173 lines
20 KiB
XML
173 lines
20 KiB
XML
<document id="76E6D2DD692CCF6A60556D8D181A9BE2" ID-DOI="10.5070/P9401057774" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913295" ID-ZooBank="11600574-2B0E-4C13-BC08-A3A5EF9EE562" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712167020321" checkinUser="felipe" docAuthor="Groves, Lindsey & Squires, Richard L." docDate="2023" docId="921FD94CFFCEFFB5FF53FF78FE65F9FF" docLanguage="en" docName="PaleoBios.40.10.1-52.pdf" docOrigin="PaleoBios 40 (10)" docSource="http://dx.doi.org/10.5070/p9401057774" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.1:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="1" docTitle="Grovesia DOLIN AND LEDON 2002" docType="treatment" docVersion="2" lastPageNumber="28" masterDocId="6E26A134FFD3FFA8FFA2FFBBFFCAFFE5" masterDocTitle="Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover" masterLastPageNumber="52" masterPageNumber="1" pageNumber="28" updateTime="1712167351276" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="32FEAEAA6B5BCF2FAF304596AAF32466">Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover</mods:title>
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<mods:namePart id="7F096FB881895192DE12D7CB37656B05">Groves, Lindsey</mods:namePart>
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<mods:namePart id="0D334EF646C6034A5E20962E941B6BBC">Squires, Richard L.</mods:namePart>
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<mods:title id="F5C02377A6BC72B2F1DD593FD233B719">PaleoBios</mods:title>
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<mods:date id="34D5C75F4CC97447D0D9AAF16B2A3A3A">2023</mods:date>
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<treatment id="921FD94CFFCEFFB5FF53FF78FE65F9FF" LSID="urn:lsid:plazi:treatment:921FD94CFFCEFFB5FF53FF78FE65F9FF" httpUri="http://treatment.plazi.org/id/921FD94CFFCEFFB5FF53FF78FE65F9FF" lastPageNumber="28" pageId="29" pageNumber="28">
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<subSubSection id="52AC3BD1FFCEFFB5FF53FF78FD55FF38" box="[241,671,195,223]" pageId="29" pageNumber="28" type="nomenclature">
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<paragraph id="1A09685AFFCEFFB5FF53FF78FD55FF38" blockId="29.[241,671,195,223]" box="[241,671,195,223]" pageId="29" pageNumber="28">
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<heading id="4141DF36FFCEFFB5FF53FF78FD55FF38" allCaps="true" box="[241,671,195,223]" centered="true" fontSize="11" level="1" pageId="29" pageNumber="28" reason="1">
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<taxonomicName id="DDB613D9FFCEFFB5FF53FF78FD55FF38" ID-CoL="7P4L7" authority="DOLIN AND LEDON, 2002" authorityName="DOLIN AND LEDON" authorityYear="2002" box="[241,671,195,223]" class="Gastropoda" family="Eocypraeidae" genus="Grovesia" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="genus">
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<emphasis id="28C2B448FFCEFFB5FF53FF78FEA4FF3A" box="[241,366,195,223]" italics="true" pageId="29" pageNumber="28">GROVESIA</emphasis>
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DOLIN AND LEDON, 2002
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="52AC3BD1FFCEFFB5FF32FEB6FD48FE74" pageId="29" pageNumber="28" type="type_taxon">
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<paragraph id="1A09685AFFCEFFB5FF32FEB6FD48FE74" blockId="29.[112,801,267,1562]" pageId="29" pageNumber="28">
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<emphasis id="28C2B448FFCEFFB5FF32FEB6FE92FEC3" bold="true" box="[144,344,268,295]" pageId="29" pageNumber="28">
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<typeStatus id="C50DD6F8FFCEFFB5FF32FEB6FF18FEC2" box="[144,210,269,295]" pageId="29" pageNumber="28">Type</typeStatus>
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Species—
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</emphasis>
