560E8C1AD97BF089175A:752203B969EA277B39AD51536D771113" country="JAPAN" county="Takeya" latitude="34.969086" longLatPrecision="1" longitude="139.89288" municipality="Ryukyu Islands" name="Akuseki Island" stateProvince="Tochigi Pref.">Akuseki Island</location>
: 
<collectingDate value="1971-04-24">24.iv.1971</collectingDate>
, leg. 
<collectorName>M. Sakai</collectorName>
(
<specimenCount type="male">2 ♂♂</specimenCount>
<specimenCount type="female">3 ♀♀</specimenCount>
4 nymphs, NSMT); 
<collectingDate value="2016-06-17">17.vi.2016</collectingDate>
, leg. 
<collectorName>H. Yoshitake</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, NIAES); Yudomari, 
<collectingDate value="2017-07-08">8.vii.2017</collectingDate>
, leg. 
<collectorName>J. Souma</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">3 ♀♀</specimenCount>
, TUA). Seven specimens collected from 
<normalizedToken originalValue="“Ohto”">&quot;Ohto&quot;</normalizedToken>
and 
<normalizedToken originalValue="“Yamogi”">&quot;Yamogi&quot;</normalizedToken>
adjacent to the type locality 
<normalizedToken originalValue="“Sakuna”">&quot;Sakuna&quot;</normalizedToken>
well match the original description of 
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">Stephanitis (Norba) aperta</taxonomicName>
(
<bibRefCitation author="Horvath, G" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="219" pagination="319 - 339" refId="B26" refString="Horvath, G, 1912. Species generis Tingitidarum Stephanitis. Annales Historico-Naturales Musei Nationalis Hungarici 10: 319 - 339" title="Species generis Tingitidarum Stephanitis." volume="10" year="1912">
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
1912
</bibRefCitation>
). In the present study, the author identified 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
based on these seven individuals
</materialsCitation>
. 
<typeStatus>Syntype</typeStatus>
(s) of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
exist(s) in the collection of HNHM (D. 
<normalizedToken originalValue="Rédei">Redei</normalizedToken>
, pers. comm. 2021).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="diagnosis">
<paragraph pageId="0" pageNumber="219">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="219">
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">Stephanitis (Norba) aperta</taxonomicName>
is recognised amongst other species of 
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
by a combination of the following characters: head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown (Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure7" httpUri="https://binary.pensoft.net/fig/782634" pageId="0" pageNumber="219">7B</figureCitation>
, 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure9" httpUri="https://binary.pensoft.net/fig/782636" pageId="0" pageNumber="219">9B</figureCitation>
, 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure11" httpUri="https://binary.pensoft.net/fig/782638" pageId="0" pageNumber="219">11B</figureCitation>
, 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure13" httpUri="https://binary.pensoft.net/fig/782640" pageId="0" pageNumber="219">13B</figureCitation>
, 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Male hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure15" httpUri="https://binary.pensoft.net/fig/782642" pageId="0" pageNumber="219">15B</figureCitation>
, 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Female hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure17" httpUri="https://binary.pensoft.net/fig/782644" pageId="0" pageNumber="219">17B</figureCitation>
, 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Rostra of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure19" httpUri="https://binary.pensoft.net/fig/782646" pageId="0" pageNumber="219">19B</figureCitation>
, 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure21" httpUri="https://binary.pensoft.net/fig/782648" pageId="0" pageNumber="219">21B</figureCitation>
, 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure23" httpUri="https://binary.pensoft.net/fig/782650" pageId="0" pageNumber="219">23B</figureCitation>
); calli dark brown; body in male 2.1 times (in female 2.0 times) as long as maximum width across hemelytra (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male habitus of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua from Honshu; B. S. (Norba) aperta from Honshu; C. S. (N.) exigua from Minamidaito Island, Daito Islands; D. S. (N.) hayashii sp. nov. from Senaga Island, central part of Ryukyu Islands; E, F. S. (N.) hiurai from Kakeroma (E) and Takara (F) islands, central part of Ryukyu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure2" httpUri="https://binary.pensoft.net/fig/782629" pageId="0" pageNumber="219">2B</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female habitus of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua from Honshu; B. S. (Norba) aperta from Honshu; C. S. (N.) exigua from Okinawa Island, central part of Ryukyu Islands; D. S. (N.) hayashii sp. nov. from Aguni Island, central part of Ryukyu Islands; E, F. S. (N.) hiurai from Okinawa (E) and Takara (F) Islands, central part of Ryukyu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure4" httpUri="https://binary.pensoft.net/fig/782631" pageId="0" pageNumber="219">4B</figureCitation>
, 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Teratological forms of two Stephanitis species from Japan, dorsal view: A. S. (Norba) aperta from Izu-Oshima Island, northern part of Izu Islands; B. S. (Stephanitis) takeyai from Kyushu. Arrows indicate antennae with segmental oligomery and malformed paranotum. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure6" httpUri="https://binary.pensoft.net/fig/782633" pageId="0" pageNumber="219">6A</figureCitation>
); rostrum not reaching metasternum; pronotum unicarinate (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Line drawings of the pronota of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Abbreviations: ca, calli; co, collar; ho, hood; lc, lateral carina; mc, median carina; pa, paranotum; pd, pronotal disc; pp, posterior process. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure25" httpUri="https://binary.pensoft.net/fig/782652" pageId="0" pageNumber="219">25B</figureCitation>
); hood pale, shorter than median carina of pronotum, as wide as vertex at widest part, not covering eye, as high as median carina of pronotum at highest part, with posterior margin not extending to middle of pronotal disc; median carina of pronotum with 2 rows of areolae at highest part; pronotal disc opaque; paranotum less erect, narrowed posteriorly, with 3 rows of areolae at widest part, with anterolateral angle protruding anteriad, with outer margin gently curved inwards at posterolateral angle, maximum height shorter than height of eye (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Line drawings of the pronota of four Stephanitis species from Japan, lateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov. Abbreviations: bu, buccula; ca, costal area; da, discoidal area; hl, hypocostal lamina; ho, hood; lc, lateral carina; mc, median carina; pa, paranotum. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure27" httpUri="https://binary.pensoft.net/fig/782654" pageId="0" pageNumber="219">27B</figureCitation>
); apices of hemelytra close to each other in rest; costal area with 3-4 rows of areolae at widest part; subcostal area in male with 2 rows (in female with 3 rows) of areolae at widest part; discoidal area with 3-4 rows of areolae at widest part; sutural area with 3-4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R+M (radiomedial) vein carinate; pygophore elevated at centre of venter, with posterior margin slightly emarginate in middle part (Fig. 
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Line drawings of the pygophore of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D, E. S. (N.) hayashii sp. nov. from Kakeroma (D) and Aguni (E) Islands, central part of Ryukyu Islands; F. S. (N.) hiurai. Scale bars: 0.1 mm." figureDoi="10.3897/dez.69.89864.figure29" httpUri="https://binary.pensoft.net/fig/782656" pageId="0" pageNumber="219">29B</figureCitation>
); and paramere stout, weakly curved inwards at apex, with outer margin not sinuate in middle part, inner margin nearly straight in basal part (Fig. 
