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<document ID-DOI="http://dx.doi.org/10.3897/dez.69.80492" ID-Pensoft-Pub="1860-1324-1-45" ID-Pensoft-UUID="0A5DABFBAA6551E194C35CEA6054B588" ID-ZooBank="C402CAE814FD402AAD0554646B1493E8" ModsDocID="1860-1324-69-1-45" checkinTime="1649136064731" checkinUser="pensoft" docAuthor="Morishita, Shunsuke &amp; Watanabe, Kyohei" docDate="2022" docId="7DD69E40C0B155A8BA15A6C5F95B6EA3" docLanguage="en" docName="DeutEntomolZeit 69(1): 45-64" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-04-04" docSource="http://dx.doi.org/10.3897/dez.69.80492" docTitle="Woldstedtius karafutensis" docType="treatment" docVersion="1" id="0A5DABFBAA6551E194C35CEA6054B588" lastPageNumber="45" masterDocId="0A5DABFBAA6551E194C35CEA6054B588" masterDocTitle="Revision of the genus Woldstedtius Carlson, 1979 (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan" masterLastPageNumber="64" masterPageNumber="45" pageId="0" pageNumber="45" updateTime="1649136064731" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genus Woldstedtius Carlson, 1979 (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Morishita, Shunsuke</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5842-4066</mods:nameIdentifier>
<mods:affiliation>Hatchodori 4 - 9, Toyohashi, Aichi 440 - 0806, Japan</mods:affiliation>
<mods:nameIdentifier type="email">ezogengoroumodoki4.23@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Watanabe, Kyohei</mods:namePart>
<mods:affiliation>Kanagawa Prefectural Museum of Natural History, Iryuda 499, Odawara, Kanagawa 250 - 0031, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-04-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>69</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>45</mods:start>
<mods:end>64</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.69.80492</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.69.80492</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-1-45</mods:identifier>
<mods:identifier type="ZooBank">C402CAE814FD402AAD0554646B1493E8</mods:identifier>
<mods:identifier type="Pensoft-UUID">0A5DABFBAA6551E194C35CEA6054B588</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:7DD69E40C0B155A8BA15A6C5F95B6EA3" httpUri="http://treatment.plazi.org/id/7DD69E40C0B155A8BA15A6C5F95B6EA3" lastPageNumber="45" pageId="0" pageNumber="45">
<subSubSection pageId="0" pageNumber="45" type="nomenclature">
<paragraph pageId="0" pageNumber="45">
<taxonomicName LSID="7DD69E40-C0B1-55A8-BA15-A6C5F95B6EA3" authority="(Uchida, 1957)" baseAuthorityName="Uchida" baseAuthorityYear="1957" class="Insecta" family="Ichneumonidae" genus="Woldstedtius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Woldstedtius karafutensis" order="Hymenoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="karafutensis">Woldstedtius karafutensis (Uchida, 1957)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Woldstedtius karafutensis (Uchida, 1957) (A-D. female; E, F. male) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure4" httpUri="https://binary.pensoft.net/fig/667924" pageId="0" pageNumber="45">Figs 4A-F</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Japanese Woldstedtius A, G. W. f. flavolineatus (Gravenhorst, 1829); B, I. W. kuroashii (Uchida, 1957); C, D, K. W. takagii (Uchida, 1957); E. W. alpicola sp. nov.; F. W. biguttatus (Gravenhorst, 1829); H. W. karafutensis (Uchida, 1957); J. W. punctatus sp. nov. (E, J. holotype; B, I. paratype) - A, B. head, lateral view; C. head, frontal view; D. propodeum, lateral view; E-K. T I, dorsal view." figureDoi="10.3897/dez.69.80492.figure9" httpUri="https://binary.pensoft.net/fig/667929" pageId="0" pageNumber="45">, 9H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="reference_group">
<paragraph pageId="0" pageNumber="45">
<taxonomicName authority="Morley, 1911" authorityName="Morley" authorityYear="1911" class="Insecta" family="Ichneumonidae" genus="Homocidus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Homocidus karafutensis" order="Hymenoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="karafutensis">Homocidus karafutensis</taxonomicName>
Uchida, 1957: 252.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="materials_examined">
<paragraph pageId="0" pageNumber="45">Materials examined.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Type series</emphasis>
:
<materialsCitation collectingDate="1934-07-25" collectorName="C. Watanabe, T. Inoue" country="RUSSIA" location="Tarandomari" specimenCount="1" stateProvince="Sakhalin" typeStatus="holotype">
<collectingCountry name="Russia">RUSSIA</collectingCountry>
: 1 F (
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">
<typeStatus>holotype</typeStatus>
</emphasis>
),
<collectingRegion country="Russia" name="Sakhalin">Sakhalin</collectingRegion>
Is.,
<location LSID="urn:lsid:plazi:treatment:7DD69E40C0B155A8BA15A6C5F95B6EA3:F0AD8D75EF8AF2EADE343D5E3B8E8770" country="RUSSIA" name="Tarandomari" stateProvince="Sakhalin">Tarandomari</location>
,
<collectingDate value="1934-07-25">25 Jul 1934</collectingDate>
,
<collectorName>C. Watanabe</collectorName>
&amp;
<collectorName>T. Inoue</collectorName>
leg. (SEHU)
</materialsCitation>
;
<materialsCitation collectingDate="1954-07-06" collectorName="Townes, Non-types" country="JAPAN" location="Sapporo" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectingCountry name="Japan">JAPAN</collectingCountry>
: [
<collectingRegion country="Japan" name="Hokkaido">Hokkaido</collectingRegion>
] 1 F (
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">
<typeStatus>paratype</typeStatus>
</emphasis>
),
<collectingRegion country="Japan" name="Hokkaido">Hokkaido</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:7DD69E40C0B155A8BA15A6C5F95B6EA3:2E6642EDB9E2333906AC7F0A81FDC3D0" country="JAPAN" name="Sapporo" stateProvince="Hokkaido">Sapporo</location>
,
<collectingDate value="1954-07-06">6 Jul 1954</collectingDate>
,
<collectorName>Townes</collectorName>
family leg. (SEHU).
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">
<collectorName>Non-types</collectorName>
</emphasis>
</materialsCitation>
:
<materialsCitation collectingDate="1998-08-24" collectingDateMax="2011-07-18" collectingDateMin="1998-08-24" collectorName="T. Muroi, Y. Matsubara, K. Fukuda, T. Sugiyama, K. Watanabe, K. Yamagishi, M. Watanabe, K. Esaki, H. Fukutomi, S. Fujie" country="JAPAN" county="Mt. Hakase" location="Honkawane Town" municipality="Mamurogawa Town" specimenCount="1" stateProvince="Fukushima Pref." typeStatus="paratype">
<collectingCountry name="Japan">JAPAN</collectingCountry>
: [Honshu] 1 F,
<collectingRegion country="Japan" name="Fukushima">Fukushima Pref.</collectingRegion>
, Showa Vil.,
<collectingCounty>Mt. Hakase</collectingCounty>
,
<collectingDate value="1998-08-24" valueMax="1998-09-19" valueMin="1998-08-24">24 Aug-19 Sep 1998</collectingDate>
,
<collectorName>T. Muroi</collectorName>
leg. (MT) (MU); 1 F,
<collectingRegion country="Japan" name="Yamagata">Yamagata Pref.</collectingRegion>
,
<collectingMunicipality>Mamurogawa Town</collectingMunicipality>
, Azusayama,
<collectingDate value="2009-09-05">5 Sep 2009</collectingDate>
,
<collectorName>Y. Matsubara</collectorName>
&amp;
<collectorName>K. Fukuda</collectorName>
leg. (MT) (KPMNH); 2 F,
<collectingRegion country="Japan" name="Shizuoka">Shizuoka Pref.</collectingRegion>
,
<collectingRegion country="Japan" name="Shizuoka">Shizuoka</collectingRegion>
City, Umegashima,
<collectingDate value="2001-06-03" valueMax="2001-07-16" valueMin="2001-06-03">3 Jun-16 Jul 2001</collectingDate>
,
<collectorName>T. Sugiyama</collectorName>
leg. (MT) (MU); 2 F, ditto,
<collectingDate value="2001-07-17" valueMax="2001-08-05" valueMin="2001-07-17">17 Jul-5 Aug 2001</collectingDate>
,
<collectorName>T. Sugiyama</collectorName>
leg. (MT) (MU); 1 M,
<collectingRegion country="Japan" name="Shizuoka">Shizuoka Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:7DD69E40C0B155A8BA15A6C5F95B6EA3:078F8C1092A61829B4A5DDD62D2D6DE7" country="JAPAN" county="Mt. Hakase" municipality="Mamurogawa Town" name="Honkawane Town" stateProvince="Fukushima Pref.">Honkawane Town</location>
, Yamainudan,
<collectingDate value="2008-06-14">14 Jun 2008</collectingDate>
,
<collectorName>K. Watanabe</collectorName>
leg. (KPMNH); 2 F,
<collectingRegion country="Japan" name="Gifu">Gifu Pref.</collectingRegion>
, Kani City, Katabira,
<collectingDate value="2004-05-08" valueMax="2004-05-14" valueMin="2004-05-08">8-14 May 2004</collectingDate>
,
<collectorName>K. Yamagishi</collectorName>
leg. (MT) (MU); 1 F,
<collectingRegion country="Japan" name="Toyama">Toyama Pref.</collectingRegion>
, Nanto City, Togamura, Kamimomose,
<collectingDate value="2009-08-04" valueMax="2009-08-11" valueMin="2009-08-04">4-11 Aug 2009</collectingDate>
,
<collectorName>M. Watanabe</collectorName>
leg. (MT) (KPMNH); 1 F,
<collectingRegion country="Japan" name="Ishikawa">Ishikawa Pref.</collectingRegion>
, Kaga City,
<location LSID="urn:lsid:plazi:treatment:7DD69E40C0B155A8BA15A6C5F95B6EA3:1112F51E048C1696D3A707C659A92D1D" country="JAPAN" county="Mt. Hakase" municipality="Mamurogawa Town" name="Mt. Kariyasuyama" stateProvince="Fukushima Pref.">Mt. Kariyasuyama</location>
,
<collectingDate value="2002-07-07" valueMax="2002-07-18" valueMin="2002-07-07">7-18 Jul 2002</collectingDate>
,
<collectorName>K. Esaki</collectorName>
leg. (MT) (TMNH); 1 F,
<collectingRegion country="Japan" name="Ishikawa">Ishikawa Pref.</collectingRegion>
, Hakusan City, Sannomiya,
<collectingDate value="2009-09-06" valueMax="2009-09-18" valueMin="2009-09-06">6-18 Sep 2009</collectingDate>
,
<collectorName>H. Fukutomi</collectorName>
leg. (MT) (MU); 1 M,
<collectingRegion country="Japan" name="Ishikawa">Ishikawa Pref.</collectingRegion>
, Hakusan City, Togadani,
<collectingDate value="2010-05-18" valueMax="2010-06-04" valueMin="2010-05-18">18 May-4 Jun 2010</collectingDate>
,
<collectorName>H. Fukutomi</collectorName>
leg. (MT) (MU); 1 F,
<collectingRegion country="Japan" name="Hyogo">Hyogo Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:7DD69E40C0B155A8BA15A6C5F95B6EA3:C03D351D1D1CB7C9421885FFB3E0723A" country="JAPAN" county="Mt. Hakase" municipality="Mamurogawa Town" name="Kami Town" stateProvince="Fukushima Pref.">Kami Town</location>
, Ojiro-ku, Niiya, Mikata-kogen,
<collectingDate value="2011-06-26" valueMax="2011-07-18" valueMin="2011-06-26">26 Jun-18 Jul 2011</collectingDate>
,
<collectorName>S. Fujie</collectorName>
leg. (MT) (KPMNH)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/dez.69.80492.figure4" httpUri="https://binary.pensoft.net/fig/667924" pageId="0" pageNumber="45" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 4.</emphasis>
<taxonomicName baseAuthorityName="Uchida" baseAuthorityYear="1957" class="Insecta" family="Ichneumonidae" genus="Woldstedtius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Woldstedtius karafutensis" order="Hymenoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="karafutensis">
<emphasis italics="true" pageId="0" pageNumber="45">Woldstedtius karafutensis</emphasis>
</taxonomicName>
(Uchida, 1957) (
<emphasis bold="true" pageId="0" pageNumber="45">A-D.</emphasis>
female;
<emphasis bold="true" pageId="0" pageNumber="45">E, F.</emphasis>
male) -
<emphasis bold="true" pageId="0" pageNumber="45">A, E.</emphasis>
Habitus;
<emphasis bold="true" pageId="0" pageNumber="45">B, F.</emphasis>
Head, frontal view;
<emphasis bold="true" pageId="0" pageNumber="45">C.</emphasis>
Mesonotum, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="45">D.</emphasis>
Propodeum, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">Female (n = 14). Body length 4.1-6.0 mm, polished, coriaceous, covered with silver setae.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Head</emphasis>
0.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. Clypeus 2.1-2.4
<normalizedToken originalValue="×">x</normalizedToken>
as broad as high, flat in lateral view. Face 1.88-2.2
<normalizedToken originalValue="×">x</normalizedToken>
as broad as high, densely punctate, convex medially in lateral view, separated from clypeus by shallow clypeal sulcus. Inner orbits almost parallel (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Woldstedtius karafutensis (Uchida, 1957) (A-D. female; E, F. male) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure4" httpUri="https://binary.pensoft.net/fig/667924" pageId="0" pageNumber="45">4B</figureCitation>
). Length of malar space 1.1-1.15
<normalizedToken originalValue="×">x</normalizedToken>
as long as basal mandibular width. POL 2.0-2.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as OD. OOL 1.0-1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as OD. POL 2.0-2.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as OOL. Antenna with 20-21 flagellomeres. FL I 1.15-1.35
<normalizedToken originalValue="×">x</normalizedToken>
as long as FL II. MP IV 1.45-1.66
<normalizedToken originalValue="×">x</normalizedToken>
as long as MP V.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Mesosoma</emphasis>
.
</emphasis>
Lateral aspect of pronotum rugulose anteriorly. Mesoscutum densely punctate (separated by ca. 0.5-1.0
<normalizedToken originalValue="×">x</normalizedToken>
their diameter) (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Woldstedtius karafutensis (Uchida, 1957) (A-D. female; E, F. male) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure4" httpUri="https://binary.pensoft.net/fig/667924" pageId="0" pageNumber="45">4C</figureCitation>
). Scutellum finely and sparsely punctate (separated by ca. 1.5-2.0
<normalizedToken originalValue="×">x</normalizedToken>
their diameter). Lower part of mesopleuron coarsely and sparsely punctate. Sternaulus indistinct. Propodeum without rugae and carinae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Woldstedtius karafutensis (Uchida, 1957) (A-D. female; E, F. male) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure4" httpUri="https://binary.pensoft.net/fig/667924" pageId="0" pageNumber="45">4D</figureCitation>
), rounded in lateral view. Fore wing length 4.0-5.0 mm. Nervellus intercepted below middle. Hind femur 3.75-4.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum depth in lateral view. Hind tibia 8.0-9.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum depth in lateral view. Ratio of length of hind first to fifth tarsomeres 1.0: 0.6: 0.5: 0.3: 0.25-0.3.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Metasoma</emphasis>
.
</emphasis>
T I square in dorsal view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Japanese Woldstedtius A, G. W. f. flavolineatus (Gravenhorst, 1829); B, I. W. kuroashii (Uchida, 1957); C, D, K. W. takagii (Uchida, 1957); E. W. alpicola sp. nov.; F. W. biguttatus (Gravenhorst, 1829); H. W. karafutensis (Uchida, 1957); J. W. punctatus sp. nov. (E, J. holotype; B, I. paratype) - A, B. head, lateral view; C. head, frontal view; D. propodeum, lateral view; E-K. T I, dorsal view." figureDoi="10.3897/dez.69.80492.figure9" httpUri="https://binary.pensoft.net/fig/667929" pageId="0" pageNumber="45">9H</figureCitation>
), 1.0-1.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum width, rugulose laterally. Latero-median carina present basal ca. 0.3 of T I (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Japanese Woldstedtius A, G. W. f. flavolineatus (Gravenhorst, 1829); B, I. W. kuroashii (Uchida, 1957); C, D, K. W. takagii (Uchida, 1957); E. W. alpicola sp. nov.; F. W. biguttatus (Gravenhorst, 1829); H. W. karafutensis (Uchida, 1957); J. W. punctatus sp. nov. (E, J. holotype; B, I. paratype) - A, B. head, lateral view; C. head, frontal view; D. propodeum, lateral view; E-K. T I, dorsal view." figureDoi="10.3897/dez.69.80492.figure9" httpUri="https://binary.pensoft.net/fig/667929" pageId="0" pageNumber="45">9H</figureCitation>
). T II 0.86-1.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum width, strigose except for posterior margin smooth.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Coloration</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Woldstedtius karafutensis (Uchida, 1957) (A-D. female; E, F. male) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure4" httpUri="https://binary.pensoft.net/fig/667924" pageId="0" pageNumber="45">4A-D</figureCitation>
).
</emphasis>
Body (excluding wings and legs) black to blackish-brown. Face with a large white median spot (this spot connected with white clypeus). Clypeus, palpi, tegula, subtegular ridge, upper mesepisternum and scutellum white. Mandible white, except for apex and base. Lateral aspect of pronotum with a white spot posteriorly. Antenna brown. Veins and pterostigma yellowish-brown to brown. Legs whitish-yellow. Hind coxa with a dorsal blackish-brown stripe. Hind femur brown. Hind tibia and tarsomere black. Base of hind tibia tinged with white.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Male (n = 2).</emphasis>
Similar to female. Inner orbits weakly divergent downward (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Woldstedtius karafutensis (Uchida, 1957) (A-D. female; E, F. male) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure4" httpUri="https://binary.pensoft.net/fig/667924" pageId="0" pageNumber="45">4F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Coloration</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Woldstedtius karafutensis (Uchida, 1957) (A-D. female; E, F. male) - A, E. Habitus; B, F. Head, frontal view; C. Mesonotum, dorsal view; D. Propodeum, dorsal view." figureDoi="10.3897/dez.69.80492.figure4" httpUri="https://binary.pensoft.net/fig/667924" pageId="0" pageNumber="45">4E, F</figureCitation>
).
</emphasis>
Body (excluding wings and legs) black to blackish-brown. Clypeus, palpi, face, ventral surface of antenna, malar space, propleuron, epicnemium, mesosternum, tegula, subtegular ridge, upper mesepisternum and scutellum white. Gena tinged with white ventrally. Mandible white except for apex. Lateral aspect of pronotum tinged with white ventrally and posteriorly. Mesoscutum with white shoulder marks. Mesopleuron with a large white marking; it is enlarged anteriorly. Wings hyaline. Veins and pterostigma yellowish-brown to brown. Legs white. Fore and mid tarsi, apex of hind trochantellus, base and apex of hind femur and hind tarsomeres brown. Hind coxa with a dorsal brown stripe. Hind tibia tinged with brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">
Japan (Hokkaido and Honshu). Outside Japan, this species has been recorded from Russia and South Korea (
<bibRefCitation DOI="https://doi.org/10.1016/j.aspen.2016.04.012" author="Balueva, EN" journalOrPublisher="Journal of Asia-Pacific Entomology" pageId="0" pageNumber="45" pagination="387 - 397" refId="B1" refString="Balueva, EN, Lee, JW, 2016. Study of the genus Woldstedtius Carlson (Hymenoptera; Ichneumonidae; Diplazontinae) with description of three new species in South Korea. Journal of Asia-Pacific Entomology 19 (2): 387 - 397, DOI: https://doi.org/10.1016/j.aspen.2016.04.012" title="Study of the genus Woldstedtius Carlson (Hymenoptera; Ichneumonidae; Diplazontinae) with description of three new species in South Korea." url="https://doi.org/10.1016/j.aspen.2016.04.012" volume="19" year="2016">Balueva and Lee 2016</bibRefCitation>
and
<bibRefCitation author="Yu, DS" journalOrPublisher="Folia Entomologica Hungarica" pageId="0" pageNumber="45" refId="B25" refString="Yu, DS, van Achterberg, K, Horstmann, K, 2016. World Ichneumonoidea 2015. Taxonomy, biology, morphology and distribution. [Database on flash drive]. Taxapad, Vancouver." title="World Ichneumonoidea 2015. Taxonomy, biology, morphology and distribution. [Database on flash drive]. Taxapad, Vancouver." year="2016">Yu et al. 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="bionomics">
<paragraph pageId="0" pageNumber="45">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="45">Host unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="remarks">
<paragraph pageId="0" pageNumber="45">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="45">This is the first record of this species from Honshu.</paragraph>
</subSubSection>
</treatment>
</document>
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