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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1014.60426" ID-GBIF-Dataset="97cd82f3-4a3a-43ca-a68f-92a5a2074d07" ID-PMC="PMC7875960" ID-Pensoft-Pub="1313-2970-1014-1" ID-Pensoft-UUID="62961664CAED5F15B9C8A9990D7D388D" ID-PubMed="33603534" ModsDocID="1313-2970-1014-1" checkinTime="1612410909014" checkinUser="pensoft" docAuthor="Achterberg, Cornelis van" docDate="2021" docId="49F43ADF66C55366AEFD8F4815B1E4D5" docLanguage="en" docName="ZooKeys 1014: 1-118" docOrigin="ZooKeys 1014" docPubDate="2021-02-03" docSource="http://dx.doi.org/10.3897/zookeys.1014.60426" docTitle="Phanerotoma mesocellata Achterberg 2021, sp. nov." docType="treatment" docVersion="4" id="62961664CAED5F15B9C8A9990D7D388D" lastPageNumber="1" masterDocId="62961664CAED5F15B9C8A9990D7D388D" masterDocTitle="The tribe Phanerotomini (Hymenoptera, Braconidae, Cheloninae) of the Arabian Peninsula, with special reference to the United Arab Emirates and Yemen" masterLastPageNumber="118" masterPageNumber="1" pageNumber="1" updateTime="1668149759257" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The tribe Phanerotomini (Hymenoptera, Braconidae, Cheloninae) of the Arabian Peninsula, with special reference to the United Arab Emirates and Yemen</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Achterberg, Cornelis van</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6495-4853</mods:nameIdentifier>
<mods:affiliation>Naturalis Biodiversity Center, Postbus 9517, 2300 RA Leiden, the Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">kees@vanachterberg.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1014</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>118</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1014.60426</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1014.60426</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1014-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">62961664CAED5F15B9C8A9990D7D388D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="177209572" LSID="urn:lsid:zoobank.org:act:7BC291B3-A91B-4EEC-AEB2-15B1767E1C31" httpUri="http://treatment.plazi.org/id/49F43ADF66C55366AEFD8F4815B1E4D5" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/7BC291B3-A91B-4EEC-AEB2-15B1767E1C31" authority="Achterberg, 2021" authorityName="Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Phanerotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanerotoma mesocellata" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mesocellata" status="sp. nov.">Phanerotoma mesocellata</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 235–238" captionStartId="F35" captionText="Figures 235 - 238. Phanerotoma mesocellata van Achterberg, sp. nov., ♀ holotype (but 236, 237 of ♂, paratype) 235 habitus lateral 236 antenna 237 hind femur and tibia lateral 238 middle femur and tibia lateral." figureDoi="10.3897/zookeys..60426.figures235-238" httpUri="https://binary.pensoft.net/fig/504658" pageId="0" pageNumber="1">Figs 235-238</figureCitation>
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">, 239-249</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
, ♀ (RMNH), &quot;
<emphasis bold="true" pageId="0" pageNumber="1">United Arab Emirates</emphasis>
, Sharjah Desert Park (15613), light trap, 1-30.iv.2007,
<geoCoordinate degrees="25" direction="north" minutes="17" orientation="latitude" precision="925" value="25.283333">25°17'N</geoCoordinate>
,
<geoCoordinate degrees="55" direction="east" minutes="42" orientation="longitude" precision="925" value="55.7">55°42'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’10”">RMNH'10&quot;</normalizedToken>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: 1♀: Idem, 22.iii.-5.iv.2009; 1♀: Idem, 29.iii.-6.iv.2005; 5♀, 1♂: Idem, 23-30.iv.2005; 13♀, 2♂: Idem, 30.iv.-7.v.2005; 11♀: Idem, 6-13.iv.2005; 1♀: Idem, 21-29.iii.2005; 1♀: &quot;United Arab Emirates, Sharjah (2279), light trap, 30.vi.-21.vii.2005,
<geoCoordinate degrees="25" direction="north" minutes="21" orientation="latitude" precision="925" value="25.35">25°21'N</geoCoordinate>
,
<geoCoordinate degrees="55" direction="east" minutes="24" orientation="longitude" precision="925" value="55.4">55°24'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’05”">RMNH'05&quot;</normalizedToken>
; 1♀: Idem, 2.v.-5.vi.2005; 3♀: &quot;United Arab Emirates, Fujairah (2438), light trap, 2.v.-5.vi.2005,
<geoCoordinate degrees="25" direction="north" minutes="08" orientation="latitude" precision="925" value="25.133333">25°08'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="east" minutes="21" orientation="longitude" precision="925" value="56.35">56°21'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 1♀: Idem, 5-24.iii.2005; 6♀: &quot;United Arab Emirates, NARC near Sweihan (1245), light trap, 28.iii.-2.iv.2005,
<geoCoordinate degrees="24" direction="north" minutes="24" orientation="latitude" precision="925" value="24.4">24°24'N</geoCoordinate>
,
<geoCoordinate degrees="55" direction="east" minutes="26" orientation="longitude" precision="925" value="55.433334">55°26'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 4♀: Idem, 2-9.iv.2005; 22♀, 3♂: Idem, 9-20.iv.2005; 1♀: Idem, 14-28.iii.2005; 8♀, 3♂: &quot;United Arab Emirates, Wadi Bih dam (11366), light trap, 24.iv.-23.v.2007,
<geoCoordinate degrees="25" direction="north" minutes="48" orientation="latitude" precision="925" value="25.8">25°48'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="east" minutes="04" orientation="longitude" precision="925" value="56.066666">56°04'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’10”">RMNH'10&quot;</normalizedToken>
; 4♀, 1♂: Idem, 13-30.iv.2008; 1♀: Idem, 19.ii.-29.iii.2007; 1♀: &quot;United Arab Emirates, Hatta (11572), light trap, 21.vi.-19.vii.2006,
<geoCoordinate degrees="24" direction="north" minutes="49" orientation="latitude" precision="925" value="24.816668">24°49'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="east" minutes="07" orientation="longitude" precision="925" value="56.116665">56°07'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’09”">RMNH'09&quot;</normalizedToken>
; 1♂: &quot;United Arab Emirates, Sharjah x Khor Kalba (11542), light trap,
<geoCoordinate degrees="24" direction="north" minutes="59" orientation="latitude" precision="925" value="24.983334">24°59'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="east" minutes="09" orientation="longitude" precision="925" value="56.15">56°09'E</geoCoordinate>
, 28.iii.-5.iv.2006, A. v. Harten,
<normalizedToken originalValue="RMNH’10”">RMNH'10&quot;</normalizedToken>
; 4♀: &quot;
<emphasis bold="true" pageId="0" pageNumber="1">Yemen</emphasis>
(5697), Al Kowd, light trap, iv.2001, A. van Harten &amp; S. Al Haruri,
<normalizedToken originalValue="RMNH’02”">RMNH'02&quot;</normalizedToken>
; 5♀: Idem, 16-20.viii.2001; 6♀: Idem, 8-12.vii.2001; 22♀, 1♂: Idem, 27-31.vii.2001; 5♀: Idem, 6-10.viii.2001; 7♀: Idem, v.-vi.2000; 14♀, 3♂: Idem, vii.1999; 2♀: Idem, ii.2000; 1♀: Idem, vii.2000; 7♀: Idem, viii.2000; 2♀: Idem, xii.2000; 1♀: Idem, viii.1999; 3♀: Idem, 21-25.viii.2001; 2♀, 1♂: Idem, vii.-ix.2001; 2♀: Idem, vi.2002; 4♀: Idem, ix.2003; 3♀: Idem, i.-iii.2003; 15♀, &quot;Yemen (6090), Al Kadan, light trap, x.2001, A. van Harten &amp; T. Abdul-Haq,
<normalizedToken originalValue="RMNH’03”">RMNH'03&quot;</normalizedToken>
; 2♀: Idem, iv.2002; 14♀, 1♂: Idem, v.2002; 2♀, 1♂: Idem, i.2003; 7♀: Idem, xi.2001; 9♀: &quot;Yemen (5404),
<normalizedToken originalValue="Hamman’Ali">Hamman'Ali</normalizedToken>
, from coffee berries (with
<taxonomicName baseAuthorityName="Wiedemann" baseAuthorityYear="1824" class="Hexapoda" family="Tephritidae" genus="Ceratitis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ceratitis capitata" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="capitata">
<emphasis italics="true" pageId="0" pageNumber="1">Ceratitis capitata</emphasis>
</taxonomicName>
?), 14.iii.2001, A. van Harten,
<normalizedToken originalValue="RMNH’02”">RMNH'02&quot;</normalizedToken>
; 4♀: &quot;Yemen (6381),
<normalizedToken originalValue="Ta’izz">Ta'izz</normalizedToken>
, light trap, ix.-x.2001, A. van Harten &amp; A.R. Al Yarimi, RMNH&quot;; 1♀: Idem, ix.1999; 3♀: Idem, x.1999; 6♀: Idem, xi.1999; 4 ♀: Idem, xii.1999; 8♀: Idem, i.2000; 12♀: Idem, v.2000; 17♀: Idem, ix.2000; 2♀, 1♂: Idem, 5.i.-2.ii.1998; 15♀: Idem, 26-28.vii.1999; 16♀, 1♂: Idem, 3-24.i.1999; 17♀, 1♂: Idem, viii.2000; 1♀: Idem, 27-31.vii.2001; 7♀: Idem, x.2001; 3♀: Idem, iii.-iv.2001; 5♀: Idem, vi.2002; 6♀, 1♂: Idem, vii.2002; 6♀: &quot;Yemen (3645),
<normalizedToken originalValue="Sana’a">Sana'a</normalizedToken>
, light trap, iii.-iv.1999, A. van Harten,
<normalizedToken originalValue="RMNH’00”">RMNH'00&quot;</normalizedToken>
; 1♀: Idem, v.1999; 1♀: &quot;Yemen (6667), 12 km NW Manakhah, Mal[aise] trap, 27.iii.-5.v.2002, A. v. Harten,
<normalizedToken originalValue="RMNH’03”">RMNH'03&quot;</normalizedToken>
; 1♀: &quot;Yemen, Seyun, light trap, 4-6.ix.2002, A. van Harten,
<normalizedToken originalValue="RMNH’03”">RMNH'03&quot;</normalizedToken>
; 3♀: &quot;Yemen (6158), Al Lahima, 17.ix.-14.xi.2001, Mal[aise] trap, A. v. Harten,
<normalizedToken originalValue="RMNH’02”">RMNH'02&quot;</normalizedToken>
.
</paragraph>
<caption doi="10.3897/zookeys..60426.figures235-238" httpUri="https://binary.pensoft.net/fig/504658" pageId="0" pageNumber="1" start="Figures 235–238" startId="F35">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 235-238.</emphasis>
<taxonomicName authorityName="Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Phanerotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanerotoma mesocellata" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mesocellata">
<emphasis italics="true" pageId="0" pageNumber="1">Phanerotoma mesocellata</emphasis>
</taxonomicName>
van Achterberg, sp. nov., ♀ holotype (but
<emphasis bold="true" pageId="0" pageNumber="1">236, 237</emphasis>
of ♂, paratype)
<emphasis bold="true" pageId="0" pageNumber="1">235</emphasis>
habitus lateral
<emphasis bold="true" pageId="0" pageNumber="1">236</emphasis>
antenna
<emphasis bold="true" pageId="0" pageNumber="1">237</emphasis>
hind femur and tibia lateral
<emphasis bold="true" pageId="0" pageNumber="1">238</emphasis>
middle femur and tibia lateral.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Eighth-tenth antennal segments from apex of ♀ moderately moniliform, stocky, matt or slightly shiny, 14th segment from apex somewhat longer than wide (Figs
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">248</figureCitation>
,
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">249</figureCitation>
); stemmaticum black or dark brown, but sometimes brownish yellow; mesosternum more or less shiny; second submarginal cell of fore wing comparatively short (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">239</figureCitation>
); POL of ♀ 0.4-0.6
<normalizedToken originalValue="×">x</normalizedToken>
width of posterior ocellus; eye 1.2-1.8
<normalizedToken originalValue="×">x</normalizedToken>
as wide as median width of temple in lateral view (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">247</figureCitation>
); vein 1-M (as usually parastigma) slightly darker than yellow M+CU1 of fore wing (Fig.
<figureCitation captionStart="Figures 235–238" captionStartId="F35" captionText="Figures 235 - 238. Phanerotoma mesocellata van Achterberg, sp. nov., ♀ holotype (but 236, 237 of ♂, paratype) 235 habitus lateral 236 antenna 237 hind femur and tibia lateral 238 middle femur and tibia lateral." figureDoi="10.3897/zookeys..60426.figures235-238" httpUri="https://binary.pensoft.net/fig/504658" pageId="0" pageNumber="1">235</figureCitation>
); ovipositor sheath narrow apically (Figs
<figureCitation captionStart="Figures 235–238" captionStartId="F35" captionText="Figures 235 - 238. Phanerotoma mesocellata van Achterberg, sp. nov., ♀ holotype (but 236, 237 of ♂, paratype) 235 habitus lateral 236 antenna 237 hind femur and tibia lateral 238 middle femur and tibia lateral." figureDoi="10.3897/zookeys..60426.figures235-238" httpUri="https://binary.pensoft.net/fig/504658" pageId="0" pageNumber="1">235</figureCitation>
,
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">242</figureCitation>
). Closely related to
<taxonomicName family="Braconidae" kingdom="Animalia" lsidName="P. ocularis" order="Hymenoptera" pageId="0" pageNumber="1" rank="species" species="ocularis">
<emphasis italics="true" pageId="0" pageNumber="1">P. ocularis</emphasis>
</taxonomicName>
and differs mainly by the shape of the apical antennal segments of the female, the more curved vein 2-SR and the size of the ocelli.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Female, holotype, length of body (excluding ovipositor) 3.6 mm; antenna 2.7 mm; fore wing 2.6 mm; visible part of ovipositor sheath 0.4 mm (erect setae mostly concentrated at apex).</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Width 1.7
<normalizedToken originalValue="×">x</normalizedToken>
median length in anterior view and part of head above eye in lateral view 0.25
<normalizedToken originalValue="×">x</normalizedToken>
height of eye (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">247</figureCitation>
); antenna with 23 cylindrical segments, slightly widened submedially and slightly longer than fore wing, seven apical antennal segments small and moniliform (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">249</figureCitation>
), with short bristles and apical segment with spine, third, fourth and penultimate segments 2.6, 2.4 and 1.4
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide in lateral view, respectively; area of stemmaticum coriaceous; OOL: diameter of posterior ocellus: POL = 15: 5: 3; length of eye 2.2
<normalizedToken originalValue="×">x</normalizedToken>
temple in dorsal view (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">245</figureCitation>
); frons with weak median carina, mainly coriaceous, rather shiny and laterally rugulose; vertex rugulose-coriaceous and rather matt, posteriorly also with some transverse rugulae and distinctly emarginate (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">245</figureCitation>
); temple mainly coriaceous and rather matt, nearly parallel-sided in lateral view (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">247</figureCitation>
), gradually narrowed behind eyes; face transversely rugose laterally, rugulose and with obsolescent median bump and with satin sheen; clypeus smooth, moderately shiny and 0.9
<normalizedToken originalValue="×">x</normalizedToken>
as wide as minimum width of face, intertentorial distance 3.6
<normalizedToken originalValue="×">x</normalizedToken>
minimum width between clypeus and eye, long erect setose and with three distinct blunt teeth medio-ventrally (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">246</figureCitation>
); eye large, strongly convex and in lateral view 1.8
<normalizedToken originalValue="×">x</normalizedToken>
wider than temple (measured medially; Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">247</figureCitation>
), in anterior view its height 0.8
<normalizedToken originalValue="×">x</normalizedToken>
minimum width of face (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">246</figureCitation>
); upper condyle of mandible above lower level of eyes (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">246</figureCitation>
); malar space mostly smooth, rather shiny and 0.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as basal width of mandible; lower tooth of mandible 0.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as apical tooth, small (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">244</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1" start="Figures 239–249" startId="F36">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 239-249.</emphasis>
<taxonomicName authorityName="Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Phanerotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanerotoma mesocellata" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mesocellata">
<emphasis italics="true" pageId="0" pageNumber="1">Phanerotoma mesocellata</emphasis>
</taxonomicName>
van Achterberg, sp. nov., ♀, holotype
<emphasis bold="true" pageId="0" pageNumber="1">239</emphasis>
fore wing
<emphasis bold="true" pageId="0" pageNumber="1">240</emphasis>
mesosoma dorsal
<emphasis bold="true" pageId="0" pageNumber="1">241</emphasis>
first-third metasomal tergites dorsal
<emphasis bold="true" pageId="0" pageNumber="1">242</emphasis>
metasoma lateral
<emphasis bold="true" pageId="0" pageNumber="1">243</emphasis>
hind leg lateral
<emphasis bold="true" pageId="0" pageNumber="1">244</emphasis>
mandible ventral
<emphasis bold="true" pageId="0" pageNumber="1">245</emphasis>
head dorsal
<emphasis bold="true" pageId="0" pageNumber="1">246</emphasis>
head anterior
<emphasis bold="true" pageId="0" pageNumber="1">247</emphasis>
head lateral
<emphasis bold="true" pageId="0" pageNumber="1">248</emphasis>
antenna lateral
<emphasis bold="true" pageId="0" pageNumber="1">249</emphasis>
apical half of antenna lateral.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
(Figs
<figureCitation captionStart="Figures 235–238" captionStartId="F35" captionText="Figures 235 - 238. Phanerotoma mesocellata van Achterberg, sp. nov., ♀ holotype (but 236, 237 of ♂, paratype) 235 habitus lateral 236 antenna 237 hind femur and tibia lateral 238 middle femur and tibia lateral." figureDoi="10.3897/zookeys..60426.figures235-238" httpUri="https://binary.pensoft.net/fig/504658" pageId="0" pageNumber="1">235</figureCitation>
,
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">240</figureCitation>
). Length 1.5
<normalizedToken originalValue="×">x</normalizedToken>
its width in lateral view; side of pronotum coriaceous dorsally and remainder rugose; posteriorly propleuron bulging near central groove; mesosternum finely granulate and with satin sheen; mesoscutum densely reticulate-rugose on granulate background, with satin sheen, notauli absent; scutellum nearly flat, finely granulate-rugulose; scutellar sulcus medium-sized, with eight carinae (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">240</figureCitation>
); metanotum with median carina and minute medio-posterior tooth, its posterior border finely serrate; propodeum coarsely reticulate-rugose, on median-sized dorsal face less coarsely rugose, with transverse carina, no median carina, and latero-posteriorly weakly tuberculate.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Wings</emphasis>
.
</emphasis>
Fore wing 2.7
<normalizedToken originalValue="×">x</normalizedToken>
longer than its maximum width; 1-R1 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as pterostigma; distance between wing apex and marginal cell apex 0.25
<normalizedToken originalValue="×">x</normalizedToken>
length of 1-R1; r issued far beyond middle of pterostigma and 0.2
<normalizedToken originalValue="×">x</normalizedToken>
3-SR; 2-SR weakly curved and slightly converging to posterior margin of pterostigma (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">239</figureCitation>
); SR1 curved; m-cu interstitial; parastigma large; 1-CU1 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as vein 2-CU1, cu-a 0.9
<normalizedToken originalValue="×">x</normalizedToken>
1-CU1, strongly inclivous; r:3-SR:SR1 = 4:19:49; 2-SR:3-SR:r-m = 28:19:9; r-m reclivous; 2-M slightly curved (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">239</figureCitation>
). Hind wing: M+CU:1-M:1r-m = 22:17:10.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Legs</emphasis>
.
</emphasis>
Hind femur rather matt, 3.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide and robust; hind tibia swollen (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">243</figureCitation>
); middle tibia with medium-sized blister; inner spur of middle tibia 0.5
<normalizedToken originalValue="×">x</normalizedToken>
its basitarsus; hind coxa largely superficially granulate, but dorsally partly smooth and rather shiny.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Metasoma</emphasis>
(Figs
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">241</figureCitation>
,
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">242</figureCitation>
). Oval in dorsal view, 1.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide and 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as mesosoma; first and second tergites irregularly and coarsely longitudinally rugose; second metasomal suture rather wide and straight; third tergite 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than second tergite and laterally curved, convex medially, rounded posteriorly in dorsal view (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">241</figureCitation>
), obtuse posteriorly in lateral view (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">242</figureCitation>
), densely reticulate-rugose and with satin sheen (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">241</figureCitation>
), lateral lamella narrow basally, but widened near apical half of third tergite and latero-apically, medio-apically somewhat barrower and concave; ovipositor sheath narrow, apically slightly widened and darkened (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">242</figureCitation>
), its visible part 0.16
<normalizedToken originalValue="×">x</normalizedToken>
as long as fore wing and 0.28
<normalizedToken originalValue="×">x</normalizedToken>
metasomal carapace, and with erect setae mainly near its apex; hypopygium of ♀ with short widely triangular and up curved apical protuberance (Fig.
<figureCitation captionStart="Figures 239–249" captionStartId="F36" captionText="Figures 239 - 249. Phanerotoma mesocellata van Achterberg, sp. nov., ♀, holotype 239 fore wing 240 mesosoma dorsal 241 first-third metasomal tergites dorsal 242 metasoma lateral 243 hind leg lateral 244 mandible ventral 245 head dorsal 246 head anterior 247 head lateral 248 antenna lateral 249 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures239-249" httpUri="https://binary.pensoft.net/fig/504659" pageId="0" pageNumber="1">242</figureCitation>
) and with mostly medium-sized setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Pale brownish yellow; stemmaticum blackish; apex of antenna, hind tibia apically and subbasally and apex of ovipositor sheath rather brown; parastigma partly brownish, remainder and vein 1-M yellow; clypeus, palpi, tegulae, remainder of legs, mesoscutum medio-posteriorly, first and second tergites and metasoma ventrally pale yellowish or ivory; pterostigma (but basally and apically pale yellowish) and most veins brown; wing membrane slightly brownish below pterostigma and near vein 1-CU1.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="male">
<paragraph pageId="0" pageNumber="1">Male.</paragraph>
<paragraph pageId="0" pageNumber="1">
Similar to female (including hind femur; Fig.
<figureCitation captionStart="Figures 235–238" captionStartId="F35" captionText="Figures 235 - 238. Phanerotoma mesocellata van Achterberg, sp. nov., ♀ holotype (but 236, 237 of ♂, paratype) 235 habitus lateral 236 antenna 237 hind femur and tibia lateral 238 middle femur and tibia lateral." figureDoi="10.3897/zookeys..60426.figures235-238" httpUri="https://binary.pensoft.net/fig/504658" pageId="0" pageNumber="1">237</figureCitation>
), but antenna slenderer (Fig.
<figureCitation captionStart="Figures 235–238" captionStartId="F35" captionText="Figures 235 - 238. Phanerotoma mesocellata van Achterberg, sp. nov., ♀ holotype (but 236, 237 of ♂, paratype) 235 habitus lateral 236 antenna 237 hind femur and tibia lateral 238 middle femur and tibia lateral." figureDoi="10.3897/zookeys..60426.figures235-238" httpUri="https://binary.pensoft.net/fig/504658" pageId="0" pageNumber="1">236</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variations.</paragraph>
<paragraph pageId="0" pageNumber="1">Length of fore wing of ♀ 2.1-2.7 mm (mainly UAE; many specimens from Yemen are larger (2.9-3.6 mm) because of using larger hosts), of ♂ 2.1-2.2 mm (UAE; Yemen 2.3-3.2 mm); vein 1-M of fore wing and parastigma pale yellowish, but sometimes more or less brown; second tergite brownish yellow or brown laterally; stemmaticum usually dark brown, but sometimes brownish yellow.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">United Arab Emirates, Yemen.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Named after its intermediate sized ocelli (
<emphasis italics="true" pageId="0" pageNumber="1">meso</emphasis>
is Greek for middle).
</paragraph>
</subSubSection>
</treatment>
</document>
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