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Line drawings of the paramere of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D, E. S. (N.) hayashii sp. nov. from Kakeroma (D) and Aguni (E) Islands, central part of Ryukyu Islands; F. S. (N.) hiurai. Scale bars: 0.1 mm. Arrows indicate important morphological differences." figureDoi="10.3897/dez.69.89864.figure31" httpUri="https://binary.pensoft.net/fig/782658" pageId="0" pageNumber="219">31B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="remarks">
<paragraph pageId="0" pageNumber="219">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="219">
Amongst the Japanese species of 
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">S. (Norba) aperta</taxonomicName>
is similar to 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
in general habitus, but it is easily distinguished by the following characters: calli dark brown (light brown in 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
) (Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure7" httpUri="https://binary.pensoft.net/fig/782634" pageId="0" pageNumber="219">7B, C</figureCitation>
, 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure9" httpUri="https://binary.pensoft.net/fig/782636" pageId="0" pageNumber="219">9B, C</figureCitation>
, 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure11" httpUri="https://binary.pensoft.net/fig/782638" pageId="0" pageNumber="219">11B, C</figureCitation>
, 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure13" httpUri="https://binary.pensoft.net/fig/782640" pageId="0" pageNumber="219">13B, C</figureCitation>
); pronotal disc opaque (lustrous in 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
); paranotum with 3 rows of areolae at widest part (2 rows in 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
), with anterolateral angle protruding anteriad (slightly protruding in 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
); and subcostal area of hemelytron in female with 3 rows of areolae at widest part (2 rows in 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
) (Figs 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Male hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure15" httpUri="https://binary.pensoft.net/fig/782642" pageId="0" pageNumber="219">15B, C</figureCitation>
, 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Female hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure17" httpUri="https://binary.pensoft.net/fig/782644" pageId="0" pageNumber="219">17B, C</figureCitation>
). The place name 
<normalizedToken originalValue="“Sakuna”">&quot;Sakuna&quot;</normalizedToken>
was considered to be a misspelling of 
<normalizedToken originalValue="“Satsuma”">&quot;Satsuma&quot;</normalizedToken>
[= Kyushu, Kagoshima-ken, former Satsuma-gun in the early 20th century (current Satsumasendai-shi and Satsuma-cho)] by 
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="65 - 84" refId="B86" refString="Takeya, C, 1931. Some Tingitidae of the Japanese Empire. Mushi 4: 65 - 84" title="Some Tingitidae of the Japanese Empire." volume="4" year="1931">Takeya (1931)</bibRefCitation>
. However, the former is the name of an actual place in Honshu. The present author confirms the occurrence of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
in 
<normalizedToken originalValue="“Ohto”">&quot;Ohto&quot;</normalizedToken>
and 
<normalizedToken originalValue="“Yamogi”">&quot;Yamogi&quot;</normalizedToken>
, adjacent to 
<normalizedToken originalValue="“Sakuna”">&quot;Sakuna&quot;</normalizedToken>
(see material examined). Therefore, 
<normalizedToken originalValue="“Sakuna”">&quot;Sakuna&quot;</normalizedToken>
seems to correspond to the type locality of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="teratological form">
<paragraph pageId="0" pageNumber="219">Teratological form.</paragraph>
<paragraph pageId="0" pageNumber="219">
The segmental oligomery of the antenna was confirmed in 
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">Stephanitis (Norba) aperta</taxonomicName>
, and one examined specimen lacks the left antennal segment IV (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Teratological forms of two Stephanitis species from Japan, dorsal view: A. S. (Norba) aperta from Izu-Oshima Island, northern part of Izu Islands; B. S. (Stephanitis) takeyai from Kyushu. Arrows indicate antennae with segmental oligomery and malformed paranotum. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure6" httpUri="https://binary.pensoft.net/fig/782633" pageId="0" pageNumber="219">6A</figureCitation>
), as reported in many tingids (
<bibRefCitation author="Stusak, JM" journalOrPublisher="Scientiae Biologicae" pageId="0" pageNumber="219" pagination="89 - 105" refId="B78" refString="Stusak, JM, Stehlik, JL, 1978. Second contribution to the teratology of Tingidae (Heteroptera) Antennal anomalies. Acta Musei Moraviae. Scientiae Biologicae 63: 89 - 105" title="Second contribution to the teratology of Tingidae (Heteroptera) Antennal anomalies. Acta Musei Moraviae." volume="63" year="1978">
<normalizedToken originalValue="Štusák">Stusak</normalizedToken>
and 
<normalizedToken originalValue="Stehlík">Stehlik</normalizedToken>
1978
</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4868.2.8" author="Souma, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="219" pagination="284 - 294" refId="B67" refString="Souma, J, 2020b. Taxonomic study of the subgenus Tropidocheila of the lace bug genus Tingis (Hemiptera: Heteroptera: Tingidae) from Japan, with description of Tingis (Tropidocheila) yamaboushi sp. nov. collected from Cornus kousa (Cornaceae). Zootaxa 4868 (2): 284 - 294, DOI: https://doi.org/10.11646/zootaxa.4868.2.8" title="Taxonomic study of the subgenus Tropidocheila of the lace bug genus Tingis (Hemiptera: Heteroptera: Tingidae) from Japan, with description of Tingis (Tropidocheila) yamaboushi sp. nov. collected from Cornus kousa (Cornaceae)." url="https://doi.org/10.11646/zootaxa.4868.2.8" volume="4868" year="2020 b">Souma 2020b</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.37520/aemnp.2020.035" author="Souma, J" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="219" pagination="527 - 536" refId="B69" refString="Souma, J, 2020d. The monocotyledon-feeding lace bugs of the genus Agramma from Japan (Hemiptera: Heteroptera: Tingidae). Acta Entomologica Musei Nationalis Pragae 60 (2): 527 - 536, DOI: https://doi.org/10.37520/aemnp.2020.035" title="The monocotyledon-feeding lace bugs of the genus Agramma from Japan (Hemiptera: Heteroptera: Tingidae)." url="https://doi.org/10.37520/aemnp.2020.035" volume="60" year="2020 d">2020d</bibRefCitation>
, 
<bibRefCitation author="Souma, J" journalOrPublisher="Rostria" pageId="0" pageNumber="219" pagination="34 - 37" refId="B70" refString="Souma, J, 2020e. Notes on habitat, 5th instar nymph, teratological form and emergence of Tingis (Tingis) lasiocera Matsumura, 1907 (Heteroptera: Tingidae). Rostria 64: 34 - 37" title="Notes on habitat, 5 th instar nymph, teratological form and emergence of Tingis (Tingis) lasiocera Matsumura, 1907 (Heteroptera: Tingidae)." volume="64" year="2020 e">2020e</bibRefCitation>
). Additionally, the right paranotum of this teratological individual is shorter than that of its normal left side, with two rows of areolae at the widest part (three rows on the left side).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="distribution">
<paragraph pageId="0" pageNumber="219">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="219">
Japan (Honshu; Izu Islands (northern part): Izu-Oshima Island; Jogashima Island; Ebisu Island; Shikoku; Okinoshima Island (Kochi Prefecture); Kyushu; Okinoshima Island (Fukuoka Prefecture); Tsushima Island; Amakusa Islands: Shimoshima Island; Goto Islands: Fukue Island; Sakura Island; Koshiki Islands: Kamikoshiki Island, Shimokoshiki Island; Ryukyu Islands (northern part): Tanegashima Island, Yakushima Island, Nakanoshima Island, Taira Island, Akuseki Island) (Fig. 
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Collection sites of Stephanitis (Stephanitis) ambigua, S. (Norba) aperta and S. (N.) exigua used in the present study. Red-filled areas = localities; circles = small isolated islands." figureDoi="10.3897/dez.69.89864.figure45" httpUri="https://binary.pensoft.net/fig/782672" pageId="0" pageNumber="219">45</figureCitation>
) (
<bibRefCitation author="Horvath, G" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="219" pagination="319 - 339" refId="B26" refString="Horvath, G, 1912. Species generis Tingitidarum Stephanitis. Annales Historico-Naturales Musei Nationalis Hungarici 10: 319 - 339" title="Species generis Tingitidarum Stephanitis." volume="10" year="1912">
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
1912
</bibRefCitation>
; 
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="65 - 84" refId="B86" refString="Takeya, C, 1931. Some Tingitidae of the Japanese Empire. Mushi 4: 65 - 84" title="Some Tingitidae of the Japanese Empire." volume="4" year="1931">Takeya 1931</bibRefCitation>
, 
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">1963</bibRefCitation>
; 
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni 2012</bibRefCitation>
; 
<bibRefCitation author="Yamada, K" editor="Hayashi, M" journalOrPublisher="Touka-shobo, Fukuoka, Japan" pageId="0" pageNumber="219" pagination="429 - 435" refId="B111" refString="Yamada, K, Ishikawa, T, 2016. Family Tingidae. In: Hayashi, M, Tomokuni, M, Yoshizawa, K, Ishikawa, T, Eds., Catalogue of the insects of Japan Volume 4 Paraneoptera. Touka-shobo, Fukuoka, Japan: 429 - 435" title="Family Tingidae." volumeTitle="Catalogue of the insects of Japan Volume 4 Paraneoptera." year="2016">Yamada and Ishikawa 2016</bibRefCitation>
; present study). Previous records from China in the 20th century (
<bibRefCitation DOI="https://doi.org/10.5479/si.03629236.243.1" author="Drake, CJ" journalOrPublisher="Iowa State College Journal of Science" pageId="0" pageNumber="219" refId="B18" refString="Drake, CJ, Ruhoff, FA, 1965. Lacebugs of the world, a catalog (Hemiptera: Tingidae). Bulletin of the United States National Museum 243: [i-viii +] 1-634[, pls. frontis piece + 1-56]. https://doi.org/10.5479/si.03629236.243.1" title="Lacebugs of the world, a catalog (Hemiptera: Tingidae). Bulletin of the United States National Museum 243: [i-viii +] 1 - 634 [, pls. frontis piece + 1 - 56]." url="https://doi.org/10.5479/si.03629236.243.1" year="1965">Drake and Ruhoff 1965</bibRefCitation>
; 
<bibRefCitation author="Jing, HL" journalOrPublisher="Niho-no-Mushi" pageId="0" pageNumber="219" refId="B31" refString="Jing, HL, 1981. Tingidae. In: Hsiao TY, Ren SZ, Zheng LY, Jing HL, Zou HG, Liu SL: A handbook for the determination of the Chinese Hemiptera-Heteroptera (Volume II). Science Press, Beijing, China, 271-368[, pls. 42-50]. [in Chinese with English summary]." title="Tingidae. In: Hsiao TY, Ren SZ, Zheng LY, Jing HL, Zou HG, Liu SL: A handbook for the determination of the Chinese Hemiptera-Heteroptera (Volume II). Science Press, Beijing, China, 271 - 368 [, pls. 42 - 50]. [in Chinese with English summary]." year="1981">Jing 1981</bibRefCitation>
) do not list the examined specimens and appear to be erroneous. Judging from the photographs, a recent record from Hong Kong (
<bibRefCitation author="Yiu, V" journalOrPublisher="Hong Kong Entomological Society, Hong Kong, China" pageId="0" pageNumber="219" refId="B115" refString="Yiu, V, Yip, CH, 2012. A photographic guide to Hong Kong true bugs. Hong Kong Entomological Society, Hong Kong, China" title="A photographic guide to Hong Kong true bugs." year="2012">Yiu and Yip 2012</bibRefCitation>
) corresponds to another species, as the pronotum has lateral carina. Therefore, the presence of 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
in China remains unconfirmed. The previous record from the central part of the Ryukyu Islands (
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="523 - 528" refId="B45" refString="Miyamoto, S, 1964b. Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera). Kontyu 32 (4): 523 - 528" title="Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 b">Miyamoto 1964b</bibRefCitation>
; 
<bibRefCitation author="Azuma, S" journalOrPublisher="The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" refId="B4" refString="Azuma, S, Kinjo, M, 1987. Check-list of the insects of Okinawa. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" title="Check-list of the insects of Okinawa." year="1987">Azuma and Kinjo 1987</bibRefCitation>
; 
<bibRefCitation author="Hayashi, M" editor="Azuma, S" journalOrPublisher=")]. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" pagination="127 - 150" refId="B24" refString="Hayashi, M, 2002. Heteroptera. In: Azuma, S, Yafuso, M, Kinjo, M, Hayashi, M, Kohama, T, Sasaki, T, Kimura, M, Kawamura, F, Eds., 沖縄産昆虫目録 [Check List of the Insect of the Ryukyu Islands (2nd Edn.)]. The Biological Society of Okinawa, Nishihara, Okinawa, Japan: 127 - 150" title="Heteroptera." volumeTitle="沖縄産昆虫目録 [Check List of the Insect of the Ryukyu Islands (2 nd Edn." year="2002">Hayashi 2002</bibRefCitation>
) corresponds to 
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="Norba">Stephanitis (Norba) exigua</taxonomicName>
, 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov. or 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" subGenus="N.">S. (N.) hiurai</taxonomicName>
. The previous records from the southern part of the Ryukyu Islands and northern Taiwan (
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">Takeya 1963</bibRefCitation>
; 
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="271 - 280" refId="B44" refString="Miyamoto, S, 1964a. Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera). Kontyu 32 (2): 271 - 280" title="Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 a">Miyamoto 1964a</bibRefCitation>
; 
<bibRefCitation author="Azuma, S" journalOrPublisher="The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" refId="B4" refString="Azuma, S, Kinjo, M, 1987. Check-list of the insects of Okinawa. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" title="Check-list of the insects of Okinawa." year="1987">Azuma and Kinjo 1987</bibRefCitation>
; 
<bibRefCitation author="Hayashi, M" editor="Azuma, S" journalOrPublisher=")]. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" pagination="127 - 150" refId="B24" refString="Hayashi, M, 2002. Heteroptera. In: Azuma, S, Yafuso, M, Kinjo, M, Hayashi, M, Kohama, T, Sasaki, T, Kimura, M, Kawamura, F, Eds., 沖縄産昆虫目録 [Check List of the Insect of the Ryukyu Islands (2nd Edn.)]. The Biological Society of Okinawa, Nishihara, Okinawa, Japan: 127 - 150" title="Heteroptera." volumeTitle="沖縄産昆虫目録 [Check List of the Insect of the Ryukyu Islands (2 nd Edn." year="2002">Hayashi 2002</bibRefCitation>
) correspond to 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="N.">S. (N.) ishikawai</taxonomicName>
sp. nov., described below. Hundreds of specimens from the central and southern parts of the Ryukyu Islands possessing the unicarinate pronotum were examined, but all of them belong to 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
, 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov., 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" subGenus="N.">S. (N.) hiurai</taxonomicName>
or 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="N.">S. (N.) ishikawai</taxonomicName>
sp. nov. Therefore, 
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
is probably not distributed in the central and southern parts of Ryukyu Islands. 
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">Stephanitis (Norba) aperta</taxonomicName>
inhabits the laurilignosa in a temperate climate of Japan proper (Honshu, Shikoku and Kyushu) and its surrounding islands and the northern part of the Izu and Ryukyu Islands, which is in the Palaearctic Region.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="host">
<paragraph pageId="0" pageNumber="219">Host plants.</paragraph>
<paragraph pageId="0" pageNumber="219">
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Cinnamomum" kingdom="Plantae" lsidName="Cinnamomum camphora" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="camphora">
<emphasis italics="true" pageId="0" pageNumber="219">Cinnamomum camphora</emphasis>
</taxonomicName>
(L.) J.Presl, 
<normalizedToken originalValue="“Kusunoki”">&quot;Kusunoki&quot;</normalizedToken>
(
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
) (Fig. 
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Lauraceous host plants of six species of Stephanitis from Japan: A. Lindera glauca from Honshu, damaged by S. (Stephanitis) ambigua; Machilus thunbergii (B), Neolitsea sericea (C) and Cinnamomum camphora (D) from Honshu, all damaged by S. (Norba) aperta; E. M. thunbergii from Okinawa Island, damaged by S. (N.) exigua; Litsea japonica (F) from Kakeroma Island and C. yabunikkei (G) from Tokashiki Island, both damaged by S. (N.) hayashii sp. nov.; H. M. thunbergii from Amami-Oshima Island, damaged by S. (N.) hiurai; I. Lit. japonica from Miyako Island, damaged by S. (N.) ishikawai sp. nov." figureDoi="10.3897/dez.69.89864.figure43" httpUri="https://binary.pensoft.net/fig/782670" pageId="0" pageNumber="219">43D</figureCitation>
) (
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">Takeya 1963</bibRefCitation>
; 
<bibRefCitation author="Yasunaga, T" journalOrPublisher=", Tokyo, Japan" pageId="0" pageNumber="219" refId="B114" refString="Yasunaga, T, Takai, M, Yamashita, I, Kawamura, M, Kawasawa, T, 1993. A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" title="A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd." year="1993">Yasunaga et al. 1993</bibRefCitation>
; 
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni 2012</bibRefCitation>
; present study); 
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Neolitsea" kingdom="Plantae" lsidName="Neolitsea sericea" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="sericea">
<emphasis italics="true" pageId="0" pageNumber="219">Neolitsea sericea</emphasis>
</taxonomicName>
(Blume) Koidz., 
<normalizedToken originalValue="“Shirodamo”">&quot;Shirodamo&quot;</normalizedToken>
(Fig. 
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Lauraceous host plants of six species of Stephanitis from Japan: A. Lindera glauca from Honshu, damaged by S. (Stephanitis) ambigua; Machilus thunbergii (B), Neolitsea sericea (C) and Cinnamomum camphora (D) from Honshu, all damaged by S. (Norba) aperta; E. M. thunbergii from Okinawa Island, damaged by S. (N.) exigua; Litsea japonica (F) from Kakeroma Island and C. yabunikkei (G) from Tokashiki Island, both damaged by S. (N.) hayashii sp. nov.; H. M. thunbergii from Amami-Oshima Island, damaged by S. (N.) hiurai; I. Lit. japonica from Miyako Island, damaged by S. (N.) ishikawai sp. nov." figureDoi="10.3897/dez.69.89864.figure43" httpUri="https://binary.pensoft.net/fig/782670" pageId="0" pageNumber="219">43C</figureCitation>
) (present study); 
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Machilus" kingdom="Plantae" lsidName="Machilus thunbergii" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="thunbergii">
<emphasis italics="true" pageId="0" pageNumber="219">Machilus thunbergii</emphasis>
</taxonomicName>
Siebold et Zucc., 
<normalizedToken originalValue="“Tabunoki”">&quot;Tabunoki&quot;</normalizedToken>
(
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
) (Fig. 
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Lauraceous host plants of six species of Stephanitis from Japan: A. Lindera glauca from Honshu, damaged by S. (Stephanitis) ambigua; Machilus thunbergii (B), Neolitsea sericea (C) and Cinnamomum camphora (D) from Honshu, all damaged by S. (Norba) aperta; E. M. thunbergii from Okinawa Island, damaged by S. (N.) exigua; Litsea japonica (F) from Kakeroma Island and C. yabunikkei (G) from Tokashiki Island, both damaged by S. (N.) hayashii sp. nov.; H. M. thunbergii from Amami-Oshima Island, damaged by S. (N.) hiurai; I. Lit. japonica from Miyako Island, damaged by S. (N.) ishikawai sp. nov." figureDoi="10.3897/dez.69.89864.figure43" httpUri="https://binary.pensoft.net/fig/782670" pageId="0" pageNumber="219">43B</figureCitation>
) (
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">Takeya 1963</bibRefCitation>
; 
<bibRefCitation author="Yasunaga, T" journalOrPublisher=", Tokyo, Japan" pageId="0" pageNumber="219" refId="B114" refString="Yasunaga, T, Takai, M, Yamashita, I, Kawamura, M, Kawasawa, T, 1993. A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" title="A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd." year="1993">Yasunaga et al. 1993</bibRefCitation>
; 
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni 2012</bibRefCitation>
; 
<bibRefCitation author="Okochi, A" journalOrPublisher="Sukashiba" pageId="0" pageNumber="219" pagination="1 - 4" refId="B54" refString="Okochi, A, 2019. 山陰地方で採集したグンバイムシ [Tingidae collected from Sanin district, Honshu, Japan]. Sukashiba 66: 1 - 4" title="山陰地方で採集したグンバイムシ [Tingidae collected from Sanin district, Honshu, Japan]." volume="66" year="2019">Okochi 2019</bibRefCitation>
; present study). 
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">Stephanitis (Norba) aperta</taxonomicName>
feeds only on lauraceous trees and is oligophagous. This lace bug was also collected from 
<taxonomicName class="Magnoliopsida" family="Symplocaceae" genus="Symplocos" kingdom="Plantae" lsidName="Symplocos glauca" order="Ericales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="glauca">
<emphasis italics="true" pageId="0" pageNumber="219">Symplocos glauca</emphasis>
</taxonomicName>
(Thunb.) Koidz., 
<normalizedToken originalValue="“Mimizubai”">&quot;Mimizubai&quot;</normalizedToken>
(
<taxonomicName class="Magnoliopsida" family="Symplocaceae" kingdom="Plantae" lsidName="" order="Ericales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="family">Symplocaceae</taxonomicName>
), without any data on its development (
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">Takeya 1963</bibRefCitation>
; 
<bibRefCitation author="Yasunaga, T" journalOrPublisher=", Tokyo, Japan" pageId="0" pageNumber="219" refId="B114" refString="Yasunaga, T, Takai, M, Yamashita, I, Kawamura, M, Kawasawa, T, 1993. A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" title="A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd." year="1993">Yasunaga et al. 1993</bibRefCitation>
; 
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni 2012</bibRefCitation>
). This lace bug sometimes occurs on plantings of 
<taxonomicName lsidName="C. camphora" pageId="0" pageNumber="219" rank="species" species="camphora">
<emphasis italics="true" pageId="0" pageNumber="219">C. camphora</emphasis>
</taxonomicName>
and 
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="M. thunbergii" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="thunbergii">
<emphasis italics="true" pageId="0" pageNumber="219">M. thunbergii</emphasis>
</taxonomicName>
in its distribution range (present study), suggesting that it can become a pest of both lauraceous trees.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="biology_ecology">
<paragraph pageId="0" pageNumber="219">Biology.</paragraph>
<paragraph pageId="0" pageNumber="219">
<taxonomicName authorityName="Horvath" authorityYear="1912" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">Stephanitis (Norba) aperta</taxonomicName>
feeds on the abaxial surface of leaves of the three known host plants (present study). Adults were collected in almost all seasons (
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="65 - 84" refId="B86" refString="Takeya, C, 1931. Some Tingitidae of the Japanese Empire. Mushi 4: 65 - 84" title="Some Tingitidae of the Japanese Empire." volume="4" year="1931">Takeya 1931</bibRefCitation>
, 
<bibRefCitation author="Takeya, C" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="219" pagination="167 - 176" refId="B90" refString="Takeya, C, 1953. Notes on the Tingidae of Shikoku, Japan (Hemiptera). Transactions of the Shikoku Entomological Society 3 (7): 167 - 176" title="Notes on the Tingidae of Shikoku, Japan (Hemiptera)." volume="3" year="1953">1953</bibRefCitation>
; 
<bibRefCitation author="Yasunaga, T" journalOrPublisher=", Tokyo, Japan" pageId="0" pageNumber="219" refId="B114" refString="Yasunaga, T, Takai, M, Yamashita, I, Kawamura, M, Kawasawa, T, 1993. A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" title="A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd." year="1993">Yasunaga et al. 1993</bibRefCitation>
; 
<bibRefCitation author="Yano, S" journalOrPublisher="Bulletin of Omogo mountain museum" pageId="0" pageNumber="219" pagination="23 - 52" refId="B113" refString="Yano, S, Takechi, L, Kikuhara, Y, 2013. Heteroptera of Ehime Prefecture, Shikoku, Japan (Insecta, Hemiptera) I. Bulletin of Omogo mountain museum 5: 23 - 52" title="Heteroptera of Ehime Prefecture, Shikoku, Japan (Insecta, Hemiptera) I." volume="5" year="2013">Yano et al. 2013</bibRefCitation>
; present study); nymphs were collected in April, May, August, September and November (present study); the overwintering stage is represented by the adult (present study).
</paragraph>
</subSubSection>
</treatment>
</document>