172 lines
19 KiB
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172 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang & Liang, Hongbin" docDate="2013" docId="92DD8AA1FC5294C12BD4C9A3E5308E62" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Anchista nubila Andrewes 1931" docType="treatment" docVersion="5" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="29" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="28" updateTime="1668155586682" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shi, Hongliang</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China & Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zhou, Hongzhang</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liang, Hongbin</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-04-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>284</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>129</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-284-1</mods:identifier>
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<mods:identifier type="ZooBank">33B15A74746D4A82A865EA1E7E55A9BB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">192EFF970E2BFFDF3161FFE53C4BFFAA</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578323</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152042278" LSID="urn:lsid:plazi:treatment:92DD8AA1FC5294C12BD4C9A3E5308E62" httpUri="http://treatment.plazi.org/id/92DD8AA1FC5294C12BD4C9A3E5308E62" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
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<subSubSection pageId="27" pageNumber="28" type="nomenclature">
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<paragraph pageId="27" pageNumber="28">
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<taxonomicName LSID="92DD8AA1-FC52-94C1-2BD4-C9A3E5308E62" authority="Andrewes, 1931" authorityName="Andrewes" authorityYear="1931" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Anchista nubila" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="nubila">Anchista nubila Andrewes, 1931</taxonomicName>
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<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Habitus, scale bars = 2.0 mm: 7 Anchista fenestrata subpubescens Chaudoir, a specimen from Pakistan 8 Anchista nubila Andrewes, a specimen from India 9 Metallanchista perlaeta (Kirschenhofer), a specimen from Sumatra 10 Metallanchista perlaeta (Kirschenhofer), a specimen from Java 11 Physodera dejeani Eschscholtz, a specimen from Yunnan 12 Physodera eschscholtzii Parry, a specimen from Yunnan." httpUri="https://binary.pensoft.net/fig/29672" pageId="27" pageNumber="28">Habitus: Figs 8</figureCitation>
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<figureCitation captionStart="Figures 37–42" captionStartId="F7" captionText="Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes." httpUri="https://binary.pensoft.net/fig/29680" pageId="27" pageNumber="28">42</figureCitation>
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<figureCitation captionStart="Figures 43–48" captionStartId="F8" captionText="Figures 43 - 48. Type materials, scale bars = 2.0 mm: 43 Holotype of Anchista pilosa sp. n. 44 Holotype of Metallanchista laticollis sp. n. 45 Holotype of Diamella kaszabi Jedlicka [= Diamella kaszabi (Jedlicka)] 46 Lectotype of Calleida cupreomicans Oberthuer [= Diamella cupreomicans (Oberthuer)] 47 Lectotype of Allocota aerata Bates [= Diamella cupreomicans (Oberthuer)] 48 Holotype of Lachnoderma arrowi Jedlicka [= Diamella arrowi (Jedlicka)]." httpUri="https://binary.pensoft.net/fig/29681" pageId="27" pageNumber="28">Habitus: male genitalia: Fig. 73</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="reference_group">
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<paragraph pageId="27" pageNumber="28">
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<taxonomicName authorityName="Andrewes" authorityYear="1931" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="nubila">Anchista nubila</taxonomicName>
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<bibRefCitation author="Andrewes, HE" journalOrPublisher="Zoologische Mededeelingen, Leiden" pageId="85" pageNumber="86" publicationUrl="10.1080/00222933108673342" refId="B15" refString="Andrewes, HE, 1931. Papers on Oriental Carabidae. XXV. The Annals and Magazine of Natural History (10) 7: 513-528. ., 10.1080/00222933108673342" title="Papers on Oriental Carabidae. XXV. The Annals and Magazine of Natural History (10) 7: 513 - 528." url="10.1080/00222933108673342" year="1931">Andrewes 1931</bibRefCitation>
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: 526 (original:
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="27" pageNumber="28">Anchista</emphasis>
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</taxonomicName>
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; type locality: Dehra Dun; lectotype deposited in NHML);
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<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen III. Carabidae III. Berlin" pageId="86" pageNumber="87" pagination="1279 - 1598" refId="B37" refString="Csiki, E, 1932. Carabidae: Harpalinae VII (Pars 124). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen III. Carabidae III. Berlin: 1279 - 1598" title="Carabidae: Harpalinae VII (Pars 124)." volumeTitle="Coleopterorum catalogus." year="1932">Csiki 1932</bibRefCitation>
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: 1456 (
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="27" pageNumber="28">Anchista</emphasis>
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</taxonomicName>
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, catalogue).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="type examined">
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<paragraph pageId="27" pageNumber="28">Type examined.</paragraph>
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<paragraph pageId="27" pageNumber="28">
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<emphasis bold="true" pageId="27" pageNumber="28">Lectotype</emphasis>
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of
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<taxonomicName authorityName="Andrewes" authorityYear="1931" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="nubila">
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<emphasis italics="true" pageId="27" pageNumber="28">Anchista nubila</emphasis>
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</taxonomicName>
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Andrewes, designated herein (NHML): female, body length = 8.5 mm, board mounted, "Dehra Dun / Rajpur, 3404 ft."; "Bought from / Staudinger & / Bang-Haas, 1930";
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<normalizedToken originalValue="“Type”">"Type"</normalizedToken>
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[red label]; "
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<emphasis italics="true" pageId="27" pageNumber="28">Anchista / nubila / Type Andr</emphasis>
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. / H. E. Andrewes det."; "H. E. Andrewes Coll. / B. M. 1945-97"; "LECTOTYPE ♀ / Anchista nubila / Andrewes, 1931 / des. SHI H. L. 2011" [Fig. 42].
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<emphasis bold="true" pageId="27" pageNumber="28">Paralectotypes</emphasis>
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of
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<taxonomicName authorityName="Andrewes" authorityYear="1931" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="nubila">
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<emphasis italics="true" pageId="27" pageNumber="28">Anchista nubila</emphasis>
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</taxonomicName>
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Andrewes: 1 male and 2 females (NHML), "Dehra Dun / Rajpur, 3404 ft."; "Bought from / Staudinger & / Bang-Haas, 1930";
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<normalizedToken originalValue="“Cotype”">"Cotype"</normalizedToken>
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[round label, with green circle]; "H. E. Andrewes Coll. / B. M.1945-97"; "PARALECTOTYPE / Anchista nubila / Andrewes, 1931 / det. SHI H. L. 2011".
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="notes on types">
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<paragraph pageId="27" pageNumber="28">Notes on types.</paragraph>
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<paragraph pageId="27" pageNumber="28">
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The original description mentioned five specimens from Dehra Dun in
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<normalizedToken originalValue="Andrewes’">Andrewes'</normalizedToken>
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collection, but did not designate a holotype. In the collection of NHML, we found only four specimens. Among them, a female specimen bears a determination label of Andrewes and a red label
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<normalizedToken originalValue="“Type”">"Type"</normalizedToken>
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, and this is herein designated as lectotype (
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<figureCitation captionStart="Figures 37–42" captionStartId="F7" captionText="Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes." httpUri="https://binary.pensoft.net/fig/29680" pageId="27" pageNumber="28">Fig. 42</figureCitation>
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) for taxonomic purpose of fixing the name to unique name-bearing type. The other three specimens have only a round label
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<normalizedToken originalValue="“co-type”">"co-type"</normalizedToken>
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, but no
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<normalizedToken originalValue="Andrewes’">Andrewes'</normalizedToken>
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determination label.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="non-type material examined">
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<paragraph pageId="27" pageNumber="28">Non-type material examined</paragraph>
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<paragraph pageId="27" pageNumber="28">(Total 16 specimens from India). 9 specimens (MNHN), "Chota Nagpore, Nowatoli, R. P. Cardon, VIII-XI 1896"[Fig. 73]. 3 specimens (MNHN), "Chota Nagpore, Nowatoli, R. P. Cardon, V.VI.1896". 1 specimen (MNHN), "Inde Angl, Ghates, R. P. F. Tabourel, VII-IX 1898". 1 specimen (MNHN), "S. India, Nedungadu, P. S. Nathan, 1936"[Fig. 8]. 1 specimen (MNHN), "Chota Nagpore"; "Ex Musaeo, H. W. Bates, 1892". 2 specimens (NHML), "India mer. Tanjore Distr. Nedungadu".</paragraph>
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</subSubSection>
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<subSubSection lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="diagnosis">
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<paragraph pageId="27" pageNumber="28">Diagnosis.</paragraph>
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<paragraph pageId="27" pageNumber="28">Head and pronotum glabrous; pronotum widest near middle, lateral margins distinctly angulate at middle; elytra uniform brownish; 3rd interval of elytra with three or four pores, 5th interval with two or three pores near base.</paragraph>
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<paragraph lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
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The glabrous surface and the number of setigerous pores on the 3rd and 5th intervals readily distinguish this species from most allied species, except
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<taxonomicName authority="fenestrata" authorityName="fenestrata (Schmidt-Goebel" authorityYear="1846" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="fenestrata" subSpecies="fenestrata">
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<emphasis italics="true" pageId="27" pageNumber="28">Anchista fenestrata fenestrata</emphasis>
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</taxonomicName>
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. Diagnosis between these two species is presented in the key. In general appearance, this species looks like
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<taxonomicName baseAuthorityName="Wiedemann" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brunnea">
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<emphasis italics="true" pageId="27" pageNumber="28">Anchista brunnea</emphasis>
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</taxonomicName>
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, but can be distinguished from the
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<pageBreakToken pageId="28" pageNumber="29" start="start">latter</pageBreakToken>
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by the wider and more angulate pronotum, and the different number of elytral setigerous pores.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="29" type="description">
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<paragraph pageId="28" pageNumber="29">Description.</paragraph>
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<paragraph pageId="28" pageNumber="29">
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Body length 7.5-8.5 mm; dorsal side uniform reddish brown, pronotum lateral explanate areas somewhat paler; elytra usually with margins darker, but not forming central patch; antennae reddish brown, 1st antennomere paler; femora yellowish, tibiae and tarsi darker; ventral side reddish brown.
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<emphasis bold="true" pageId="28" pageNumber="29">Head</emphasis>
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glabrous, without or with very fine isodiametric microsculpture.
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<emphasis bold="true" pageId="28" pageNumber="29">Pronotum</emphasis>
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glabrous,cordiform, widest slightly before middle; ratio PW/PL 1.30 to 1.41; pronotal base briefly but distinctly lobed; disc moderately convex, microsculpture indistinct, without punctures; front angles with few short setae (distinctly shorter than in
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<taxonomicName authority="fenestrata" authorityName="fenestrata (Schmidt-Goebel" authorityYear="1846" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="fenestrata" subSpecies="fenestrata">
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<emphasis italics="true" pageId="28" pageNumber="29">Anchista fenestrata fenestrata</emphasis>
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</taxonomicName>
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); lateral margins distinctly angulate in middle, strongly sinuate before hind angles; lateral explanate areas wide and flat, with few coarse punctures; hind angles sharp, rectangular or slightly acute, with few very short accessory setae; basal foveae moderately deep, with few punctures; median line distinct, strongly depressed and punctate anteriorly and subbasally.
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<emphasis bold="true" pageId="28" pageNumber="29">Elytra</emphasis>
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with striae shallow, distinct, finely punctate; intervals slightly convex, without accessory setae, finely and sparsely punctate; microsculpture distinct, isodiametric; 3rd interval usually with three setigerous pores, sometimes with four, all adjacent to 2nd interval, basal one usually placed at basal one-fourth approximately; 5th interval usually with two setigerous pores, sometimes with three, basal one near to interval base, all pores adjacent to 5th stria; basal margination nearly complete. Males with adhesive hairs rudimentary (two rows weakly present near apex) on 1st mesotarsomere, well developed (two whole rows) on 2nd and 3rd mesotarsomeres.
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<emphasis bold="true" pageId="28" pageNumber="29">Male genitalia</emphasis>
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with median lobe of aedeagus stout, bent to right side near apex; in lateral view, distinctly expanded in middle, gradually narrowed to apex; left lateral margin not curved in middle; apical orifice opened dorsally, basal margin of apical orifice with long setae; apical lamella placed on right side, flat, wide and rounded in dorsal view, not curved up dorsally; internal sac with a weakly sclerotized sclerite near middle, two weakly sclerotized elongate and sinuate sclerites near apical orifice, some parts of internal sac irregularly scaled (
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<figureCitation captionStart="Figure 73" captionStartId="F18" captionText="Figure 73. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Anchista nubila Andrewes, a specimen from Chota Nagpore." httpUri="https://binary.pensoft.net/fig/29691" pageId="28" pageNumber="29">Fig. 73</figureCitation>
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). Female genitalia not studied.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="29" type="distribution">
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<paragraph pageId="28" pageNumber="29">Distribution</paragraph>
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<paragraph pageId="28" pageNumber="29">
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(
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<figureCitation captionStart="Map 3" captionStartId="F47" captionText="Map 3. Known distribution of genus Anchista: ◆ Anchista brunnea (Wiedemann) ★ Anchista nubila Andrewes ● Anchista pilosa sp. n. △ Anchista fenestrata fenestrata (Schmidt-Goebel) ▼ Anchista fenestrata subpubescens Chaudoir." httpUri="https://binary.pensoft.net/fig/29725" pageId="28" pageNumber="29">Map 3</figureCitation>
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). India.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |