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<document id="2C3BB3484997F9169E6E2392366C7B74" ID="10.11646/zootaxa.4273.3.11SLASH11252" ID-CLB-Dataset="32898" ID-DOI="10.11646/zootaxa.4273.3.11" ID-GBIF-Dataset="19e53c4e-0f3c-44bc-bae0-fe57eae79b12" ID-ISSN="1175-5326" ID-Zenodo-Dep="803219" ID-ZooBank="CA2F4687-DBED-43A4-837F-2E57D43D2959" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496738786438" checkinUser="plazi" docAuthor="Gaiani, Marco A." docDate="2017" docId="494787A12D7EDE50FF798A8EFDE6FC71" docLanguage="en" docName="zootaxa.4273.3.11SLASH11252.pdf" docOrigin="Zootaxa 4273 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bythonia freytagi Gaiani, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="449" masterDocId="B57EFFD92D7FDE52FFEE8A19FFE9FFF6" masterDocTitle="Review of the genus Bythonia Oman 1936 with description of a new species and new record from Venezuela (Hemiptera, Cicadellidae, Iassinae)" masterLastPageNumber="450" masterPageNumber="447" pageNumber="448" updateTime="1698440223992" updateUser="ExternalLinkService">
<mods:mods id="DB918B7C0830BAF3C299CA87FD32717B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="BF87F1BF66FC25F19D1564152EC1DC3D">Review of the genus Bythonia Oman 1936 with description of a new species and new record from Venezuela (Hemiptera, Cicadellidae, Iassinae)</mods:title>
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<mods:name id="37B7BF48F6447C73BF801505D256423C" type="personal">
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<mods:namePart id="90997275D90ABAAE63A3F9098FC66A71">Gaiani, Marco A.</mods:namePart>
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<mods:title id="AF7240803F2D86BC121369519078AEC6">Zootaxa</mods:title>
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<mods:part id="606AD3356FDE560BE2FBAAC1BBDCD1D5">
<mods:date id="AB1E0EA98F68D19401F3DD7562124E35">2017</mods:date>
<mods:detail id="209E48E0A3FE397B2A6FE4F665F84EC6" type="volume">
<mods:number id="80BF5712C7EA1D2BF725AA49D77661DC">4273</mods:number>
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<mods:classification id="2B861F7416743C69C711185E69580FAE">journal article</mods:classification>
<mods:identifier id="9814671EE37CBE2180344071340131B0" type="CLB-Dataset">32898</mods:identifier>
<mods:identifier id="DB3C148329BD33DB2D0770B125597628" type="DOI">10.11646/zootaxa.4273.3.11</mods:identifier>
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<treatment id="494787A12D7EDE50FF798A8EFDE6FC71" ID-DOI="http://doi.org/10.5281/zenodo.6053244" ID-GBIF-Taxon="131812387" ID-Zenodo-Dep="6053244" LSID="urn:lsid:plazi:treatment:494787A12D7EDE50FF798A8EFDE6FC71" httpUri="http://treatment.plazi.org/id/494787A12D7EDE50FF798A8EFDE6FC71" lastPageId="2" lastPageNumber="449" pageId="1" pageNumber="448">
<subSubSection id="89F4653C2D7EDE53FF798A8EFE4FFF27" pageId="1" pageNumber="448" type="nomenclature">
<paragraph id="C15136B72D7EDE53FF798A8EFE27FF44" blockId="1.[151,1436,151,521]" box="[151,462,151,178]" pageId="1" pageNumber="448">
<heading id="9A1981DB2D7EDE53FF798A8EFE27FF44" bold="true" box="[151,462,151,178]" fontSize="11" level="1" pageId="1" pageNumber="448" reason="1">
<emphasis id="F39AEAA52D7EDE53FF798A8EFE27FF44" bold="true" box="[151,462,151,178]" pageId="1" pageNumber="448">
<taxonomicName id="06EE4D342D7EDE53FF798A8EFE80FF47" ID-CoL="8JQHM" box="[151,361,151,177]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="freytagi" status="sp. nov.">
<emphasis id="F39AEAA52D7EDE53FF798A8EFE80FF47" bold="true" box="[151,361,151,177]" italics="true" pageId="1" pageNumber="448">Bythonia freytagi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8A957DE2D7EDE53FE9F8A81FE27FF44" box="[369,462,152,178]" pageId="1" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C15136B72D7EDE53FF798AA3FE4FFF27" blockId="1.[151,1436,151,521]" box="[151,422,186,209]" pageId="1" pageNumber="448">
Length of male
<quantity id="06169B522D7EDE53FEAE8AA3FE4AFF27" box="[320,419,186,209]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" pageId="1" pageNumber="448" unit="mm" value="6.15">6.15 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="89F4653C2D7EDE50FF298AC4FE70FEAA" lastPageId="2" lastPageNumber="449" pageId="1" pageNumber="448" type="description">
<paragraph id="C15136B72D7EDE53FF298AC4FC38FE89" blockId="1.[151,1436,151,521]" pageId="1" pageNumber="448">
Body wedge shaped, uniformly reddish-brown (
<figureCitation id="59D52A322D7EDE53FD3A8AC4FCDBFF02" box="[724,818,221,244]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="1" pageNumber="448">Figure 1</figureCitation>
). Head, including eyes, distinctly wider than pronotum. Crown, ocello-ocular area, upper margin and vertical median band of frons very dark brown, almost black. Lateral areas of frons, at each side of vertical dark band, lighter, brown stramineous. Forewings evenly translucid brown, with lighter area on base of fourth apical cell extending to approximately half its length. Legs brown. Femoral setal formula 2:1. Body covered by many short sparse yellow hairs, more evident on forewings.
</paragraph>
<paragraph id="C15136B72D7EDE53FF298B93FA80FDFF" blockId="1.[151,1436,151,521]" pageId="1" pageNumber="448">
Head. Vertex mostly smooth, with few transverse rugae between ocelli on posterior margin. Frons, clypeus, lora and lower genae with strong transverse irregular rugae (
<figureCitation id="59D52A322D7EDE53FD2D8BB4FCCAFE32" box="[707,803,429,452]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="1" pageNumber="448">Figure 2</figureCitation>
). Pronotum with numerous strong transverse striae, except on apical 1/5, which is smooth to faintly microreticulated. Thorax. Scutellum with transverse striations not as strong as on pronotum; with two semicircular anterolateral and two elongated centromedial microreticulated areas (
<figureCitation id="59D52A322D7EDE53FB108BEBFAB0FDFF" box="[1278,1369,498,521]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="1" pageNumber="448">Figure 1</figureCitation>
).
</paragraph>
<caption id="9591663F2D7EDE53FF798CF4FDB0F8CB" httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="1" pageNumber="448" targetBox="[354,1180,603,1733]" targetPageId="1">
<paragraph id="C15136B72D7EDE53FF798CF4FDB0F8CB" blockId="1.[151,1436,1773,1853]" pageId="1" pageNumber="448">
<emphasis id="F39AEAA52D7EDE53FF798CF4FEDDF8F5" bold="true" box="[151,308,1773,1795]" pageId="1" pageNumber="448">FIGURES 17.</emphasis>
<taxonomicName id="06EE4D342D7EDE53FED28CF4FE0FF8F4" box="[316,486,1773,1794]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="freytagi" status="sp. nov.">
<emphasis id="F39AEAA52D7EDE53FED28CF4FE0FF8F4" box="[316,486,1773,1794]" italics="true" pageId="1" pageNumber="448">Bythonia freytagi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8A957DE2D7EDE53FE008CF4FDCEF8F5" box="[494,551,1773,1795]" pageId="1" pageNumber="448" rank="species">
<emphasis id="F39AEAA52D7EDE53FE008CF4FDCEF8F5" bold="true" box="[494,551,1773,1795]" pageId="1" pageNumber="448">sp. n.</emphasis>
</taxonomicNameLabel>
Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view.
</paragraph>
</caption>
<paragraph id="C15136B72D7EDE50FF298D71FACCFECF" blockId="1.[151,1436,1896,2023]" lastBlockId="2.[151,1436,152,903]" lastPageId="2" lastPageNumber="449" pageId="1" pageNumber="448">
Male genitalia. Aedeagus (
<figureCitation id="59D52A322D7EDE53FE098D71FDADF889" box="[487,580,1896,1919]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="1" pageNumber="448">Figure 4</figureCitation>
) long and slender, produced caudally and then dorsally from base; apex curved ventrally, rounded, weakly sclerotized; with two parallel spiniform processes (
<figureCitation id="59D52A322D7EDE53FC328D92FBD0F854" box="[988,1081,1931,1954]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="1" pageNumber="448">Figure 4</figureCitation>
a) strongly adpressed to shaft 2/3 from base. Pygofer (
<figureCitation id="59D52A322D7EDE53FE988DB4FE3AF832" box="[374,467,1965,1988]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="1" pageNumber="448">Figure 3</figureCitation>
) with micro- and macrosetae; macrosetae restricted to posteroventral margin, microsetae well distributed over entire surface; apex truncated; without ventral processes. Genital plates elongated, parallel-sided, separated from each other throughout their length, almost as long as pygofer (
<figureCitation id="59D52A322D7DDE50FC228A81FBC1FF59" box="[972,1064,152,175]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="2" pageNumber="449">Figure 3</figureCitation>
), truncated at apex, ventral surface with short setae; lateral outer margin with fine setae on basal half, very long setae on apical half (
<figureCitation id="59D52A322D7DDE50FB0C8AA3FAAAFF27" box="[1250,1347,186,209]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="2" pageNumber="449">Figure 7</figureCitation>
). Styles (
<figureCitation id="59D52A322D7DDE50FF708AC4FF15FF02" box="[158,252,221,244]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="2" pageNumber="449">Figure 5</figureCitation>
) elongated, longer than genital plates; apical 1/5th laterally expanded into thin flange, ending in sharp point displaced medially from central axis of style, area at base of spine with row of piliform setae (
<figureCitation id="59D52A322D7DDE50FB6A8B19FB09FEE1" box="[1156,1248,256,279]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="2" pageNumber="449">Figure 5</figureCitation>
a). Connective Yshaped, well sclerotized. Seventh sternite roundly concave at middle, with small central convexity (
<figureCitation id="59D52A322D7DDE50FB548B3BFAF1FECF" box="[1210,1304,290,313]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="2" pageNumber="449">Figure 6</figureCitation>
).
</paragraph>
<paragraph id="C15136B72D7DDE50FF298B5CFE70FEAA" blockId="2.[151,1436,152,903]" box="[199,409,325,348]" pageId="2" pageNumber="449">
<emphasis id="F39AEAA52D7DDE50FF298B5CFEF3FEAA" bold="true" box="[199,282,325,348]" pageId="2" pageNumber="449">Female</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="89F4653C2D7DDE50FF298B71FDBCFE32" pageId="2" pageNumber="449" type="materials_examined">
<paragraph id="C15136B72D7DDE50FF298B71FE8CFE89" blockId="2.[151,1436,152,903]" box="[199,357,360,383]" pageId="2" pageNumber="449">
<emphasis id="F39AEAA52D7DDE50FF298B71FE8CFE89" bold="true" box="[199,357,360,383]" pageId="2" pageNumber="449">Type material</emphasis>
</paragraph>
<paragraph id="C15136B72D7DDE50FF298B93FDBCFE32" blockId="2.[151,1436,152,903]" pageId="2" pageNumber="449">
<materialsCitation id="71863CEA2D7DDE50FF298B93FDBCFE32" ID-GBIF-Occurrence="1563404911" collectingDate="1975-05-05" collectingDateMax="1975-05-09" collectingDateMin="1975-05-05" collectionCode="MIZA" collectorName="Km Norte Altagracia de Orituco &amp; J. Salcedo &amp; R. E. Dietz" country="Venezuela" elevation="640" location="Km Norte Altagracia de Orituco" municipality="Guatopo" pageId="2" pageNumber="449" specimenCount="1" stateProvince="Miranda" typeStatus="holotype">
<typeStatus id="1E5588152D7DDE50FF298B93FEC3FE57" box="[199,298,394,417]" pageId="2" pageNumber="449" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="B9F976272D7DDE50FE9B8B93FE0CFE57" box="[373,485,394,417]" name="Venezuela" pageId="2" pageNumber="449">Venezuela</collectingCountry>
,
<collectingRegion id="032AF8552D7DDE50FE1E8B93FDA4FE57" box="[496,589,394,417]" country="Venezuela" name="Miranda" pageId="2" pageNumber="449">Miranda</collectingRegion>
, Parque Nac.[ional]
<collectingMunicipality id="2135ACCD2D7DDE50FCC28B93FC63FE57" box="[812,906,394,417]" pageId="2" pageNumber="449">Guatopo</collectingMunicipality>
, 24
<collectorName id="6C1B53612D7DDE50FC568B93FAF1FE57" box="[952,1304,394,417]" pageId="2" pageNumber="449">
<location id="C431606C2D7DDE50FC568B93FAF1FE57" LSID="urn:lsid:plazi:treatment:494787A12D7EDE50FF798A8EFDE6FC71:C431606C2D7DDE50FC568B93FAF1FE57" box="[952,1304,394,417]" country="Venezuela" municipality="Guatopo" name="Km Norte Altagracia de Orituco" pageId="2" pageNumber="449" stateProvince="Miranda">Km Norte Altagracia de Orituco</location>
</collectorName>
,
<elevation id="4AC3D1842D7DDE50FACA8B93FA8AFE57" box="[1316,1379,394,417]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="449" unit="m" value="640.0">
<quantity id="06169B522D7DDE50FACA8B93FA8AFE57" box="[1316,1379,394,417]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="449" unit="m" value="640.0">640m</quantity>
</elevation>
,
<collectingDate id="A514E99F2D7DDE50FA818B93FF00FE32" pageId="2" pageNumber="449" value="1975-05-05" valueMax="1975-05-09" valueMin="1975-05-05">
<date id="B55010772D7DDE50FA818B93FF00FE32" pageId="2" pageNumber="449" value="1975-05-05" valueMax="1975-05-09" valueMin="1975-05-05">5-9- V-1975</date>
</collectingDate>
.
<collectorName id="6C1B53612D7DDE50FF1D8BB4FEB6FE32" box="[243,351,429,452]" pageId="2" pageNumber="449">J. Salcedo</collectorName>
&amp;
<collectorName id="6C1B53612D7DDE50FE6F8BB4FE18FE32" box="[385,497,429,452]" pageId="2" pageNumber="449">R.E. Dietz</collectorName>
(
<collectionCode id="A7FFAE722D7DDE50FE118BB4FDA0FE32" box="[511,585,429,452]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="2" pageNumber="449">MIZA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="89F4653C2D7DDE50FF298BC9FD8CFDDA" pageId="2" pageNumber="449" type="diagnosis">
<paragraph id="C15136B72D7DDE50FF298BC9FD8CFDDA" blockId="2.[151,1436,152,903]" pageId="2" pageNumber="449">
<emphasis id="F39AEAA52D7DDE50FF298BC9FED5FE11" bold="true" box="[199,316,464,487]" pageId="2" pageNumber="449">Diagnosis.</emphasis>
This species can be distinguished from all other species of
<taxonomicName id="06EE4D342D7DDE50FC098BD6FBA1FE11" box="[999,1096,463,487]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="F39AEAA52D7DDE50FC098BD6FBA1FE11" box="[999,1096,463,487]" italics="true" pageId="2" pageNumber="449">Bythonia</emphasis>
</taxonomicName>
by the presence of the paired adpressed subapical aedeagal processes (
<figureCitation id="59D52A322D7DDE50FDB18BEBFD4DFDFF" box="[607,676,498,521]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="2" pageNumber="449">Fig. 4</figureCitation>
a), the shape of sternite VII (
<figureCitation id="59D52A322D7DDE50FC1D8BEBFBD0FDFF" box="[1011,1081,498,521]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="2" pageNumber="449">Fig. 6</figureCitation>
), and the head, including eyes, distinctly wider than the pronotum (
<figureCitation id="59D52A322D7DDE50FDF8880CFDBCFDDA" box="[534,597,533,556]" captionStart="FIGURES 1 7" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[354,1180,603,1733]" captionTargetId="figure@1.[340,1247,575,1749]" captionTargetPageId="1" captionText="FIGURES 1 7. Bythonia freytagi sp. n. Holotype. 1) Dorsal habitus, 2) Head ventral view, 3) Pygofer lateral view, 4) Aedeagus lateral view, 4 a) Aedeagus, detail of the sub-apical processes, 5) Right style, ventral view, 5 a) Right style apex, dorsal view 6) VII sternite, 7) Left sub-genital plate, ventral view." httpUri="https://zenodo.org/record/803221/files/figure.png" pageId="2" pageNumber="449">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89F4653C2D7DDE50FF298821FD7BFD87" pageId="2" pageNumber="449" type="etymology">
<paragraph id="C15136B72D7DDE50FF298821FD7BFD87" blockId="2.[151,1436,152,903]" pageId="2" pageNumber="449">
<emphasis id="F39AEAA52D7DDE50FF298821FEAEFDB9" bold="true" box="[199,327,568,591]" pageId="2" pageNumber="449">Etymology.</emphasis>
I take great pleasure in naming this species after my mentor in leafhopper systematics, retired professor Dr. Paul Freytag of the University of Kentucky.
</paragraph>
</subSubSection>
<subSubSection id="89F4653C2D7DDE50FF298864FDE6FC71" pageId="2" pageNumber="449" type="discussion">
<paragraph id="C15136B72D7DDE50FF298864FDE6FC71" blockId="2.[151,1436,152,903]" pageId="2" pageNumber="449">
<emphasis id="F39AEAA52D7DDE50FF298864FEAFFD62" bold="true" box="[199,326,637,660]" pageId="2" pageNumber="449">Discussion.</emphasis>
Based on the few specimens at hand and the published descriptions it seems that there are two species groups in
<taxonomicName id="06EE4D342D7DDE50FEE78886FE80FD41" box="[265,361,671,695]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="F39AEAA52D7DDE50FEE78886FE80FD41" box="[265,361,671,695]" italics="true" pageId="2" pageNumber="449">Bythonia</emphasis>
</taxonomicName>
that eventually might be considered subgenera. The “
<taxonomicName id="06EE4D342D7DDE50FC2A88B9FBF1FD41" box="[964,1048,672,695]" class="Insecta" family="Cicadellidae" genus="Nionia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="rugosa">Rugosa</taxonomicName>
” species group is characterized by aedeagi with lateral or ventral spine-like processes on the apical 1/4th of the shaft and with the shaft apex curved caudally; species included are
<taxonomicName id="06EE4D342D7DDE50FE0788FCFDBDFD0A" box="[489,596,741,764]" class="Insecta" family="Cicadellidae" genus="Nionia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="F39AEAA52D7DDE50FE0788FCFDBDFD0A" box="[489,596,741,764]" italics="true" pageId="2" pageNumber="449">B. rugosa</emphasis>
</taxonomicName>
,
<taxonomicName id="06EE4D342D7DDE50FD8A88FCFD0EFD0A" box="[612,743,741,764]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="consensa">
<emphasis id="F39AEAA52D7DDE50FD8A88FCFD0EFD0A" box="[612,743,741,764]" italics="true" pageId="2" pageNumber="449">B. consensa</emphasis>
</taxonomicName>
,
<taxonomicName id="06EE4D342D7DDE50FD1988FCFC60FD0A" box="[759,905,740,764]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="F39AEAA52D7DDE50FD1988FCFC60FD0A" box="[759,905,740,764]" italics="true" pageId="2" pageNumber="449">B. ferruginea</emphasis>
</taxonomicName>
and
<taxonomicName id="06EE4D342D7DDE50FC2D88FCFBDFFD0A" box="[963,1078,740,764]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="freytagi" status="sp. nov.">
<emphasis id="F39AEAA52D7DDE50FC2D88FCFBDFFD0A" box="[963,1078,740,764]" italics="true" pageId="2" pageNumber="449">B. freytagi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8A957DE2D7DDE50FBD188FCFB71FD0A" box="[1087,1176,741,764]" pageId="2" pageNumber="449" rank="species">
<emphasis id="F39AEAA52D7DDE50FBD188FCFB71FD0A" bold="true" box="[1087,1176,741,764]" pageId="2" pageNumber="449">sp. nov.</emphasis>
</taxonomicNameLabel>
The “
<taxonomicName id="06EE4D342D7DDE50FB3088FCFAD2FD0A" box="[1246,1339,741,764]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="kalypso">Kalypso</taxonomicName>
” species group contains only
<taxonomicName id="06EE4D342D7DDE50FE998911FE01FCE9" box="[375,488,775,799]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="kalypso">
<emphasis id="F39AEAA52D7DDE50FE998911FE01FCE9" box="[375,488,775,799]" italics="true" pageId="2" pageNumber="449">B. kalypso</emphasis>
</taxonomicName>
and is characterized by the lack of ventral/lateral spine-like processes on the aedeagus and by the shaft apex curved mesally. Unfortunately, the
<typeStatus id="1E5588152D7DDE50FCE68933FCDFFCB7" box="[776,822,810,833]" pageId="2" pageNumber="449">type</typeStatus>
of
<taxonomicName id="06EE4D342D7DDE50FCB08933FC39FCB7" box="[862,976,809,833]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="kalypso">
<emphasis id="F39AEAA52D7DDE50FCB08933FC39FCB7" box="[862,976,809,833]" italics="true" pageId="2" pageNumber="449">B. kalypso</emphasis>
</taxonomicName>
is apparently lost and to the extent of my knowledge no other specimens have been collected. Therefore, it is very difficult to assess the relationship of
<taxonomicName id="06EE4D342D7DDE50FAC28954FA75FC92" box="[1324,1436,844,868]" class="Insecta" family="Cicadellidae" genus="Bythonia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="kalypso">
<emphasis id="F39AEAA52D7DDE50FAC28954FA75FC92" box="[1324,1436,844,868]" italics="true" pageId="2" pageNumber="449">B. kalypso</emphasis>
</taxonomicName>
with the other species of the genus.
</paragraph>
</subSubSection>
</treatment>
</document>