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<taxonomicName id="DDB613D9FFCEFFB5FEFAFEB0FE8EFEAF" authority="Dolin and Ledon, 2002" authorityName="Dolin and Ledon" authorityYear="2002" class="Gastropoda" family="Ovulidae" genus="Prionovolva" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="species" species="ganensis" subGenus="Grovesia">
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<emphasis id="28C2B448FFCEFFB5FEFAFEB0FE23FEC2" box="[344,489,267,295]" italics="true" pageId="29" pageNumber="28">Prionovolva</emphasis>
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(
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<emphasis id="28C2B448FFCEFFB5FE5AFEB6FDAAFECC" box="[504,608,269,297]" italics="true" pageId="29" pageNumber="28">Grovesia</emphasis>
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)
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<emphasis id="28C2B448FFCEFFB5FDCDFEB6FD12FECC" box="[623,728,269,297]" italics="true" pageId="29" pageNumber="28">ganensis</emphasis>
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<bibRefCitation id="7E2715ABFFCEFFB5FD7FFEB7FE8EFEAF" author="Dolin, L. & D. Ledon" pageId="29" pageNumber="28" pagination="329 - 347" refId="ref36561" refString="Dolin, L. and D. Ledon. 2002. Nouveaux taxons et discus- sion de la systematique des genres correspondants d'Ovulidae (Mollusca, Caenogastropoda) de l'Eocene inferieur de Gan (France). Geodiversitas 24 (2): 329 - 347, figs. 1 - 5." type="journal article" year="2002">Dolin and Ledon, 2002</bibRefCitation>
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</taxonomicName>
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, by original designation (of
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<bibRefCitation id="7E2715ABFFCEFFB5FD04FE8BFED8FE88" author="Dolin, L. & D. Ledon" pageId="29" pageNumber="28" pagination="329 - 347" refId="ref36561" refString="Dolin, L. and D. Ledon. 2002. Nouveaux taxons et discus- sion de la systematique des genres correspondants d'Ovulidae (Mollusca, Caenogastropoda) de l'Eocene inferieur de Gan (France). Geodiversitas 24 (2): 329 - 347, figs. 1 - 5." type="journal article" year="2002">Dolin and Ledon, 2002</bibRefCitation>
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: p. 334). Lower Eocene (Ypresian), Gan, Pyrénées-Atlantiques Department,
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<collectingCountry id="62A128CAFFCEFFB5FD85FECCFDB4FE74" box="[551,638,375,401]" name="France" pageId="29" pageNumber="28">France</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="52AC3BD1FFCEFFB5FF32FE22FD0AFC9B" pageId="29" pageNumber="28" type="diagnosis">
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<paragraph id="1A09685AFFCEFFB5FF32FE22FEB3FD4F" blockId="29.[112,801,267,1562]" pageId="29" pageNumber="28">
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<emphasis id="28C2B448FFCEFFB5FF32FE22FEFCFE56" bold="true" box="[144,310,409,435]" pageId="29" pageNumber="28">Diagnosis—</emphasis>
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Size small. Shell moderately inflated, can be somewhat narrow and tapered. Dorsum smooth. Fossula auriform. Terminal fold short. Columellar teeth carinated, extending partially across basal face, especially on posterior half of base. Outer lip coarsely denticulate, with short but stout denticles. Siphonal canal open. Posterior (exhalant) canal poorly delimited (
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<bibRefCitation id="7E2715ABFFCEFFB5FD07FDD7FEC6FD4F" author="Dolin, L. & D. Ledon" pageId="29" pageNumber="28" pagination="329 - 347" refId="ref36561" refString="Dolin, L. and D. Ledon. 2002. Nouveaux taxons et discus- sion de la systematique des genres correspondants d'Ovulidae (Mollusca, Caenogastropoda) de l'Eocene inferieur de Gan (France). Geodiversitas 24 (2): 329 - 347, figs. 1 - 5." type="journal article" year="2002">Dolin and Ledon, 2002</bibRefCitation>
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: p. 334).
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</paragraph>
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<paragraph id="1A09685AFFCEFFB5FF32FD08FD0AFC9B" blockId="29.[112,801,267,1562]" pageId="29" pageNumber="28">
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<emphasis id="28C2B448FFCEFFB5FF32FD08FE4AFD28" bold="true" box="[144,384,691,717]" pageId="29" pageNumber="28">Geologic Range—</emphasis>
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<materialsCitation id="AADE6207FFCEFFB5FE22FD08FD76FC9B" location="upper lower Eocene" pageId="29" pageNumber="28" specimenCount="1" stateProvince="Washington">
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Early Eocene (Yepresian) to late Eocene (Priabonian) (
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<bibRefCitation id="7E2715ABFFCEFFB5FE24FD6DFDD7FD15" author="Fehse, D." box="[390,541,726,752]" pageId="29" pageNumber="28" pagination="1 - 12" refId="ref37236" refString="Fehse, D. 2021. Zur systemstischen stellung der Eocy- praeidae F. A. Schilder 1924. Part 2. (Mollusca: Gas- tropoda: Cypraeoidea). Privately published online, Berlin, Germany 46 p., pls. 1 - 12." type="book chapter" year="2021">Fehse, 2021</bibRefCitation>
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, in part). In the NEP
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<taxonomicName id="DDB613D9FFCEFFB5FFD2FD41FF12FCF3" authorityName="DOLIN AND LEDON" authorityYear="2002" box="[112,216,762,790]" class="Gastropoda" family="Eocypraeidae" genus="Grovesia" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="genus">
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<emphasis id="28C2B448FFCEFFB5FFD2FD41FF12FCF3" box="[112,216,762,790]" italics="true" pageId="29" pageNumber="28">Grovesia</emphasis>
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</taxonomicName>
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ranges from the
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<location id="1F693E81FFCEFFB5FE0AFD41FD6AFCF1" LSID="urn:lsid:plazi:treatment:921FD94CFFCEFFB5FF53FF78FE65F9FF:1F693E81FFCEFFB5FE0AFD41FD6AFCF1" box="[424,672,761,788]" name="upper lower Eocene" pageId="29" pageNumber="28" stateProvince="Washington">upper lower Eocene</location>
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(“Domen- gine Stage”) to middle Eocene (“Tejon Stage”) (new information). Also questionably upper middle Eocene (Priabonian) in
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<collectingRegion id="D872A6B8FFCEFFB5FE94FCD8FE01FC98" box="[310,459,867,893]" country="United States of America" name="Washington" pageId="29" pageNumber="28">Washington</collectingRegion>
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(new information)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="52AC3BD1FFCEFFB5FF32FC3DFE65F9FF" pageId="29" pageNumber="28" type="discussion">
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<paragraph id="1A09685AFFCEFFB5FF32FC3DFCCCFAC1" blockId="29.[112,801,267,1562]" pageId="29" pageNumber="28">
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<emphasis id="28C2B448FFCEFFB5FF32FC3DFEEDFC45" bold="true" box="[144,295,902,928]" pageId="29" pageNumber="28">Remarks—</emphasis>
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||
<bibRefCitation id="7E2715ABFFCEFFB5FE85FC3DFD9EFC45" author="Dolin, L. & D. Ledon" box="[295,596,902,928]" pageId="29" pageNumber="28" pagination="329 - 347" refId="ref36561" refString="Dolin, L. and D. Ledon. 2002. Nouveaux taxons et discus- sion de la systematique des genres correspondants d'Ovulidae (Mollusca, Caenogastropoda) de l'Eocene inferieur de Gan (France). Geodiversitas 24 (2): 329 - 347, figs. 1 - 5." type="journal article" year="2002">Dolin and Ledon (2002</bibRefCitation>
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||
: p. 334) placed their new subgenus,
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<taxonomicName id="DDB613D9FFCEFFB5FEC8FC11FE1BFC23" authorityName="DOLIN AND LEDON" authorityYear="2002" box="[362,465,938,966]" class="Gastropoda" family="Eocypraeidae" genus="Grovesia" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="genus">
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<emphasis id="28C2B448FFCEFFB5FEC8FC11FE1BFC23" box="[362,465,938,966]" italics="true" pageId="29" pageNumber="28">Grovesia</emphasis>
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</taxonomicName>
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, in the extant genus
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<taxonomicName id="DDB613D9FFCEFFB5FD64FC13FE9CFC02" authority="Iredale, 1930" authorityName="Iredale" authorityYear="1930" class="Gastropoda" family="Ovulidae" genus="Prionovolva" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="genus">
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<emphasis id="28C2B448FFCEFFB5FD64FC13FF7AFC02" italics="true" pageId="29" pageNumber="28">Prionovolva</emphasis>
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<bibRefCitation id="7E2715ABFFCEFFB5FF16FC77FE9CFC02" author="Iredale, T." box="[180,342,972,999]" pageId="29" pageNumber="28" pagination="73 - 88" refId="ref39159" refString="Iredale, T. 1930. Queensland molluscan notes, no. 2. Memoirs of the Queensland Museum 10 (10): 73 - 88, pl. 9." type="journal article" year="1930">Iredale, 1930</bibRefCitation>
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</taxonomicName>
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, but
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<bibRefCitation id="7E2715ABFFCEFFB5FE2DFC77FD78FC02" author="Dolin, L. & J-M. Pacaud" box="[399,690,972,999]" pageId="29" pageNumber="28" pagination="277 - 314" refId="ref36679" refString="Dolin, L. and J-M. Pacaud. 2009. Les Cypraeoidea et Ve- lutinoidea (Mollusca, Caenogastropoda) du Lutetien inferieur du Vicentin et dus Veronais (nord-est de l'Italie). Revue de Paleobiologie, Geneve 28 (2): 277 - 314, pls. 1 - 8." type="journal article" year="2009">Dolin and Pacaud (2009</bibRefCitation>
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||
: pp. 288, 289) recognized
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<taxonomicName id="DDB613D9FFCEFFB5FEEAFC4BFE7AFBE9" authorityName="DOLIN AND LEDON" authorityYear="2002" box="[328,432,1008,1036]" class="Gastropoda" family="Eocypraeidae" genus="Grovesia" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="genus">
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||
<emphasis id="28C2B448FFCEFFB5FEEAFC4BFE7AFBE9" box="[328,432,1008,1036]" italics="true" pageId="29" pageNumber="28">Grovesia</emphasis>
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</taxonomicName>
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as a distinct genus.
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<taxonomicName id="DDB613D9FFCEFFB5FD1AFC4BFEC7FBC8" authorityName="WA, OR, So CA" baseAuthorityName="Gabb" baseAuthorityYear="1869" class="Gastropoda" family="Eocypraeidae" genus="Grovesia" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="species" species="mathewsonii">
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<emphasis id="28C2B448FFCEFFB5FD1AFC4BFEC7FBC8" italics="true" pageId="29" pageNumber="28">Grovesia mathewsonii</emphasis>
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</taxonomicName>
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(see below) and
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<taxonomicName id="DDB613D9FFCEFFB5FE48FBA8FD58FBC8" box="[490,658,1041,1071]" class="Gastropoda" family="Cypraeidae" genus="Cypraea" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="species" species="castacensis">
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<emphasis id="28C2B448FFCEFFB5FE48FBA8FD58FBC8" box="[490,658,1041,1071]" italics="true" pageId="29" pageNumber="28">G. castacensis</emphasis>
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</taxonomicName>
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form a lin- eage (Fig. 1), ranging from the early middle (“Domengine Stage”) to late middle Eocene (“Tejon Stage”) in the NEP region (new information).
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<bibRefCitation id="7E2715ABFFCEFFB5FE0FFBC7FD84FB72" author="Fehse, D." box="[429,590,1148,1175]" pageId="29" pageNumber="28" pagination="1 - 12" refId="ref37236" refString="Fehse, D. 2021. Zur systemstischen stellung der Eocy- praeidae F. A. Schilder 1924. Part 2. (Mollusca: Gas- tropoda: Cypraeoidea). Privately published online, Berlin, Germany 46 p., pls. 1 - 12." type="book chapter" year="2021">Fehse (2021)</bibRefCitation>
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noted that
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<taxonomicName id="DDB613D9FFCEFFB5FD70FBC6FF58FB5F" authorityName="DOLIN AND LEDON" authorityYear="2002" class="Gastropoda" family="Eocypraeidae" genus="Grovesia" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="genus">
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<emphasis id="28C2B448FFCEFFB5FD70FBC6FF58FB5F" italics="true" pageId="29" pageNumber="28">Grovesia</emphasis>
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</taxonomicName>
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ranges from the Late Cretaceous (Cenomanian) to late Eocene (Priabonian). He included the Cretaceous species
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<taxonomicName id="DDB613D9FFCEFFB5FFD2FB5DFD8CFAE5" authority="Schilder, 1932 b" authorityName="Schilder" authorityYear="1932" box="[112,582,1254,1282]" class="Gastropoda" family="Pediculariidae" genus="Cyproglobina" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="species" species="aegyptia">
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<emphasis id="28C2B448FFCEFFB5FFD2FB5DFE48FAE7" box="[112,386,1254,1282]" italics="true" pageId="29" pageNumber="28">Cyproglobina aegyptia</emphasis>
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<bibRefCitation id="7E2715ABFFCEFFB5FE25FB5DFDFCFAE5" author="Schilder, F. A." box="[391,566,1254,1280]" pageId="29" pageNumber="28" refId="ref41900" refString="Schilder, F. A. 1932. Cypraeacea (Mollusca Gastropoda Prosobranchia) (Familien Eratoidae, Cypraeidae und Amphiperatidae). Pp. 1 - 276 in W. Quenstedt (ed.). Fossilium Catalogus, 1: Animalia, pt. 55. W. Junk, Berlin." type="book" year="1932">Schilder, 1932</bibRefCitation>
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b
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</taxonomicName>
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and
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<taxonomicName id="DDB613D9FFCEFFB5FDDDFB5DFEEBFAC6" authority="Schilder, 1922" authorityName="Schilder" authorityYear="1922" class="Gastropoda" family="Pediculariidae" genus="Cyproglobina" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="species" species="strombecki">
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<emphasis id="28C2B448FFCEFFB5FDDDFB5DFCEAFAE4" box="[639,800,1253,1282]" italics="true" pageId="29" pageNumber="28">C. strombecki</emphasis>
|
||
<bibRefCitation id="7E2715ABFFCEFFB5FFD2FAB2FEEBFAC6" author="Schilder, F. A." box="[112,289,1289,1315]" pageId="29" pageNumber="28" pagination="98 - 122" refId="ref41740" refString="Schilder, F. A. 1922. Contributions to the knowledge of tlhe genera Cypraea and Trivia. Proceedings of the Malacological Society of London 15: 98 - 122." type="journal article" year="1922">Schilder, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
, which are both questionably
|
||
<taxonomicName id="DDB613D9FFCEFFB5FD35FAB1FD35FAC3" authorityName="DOLIN AND LEDON" authorityYear="2002" box="[663,767,1290,1318]" class="Gastropoda" family="Eocypraeidae" genus="Grovesia" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="genus">
|
||
<emphasis id="28C2B448FFCEFFB5FD35FAB1FD35FAC3" box="[663,767,1290,1318]" italics="true" pageId="29" pageNumber="28">Grovesia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1A09685AFFCEFFB5FF32FA97FE65F9FF" blockId="29.[112,801,267,1562]" pageId="29" pageNumber="28">
|
||
Specimens of
|
||
<taxonomicName id="DDB613D9FFCEFFB5FE9AFA96FDE5FAA2" baseAuthorityName="Stewart" baseAuthorityYear="1926" box="[312,559,1323,1353]" class="Gastropoda" family="Eocypraeidae" genus="Grovesia" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="species" species="castacensis">
|
||
<emphasis id="28C2B448FFCEFFB5FE9AFA96FDE5FAA2" box="[312,559,1323,1353]" italics="true" pageId="29" pageNumber="28">Grovesia castacensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DDB613D9FFCEFFB5FDC5FA96FCEAFAA2" authorityName="WA, OR, So CA" baseAuthorityName="Gabb" baseAuthorityYear="1869" box="[615,800,1323,1353]" class="Gastropoda" family="Eocypraeidae" genus="Grovesia" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="28" phylum="Mollusca" rank="species" species="mathewsonii">
|
||
<emphasis id="28C2B448FFCEFFB5FDC5FA96FCEAFAA2" box="[615,800,1323,1353]" italics="true" pageId="29" pageNumber="28">G. mathewsonii</emphasis>
|
||
</taxonomicName>
|
||
(below) are many times poorly preserved and identifi- cations were made as ‘best judgements’ by the authors. Throughout the duration of the pandemic it was impos- sible to verify nonfigured identifications cited by Dick- erson (1915, 1916),
|
||
<bibRefCitation id="7E2715ABFFCEFFB5FED4FA67FD28FA12" author="Anderson, F. M. & D. Hanna" box="[374,738,1500,1527]" pageId="29" pageNumber="28" pagination="1 - 249" refId="ref35137" refString="Anderson, F. M. and G D. Hanna. 1925. Fauna and strati- graphic relations of the Tejon Eocene at the type local- ity in Kern County, California. Occasional Papers of the California Academy of Sciences 11: 1 - 249, pls. 1 - 16." type="journal article" year="1925">Anderson and Hanna (1925)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="7E2715ABFFCEFFB5FFD2F9BBFEE6F9FF" author="Weaver, C. E." box="[112,300,1535,1562]" pageId="29" pageNumber="28" refId="ref44451" refString="Weaver, C. E. 1953. Eocene and Paleocene deposits at" type="book" year="1953">Weaver (1953)</bibRefCitation>
|
||
in person.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |