503 lines
53 KiB
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503 lines
53 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.46.105798" ID-Pensoft-Pub="1314-2615-46-1" ID-Pensoft-UUID="73A944AECDDB5130BA73E57C7CB6D83F" ID-ZooBank="1A679D4370004DD1BA0D69DA2057401E" ModsDocID="1314-2615-46-1" checkinTime="1690300902025" checkinUser="pensoft" docAuthor="Huber, Bernhard A., Meng, Guanliang, Clark, Huon L. & Cazanove, Gregory" docDate="2023" docId="4A1DB263C11D5817950E606BA2C17E7D" docLanguage="en" docName="SubterBiol 46: 1-19" docOrigin="Subterranean Biology 46" docPubDate="2023-07-24" docSource="http://dx.doi.org/10.3897/subtbiol.46.105798" docTitle="Belisana coblynau Huber & Clark 2023, sp. nov." docType="treatment" docUuid="50A34349-C82F-49FA-8679-77C8CCC86D1B" docUuidSource="ZooBank" docVersion="2" id="73A944AECDDB5130BA73E57C7CB6D83F" lastPageNumber="1" masterDocId="73A944AECDDB5130BA73E57C7CB6D83F" masterDocTitle="First blind daddy long-legs spiders from Australia and Reunion (Araneae, Pholcidae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="1" updateTime="1690301203058" updateUser="ExternalLinkService">
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<mods:titleInfo id="F706DB4DF83ECB745AEB2E933CFC6203">
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<mods:title id="D1659D76DF5AF6F9F3A675B389ABA7C2">First blind daddy long-legs spiders from Australia and Reunion (Araneae, Pholcidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="1DD39E37F60A9D605013B43C675F8521" type="personal">
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<mods:role id="B7C156DBAF9BF874F526FE405A4D0BE8">
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<mods:roleTerm id="D9AB37E2992A279C6C003D25DC2E31BF">Author</mods:roleTerm>
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<mods:namePart id="DC6DDC6CA47A5A155D309D1E4CEF75AF">Huber, Bernhard A.</mods:namePart>
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<mods:nameIdentifier id="622F678C0A4EB4A0FF09344D8C75784C" type="ORCID">https://orcid.org/0000-0002-7566-5424</mods:nameIdentifier>
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<mods:affiliation id="F3D1D6991B3337D55099B5557F50A17D">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany</mods:affiliation>
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<mods:nameIdentifier id="90551964CBBB9C4E2C8FCBC88B3ABB08" type="email">b.huber@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:name id="D235FD7B8A0F05AA9B0C2B18A508C545" type="personal">
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<mods:roleTerm id="E64E1C8E2DDF9A7481FF227CE2CE029A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DD9FD382CF7E80C58FD4CD69378ADCD2">Meng, Guanliang</mods:namePart>
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<mods:affiliation id="72420923583C9C33D21C85FA1EC01370">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany</mods:affiliation>
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</mods:name>
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<mods:name id="8CC5EDD1F8F27A329B47BBA0F809927D" type="personal">
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<mods:roleTerm id="26E6D66D6D292197CB05130118A99710">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="526AFE523AA4A5ADB8846F6A75367B2C">Clark, Huon L.</mods:namePart>
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<mods:affiliation id="8D23512CA870732642347E9412A8B46F">Bennelongia Environmental Consultants, 5 Bishop Street, Jolimont WA 6014, Australia & Collections and Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia</mods:affiliation>
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</mods:name>
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<mods:name id="01167DCB3B1B579B879EF1CBB2D86546" type="personal">
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<mods:roleTerm id="EF2B80854EE182CB0BF1E659518550D5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7BCA47FA4E44D112EC53C0D8E7C7CE36">Cazanove, Gregory</mods:namePart>
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<mods:affiliation id="D508E2CED2A6AA3F7FE0591EEAE05AA0">Museum d'Histoire Naturelle, Saint Denis, Reunion</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="3520EEBD3A50CB10A0AEEDA4020378BF">text</mods:typeOfResource>
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<mods:titleInfo id="AC6978A6F0C17801DC9E8BCF7AED3E2C">
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<mods:title id="7B80576D18CE415832E861583111774E">Subterranean Biology</mods:title>
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<mods:part id="750B54F7784F4697FBD291DB4F312719">
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<mods:date id="A47E023FED115FA33334365211A38A26">2023</mods:date>
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<mods:detail id="79404FC4BDB7AAB49840F2A9FAB44335" type="pubDate">
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<mods:number id="EEB0A09719BB99940AA43C393CC85526">2023-07-24</mods:number>
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<mods:detail id="C37A0FCC4672B70B0636154053357646" type="volume">
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<mods:number id="6BB72781505DB894831455A4C5530E59">46</mods:number>
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<mods:start id="2FB07246AD8447660A3EE80CF34CB796">1</mods:start>
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<mods:end id="A35910988A6D2EBED7C419A7A0929228">19</mods:end>
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<mods:url id="C07EA5C7B979D838DBA7DFC76D9506BF">http://dx.doi.org/10.3897/subtbiol.46.105798</mods:url>
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<mods:classification id="18CA4ED049ED21CC64A1EB44FE2DC183">journal article</mods:classification>
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<mods:identifier id="43DABE05C3322C164E0785FEDADD8EFC" type="DOI">http://dx.doi.org/10.3897/subtbiol.46.105798</mods:identifier>
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<mods:identifier id="04389CDA7A4AFD3CD0F9BB6EE37E1FA9" type="Pensoft-Pub">1314-2615-46-1</mods:identifier>
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<mods:identifier id="71F8E37AB2FDB5D3EF5C962999A9AC41" type="ZooBank">1A679D4370004DD1BA0D69DA2057401E</mods:identifier>
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<mods:identifier id="211F046EA5BB540DA869DD929A2D2657" type="Pensoft-UUID">73A944AECDDB5130BA73E57C7CB6D83F</mods:identifier>
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</mods:mods>
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<treatment id="4A1DB263C11D5817950E606BA2C17E7D" LSID="urn:lsid:zoobank.org:act:50A34349-C82F-49FA-8679-77C8CCC86D1B" httpUri="http://treatment.plazi.org/id/4A1DB263C11D5817950E606BA2C17E7D" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="DF3876EC95824E2E00C4E6450A58EBBB" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="8D3880D0A46C4D922BD3134E837DFC73" pageId="0" pageNumber="1">
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<taxonomicName id="D46BDBD7A37114BD6A64CE2BC1D9F3B1" LSID="https://zoobank.org/50A34349-C82F-49FA-8679-77C8CCC86D1B" authority="Huber & Clark" authorityName="Huber & Clark" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana coblynau" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coblynau" status="sp. nov.">Belisana coblynau Huber & Clark</taxonomicName>
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<taxonomicNameLabel id="2A80831262A1A99AE205CD773859DF33" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="52A108B48DBBF9DB899A48257F9BB1E8" pageId="0" pageNumber="1" type="description">
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<paragraph id="AFC9F76A3DF5E7D9F3AAEC6BC41C6853" pageId="0" pageNumber="1">
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<figureCitation id="D75A0CD7364C706C4BF6FF463E4012E7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A male habitus, lateral view B female abdomen, ventral view C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ep, epigynal pockets; lp, lateral pocket; pp, pore plate; sp, spinnerets. Scale bars: 0.5 mm (A); 0.1 mm (B-D)." figureDoi="10.3897/subtbiol.46.105798.figure1" httpUri="https://binary.pensoft.net/fig/883297" pageId="0" pageNumber="1">Figs 1</figureCitation>
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<figureCitation id="F78F5018ED2208DB3CB84C6486F6F7E9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A, B left male palp, prolateral and retrolateral views C male chelicerae, frontal view D, E left procursus, prolateral and retrolateral views F cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; ep, epigynal pocket; f, femur; lp, lateral pocket; pp, pore plate; pr, procursus; ta, tarsus; ti, tibia. Scale bars: 0.2 mm (A, B, F); 0.1 mm (C-E)." figureDoi="10.3897/subtbiol.46.105798.figure2" httpUri="https://binary.pensoft.net/fig/883298" pageId="0" pageNumber="1">, 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="FFAE3B879AEADAF7A98F7B093980F94E" pageId="0" pageNumber="1" type="type material">
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<paragraph id="7C36914620EFC6798F2DED6C2866C8BD" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="918E9B4C220DBCC07CD4EF9C262CE7A9" pageId="0" pageNumber="1">
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Australia - Western Australia • ♂ holotype; ~85 km ESE Pannawonica;
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<geoCoordinate id="AC3D15785B64664FABD60493C926DB3B" degrees="21.8836" direction="south" orientation="latitude" precision="5" value="-21.8836">21.8836°S</geoCoordinate>
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,
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<geoCoordinate id="6B70FBD4A7EE2B17A45A0A4D975B9875" degrees="117.1211" direction="east" orientation="longitude" precision="5" value="117.1211">117.1211°E</geoCoordinate>
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; 590 m a.s.l.; 25 Jun. 2019; M.D. Scanlon and H.L. Clark leg.; WAM T158896.
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</paragraph>
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</subSubSection>
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<subSubSection id="D3DF7D1C8E8CAFAB861D109FEB5E4F0F" pageId="0" pageNumber="1" type="type locality">
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<paragraph id="D773F01469DA8A8AB2D6602C7ED2F6F5" pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph id="34DB5E6F8F7DD45207D8B7268E2D14E4" pageId="0" pageNumber="1">
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Australia, Western Australia, ~85 km ESE Pannawonica,
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<geoCoordinate id="C53A128D46F831A5AA488871FAC0DEFD" degrees="21.8836" direction="south" orientation="latitude" precision="5" value="-21.8836">21.8836°S</geoCoordinate>
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,
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<geoCoordinate id="151FE76FC6CBE558BB6736A63C06A6E8" degrees="117.1211" direction="east" orientation="longitude" precision="5" value="117.1211">117.1211°E</geoCoordinate>
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, 590 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection id="FA66F041631B8D3CB558D1ED57DD306D" pageId="0" pageNumber="1" type="type specimen">
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<paragraph id="8C9A369104D134AF60E6A54C18134B9E" pageId="0" pageNumber="1">Type specimen.</paragraph>
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<paragraph id="F137164D46A6B5F239BE0E1024421E91" pageId="0" pageNumber="1">
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Holotype male, in ethanol. Original labels: "WA: JSE, ca 87 km ESE Pannawonica,
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<geoCoordinate id="6CFD3A963225FDC72FF47E8E8B64EBFD" degrees="21" direction="south" minutes="53" orientation="latitude" precision="1" seconds="43.450" value="-21.895403">21°53'43.450"S</geoCoordinate>
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,
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<geoCoordinate id="B3F2E23730F59F096C9ACE260253DECD" degrees="117" direction="east" minutes="07" orientation="longitude" precision="1" seconds="48.63" value="117.13018">117°07'48.63"E</geoCoordinate>
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(GDA 94) 25. Jun. 2019, Scanlon MD, Clark HL (KRC 0364), Trog scrape, 35 m",
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<normalizedToken id="C9BC7CFD5C85C312483882843F593BDD" originalValue="“KRC0364”">"KRC0364"</normalizedToken>
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, " WAM T 158896", "100% EtOH".
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</paragraph>
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</subSubSection>
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<subSubSection id="77D47797D833170883ED614F4D56FCB2" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="3FC65CDC74378D785AB5C824477C61B2" pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph id="8264792ECF7217F944A0DEC8E9035BB8" pageId="0" pageNumber="1">
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<materialsCitation id="1A150ABFFD589466870D63B6E5EB5B4E" collectingDate="2019-04-25" collectorName="M. D. Scanlon, H. L. Clark" country="Australia" elevation="540" latitude="-21.8583" location="Remark" longLatPrecision="7" longitude="117.0972" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
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<collectingCountry id="ED9EDAD144F6E69C952E67C4BB8FBEA3" name="Australia">Australia</collectingCountry>
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-
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<collectingRegion id="28997DF1B1ECF8975105733A7855169E" country="Australia" name="Western Australia">Western Australia</collectingRegion>
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•
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<specimenCount id="1DA8488FA7DF577FDE07FDDEF452B4C7" type="female">1 ♀</specimenCount>
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, assigned tentatively, see
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<location id="A5A17F45F9E732B5D2433BECF513C88B" LSID="urn:lsid:plazi:treatment:4A1DB263C11D5817950E606BA2C17E7D:A5A17F45F9E732B5D2433BECF513C88B" country="Australia" latitude="-21.8583" longLatPrecision="7" longitude="117.0972" name="Remark" stateProvince="Western Australia">Remark</location>
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below; same locality as holotype but
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<quantity id="1D11DD0BC9635EDDEF91CD5B8680494F" metricMagnitude="3" metricUnit="m" metricValue="3.7" unit="km" value="3.7">3.7 km</quantity>
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NW;
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<geoCoordinate id="54DB1D7120E80C1380606E24747E4F4C" degrees="21.8583" direction="south" orientation="latitude" precision="5" value="-21.8583">21.8583°S</geoCoordinate>
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,
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<geoCoordinate id="76922F2CBE77A780F063F49149433155" degrees="117.0972" direction="east" orientation="longitude" precision="5" value="117.0972">117.0972°E</geoCoordinate>
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;
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<elevation id="C589C607A1623F43525D0850DF5F4378" metricMagnitude="2" metricUnit="m" metricValue="5.4" unit="m" value="540.0">
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<quantity id="E5E0E539DD12B09C01C4DA14BD2F1604" metricMagnitude="2" metricUnit="m" metricValue="5.4" unit="m" value="540.0">540 m</quantity>
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a.s.l.
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</elevation>
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;
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<collectingDate id="7A717F05375C6F020E3B7779A1B782CB" value="2019-04-25">25 Apr. 2019</collectingDate>
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;
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<collectorName id="F42D99DE3132C04B04F481ED7EC9115A">M.D. Scanlon</collectorName>
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and
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<collectorName id="4535B8F292CAEF482E5044FD73CBA00A">H.L. Clark</collectorName>
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leg.;
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<normalizedToken id="C84D1F45360C407D76E0A3EA9F3C5CA4" originalValue="“KRC0226”">"KRC0226"</normalizedToken>
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, WAM T158897
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</materialsCitation>
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•
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<materialsCitation id="75B98B0D26C55AF3D666C57F87446FDA" collectingDate="2019-04-26" collectorName="M. D. Scanlon, H. L. Clark" country="Australia" elevation="570" latitude="-21.8954" location="Remark" longLatPrecision="7" longitude="117.1302" specimenCount="1" specimenCount-juvenile="1" stateProvince="Western Australia">
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<specimenCount id="632CFC7EB31329270F0B053EC208C489" type="generic">1 juv.</specimenCount>
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; same locality as holotype but
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<quantity id="4CDB7D348567A8BD04CB2147364F88D0" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="km" value="1.6">1.6 km</quantity>
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SE;
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<geoCoordinate id="212F6A5FA2606970B0318001B649E9FA" degrees="21.8954" direction="south" orientation="latitude" precision="5" value="-21.8954">21.8954°S</geoCoordinate>
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,
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<geoCoordinate id="04FCF14FAAA8635E6B2144257AF16A87" degrees="117.1302" direction="east" orientation="longitude" precision="5" value="117.1302">117.1302°E</geoCoordinate>
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;
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<elevation id="6A12FBE5A0B54ABF06DD862A3428416B" metricMagnitude="2" metricUnit="m" metricValue="5.7" unit="m" value="570.0">
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<quantity id="7C150E24141D731EE9A3F09237C63887" metricMagnitude="2" metricUnit="m" metricValue="5.7" unit="m" value="570.0">570 m</quantity>
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a.s.l.
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</elevation>
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;
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<collectingDate id="ADEEDFC2DB0815A21B7A4246DF70BF98" value="2019-04-26">26 Apr. 2019</collectingDate>
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;
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<collectorName id="ABD91DBBCF44B6227E16F37FAA0F53F7">M.D. Scanlon</collectorName>
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and
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<collectorName id="CFF1A0AB6E43F1500521A3761AB391AB">H.L. Clark</collectorName>
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leg.;
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<normalizedToken id="6C89795C434C3A7DA38AE5266B24D0C0" originalValue="“KRC0254”">"KRC0254"</normalizedToken>
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, WAM T158895
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B48946543EA4CB3CB9E05631B5441013" pageId="0" pageNumber="1" type="remarks">
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<paragraph id="ECC878646B14D66BB073C5F415A4E4B6" pageId="0" pageNumber="1">Remark.</paragraph>
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<paragraph id="B77CC1DE6BCB531788702EE2FA86DC6A" pageId="0" pageNumber="1">
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The genetic distance (Table
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<tableCitation id="E2F94B33E323F4E3F5BB6AD30D110328" captionStart="Table 1" captionStartId="T1" captionText="Table 1. GenBank accession numbers and genetic distances. CO 1 accession numbers of the three Belisana coblynau specimens and genetic distances (K 2 P) among them and other Belisana species taken from Eberle et al. (2018)." httpUri="http://table.plazi.org/id/EA1F0C3BC18DB8D9FDDB6C0125BE16CC" pageId="0" pageNumber="1" tableUuid="EA1F0C3BC18DB8D9FDDB6C0125BE16CC">1</tableCitation>
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) between the male holotype and the juvenile specimen was 0.5%, strongly suggesting that the two are conspecific. The distances between these two specimens and the female was much higher, at 8.5 and 8.8%, respectively. This is within a range that in
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<taxonomicName id="468877266A12BF0518839BC422524291" class="Arachnida" family="Pholcidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
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is always problematic due to the overlap of intra- and interspecific genetic (CO1) variation between approximately 8 and 11% (
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<bibRefCitation id="6FBF064522B53668B03A896FC540C23B" DOI="https://doi.org/10.1111/j.1463-6409.2006.00239.x" author="Astrin, JJ" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="1" pagination="441 - 457" refId="B1" refString="Astrin, JJ, Huber, BA, Misof, B, Kluetsch, CFC, 2006. Molecular taxonomy in pholcid spiders (Pholcidae, Araneae): evaluations of species identification methods using CO1 and 16S rRNA. Zoologica Scripta 35: 441 - 457, DOI: https://doi.org/10.1111/j.1463-6409.2006.00239.x" title="Molecular taxonomy in pholcid spiders (Pholcidae, Araneae): evaluations of species identification methods using CO 1 and 16 S rRNA." url="https://doi.org/10.1111/j.1463-6409.2006.00239.x" volume="35" year="2006">Astrin et al. 2006</bibRefCitation>
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;
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|
<bibRefCitation id="DE31C38CEDA070A184246310F715BD4B" author="Huber, BA" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="1" refId="B34" refString="Huber, BA, Meng, G, Duperre, N, Astrin, J, Herrera, M, in press. Andean giants: Priscula spiders from Ecuador, with notes on species groups and egg-sac troglomorphism (Araneae: Pholcidae). European Journal of Taxonomy." title="Andean giants: Priscula spiders from Ecuador, with notes on species groups and egg-sac troglomorphism (Araneae: Pholcidae). European Journal of Taxonomy." year="in press">Huber et al. in press</bibRefCitation>
|
|
). In the case of the female, we are thus unsure about its taxonomic status.
|
|
</paragraph>
|
|
<caption id="EA1F0C3BC18DB8D9FDDB6C0125BE16CC" ID-Table-UUID="EA1F0C3BC18DB8D9FDDB6C0125BE16CC" httpUri="http://table.plazi.org/id/EA1F0C3BC18DB8D9FDDB6C0125BE16CC" pageId="0" pageNumber="1" start="Table 1" startId="T1">
|
|
<paragraph id="B70DFAE4D4031A695E6D21B2D9047BE8" pageId="0" pageNumber="1">
|
|
<emphasis id="D24C8719F1338683331908043469D335" bold="true" pageId="0" pageNumber="1">Table 1.</emphasis>
|
|
GenBank accession numbers and genetic distances. CO1 accession numbers of the three
|
|
<taxonomicName id="CFAAFC12E6AF4E74B362EC0C4B0080C8" authorityName="Huber & Clark" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana coblynau" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coblynau">
|
|
<emphasis id="390295E008C6C8441D39C97D49AC82BC" italics="true" pageId="0" pageNumber="1">Belisana coblynau</emphasis>
|
|
</taxonomicName>
|
|
specimens and genetic distances (K2P) among them and other
|
|
<taxonomicName id="59CE8CBD2B2B83821650F9CD5BF5ABCF" authorityName="Thorell" authorityYear="1898" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C579D48E7E1820E81E5A6582367BD5AF" italics="true" pageId="0" pageNumber="1">Belisana</emphasis>
|
|
</taxonomicName>
|
|
species taken from
|
|
<bibRefCitation id="6AC7B8CDBA014E5D67450E6556629450" DOI="https://doi.org/10.1186/s12862-018-1244-8" author="Eberle, J" journalOrPublisher="L'Entomologiste" pageId="0" pageNumber="1" refId="B13" refString="Eberle, J, Dimitrov, D, Valdez-Mondragon, A, Huber, BA, 2018. Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology 18: e141. https://doi.org/10.1186/s12862-018-1244-8" title="Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology 18: e 141." url="https://doi.org/10.1186/s12862-018-1244-8" year="2018">Eberle et al. (2018)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="CFFA950FC5A07E51B2C548E193D4A936" pageId="0" pageNumber="1">
|
|
<table id="EB5F200838942A28198B858CD96C6A93" pageId="0" pageNumber="1">
|
|
<tr id="1AE8C89B0B988FBD7B8719CD4375471C" pageId="0" pageNumber="1">
|
|
<th id="4F9D624DDED7DA8A6F93D5279A3E1100" colspan="1" pageId="0" pageNumber="1" rowspan="1">Taxon</th>
|
|
<th id="085634C4803BA514440487309F8E639F" colspan="1" pageId="0" pageNumber="1" rowspan="1">OQ525972</th>
|
|
<th id="008B0CBF2BA8D5BD67FB044571441508" colspan="1" pageId="0" pageNumber="1" rowspan="1">OQ525971</th>
|
|
<th id="4CF047D45BA144EE133E73D733129669" colspan="1" pageId="0" pageNumber="1" rowspan="1">OQ525973</th>
|
|
</tr>
|
|
<tr id="4B873FF06691B68974AE621A2F04FA1C" pageId="0" pageNumber="1">
|
|
<td id="1958B0FA76CB8BEA9789F69EEAF2C576" colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
OQ525972,
|
|
<taxonomicName id="D1E5D4C96BF80BF7628844BEFDC08226" authorityName="Huber & Clark" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana coblynau" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coblynau">
|
|
<emphasis id="BD6DB927677FBE564EB52770F5A4FA3E" italics="true" pageId="0" pageNumber="1">Belisana coblynau</emphasis>
|
|
</taxonomicName>
|
|
male
|
|
</td>
|
|
<td id="060A8673D9AB49C293D75976B8C4E6BE" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td id="58BE624005F1530FFF135AA8A380CDA0" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td id="F828A1B3F6154705F66CFA0692C9D6F4" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
</tr>
|
|
<tr id="CC0C78A1D622C5214F7FD5AA0BA71E7A" pageId="0" pageNumber="1">
|
|
<td id="07E647412F8F40138270FD77823AAD6A" colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
OQ525971,
|
|
<taxonomicName id="7D9794C7657F53AED7CC2BBDE0FAE080" authorityName="Huber & Clark" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana coblynau" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coblynau">
|
|
<emphasis id="E981FD8C7C0749FAB2DAFED7FE1864FF" italics="true" pageId="0" pageNumber="1">Belisana coblynau</emphasis>
|
|
</taxonomicName>
|
|
female
|
|
</td>
|
|
<td id="F559059A48832C6E33D84BE1605DABCD" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.088</td>
|
|
<td id="667473CA698100B9AFE65F862BFE4781" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td id="2BEEDEBF3E674B1AF0654761DF955A6E" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
</tr>
|
|
<tr id="ADFD48F03095E8CEB3D57E3F3BA8E085" pageId="0" pageNumber="1">
|
|
<td id="7BB353034927657440CB84D9A2793473" colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
OQ525973,
|
|
<taxonomicName id="AA77203AF2B2064AE1A99D1803EAE173" authorityName="Huber & Clark" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana coblynau" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coblynau">
|
|
<emphasis id="396DA2F3D1637627726980035DF84646" italics="true" pageId="0" pageNumber="1">Belisana coblynau</emphasis>
|
|
</taxonomicName>
|
|
juvenile
|
|
</td>
|
|
<td id="465CF8260E47B3B24CB289C05E0B0205" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.005</td>
|
|
<td id="1AB6CB5C1FAF0341AD46C5050BFF8629" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.085</td>
|
|
<td id="073DF3DF689CC485F5449C3AB4AE4A3B" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
</tr>
|
|
<tr id="2E840823004F37449490A3A006656B64" pageId="0" pageNumber="1">
|
|
<td id="7E9EB5417409706A69C798A435DE82A6" colspan="1" pageId="0" pageNumber="1" rowspan="1">S127_Belisana_Bor152_Bor166</td>
|
|
<td id="02DEB0CF5627D135834F568A8D744715" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.277</td>
|
|
<td id="9B376AE57641D5124457CCA0A1A12170" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.261</td>
|
|
<td id="9504181512B0AF320D1CF8E4D7E15A7B" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.277</td>
|
|
</tr>
|
|
<tr id="826C1378373473583B509660C42BB37F" pageId="0" pageNumber="1">
|
|
<td id="769CC90C8791CF667484229F609FE6C6" colspan="1" pageId="0" pageNumber="1" rowspan="1">S249_Belisana_Mal91_Mal290</td>
|
|
<td id="7E98510AE858ADA1B1DC19ED4A2AC404" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.285</td>
|
|
<td id="C74FA4FBD34871BE7182CE65FC245B08" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.283</td>
|
|
<td id="6338DF9AA7E3E523C79509E67C45ED19" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.287</td>
|
|
</tr>
|
|
<tr id="0AB448E7F4375100C587BF7A07602BD5" pageId="0" pageNumber="1">
|
|
<td id="83DF07E384F1B713D4376CCD16CDA292" colspan="1" pageId="0" pageNumber="1" rowspan="1">S242_Belisana_leuser_Mal304</td>
|
|
<td id="979B33E2072E944D2A1D70AE856B057A" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.233</td>
|
|
<td id="9B05A77129B01AAE7C7087D5B2A587EE" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.241</td>
|
|
<td id="9D9A6728766659FC6718D54940E549E7" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.233</td>
|
|
</tr>
|
|
<tr id="3DAAA6FE906C6135CCBB5832745A6268" pageId="0" pageNumber="1">
|
|
<td id="FDFF78F01DEFFCE59D5F74FB46080B2E" colspan="1" pageId="0" pageNumber="1" rowspan="1">S247_Belisana_Mal76_Mal278</td>
|
|
<td id="B8C4F808689D58BC075043F1F674F831" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.274</td>
|
|
<td id="750B5317AA947505134CABDE4A74F3B1" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.299</td>
|
|
<td id="758F72B910E512E8122DEB4DA3F46DCC" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.276</td>
|
|
</tr>
|
|
<tr id="D0871676B834748E0F921765D7AACCEF" pageId="0" pageNumber="1">
|
|
<td id="41A72D054AFF1091B22D4391F811F28D" colspan="1" pageId="0" pageNumber="1" rowspan="1">S498_Belisana_minneriya_SL123</td>
|
|
<td id="30A9DC12748C22578FCB79654D3995F6" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.238</td>
|
|
<td id="17F824150EA2D33E00396478581DCD3D" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.245</td>
|
|
<td id="90514D02CB7392B2E78B9A160FA85A7A" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.236</td>
|
|
</tr>
|
|
<tr id="478350F520621595C584B1127CA219E4" pageId="0" pageNumber="1">
|
|
<td id="11954A1E61965FA2950AF217790B3B8D" colspan="1" pageId="0" pageNumber="1" rowspan="1">S248_Belisana_Mal77_Mal279</td>
|
|
<td id="14F9818451772B9EABEB452F9EBCC9B4" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.277</td>
|
|
<td id="CBA61C72E294017FD869257E604F5B00" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.274</td>
|
|
<td id="34268EA48D7E4BF66F52D04BDE97B617" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.275</td>
|
|
</tr>
|
|
<tr id="1706E77767A3C8EDDA8198E64C3B9A61" pageId="0" pageNumber="1">
|
|
<td id="B630C8BC1B33C2E37D2C4700982ED7E7" colspan="1" pageId="0" pageNumber="1" rowspan="1">S103_Belisana_Tai4_Tai67</td>
|
|
<td id="95135607FB9A9B4C704EA982A7245EA6" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.233</td>
|
|
<td id="49B0B36BB516504AAF0913FC9B38F3C3" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.264</td>
|
|
<td id="6FE7BB238176D277C7E851358858C973" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.231</td>
|
|
</tr>
|
|
<tr id="E999698E5F0B31860D3CB17A7B494C8C" pageId="0" pageNumber="1">
|
|
<td id="8F861F728251E0F88C163FAEE836190A" colspan="1" pageId="0" pageNumber="1" rowspan="1">S230_Belisana_australis_Phi280</td>
|
|
<td id="57F242831DE9982FC693D4406D2D2D05" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.244</td>
|
|
<td id="C16DF487A7CF9B974B70B18754011FDD" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.264</td>
|
|
<td id="016B91B1F6018EB6CA299122666B5999" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.244</td>
|
|
</tr>
|
|
<tr id="0F6566EDE7F8494F30168E68766204C3" pageId="0" pageNumber="1">
|
|
<td id="26C09955D5988BF1B04426037409BB3D" colspan="1" pageId="0" pageNumber="1" rowspan="1">S125_Belisana_Bor121_Bor198</td>
|
|
<td id="C2B687F5A792DAA454FE973209A6BCD3" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.227</td>
|
|
<td id="68F2099650523681DF4C4F647D7FD486" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.241</td>
|
|
<td id="D0A9C84957E5E96745D2AD2308A1EFE2" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.225</td>
|
|
</tr>
|
|
<tr id="1553A4D25093192719D7536FE8D7D1CE" pageId="0" pageNumber="1">
|
|
<td id="AD5E7C4054F761D076647B10335938A5" colspan="1" pageId="0" pageNumber="1" rowspan="1">S244_Belisana_Mal30_Mal241</td>
|
|
<td id="CE69E23E23DA48F3E2C12B88D50D4592" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.255</td>
|
|
<td id="C1F862FE0938F608BEB8098F3C50F177" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.269</td>
|
|
<td id="6E9FA7BE87033E5B64677562DF9CE1E1" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.259</td>
|
|
</tr>
|
|
<tr id="031B694C9BABBD0F289C07353395FEC3" pageId="0" pageNumber="1">
|
|
<td id="41D3740897866238C7E3149CAA40C001" colspan="1" pageId="0" pageNumber="1" rowspan="1">S333_Belisana_bohorok_Ind127</td>
|
|
<td id="64BF2141AD7B9E0A9BDACDCC047A4811" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.250</td>
|
|
<td id="F2AD9206CEAE0FA4296CADDD95EB0727" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.250</td>
|
|
<td id="2857A7A4EC0568F635FF6D991B08D0BB" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.248</td>
|
|
</tr>
|
|
<tr id="C8D52E826BB80BBF96C3647167F00362" pageId="0" pageNumber="1">
|
|
<td id="424CF0177C57BDB95AAB479A8622A140" colspan="1" pageId="0" pageNumber="1" rowspan="1">S336_Belisana_nahtanoj_Ind181</td>
|
|
<td id="14F56911F1ECB452C4B28377CE65FD9C" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.209</td>
|
|
<td id="53CCB1078525229EC984FFD12B5EF240" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.212</td>
|
|
<td id="E791B9419FB0665D68A5FB83F8AF65CC" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.208</td>
|
|
</tr>
|
|
<tr id="F5EA68605C32EB3346B2D08FE1904A18" pageId="0" pageNumber="1">
|
|
<td id="6FB941110385DD866353DDD1B94C2044" colspan="1" pageId="0" pageNumber="1" rowspan="1">S338_Belisana_tambligan_Ind213</td>
|
|
<td id="531931FF1C27D087A2F687C0870B7E27" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.237</td>
|
|
<td id="9E81E5AA13FD8D6B297A3DF1E5CE0725" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.234</td>
|
|
<td id="4BF668F1A7E45BD2304DB7466895C8B2" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.234</td>
|
|
</tr>
|
|
<tr id="EC6047586FA535DB17B22304EA07F0F7" pageId="0" pageNumber="1">
|
|
<td id="A4634C6E60478479F030E1478E12646D" colspan="1" pageId="0" pageNumber="1" rowspan="1">S339_Buitinga_buhoma_Uga124</td>
|
|
<td id="5D4EF1CEA9E132ED9E55716EC36CCD62" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.254</td>
|
|
<td id="F5FE21A0115B22840862A991C1DC5EAD" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.269</td>
|
|
<td id="20CC5DA7C9BF4A057276F618D9AEA917" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.251</td>
|
|
</tr>
|
|
<tr id="64E78B25DC0D16664B5554F667D49137" pageId="0" pageNumber="1">
|
|
<td id="6AAC9C335FBCF4B230EEFC30697EA9C8" colspan="1" pageId="0" pageNumber="1" rowspan="1">S340_Buitinga_ruhiza_Uga122</td>
|
|
<td id="53D90F5C21454737E1211FAD792CF4B5" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.252</td>
|
|
<td id="61B9C681D77B6A75208116717694535B" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.262</td>
|
|
<td id="949B0672B37AD84C5E0E33C139D8BCD3" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.247</td>
|
|
</tr>
|
|
<tr id="413388ADEA76EA72001304E779D9A72F" pageId="0" pageNumber="1">
|
|
<td id="D1C16937D3BC3734D3B848737CBCEEFD" colspan="1" pageId="0" pageNumber="1" rowspan="1">S341_Buitinga_ruwenzori_Uga156</td>
|
|
<td id="0A14627C589E05F80DEEE04340D3EE8E" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.255</td>
|
|
<td id="2EC439B2BF2CE317A9BE33B28B9064BE" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.269</td>
|
|
<td id="613A762C4BEDECF7B527F3AAB33B4144" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.249</td>
|
|
</tr>
|
|
<tr id="5B6C7E7FD1EF0E814B10F68807726AC9" pageId="0" pageNumber="1">
|
|
<td id="7754FE960BF5FD5A8E07EE0773D01951" colspan="1" pageId="0" pageNumber="1" rowspan="1">S334_Belisana_Ind15_Ind140</td>
|
|
<td id="979009E9848D7C84F4C20D98572D6DA2" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.274</td>
|
|
<td id="17B1C58DE658163C9E9A53EC915684CA" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.256</td>
|
|
<td id="642885CFC8CD3A4933860E93783764EB" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.274</td>
|
|
</tr>
|
|
<tr id="3AD83383713F700B08425AE8BCCCC6DA" pageId="0" pageNumber="1">
|
|
<td id="DEB37D4C8AFB1600DCA7DBBC4760EE59" colspan="1" pageId="0" pageNumber="1" rowspan="1">S386_Belisana_sabah_Bor214</td>
|
|
<td id="B9FBD1718CB8FA8C8F67976EB9DA78E0" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.235</td>
|
|
<td id="03F45D1D5CADF4378FA7F66A96384946" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.250</td>
|
|
<td id="003C5603C81681BDAA392637FE6B7CBC" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.233</td>
|
|
</tr>
|
|
<tr id="56FFA3D68DF1648F0D66B990A228A3EF" pageId="0" pageNumber="1">
|
|
<td id="D6C5970B103B1AA803863A2FB681513C" colspan="1" pageId="0" pageNumber="1" rowspan="1">S417_Belisana_ranong_Mal361</td>
|
|
<td id="A99BBACF39FA16DDF1E68549C23A6B50" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.210</td>
|
|
<td id="A8B7F831CBDA99099EA537DFA13C99A6" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.215</td>
|
|
<td id="02FC30DD0C3411A99D9C2465609D8EA6" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.210</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="DBE72FF2CAAD0EE8096FC7617C13E972" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="E1CB4FCDC2DCCB145B59F50194D9E796" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph id="1B757BD6AC9C970D495885D4F1462522" pageId="0" pageNumber="1">
|
|
Distinguished from known congeners (and other
|
|
<taxonomicName id="568645A1B3FA85200FC69D19F3B637F6" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
|
|
) by details of procursus (Fig.
|
|
<figureCitation id="B3335A93D9E18E5EFF73542B69DA6E3B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A, B left male palp, prolateral and retrolateral views C male chelicerae, frontal view D, E left procursus, prolateral and retrolateral views F cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; ep, epigynal pocket; f, femur; lp, lateral pocket; pp, pore plate; pr, procursus; ta, tarsus; ti, tibia. Scale bars: 0.2 mm (A, B, F); 0.1 mm (C-E)." figureDoi="10.3897/subtbiol.46.105798.figure2" httpUri="https://binary.pensoft.net/fig/883298" pageId="0" pageNumber="1">2D, E</figureCitation>
|
|
; straight, with simple retrolateral process, bifid prolateral process, and widened membranous tip), genital bulb (Fig.
|
|
<figureCitation id="4AEC35BBC50588130837DA860891A85C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A, B left male palp, prolateral and retrolateral views C male chelicerae, frontal view D, E left procursus, prolateral and retrolateral views F cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; ep, epigynal pocket; f, femur; lp, lateral pocket; pp, pore plate; pr, procursus; ta, tarsus; ti, tibia. Scale bars: 0.2 mm (A, B, F); 0.1 mm (C-E)." figureDoi="10.3897/subtbiol.46.105798.figure2" httpUri="https://binary.pensoft.net/fig/883298" pageId="0" pageNumber="1">2A, B</figureCitation>
|
|
; bulbal apophysis short and hooked, embolus straight and pointed at tip), male chelicerae (Fig.
|
|
<figureCitation id="CD65ECDC2C1547FE787C0ACD3618218E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A, B left male palp, prolateral and retrolateral views C male chelicerae, frontal view D, E left procursus, prolateral and retrolateral views F cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; ep, epigynal pocket; f, femur; lp, lateral pocket; pp, pore plate; pr, procursus; ta, tarsus; ti, tibia. Scale bars: 0.2 mm (A, B, F); 0.1 mm (C-E)." figureDoi="10.3897/subtbiol.46.105798.figure2" httpUri="https://binary.pensoft.net/fig/883298" pageId="0" pageNumber="1">2C</figureCitation>
|
|
; pair of light proximal processes; pair of simple distal frontal apophyses without modified hairs), epigynum (Fig.
|
|
<figureCitation id="C86061ED200A5CEFBCBE5E7C11BCF1A0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A male habitus, lateral view B female abdomen, ventral view C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ep, epigynal pockets; lp, lateral pocket; pp, pore plate; sp, spinnerets. Scale bars: 0.5 mm (A); 0.1 mm (B-D)." figureDoi="10.3897/subtbiol.46.105798.figure1" httpUri="https://binary.pensoft.net/fig/883297" pageId="0" pageNumber="1">1B</figureCitation>
|
|
; protruding, with pair of pockets close together), and female internal genitalia (Figs
|
|
<figureCitation id="0F9A2B4149AC12476CBEF9595873767D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A male habitus, lateral view B female abdomen, ventral view C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ep, epigynal pockets; lp, lateral pocket; pp, pore plate; sp, spinnerets. Scale bars: 0.5 mm (A); 0.1 mm (B-D)." figureDoi="10.3897/subtbiol.46.105798.figure1" httpUri="https://binary.pensoft.net/fig/883297" pageId="0" pageNumber="1">1C, D</figureCitation>
|
|
,
|
|
<figureCitation id="EF3DA1FA8D414D30F933BC55C6B08FE3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A, B left male palp, prolateral and retrolateral views C male chelicerae, frontal view D, E left procursus, prolateral and retrolateral views F cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; ep, epigynal pocket; f, femur; lp, lateral pocket; pp, pore plate; pr, procursus; ta, tarsus; ti, tibia. Scale bars: 0.2 mm (A, B, F); 0.1 mm (C-E)." figureDoi="10.3897/subtbiol.46.105798.figure2" httpUri="https://binary.pensoft.net/fig/883298" pageId="0" pageNumber="1">2F</figureCitation>
|
|
; pore plates wide apart; distinctive pair of additional internal lateral pockets).
|
|
</paragraph>
|
|
<caption id="21CE728BCD4AFF895EC5453FD81BB362" doi="10.3897/subtbiol.46.105798.figure1" httpUri="https://binary.pensoft.net/fig/883297" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
|
|
<paragraph id="D6E6C75BBA1F806B49A75E910ADBDD59" pageId="0" pageNumber="1">
|
|
<emphasis id="D9FA18B0499674C99A92774A18935B33" bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
|
|
<taxonomicName id="96776986459F068B42BFF81E29584E7E" authorityName="Huber & Clark" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana coblynau" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coblynau">
|
|
<emphasis id="758929544AEDBEE74546F600554B9667" italics="true" pageId="0" pageNumber="1">Belisana coblynau</emphasis>
|
|
</taxonomicName>
|
|
Huber & Clark, sp. nov., male holotype and female, WAM T158896-7
|
|
<emphasis id="A44CF77DFDFAC8CB70684017B9A9B880" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
male habitus, lateral view
|
|
<emphasis id="F93FE9FD383A926506014B8095F29361" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
female abdomen, ventral view
|
|
<emphasis id="36A838BBF21586293DDE76357D45988D" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
|
cleared female genitalia, ventral and dorsal views. Abbreviations: ep, epigynal pockets; lp, lateral pocket; pp, pore plate; sp, spinnerets. Scale bars: 0.5 mm (
|
|
<emphasis id="322B9C33DDDD2CEFB6AAB576EB89A173" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis id="74B962720A9DFE1E2E8E8B71F33A316D" bold="true" pageId="0" pageNumber="1">B-D</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="4E0FC65B61C57220BF572AF0BBD2F1E5" doi="10.3897/subtbiol.46.105798.figure2" httpUri="https://binary.pensoft.net/fig/883298" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
|
|
<paragraph id="A0BA2120BAD782C0D2AE47CF47DD18A9" pageId="0" pageNumber="1">
|
|
<emphasis id="98D9CD304093B61AA13D6BCA977CE949" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
|
|
<taxonomicName id="225319A9CFCBBD07E442FEDC7BEE553D" authorityName="Huber & Clark" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana coblynau" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coblynau">
|
|
<emphasis id="A11262CB203627783DA06C9A272000D8" italics="true" pageId="0" pageNumber="1">Belisana coblynau</emphasis>
|
|
</taxonomicName>
|
|
Huber & Clark, sp. nov., male holotype and female, WAM T158896-7
|
|
<emphasis id="68039D0BDD3B4967C476284558BA7994" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
|
left male palp, prolateral and retrolateral views
|
|
<emphasis id="1BFA3239CA03262EF6AC0DE4345EAF49" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
male chelicerae, frontal view
|
|
<emphasis id="33561D1DA234DC6529907DE198C79E8C" bold="true" pageId="0" pageNumber="1">D, E</emphasis>
|
|
left procursus, prolateral and retrolateral views
|
|
<emphasis id="767471C71D6BED92DD8027077D508FA2" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; ep, epigynal pocket; f, femur; lp, lateral pocket; pp, pore plate; pr, procursus; ta, tarsus; ti, tibia. Scale bars: 0.2 mm (
|
|
<emphasis id="04FA7BF2B021A410B6D1E0C12F315966" bold="true" pageId="0" pageNumber="1">A, B, F</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis id="9DC56FEAA131348D6F8D474565990A40" bold="true" pageId="0" pageNumber="1">C-E</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="E96BB4962D5A37259A10C250B01E9391" pageId="0" pageNumber="1" type="description">
|
|
<paragraph id="E32D7E7DA938114AF40019578C7B40CC" pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph id="5D84DBD375482BD0BE55646A47FAE9C0" pageId="0" pageNumber="1">
|
|
<emphasis id="688481E5A2EF1AC8AC6E1D2C50077DDE" bold="true" pageId="0" pageNumber="1">Male</emphasis>
|
|
(holotype).
|
|
<emphasis id="11CEA4765CAE2F0A816B4FA6BCBCBC6D" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="8ED7DFD97EE8438A604B6ECFAF68ABFB" italics="true" pageId="0" pageNumber="1">Measurements</emphasis>
|
|
.
|
|
</emphasis>
|
|
Total body length ~1.5-1.6 (abdomen detached), carapace width 0.60. Leg 1: 11.5 (2.9 + 0.2 + 3.0 + 4.4 + 1.0); legs 2 and 3 missing; tibia 4: 2.3; tibia 1 L/d: 50.
|
|
</paragraph>
|
|
<paragraph id="63A5C6DF51E2FAE328811D179669A986" pageId="0" pageNumber="1">
|
|
<emphasis id="815CF5F000F10BB1F5FA4D4A86D5B628" bold="true" italics="true" pageId="0" pageNumber="1">Color</emphasis>
|
|
(in ethanol). Entire specimen pale ochre to whitish (Fig.
|
|
<figureCitation id="99AC90F77B98AB7CA6676A95FA42832B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A male habitus, lateral view B female abdomen, ventral view C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ep, epigynal pockets; lp, lateral pocket; pp, pore plate; sp, spinnerets. Scale bars: 0.5 mm (A); 0.1 mm (B-D)." figureDoi="10.3897/subtbiol.46.105798.figure1" httpUri="https://binary.pensoft.net/fig/883297" pageId="0" pageNumber="1">1A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="AD34C2804BDF3BFF59ADA0935035FFD2" pageId="0" pageNumber="1">
|
|
<emphasis id="9A631E03BC281726983E4D8423BE7C6F" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="F7A6EDF5FA8813C6C3D9785E7FE3F2F5" italics="true" pageId="0" pageNumber="1">Body</emphasis>
|
|
.
|
|
</emphasis>
|
|
Habitus as in Fig.
|
|
<figureCitation id="FD048E4F2C1AFD2CEFB14EBB5A89D185" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A male habitus, lateral view B female abdomen, ventral view C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ep, epigynal pockets; lp, lateral pocket; pp, pore plate; sp, spinnerets. Scale bars: 0.5 mm (A); 0.1 mm (B-D)." figureDoi="10.3897/subtbiol.46.105798.figure1" httpUri="https://binary.pensoft.net/fig/883297" pageId="0" pageNumber="1">1A</figureCitation>
|
|
. Ocular area not raised, without any remnants of eye pigment and lenses. Without thoracic groove. Clypeus unmodified. Sternum wider than long (0.40/0.30), unmodified. Abdomen globular, conical at spinnerets.
|
|
</paragraph>
|
|
<paragraph id="C0312DA11ED90DFC45006208E70477B1" pageId="0" pageNumber="1">
|
|
<emphasis id="D03B26D76CF21FF59FD8F9A6A7D9E954" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="A774CF43BAB853746A793CB3115D85EF" italics="true" pageId="0" pageNumber="1">Chelicerae</emphasis>
|
|
.
|
|
</emphasis>
|
|
As in Fig.
|
|
<figureCitation id="E36EB4C0F3A0ABE38A9CDA733E5BB1E1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A, B left male palp, prolateral and retrolateral views C male chelicerae, frontal view D, E left procursus, prolateral and retrolateral views F cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; ep, epigynal pocket; f, femur; lp, lateral pocket; pp, pore plate; pr, procursus; ta, tarsus; ti, tibia. Scale bars: 0.2 mm (A, B, F); 0.1 mm (C-E)." figureDoi="10.3897/subtbiol.46.105798.figure2" httpUri="https://binary.pensoft.net/fig/883298" pageId="0" pageNumber="1">2C</figureCitation>
|
|
, with pair of light proximal processes, pair of simple distal frontal apophyses without modified hairs (distance between tips: 50
|
|
<normalizedToken id="54711289500746C2703C986F592BD0CC" originalValue="µm">µm</normalizedToken>
|
|
), without stridulatory files.
|
|
</paragraph>
|
|
<paragraph id="94243008CDF9E074696E04778EF704EB" pageId="0" pageNumber="1">
|
|
<emphasis id="3CA7DF7A68406ABFBC3D0213095FBFC7" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="50F816FA8641AF39B90660BBFB0F00DD" italics="true" pageId="0" pageNumber="1">Palps</emphasis>
|
|
.
|
|
</emphasis>
|
|
As in Fig.
|
|
<figureCitation id="15CB91ACCF65F591CF69C940A8BE6E49" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A, B left male palp, prolateral and retrolateral views C male chelicerae, frontal view D, E left procursus, prolateral and retrolateral views F cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; ep, epigynal pocket; f, femur; lp, lateral pocket; pp, pore plate; pr, procursus; ta, tarsus; ti, tibia. Scale bars: 0.2 mm (A, B, F); 0.1 mm (C-E)." figureDoi="10.3897/subtbiol.46.105798.figure2" httpUri="https://binary.pensoft.net/fig/883298" pageId="0" pageNumber="1">2A, B</figureCitation>
|
|
; coxa unmodified, trochanter with low rounded retrolateral ventral protrusion, femur cylindrical, without process; tibia with only one (retrolateral distal) trichobothrium; procursus (Fig.
|
|
<figureCitation id="C872BAA572E4C962D3ADCE25D22FD1BF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A, B left male palp, prolateral and retrolateral views C male chelicerae, frontal view D, E left procursus, prolateral and retrolateral views F cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; ep, epigynal pocket; f, femur; lp, lateral pocket; pp, pore plate; pr, procursus; ta, tarsus; ti, tibia. Scale bars: 0.2 mm (A, B, F); 0.1 mm (C-E)." figureDoi="10.3897/subtbiol.46.105798.figure2" httpUri="https://binary.pensoft.net/fig/883298" pageId="0" pageNumber="1">2D, E</figureCitation>
|
|
) straight, with simple retrolateral process, bifid prolateral process, and widened membranous tip; genital bulb with short hooked bulbal apophysis and straight embolus pointed at tip.
|
|
</paragraph>
|
|
<paragraph id="DE2BB1E1C1C145D9C6E99B7D6E75710C" pageId="0" pageNumber="1">
|
|
<emphasis id="FB79CB0EA1AEC1320C664CFA4C82C437" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="FD66A3FF3756472AE9E0D32A32ED8879" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
|
|
.
|
|
</emphasis>
|
|
Apparently without spines and curved hairs (most hairs on available legs missing); leg trichobothria not seen; tarsus 1 with ~10 pseudosegments, barely visible in dissecting microscope.
|
|
</paragraph>
|
|
<paragraph id="0FBE18E2C9A753C9A535BD11B15B15A9" pageId="0" pageNumber="1">
|
|
<emphasis id="54A04C79AB0C4A2E6FA0EF8E14FC49BB" bold="true" pageId="0" pageNumber="1">Female.</emphasis>
|
|
In general similar to male, but larger (in particular the abdomen); total body length 1.8; carapace width 0.62; most legs detached (tibia 4: 2.6). Epigynum (Fig.
|
|
<figureCitation id="1305B2F6E8C53530939E5044D8EF546E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A male habitus, lateral view B female abdomen, ventral view C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ep, epigynal pockets; lp, lateral pocket; pp, pore plate; sp, spinnerets. Scale bars: 0.5 mm (A); 0.1 mm (B-D)." figureDoi="10.3897/subtbiol.46.105798.figure1" httpUri="https://binary.pensoft.net/fig/883297" pageId="0" pageNumber="1">1B</figureCitation>
|
|
) main anterior plate semicircular, with distinct pair of pockets (distance between pockets: 30
|
|
<normalizedToken id="FD21497FC586F6537415510367C1947B" originalValue="µm">µm</normalizedToken>
|
|
), in lateral view protruding. Internal genitalia (Figs
|
|
<figureCitation id="BB0E8E4B9BC07A08080FADEE7A9C2B65" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A male habitus, lateral view B female abdomen, ventral view C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ep, epigynal pockets; lp, lateral pocket; pp, pore plate; sp, spinnerets. Scale bars: 0.5 mm (A); 0.1 mm (B-D)." figureDoi="10.3897/subtbiol.46.105798.figure1" httpUri="https://binary.pensoft.net/fig/883297" pageId="0" pageNumber="1">1C, D</figureCitation>
|
|
,
|
|
<figureCitation id="503992FAF99B4754A035BC63D1427524" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana coblynau Huber & Clark, sp. nov., male holotype and female, WAM T 158896 - 7 A, B left male palp, prolateral and retrolateral views C male chelicerae, frontal view D, E left procursus, prolateral and retrolateral views F cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; ep, epigynal pocket; f, femur; lp, lateral pocket; pp, pore plate; pr, procursus; ta, tarsus; ti, tibia. Scale bars: 0.2 mm (A, B, F); 0.1 mm (C-E)." figureDoi="10.3897/subtbiol.46.105798.figure2" httpUri="https://binary.pensoft.net/fig/883298" pageId="0" pageNumber="1">2F</figureCitation>
|
|
) with pair of pore plates wide apart, distinctive pair of additional internal lateral pockets. Each ALS with 3-4 small cylindrical spigots in addition to large widened spigot and long pointed spigot (checked in cleared abdomen).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="F2D47CB9AFCAB5218DA5CEA2E75147C0" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="534E3281A5F1BCDECE4615B2DB4C2E34" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="D335B9D2FFF46A506DFC8F337D4885B5" pageId="0" pageNumber="1">The species epithet refers to the mythical gnome-like creatures that are said to haunt mines and quarries (of Wales and beyond); noun in apposition.</paragraph>
|
|
</subSubSection>
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<subSubSection id="F905A1441B57E8F5388F7AE1354C6A6A" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="A51EBAB64EE25B6F95F322753E211962" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="277BF5A966ABBCE0D009FB926754E3BD" pageId="0" pageNumber="1">Known from three neighboring (within 5 km) localities in Western Australia.</paragraph>
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</subSubSection>
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<subSubSection id="EF01EA9F31F66650EE65AFB5607C8D4D" pageId="0" pageNumber="1" type="natural history">
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<paragraph id="0C8C308EFB92348CA9D0625D07093736" pageId="0" pageNumber="1">Natural history.</paragraph>
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<paragraph id="2CFFE5B270A0B512B679A1D64F2BC542" pageId="0" pageNumber="1">
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The spiders were extracted from mining bores in the Hamersley Range using scrape sampling (Fig.
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<figureCitation id="FF0CED5F4CE632F8C918CB59279302FD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epigean and hypogean habitats and collecting method A Hamersley Range in Western Australia; the area of the sampled boreholes B weighted net (arrow) above a borehole C weighted net used for scrape sampling D main entrance of the Grotte de La Tortue (photo T. Percheron, 2015) E gallery of the lost goat in Caverne de La Tortue (photo T. Percheron, 2022)." figureDoi="10.3897/subtbiol.46.105798.figure5" httpUri="https://binary.pensoft.net/fig/883301" pageId="0" pageNumber="1">5A-C</figureCitation>
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). This collecting method precludes gaining detailed natural history data. The specimens were collected between ground level and approximately 35 m below ground level. The temperature in these holes at 15 m below ground level was constant at 32-33 °C. For a general characterization of the Pilbara troglofauna, see Discussion.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |