325 lines
38 KiB
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325 lines
38 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.79.e66321" ID-GBIF-Dataset="df495ca3-4f40-481d-b6e6-c1835d81b0a0" ID-GBIF-Taxon="189469082" ID-Pensoft-Pub="1864-8312-79-485" ID-Pensoft-UUID="12D8C5F9E3F15CBCA235F81E56FA0BC8" ID-ZooBank="3C3B731E89B44D1B9E0347BED83F8530" LSID="urn:lsid:zoobank.org:act:EA58E870-AE35-4802-AB14-665CE6ACDB39" ModsDocID="1864-8312-79-485" checkinTime="1633780201340" checkinUser="pensoft" docAuthor="Saraiva, Nicolas Eugenio de Vasconcelos, Hara, Marcos Ryotaro & DaSilva, Marcio Bernardino" docDate="2021" docId="4A856BC3DDED52B3AA407B7662A38AAC" docLanguage="en" docName="ArthropodSystPhyl 79: 485-507" docOrigin="Arthropod Systematics & amp; Phylogeny 79" docPubDate="2021-10-08" docSource="http://dx.doi.org/10.3897/asp.79.e66321" docTitle="Sertaneja crassitibialis Saraiva & Hara & DaSilva 2021, sp. nov." docType="treatment" docUuid="EA58E870-AE35-4802-AB14-665CE6ACDB39" docUuidSource="ZooBank" docVersion="3" id="12D8C5F9E3F15CBCA235F81E56FA0BC8" lastPageNumber="485" masterDocId="12D8C5F9E3F15CBCA235F81E56FA0BC8" masterDocTitle="Harvestmen in the semiarid: a new genus and three new species of Pachylinae (Opiliones: Gonyleptidae) from Caatinga dry vegetation, with a cladistic analysis" masterLastPageNumber="507" masterPageNumber="485" pageId="0" pageNumber="485" updateTime="1635284737390" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Harvestmen in the semiarid: a new genus and three new species of Pachylinae (Opiliones: Gonyleptidae) from Caatinga dry vegetation, with a cladistic analysis</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Saraiva, Nicolas Eugenio de Vasconcelos</mods:namePart>
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<mods:affiliation>Universidade Federal da Paraiba, Centro de Ciencias Exatas e da Natureza, Departamento de Sistematica e Ecologia, Programa de Pos-Graduacao em Ciencias Biologicas (Zoologia), Joao Pessoa, PB, Brazil; Nicolas E. de V. Saraiva [nicools. eugenio @ gmail. com]; Marcio B. DaSilva [1940 @ uol. com. br]</mods:affiliation>
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<mods:nameIdentifier type="email">nicools.eugenio@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hara, Marcos Ryotaro</mods:namePart>
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<mods:affiliation>Universidade de Sao Paulo, Escola de Artes, Ciencias e Humanidades (EACH), Sao Paulo, SP, Brazil; Marcos R. Hara [marcosrh @ usp. br]</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>DaSilva, Marcio Bernardino</mods:namePart>
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<mods:affiliation>Universidade Federal da Paraiba, Centro de Ciencias Exatas e da Natureza, Departamento de Sistematica e Ecologia, Programa de Pos-Graduacao em Ciencias Biologicas (Zoologia), Joao Pessoa, PB, Brazil; Nicolas E. de V. Saraiva [nicools. eugenio @ gmail. com]; Marcio B. DaSilva [1940 @ uol. com. br]</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-10-08</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>79</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>485</mods:start>
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<mods:end>507</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.79.e66321</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.79.e66321</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-79-485</mods:identifier>
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<mods:identifier type="ZooBank">3C3B731E89B44D1B9E0347BED83F8530</mods:identifier>
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<mods:identifier type="Pensoft-UUID">12D8C5F9E3F15CBCA235F81E56FA0BC8</mods:identifier>
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<subSection pageId="0" pageNumber="485" type="3. results">
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<treatment ID-GBIF-Taxon="189469082" LSID="urn:lsid:zoobank.org:act:EA58E870-AE35-4802-AB14-665CE6ACDB39" httpUri="http://treatment.plazi.org/id/4A856BC3DDED52B3AA407B7662A38AAC" lastPageNumber="485" pageId="0" pageNumber="485">
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<subSubSection pageId="0" pageNumber="485" type="nomenclature">
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<paragraph pageId="0" pageNumber="485">
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3.3.4.
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<taxonomicName LSID="http://zoobank.org/EA58E870-AE35-4802-AB14-665CE6ACDB39" authority="Saraiva & Hara & DaSilva, 2021" authorityName="Saraiva & Hara & DaSilva" authorityYear="2021" family="Gonyleptidae" genus="Sertaneja" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Sertaneja crassitibialis" order="Opiliones" pageId="0" pageNumber="485" rank="species" species="crassitibialis" status="sp. nov.">Sertaneja crassitibialis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="485">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="485" type="description">
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<paragraph pageId="0" pageNumber="485">
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">Figs 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">, 7</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sertaneja crassitibialis sp. nov. (UFPB OP- 171), apical portion of penis. A: dorsal view; B: right lateral view." figureDoi="10.3897/asp.79.e66321.figure8" httpUri="https://binary.pensoft.net/fig/596981" pageId="0" pageNumber="485">, 8</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Photographs of live Sertaneja gen. nov. A: Female of S. crassitibialis sp. nov. (courtesy of Leonardo Sousa Carvalho); B: Male of S. bicuspidata sp. nov.; C, E: Male of S. falcata sp. nov.; D: Female of S. falcata sp. nov." figureDoi="10.3897/asp.79.e66321.figure12" httpUri="https://binary.pensoft.net/fig/596985" pageId="0" pageNumber="485">, 12A</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Geographic distribution of Sertaneja gen. nov. and biomes. A: northeastern Brazil detail; B: distribution compared to Pachylinae groups." figureDoi="10.3897/asp.79.e66321.figure13" httpUri="https://binary.pensoft.net/fig/596986" pageId="0" pageNumber="485">, 13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="485" type="type locality">
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<paragraph pageId="0" pageNumber="485">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="485">
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BRAZIL,
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<normalizedToken originalValue="Ceará">Ceara</normalizedToken>
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:
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<normalizedToken originalValue="Quixadá">Quixada</normalizedToken>
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, Serra do Urucu,
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<normalizedToken originalValue="Santuário">Santuario</normalizedToken>
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Nossa Senhora Imaculada Rainha do
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<normalizedToken originalValue="Sertão">Sertao</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="485" type="type material">
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<paragraph pageId="0" pageNumber="485">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="485">
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<emphasis bold="true" pageId="0" pageNumber="485">Holotype.</emphasis>
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BRAZIL.
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<normalizedToken originalValue="Ceará">Ceara</normalizedToken>
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:
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<normalizedToken originalValue="Quixadá">Quixada</normalizedToken>
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, Serra do Urucu,
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<normalizedToken originalValue="Santuário">Santuario</normalizedToken>
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Nossa Senhora Imaculada Rainha do
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<normalizedToken originalValue="Sertão">Sertao</normalizedToken>
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, elev. 344 m, 19.iv.2014, DaSilva M.B. and Saraiva N.E.V., ♂ (UFPB OP-487).
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<emphasis bold="true" pageId="0" pageNumber="485">Paratypes.</emphasis>
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BRAZIL.
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<normalizedToken originalValue="Ceará">Ceara</normalizedToken>
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: Ubajara, PARNA de Ubajara, Brejo de Altitude, 700 m, 27.i.2014, DaSilva M.B., Saraiva N.E.V. and Sampaio C., 1 ♂ (MNRJ 60614) [genitalia used for SEM]; Ubajara, PARNA de Ubajara, Rio Gameleira, 840 m, 27.i.2014, DaSilva M.B.; Saraiva N.E.V. and Sampaio C., 2 ♀ (UFPB OP-480), 1 ♀ (UFPB OP-727) [female description]; Ubajara, PARNA de Ubajara, Gr. Morcego Branco, Gr. Ubajara, Trilha
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<normalizedToken originalValue="Portão">Portao</normalizedToken>
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Planalto, 22.x.2011, DeSouza A.M., Araujo E.S., Carvalho L., 1 ♀ (MNRJ 60615); Ubajara, PARNA de Ubajara, Trilha da Samambaia, 30.iv.2018, Silvino A.C.S. and Saraiva N.E.V., 1 ♂ 1 ♀ 1 imm. (MZUSP 76646) [male dissected].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="485" type="diagnosis">
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<paragraph pageId="0" pageNumber="485">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="485">
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Differs from the other species by the lower density of tubercles and armature (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6A</figureCitation>
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); ocularium is the tallest in the genus and bears a robust, single spine (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6C</figureCitation>
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); lateral margin of DS with a large acuminate tubercle on the external row, placed close to scute area III (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6C</figureCitation>
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); scute area III paramedian pair of spines slender when compared to
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<taxonomicName genus="S." lsidName="S. bicuspidata" pageId="0" pageNumber="485" rank="species" species="bicuspidata">
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<emphasis italics="true" pageId="0" pageNumber="485">S. bicuspidata</emphasis>
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</taxonomicName>
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sp. nov. (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6C</figureCitation>
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); coxa IV with an unbranched, robust prodorsal apical apophysis, missing a retro-apical apophysis (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6D</figureCitation>
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); trochanter IV short, square-shaped with a robust, blunt central apophysis (Figs
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6D</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">7</figureCitation>
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); femur IV less armed; tibia IV sexually dimorphic, swollen in males (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">7</figureCitation>
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); penis
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<normalizedToken originalValue="glans’">glans'</normalizedToken>
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stylus with an angular dorso-apical projection (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sertaneja crassitibialis sp. nov. (UFPB OP- 171), apical portion of penis. A: dorsal view; B: right lateral view." figureDoi="10.3897/asp.79.e66321.figure8" httpUri="https://binary.pensoft.net/fig/596981" pageId="0" pageNumber="485">8B</figureCitation>
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); VP macrosetae spatulated, except for the basal MS D pair and MS E; MS B inserted very basally; and MS D pairs much apart from each other (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sertaneja crassitibialis sp. nov. (UFPB OP- 171), apical portion of penis. A: dorsal view; B: right lateral view." figureDoi="10.3897/asp.79.e66321.figure8" httpUri="https://binary.pensoft.net/fig/596981" pageId="0" pageNumber="485">8</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="485">
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<emphasis bold="true" pageId="0" pageNumber="485">Figure 6.</emphasis>
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<taxonomicName authorityName="Saraiva & Hara & DaSilva" authorityYear="2021" family="Gonyleptidae" genus="Sertaneja" kingdom="Animalia" lsidName="Sertaneja crassitibialis" order="Opiliones" pageId="0" pageNumber="485" rank="species" species="crassitibialis">
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<emphasis italics="true" pageId="0" pageNumber="485">Sertaneja crassitibialis</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis bold="true" pageId="0" pageNumber="485">A</emphasis>
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,
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<emphasis bold="true" pageId="0" pageNumber="485">C</emphasis>
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-
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<emphasis bold="true" pageId="0" pageNumber="485">G</emphasis>
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) Male holotype (UFPB OP-487),
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<emphasis bold="true" pageId="0" pageNumber="485">A</emphasis>
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: habitus, dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="485">C</emphasis>
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: same, right lateral view;
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<emphasis bold="true" pageId="0" pageNumber="485">D</emphasis>
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: right coxa and trochanter IV in dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="485">E</emphasis>
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: right pedipalp in lateral view;
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<emphasis bold="true" pageId="0" pageNumber="485">F</emphasis>
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: same, mesal view;
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<emphasis bold="true" pageId="0" pageNumber="485">G</emphasis>
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: ocularium frontal view. (
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<emphasis bold="true" pageId="0" pageNumber="485">B</emphasis>
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) Female paratype (UFPB OP-727) habitus, dorsal view. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="485">
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<emphasis bold="true" pageId="0" pageNumber="485">Figure 7.</emphasis>
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<taxonomicName authorityName="Saraiva & Hara & DaSilva" authorityYear="2021" family="Gonyleptidae" genus="Sertaneja" kingdom="Animalia" lsidName="Sertaneja crassitibialis" order="Opiliones" pageId="0" pageNumber="485" rank="species" species="crassitibialis">
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<emphasis italics="true" pageId="0" pageNumber="485">Sertaneja crassitibialis</emphasis>
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</taxonomicName>
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sp. nov. Male holotype (UFPB OP-487) right trochanter-tibia IV.
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<emphasis bold="true" pageId="0" pageNumber="485">A</emphasis>
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: dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="485">B</emphasis>
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: prolateral view;
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<emphasis bold="true" pageId="0" pageNumber="485">C</emphasis>
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: ventral view;
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<emphasis bold="true" pageId="0" pageNumber="485">D</emphasis>
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: retro-lateral view. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/asp.79.e66321.figure8" httpUri="https://binary.pensoft.net/fig/596981" pageId="0" pageNumber="485" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="485">
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<emphasis bold="true" pageId="0" pageNumber="485">Figure 8.</emphasis>
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<taxonomicName authorityName="Saraiva & Hara & DaSilva" authorityYear="2021" family="Gonyleptidae" genus="Sertaneja" kingdom="Animalia" lsidName="Sertaneja crassitibialis" order="Opiliones" pageId="0" pageNumber="485" rank="species" species="crassitibialis">
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<emphasis italics="true" pageId="0" pageNumber="485">Sertaneja crassitibialis</emphasis>
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</taxonomicName>
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sp. nov. (UFPB OP-171), apical portion of penis.
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<emphasis bold="true" pageId="0" pageNumber="485">A</emphasis>
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: dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="485">B</emphasis>
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: right lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="485" type="description">
|
||
<paragraph pageId="0" pageNumber="485">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="485">
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Holotype MALE</emphasis>
|
||
(UFPB OP-487; Figs
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">7</figureCitation>
|
||
):
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Measurements</emphasis>
|
||
: DSL: 4.91; DSW: 4.67; LI: 9.35; LII: 16.17; LIII: 12.44; LIV: 16.14.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Dorsum</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6A, C, G</figureCitation>
|
||
): Prosoma sparsely covered by small granule-like tubercles. Ocularium twice as tall as the eye diameter, with a robust, conical spine projected anteriorly, slightly larger than the ocularium (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6G</figureCitation>
|
||
). Two pair of ozopores placed on the lateral surface, both slit shaped; lateral channel well developed, conspicuous and with small sensorial pegs. Scute area II, III and IV with a central transversal row of tubercles (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6A</figureCitation>
|
||
). Scute area III paramedian pair of spines slightly curved posteriorly in all its extension, barely reaching scute area IV (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6A</figureCitation>
|
||
). Lateral margin of DS with an external row of tubercles increasing in size posteriorly, becoming acuminated, largest one placed near scute area III (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6C</figureCitation>
|
||
). Posterior margin of DS and free tergites I-III each with a row of similar sized round tubercles. Anal operculum tuberculate.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Venter</emphasis>
|
||
: Coxae I-IV with low tubercles; coxa I with median row of enlarged tubercles. Coxa IV covered by granule-like tubercles.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Chelicerae</emphasis>
|
||
: Segment I with the posterior portion of the bulla bearing a transversal row of four tubercles. Fixed and movable fingers each with five and five to six teeth, respectively.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Pedipalps</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6E, F</figureCitation>
|
||
): Trochanter inflated dorsally; ventral face with two setiferous tubercles, mesal tubercle largest. Femur with a dorsal median row of six tubercles (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6E</figureCitation>
|
||
); ventral surface with a large basal setiferous tubercle and a retro-lateral row of three tubercles on apical half. Tibial setation: mesal and lateral IiIi. Tarsal setation: mesal IIi, lateral IiIi.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Legs</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">7</figureCitation>
|
||
): Coxae I-III each with one prodorsal and 1 retro-dorsal apophysis; coxa II retro-dorsal apophysis trifid and fused in the apex with coxa III prodorsal one. Coxa IV large, tuberculate, bearing a robust, unbranched prodorsal apical apophysis, this almost transversally inserted, slightly curved downwards and posteriorly (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6A, C, D</figureCitation>
|
||
). Trochanter IV with a robust, slightly bifid, blunt central apophysis, a prodorsal sub-apical enlarged tubercle, and an acuminated, straight retro-apical apophysis (Figs
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">7</figureCitation>
|
||
). Femur III with an enlarged retro-apical dorsal spine; ventral face with a pro- and retro-lateral rows of tubercles increasing in size apically on apical half. Tibia III sub-apical surface slightly swollen, with retro-ventral sub-apical row of enlarged tubercles. Femur IV slightly curved inwards and upwards; dorsal surface covered by small tubercles, with a pair of paralateral apical spines slightly curved upwards (prolateral spine largest) (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">7A</figureCitation>
|
||
); lateral rows of tubercles slightly increasing in size apically, retro-lateral row largest; with pro and retro-ventral rows that increase in size apically, proventral row largest and with tubercles more close to each other (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">7C</figureCitation>
|
||
); apically with a pair of paralateral spines, prolateral largest (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">7C</figureCitation>
|
||
). Patella IV tuberculate, with a slightly swollen retro-dorsal apical spine (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">7A</figureCitation>
|
||
). Tibia IV swollen at apical 3/4 of its length, with a proventral and retro-ventral row of enlarged sub-apical tubercles, and a pair of paralateral ventro-apical spines (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sertaneja crassitibialis sp. nov. Male holotype (UFPB OP- 487) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure7" httpUri="https://binary.pensoft.net/fig/596980" pageId="0" pageNumber="485">7B, C, D</figureCitation>
|
||
). Tarsal counts: 6, 8, 6, 6.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Penis</emphasis>
|
||
(UFPB OP-171; Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sertaneja crassitibialis sp. nov. (UFPB OP- 171), apical portion of penis. A: dorsal view; B: right lateral view." figureDoi="10.3897/asp.79.e66321.figure8" httpUri="https://binary.pensoft.net/fig/596981" pageId="0" pageNumber="485">8</figureCitation>
|
||
): VP apex slightly concave, lateral margin sub-straight. Proximal portion of VP with four pairs MS A and one pair MS B placed far from A4 and placed more basally. MS C set apically curved and showing asymmetry between left (3) and right (2) sides (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sertaneja crassitibialis sp. nov. (UFPB OP- 171), apical portion of penis. A: dorsal view; B: right lateral view." figureDoi="10.3897/asp.79.e66321.figure8" httpUri="https://binary.pensoft.net/fig/596981" pageId="0" pageNumber="485">8A</figureCitation>
|
||
). MS D apical pair flattened and acuminate, basal spine conical and blunt; and two pairs of MS E placed more ventrally and close to MS C and MS D (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sertaneja crassitibialis sp. nov. (UFPB OP- 171), apical portion of penis. A: dorsal view; B: right lateral view." figureDoi="10.3897/asp.79.e66321.figure8" httpUri="https://binary.pensoft.net/fig/596981" pageId="0" pageNumber="485">8B</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Coloration</emphasis>
|
||
: DS predominantly dark grayish yellowish brown (81) with scute areas limits, area III paramedian spines, and specks on prosoma dark orange yellow (72). Chelicerae, pedipalps, coxae I-III, and legs I-III dark yellow (88). Leg IV dark grayish yellowish brown (81).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="485">
|
||
<emphasis bold="true" pageId="0" pageNumber="485">FEMALE</emphasis>
|
||
(UFPB OP-727; Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6B</figureCitation>
|
||
):
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Measurements</emphasis>
|
||
: DSL: 4.70; DSW: 4.26; LI: 8.88; LII: 15.36; LIII: 11.19; LIV: 14.81.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Dorsum</emphasis>
|
||
: Overall less developed armature and tubercles. DS alpha (α), coda slightly longer. Ocularium spine slender. Scute area III central armature small. Lateral margin of DS with external row of smaller tubercles than in male.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Legs</emphasis>
|
||
: Coxa IV narrower and shorter than in male, with an obliquely inserted, slender, conical, prodorsal apical apophysis, slightly curved downwards. Trochanter III retro-apical apophysis reduced. Trochanter IV narrower than in male, prolateral face unarmed, retro-lateral face as in males but with smaller apical apophysis. Femur III with ventral rows of small tubercles. Femur IV less armed, ventral rows of tubercles slightly increasing in size apically, dorso- and ventro-apical pair of spines small. Patella IV prodorsal apical projection reduced. Tibiae III-IV unarmed, not inflated. Tarsal counts: 6, 7, 6, 6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="485" type="variation">
|
||
<paragraph pageId="0" pageNumber="485">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="485">
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Males</emphasis>
|
||
(n = 3):
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Measurements</emphasis>
|
||
: DSL: 4.91-4.99; DSW: 4.67-4.90; LI: 8.98-10.08; LII: 15.31-16.17; LIII: 11.58-12.44; LIV: 16.14-16.34.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Dorsum</emphasis>
|
||
: Ocularium spine more, or less acute. DS lateral margin external row of rounded to acuminate tubercles, becoming almost spine-like. Scute area II with tubercles placed near the lateral margins.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Legs</emphasis>
|
||
: Tibia IV retro-ventral apical spine single or bifid. Tarsal counts: 6, 8-9, 6, 6.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Females</emphasis>
|
||
(n = 5):
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Measurements</emphasis>
|
||
: DSL: 4.61-4.75; DSW: 4.20-4.55; LI: 8.56-9.45; LII: 13.93-15.97; LIII: 10.25-11.28; LIV: 14.80-15.52.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Pedipalps</emphasis>
|
||
: Pedipalp tibial setation: mesal IiIi/IIi, lateral IiIi; tarsal setation: mesal IIi, lateral IiIi/Iiii.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Legs</emphasis>
|
||
: Femur IV proventral row of tubercles slightly enlarged or not. Patella IV with or without prodorsal apical projection. Tarsal counts: 5-6, 8-9, 6, 6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="485" type="etymology">
|
||
<paragraph pageId="0" pageNumber="485">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="485">
|
||
<normalizedToken originalValue="‘Crassi-’">'Crassi-'</normalizedToken>
|
||
is a Greek adjective, meaning thick, and
|
||
<normalizedToken originalValue="‘tibia’">'tibia'</normalizedToken>
|
||
is the Latin noun meaning the podomere; combined to
|
||
<normalizedToken originalValue="‘crassitibia’">'crassitibia'</normalizedToken>
|
||
(nom. sing. fem.) in reference to the diagnostic male tibia IV of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="485" type="distribution">
|
||
<paragraph pageId="0" pageNumber="485">
|
||
Distribution (Fig.
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Geographic distribution of Sertaneja gen. nov. and biomes. A: northeastern Brazil detail; B: distribution compared to Pachylinae groups." figureDoi="10.3897/asp.79.e66321.figure13" httpUri="https://binary.pensoft.net/fig/596986" pageId="0" pageNumber="485">13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="485">
|
||
Brejos de Altitude humid montane forests of Ibiapaba mountains and Caatinga shrublands and dry forests of northern interior
|
||
<normalizedToken originalValue="Ceará">Ceara</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |