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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1024.56927" ID-GBIF-Dataset="7a0ded71-ddb1-49e5-9c2c-10c01fd03a1f" ID-PMC="PMC7990858" ID-Pensoft-Pub="1313-2970-1024-157" ID-Pensoft-UUID="086BAB631B8D5B888571137F0CEFF686" ID-PubMed="33786006" ID-ZooBank="FDD37EA991214385B67D51AD313CB49E" ModsDocID="1313-2970-1024-157" checkinTime="1615979483707" checkinUser="pensoft" docAuthor="Nestor-Arriola, Jorge Ismael, Toledo-Hernandez, Victor Hugo, Solis, Angel, Gonzalez, Guillermo &amp; Vetrovec, Jaroslav" docDate="2021" docId="477D70860D1E55ED819E0194624AC020" docLanguage="en" docName="ZooKeys 1024: 157-196" docOrigin="ZooKeys 1024" docPubDate="2021-03-16" docSource="http://dx.doi.org/10.3897/zookeys.1024.56927" docTitle="Brachiacantha invertita Nestor-Arriola, Toledo-Hernandez &amp; Solis 2021, sp. nov." docType="treatment" docUuid="2665461E-6DF3-4ACF-B4E7-D00FA2BEBFBC" docUuidSource="ZooBank" docVersion="5" id="086BAB631B8D5B888571137F0CEFF686" lastPageNumber="157" masterDocId="086BAB631B8D5B888571137F0CEFF686" masterDocTitle="The Brachiacantha Dejean, 1837 (Coleoptera, Coccinellidae) of Central America" masterLastPageNumber="196" masterPageNumber="157" pageNumber="157" updateTime="1668149905347" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Brachiacantha Dejean, 1837 (Coleoptera, Coccinellidae) of Central America</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nestor-Arriola, Jorge Ismael</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2394-1586</mods:nameIdentifier>
<mods:affiliation>Centro de Investigacion en Biodiversidad y Conservacion (CIByC), Universidad Autonoma del Estado de Morelos. Av. Universidad # 1001, Col. Chamilpa, Cuernavaca, Morelos, C. P. 62209, Mexico</mods:affiliation>
<mods:nameIdentifier type="email">jorge.nestorarr@uaem.edu.mx</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Toledo-Hernandez, Victor Hugo</mods:namePart>
<mods:affiliation>Centro de Investigacion en Biodiversidad y Conservacion (CIByC), Universidad Autonoma del Estado de Morelos. Av. Universidad # 1001, Col. Chamilpa, Cuernavaca, Morelos, C. P. 62209, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Solis, Angel</mods:namePart>
<mods:affiliation>Departamento de Historia Natural, Museo Nacional de Costa Rica. Santo Domingo de Heredia, Costa Rica</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gonzalez, Guillermo</mods:namePart>
<mods:affiliation>La Reina, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vetrovec, Jaroslav</mods:namePart>
<mods:affiliation>Buzulucka, Hradec Kralove, Ceska Republika</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1024</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>157</mods:start>
<mods:end>196</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1024.56927</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1024.56927</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1024-157</mods:identifier>
<mods:identifier type="ZooBank">FDD37EA991214385B67D51AD313CB49E</mods:identifier>
<mods:identifier type="Pensoft-UUID">086BAB631B8D5B888571137F0CEFF686</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="180585857" LSID="urn:lsid:zoobank.org:act:2665461E-6DF3-4ACF-B4E7-D00FA2BEBFBC" httpUri="http://treatment.plazi.org/id/477D70860D1E55ED819E0194624AC020" lastPageNumber="157" pageId="0" pageNumber="157">
<subSubSection pageId="0" pageNumber="157" type="nomenclature">
<paragraph pageId="0" pageNumber="157">
<taxonomicName LSID="http://zoobank.org/2665461E-6DF3-4ACF-B4E7-D00FA2BEBFBC" authority="Nestor-Arriola, Toledo-Hernández &amp; Solís" authorityName="Nestor-Arriola, Toledo-Hernandez &amp; Solis" authorityYear="2021" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha invertita" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="invertita" status="sp. nov.">
Brachiacantha invertita Nestor-Arriola,
<normalizedToken originalValue="Toledo-Hernández">Toledo-Hernandez</normalizedToken>
&amp;
<normalizedToken originalValue="Solís">Solis</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="157">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 5964" captionStartId="F14" captionText="Figures 59 - 64. Brachiacantha invertita sp. nov., male 59 Dorsal view 60 Abdomen 61 Tegmen 62 Tegmen lateral view 63 Penis 64 Penis apex." figureDoi="10.3897/zookeys.1024.56927.figures59-64" httpUri="https://binary.pensoft.net/fig/520879" pageId="0" pageNumber="157">Figures 59-64</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="holotype">
<paragraph pageId="0" pageNumber="157">
<typeStatus>Holotype</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<materialsCitation ID-GBIF-Occurrence="3063616380" collectingDate="1994-05" collectionCode="MNCR" collectorName="A. C. Arenal &amp; Z. Fuentes" country="Costa Rica" elevation="1175" location="Original" municipality="Male" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<collectingMunicipality>Male</collectingMunicipality>
, pinned, with genitalia in a separate microvial.
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:AC78765E57EE926A54ADF6563394FDD3" country="Costa Rica" municipality="Male" name="Original" stateProvince="Puntarenas">Original</location>
labels: &quot;
<collectingCountry name="Costa Rica">COSTA RICA</collectingCountry>
,
<collectingRegion country="Costa Rica" name="Puntarenas">PUNTARENAS</collectingRegion>
,
<collectorName>A. C. Arenal</collectorName>
, Buen Amigo, San Luis,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.35" metricValueMin="1.0" unit="m" value="1175.0" valueMax="1350.0" valueMin="1000.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.35" metricValueMin="1.0" unit="m" value="1175.0" valueMax="1350.0" valueMin="1000.0">1000-1350 m</elevation>
</quantity>
,
<date value="1994-05">
<collectingDate value="1994-05">May 1994</collectingDate>
</date>
, col.
<collectorName>Z. Fuentes</collectorName>
, L N 250850_449250, #2926&quot; (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="paratypes">
<paragraph pageId="0" pageNumber="157">
<typeStatus>Paratypes</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<materialsCitation ID-GBIF-Occurrence="3063616353" collectingDate="1980-08-07" collectionCode="USNMNH" collectorName="D. R. Whitehead" country="Costa Rica" elevation="1400" location="Birrisito" specimenCount="1" specimenCount-male="1" stateProvince="Cartago" typeStatus="paratype">
<collectingCountry name="Costa Rica">Costa Rica</collectingCountry>
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Costa Rica" name="Cartago">Cartago</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:166D409734D24CEA076E0ADBB33E9511" country="Costa Rica" name="Birrisito" stateProvince="Cartago">Birrisito</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</elevation>
</quantity>
;
<date value="1980-08-07">
<collectingDate value="1980-08-07">07 Aug. 1980</collectingDate>
</date>
;
<collectorName>D. R. Whitehead</collectorName>
leg. (
<collectionCode>USNMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616344" collectingDate="2001-06-22" collectingDateMax="2001-07-22" collectingDateMin="2001-06-22" collectionCode="MNCR" collectorName="Tayutic, P. N &amp; W. Arana" country="Costa Rica" county="Turrialba" elevation="1550" location="Sector Cerro Tigre" municipality="Barbilla" specimenCount="1" specimenCount-female="1" stateProvince="Cartago" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Costa Rica" name="Cartago">Cartago</collectingRegion>
,
<collectingCounty>Turrialba</collectingCounty>
,
<collectorName>Tayutic, P. N</collectorName>
.
<collectingMunicipality>Barbilla</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:FCBB3C88C1CAA4D4324C64A66DF30275" country="Costa Rica" county="Turrialba" municipality="Barbilla" name="Sector Cerro Tigre" stateProvince="Cartago">Sector Cerro Tigre</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">1500-1600 m</elevation>
</quantity>
;
<date value="2001-06-22" valueMax="2001-07-22" valueMin="2001-06-22">
<collectingDate value="2001-06-22" valueMax="2001-07-22" valueMin="2001-06-22">22 June-22 July 2001</collectingDate>
</date>
;
<collectorName>W. Arana</collectorName>
leg.;
<collectingMethod>malaise trap</collectingMethod>
; L_N_211700_602500, #64679 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063616306" collectingDate="1992-10" collectionCode="MNCR" collectorName="P. N. Tapanti &amp; G. Mora" country="Costa Rica" elevation="1250" location="Quebrada Segunda" specimenCount="1" specimenCount-female="1" stateProvince="Cartago" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Costa Rica" name="Cartago">Cartago</collectingRegion>
,
<collectorName>
P. N.
<normalizedToken originalValue="Tapantí">Tapanti</normalizedToken>
</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:C82D419D46A0EB2C140F0C30E8D34929" country="Costa Rica" name="Quebrada Segunda" stateProvince="Cartago">Quebrada Segunda</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.25" unit="m" value="1250.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.25" unit="m" value="1250.0">1250 m</elevation>
</quantity>
;
<date value="1992-10">
<collectingDate value="1992-10">Oct. 1992</collectingDate>
</date>
;
<collectorName>G. Mora</collectorName>
leg.; L-N 194000_560000 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616323" collectingDate="1991-12" collectionCode="MNCR" collectorName="R. F. Cord. &amp; C. Alvarado" country="Costa Rica" elevation="1050" location="Tierras Morenas" municipality="Rio San Lorenzo" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
,
<collectorName>R. F. Cord.</collectorName>
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
,
<collectingMunicipality>Rio San Lorenzo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:FC69DF57AEF2BFBA8F14143B4B292684" country="Costa Rica" municipality="Rio San Lorenzo" name="Tierras Morenas" stateProvince="Guanacaste">Tierras Morenas</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.05" unit="m" value="1050.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.05" unit="m" value="1050.0">1050 m</elevation>
</quantity>
;
<date value="1991-12">
<collectingDate value="1991-12">Dec. 1991</collectingDate>
</date>
;
<collectorName>C. Alvarado</collectorName>
leg.; L-N 287800_427600 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616308" collectingDate="1992-01" collectionCode="MNCR" collectorName="R. F. Cord. &amp; C. Alvarado" country="Costa Rica" elevation="1050" location="Tierras Morenas" municipality="Rio San Lorenzo" specimenCount="2" specimenCount-male="2" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="male">2♂</specimenCount>
; same data;
<date value="1992-01">
<collectingDate value="1992-01">ene.1992</collectingDate>
</date>
(
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616362" collectingDate="1988-04" collectionCode="MNCR" collectorName="Est. Cacao, SW &amp; Biodiv. Survey" country="Costa Rica" elevation="1200" location="Volcan Cacao" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
,
<collectorName>Est. Cacao, SW</collectorName>
side
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:3B72131D0DBC5850602DE7E530FB282E" country="Costa Rica" name="Volcan Cacao" stateProvince="Guanacaste">
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
Cacao
</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">1000-1400 m</elevation>
</quantity>
;
<date value="1988-04">
<collectingDate value="1988-04">Apr. 1988</collectingDate>
</date>
; GNP.
<collectorName>Biodiv. Survey</collectorName>
exped.; 323300_375700 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616304" collectingDate="1988" collectingDateMax="1989" collectingDateMin="1988" collectionCode="MNCR" collectorName="Est. Cacao, SW &amp; Biodiv. Survey" country="Costa Rica" elevation="1200" location="Volcan Cacao" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
; same data;
<collectingMethod>malaise trap</collectingMethod>
;
<date value="1988" valueMax="1989">
<collectingDate value="1988" valueMax="1989">1988-1989</collectingDate>
</date>
(
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616310" collectingDate="1994-06-06" collectingDateMax="1994-06-26" collectingDateMin="1994-06-06" collectionCode="MNCR" collectorName="K. Taylor" country="Costa Rica" elevation="800" location="Sect. Las Pailas" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
,
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
, A. C.
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:6CA8B4202547E3FEAD9A1AB81E50D0FA" country="Costa Rica" name="Sect. Las Pailas" stateProvince="Guanacaste">Sect. Las Pailas</location>
;
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
</quantity>
;
<date value="1994-06-06" valueMax="1994-06-26" valueMin="1994-06-06">
<collectingDate value="1994-06-06" valueMax="1994-06-26" valueMin="1994-06-06">6-26 June 1994</collectingDate>
</date>
;
<collectorName>K. Taylor</collectorName>
leg.; L N 309500_389500, #3063 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616374" collectingDate="1994-02-08" collectingDateMax="1994-02-10" collectingDateMin="1994-02-08" collectionCode="MNCR" collectorName="Z. Fuentes" country="Costa Rica" elevation="650" location="Tierras Morenas" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:7167F64B86B6CFD7A094F486098038D7" country="Costa Rica" name="Tierras Morenas" stateProvince="Guanacaste">Tierras Morenas</location>
;
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.5" unit="m" value="650.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.5" unit="m" value="650.0">650 m</elevation>
</quantity>
;
<date value="1994-02-08" valueMax="1994-02-10" valueMin="1994-02-08">
<collectingDate value="1994-02-08" valueMax="1994-02-10" valueMin="1994-02-08">8-10 Feb 1994</collectingDate>
</date>
;
<collectorName>Z. Fuentes</collectorName>
leg.; L S 283950_424500, #2616 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616383" collectingDate="1993-11-10" collectingDateMax="1993-11-21" collectingDateMin="1993-11-10" collectionCode="MNCR" collectorName="P. N. Rincon de la Vieja &amp; M. A. Zumbado &amp; D. G. Garcia" country="Costa Rica" elevation="800" location="Est. Las Pailas" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
,
<collectorName>P. N. Rincon de la Vieja</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:356A3749E5EFD87BF3B7A0D0502DA075" country="Costa Rica" name="Est. Las Pailas" stateProvince="Guanacaste">Est. Las Pailas</location>
;
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
</quantity>
;
<date value="1993-11-10" valueMax="1993-11-21" valueMin="1993-11-10">
<collectingDate value="1993-11-10" valueMax="1993-11-21" valueMin="1993-11-10">10-21 Nov. 1993</collectingDate>
</date>
;
<collectorName>M. A. Zumbado</collectorName>
and
<collectorName>
D. G.
<normalizedToken originalValue="García">Garcia</normalizedToken>
</collectorName>
leg.; L N 306300_388600, #2559 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616370" collectingDate="1995-04-12" collectingDateMax="1995-05-04" collectingDateMin="1995-04-12" collectionCode="MNCR" collectorName="K. Taylor" country="Costa Rica" elevation="800" location="Las Pailas" specimenCount="8" specimenCount-female="2" specimenCount-male="6" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="female">2♀</specimenCount>
,
<specimenCount type="male">6♂</specimenCount>
;
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
, sect.
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:09B3944EE6EB41B6705AC622E8C6F6C1" country="Costa Rica" name="Las Pailas" stateProvince="Guanacaste">Las Pailas</location>
,
<locationDeviation location="Rincon de la Vieja">
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.5" unit="km" value="4.5">4.5 km</quantity>
SW
<normalizedToken originalValue="Rincón">Rincon</normalizedToken>
de la Vieja
</locationDeviation>
volcano;
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
</quantity>
;
<date value="1995-04-12" valueMax="1995-05-04" valueMin="1995-04-12">
<collectingDate value="1995-04-12" valueMax="1995-05-04" valueMin="1995-04-12">12 Apr.-4 May 1995</collectingDate>
</date>
;
<collectorName>K. Taylor</collectorName>
leg.; L_N_306300_388600, #6194 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616378" collectingDate="1992-08" collectionCode="MNCR" collectorName="Z. P. Tenorio &amp; G. Rodriguez" country="Costa Rica" elevation="1050" location="Tierras Morenas" municipality="Rio San Lorenzo" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
,
<collectorName>Z. P. Tenorio</collectorName>
,
<collectingMunicipality>Rio San Lorenzo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:48FEDB30D1E5857E93E912F617A27686" country="Costa Rica" municipality="Rio San Lorenzo" name="Tierras Morenas" stateProvince="Guanacaste">Tierras Morenas</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.05" unit="m" value="1050.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.05" unit="m" value="1050.0">1050 m</elevation>
</quantity>
;
<date value="1992-08">
<collectingDate value="1992-08">Aug. 1992</collectingDate>
</date>
;
<collectorName>G. Rodriguez</collectorName>
leg.; L-N 287800_427600 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616334" collectingDate="1992-10" collectionCode="MNCR" collectorName="Z. P. Tenorio &amp; G. Rodriguez" country="Costa Rica" elevation="1050" location="Tierras Morenas" municipality="Rio San Lorenzo" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
; same data,
<date value="1992-10">
<collectingDate value="1992-10">Oct. 1992</collectingDate>
</date>
(
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616355" collectingDate="1992-03-28" collectingDateMax="1992-04-21" collectingDateMin="1992-03-28" collectionCode="MNCR" collectorName="Z. P. Tenorio &amp; M. Segura" country="Costa Rica" elevation="1050" location="Tierras Morenas" municipality="Rio San Lorenzo" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
,
<collectorName>Z. P. Tenorio</collectorName>
,
<collectingMunicipality>Rio San Lorenzo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:FD7E473F54CD021F7883B7AFF65BF2BC" country="Costa Rica" municipality="Rio San Lorenzo" name="Tierras Morenas" stateProvince="Guanacaste">Tierras Morenas</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.05" unit="m" value="1050.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.05" unit="m" value="1050.0">1050 m</elevation>
</quantity>
;
<date value="1992-03-28" valueMax="1992-04-21" valueMin="1992-03-28">
<collectingDate value="1992-03-28" valueMax="1992-04-21" valueMin="1992-03-28">28 Mar.-21 Apr. 1992</collectingDate>
</date>
;
<collectorName>M. Segura</collectorName>
leg.; L-N 287800_427600 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616345" collectingDate="1994-02" collectionCode="MNCR" collectorName="Z. Fuentes" country="Costa Rica" elevation="1175" location="San Luis" municipality="Monteverde" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
,
<collectingMunicipality>Monteverde</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:533EE108E7F9C947956C0A2F90ADDC41" country="Costa Rica" municipality="Monteverde" name="San Luis" stateProvince="Puntarenas">San Luis</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.35" metricValueMin="1.0" unit="m" value="1175.0" valueMax="1350.0" valueMin="1000.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.35" metricValueMin="1.0" unit="m" value="1175.0" valueMax="1350.0" valueMin="1000.0">1000-1350 m</elevation>
</quantity>
,
<date value="1994-02">
<collectingDate value="1994-02">Feb. 1994</collectingDate>
</date>
;
<collectorName>Z. Fuentes</collectorName>
leg.; L-N 449250_250850, #2615 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616357" collectingDate="1992-08-24" collectingDateMax="1992-09-15" collectingDateMin="1992-08-24" collectionCode="MNCR" collectorName="F. A. Quesada" country="Costa Rica" elevation="1040" location="San Luis" municipality="Monteverde" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
,
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
,
<collectingMunicipality>Monteverde</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:9F3B59490A0C1AE5ADA7EF16BB822951" country="Costa Rica" municipality="Monteverde" name="San Luis" stateProvince="Puntarenas">San Luis</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.04" unit="m" value="1040.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.04" unit="m" value="1040.0">1040 m</elevation>
</quantity>
;
<date value="1992-08-24" valueMax="1992-09-15" valueMin="1992-08-24">
<collectingDate value="1992-08-24" valueMax="1992-09-15" valueMin="1992-08-24">24 Aug.-15 Sep. 1992</collectingDate>
</date>
;
<collectorName>F. A. Quesada</collectorName>
leg.; L-N 250850_449250 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616391" collectingDate="1992-07" collectionCode="MNCR" collectorName="R. B. Monteverde &amp; N. Obando" country="Costa Rica" elevation="1520" location="La Casona" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
,
<collectorName>R. B. Monteverde</collectorName>
, est.
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:384458ADEC4A90E449EAB3A4094B2230" country="Costa Rica" name="La Casona" stateProvince="Puntarenas">La Casona</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.52" unit="m" value="1520.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.52" unit="m" value="1520.0">1520 m</elevation>
</quantity>
;
<date value="1992-07">
<collectingDate value="1992-07">July 1992</collectingDate>
</date>
;
<collectorName>N. Obando</collectorName>
leg.; L N 253200_449700 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063616309" collectingDate="2008-12-05" collectionCode="MNCR" collectorName="A. S. Saenz" country="Costa Rica" elevation="1400" location="San Jeronimo" municipality="Moravia" specimenCount="1" specimenCount-female="1" stateProvince="San Jose" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Costa Rica" name="San Jose">
San
<normalizedToken originalValue="José">Jose</normalizedToken>
</collectingRegion>
,
<collectingMunicipality>Moravia</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:FC48067DDD2D539009F86EF402D3D4FE" country="Costa Rica" municipality="Moravia" name="San Jeronimo" stateProvince="San Jose">San Jeronimo</location>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</elevation>
</quantity>
;
<date value="2008-12-05">
<collectingDate value="2008-12-05">05 Dec. 2008</collectingDate>
</date>
;
<collectorName>A. S. Saenz</collectorName>
leg.; L N 222050_535800, #95353 (
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="diagnosis">
<paragraph pageId="0" pageNumber="157">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="157">Dorsal color yellow with black spots. Male abdomen with several ventrites depressed and emarginate. Penis guide of male genitalia with an apical triangular projection, penis with long slender alae.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="description">
<paragraph pageId="0" pageNumber="157">Description.</paragraph>
<paragraph pageId="0" pageNumber="157">
<emphasis bold="true" pageId="0" pageNumber="157">
<emphasis italics="true" pageId="0" pageNumber="157">
<typeStatus>Holotype</typeStatus>
</emphasis>
.
</emphasis>
Male. Length
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.6" unit="mm" value="3.6">3.6 mm</quantity>
, width 2.0 mm; body oval, convex. Dorsal color yellow except pronotum, with a small basal black spot on each side of the disk center, a small black basal spot and a black subquadrate spot on each side at
<normalizedToken originalValue="½">1/2</normalizedToken>
from the lateral margin; scutellar shield black; each elytron with five black spots: scutellar spot slightly oval, sub-humeral spot large and oval, mid-lateral spot at the apical half of the elytron, discal and sub-apical spots rounded, the spots do not touch the elytral margin, the elytral suture or the anterior border (Fig.
<figureCitation captionStart="Figures 5964" captionStartId="F14" captionText="Figures 59 - 64. Brachiacantha invertita sp. nov., male 59 Dorsal view 60 Abdomen 61 Tegmen 62 Tegmen lateral view 63 Penis 64 Penis apex." figureDoi="10.3897/zookeys.1024.56927.figures59-64" httpUri="https://binary.pensoft.net/fig/520879" pageId="0" pageNumber="157">59</figureCitation>
). Ventral surface with head, prosternum, metaventrite and abdomen black; legs, mouthparts, and antennae yellow. Head punctures small, separated by their diameter, each puncture slightly larger than an eye facet; pronotal punctures larger than head punctures, separated by their diameter; elytral punctures as large as pronotal punctures, separated by their diameter; metaventral punctures larger than elytral punctures, separated by more than three diameters at center, about one diameter on sides. Clypeus straight. Carinae on prosternal process curved, divergent, not joined, larger than half the prosternum. Protibia not flanged, basal tooth small. Basal abdominal ventrite without setal tuft. Abdomen with postcoxal line on basal abdominal ventrite slightly flattened along posterior ventrite margin; ventrite with sparse, long pubescence and small, dense punctures. Ventrites III-VI depressed and emarginated (Fig.
<figureCitation captionStart="Figures 5964" captionStartId="F14" captionText="Figures 59 - 64. Brachiacantha invertita sp. nov., male 59 Dorsal view 60 Abdomen 61 Tegmen 62 Tegmen lateral view 63 Penis 64 Penis apex." figureDoi="10.3897/zookeys.1024.56927.figures59-64" httpUri="https://binary.pensoft.net/fig/520879" pageId="0" pageNumber="157">60</figureCitation>
). Ventrites II-VI pubescent throughout, punctures fine, dense; third ventrite with a tubercle on each side of middle. Genitalia with penis guide as long as parameres, symmetrical, sides slightly convergent, apex almost truncated but with a triangular appendage (Fig.
<figureCitation captionStart="Figures 5964" captionStartId="F14" captionText="Figures 59 - 64. Brachiacantha invertita sp. nov., male 59 Dorsal view 60 Abdomen 61 Tegmen 62 Tegmen lateral view 63 Penis 64 Penis apex." figureDoi="10.3897/zookeys.1024.56927.figures59-64" httpUri="https://binary.pensoft.net/fig/520879" pageId="0" pageNumber="157">61</figureCitation>
); parameres narrowed to apex, apex acute, setae rising from the convex side border (Fig.
<figureCitation captionStart="Figures 5964" captionStartId="F14" captionText="Figures 59 - 64. Brachiacantha invertita sp. nov., male 59 Dorsal view 60 Abdomen 61 Tegmen 62 Tegmen lateral view 63 Penis 64 Penis apex." figureDoi="10.3897/zookeys.1024.56927.figures59-64" httpUri="https://binary.pensoft.net/fig/520879" pageId="0" pageNumber="157">62</figureCitation>
); penis curved in basal
<normalizedToken originalValue="½">1/2</normalizedToken>
, apex with long slender alae, basal capsule crested, inner arm of basal capsule long and slender (Figs
<figureCitation captionStart="Figures 5964" captionStartId="F14" captionText="Figures 59 - 64. Brachiacantha invertita sp. nov., male 59 Dorsal view 60 Abdomen 61 Tegmen 62 Tegmen lateral view 63 Penis 64 Penis apex." figureDoi="10.3897/zookeys.1024.56927.figures59-64" httpUri="https://binary.pensoft.net/fig/520879" pageId="0" pageNumber="157">63</figureCitation>
,
<figureCitation captionStart="Figures 5964" captionStartId="F14" captionText="Figures 59 - 64. Brachiacantha invertita sp. nov., male 59 Dorsal view 60 Abdomen 61 Tegmen 62 Tegmen lateral view 63 Penis 64 Penis apex." figureDoi="10.3897/zookeys.1024.56927.figures59-64" httpUri="https://binary.pensoft.net/fig/520879" pageId="0" pageNumber="157">64</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="157">Female. Similar to male except abdominal characters. Genitalia not examined.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="variation">
<paragraph pageId="0" pageNumber="157">Variation.</paragraph>
<paragraph pageId="0" pageNumber="157">
Males. Length
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.9-3.1 mm</quantity>
, width
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2-2.7 mm</quantity>
. Females. Length
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.8" metricValueMin="2.6" unit="mm" value="3.2" valueMax="3.8" valueMin="2.6">2.6-3.8 mm</quantity>
, width
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.9" metricValueMin="2.0" unit="mm" value="2.45" valueMax="2.9" valueMin="2.0">2-2.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="etymology">
<paragraph pageId="0" pageNumber="157">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="157">
From the Latin
<emphasis italics="true" pageId="0" pageNumber="157">inversos</emphasis>
(= invert). The name refers to the dorsal color pattern, yellow with five black spots, the inverse of that of many
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha</emphasis>
</taxonomicName>
species (black with five yellow spots).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="type locality">
<paragraph pageId="0" pageNumber="157">
<typeStatus>Type</typeStatus>
locality.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<materialsCitation ID-GBIF-Occurrence="3063616313" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Costa Rica" latitude="10.286389" location="San Luis" longLatPrecision="21" longitude="-84.80139" municipality="Monteverde" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<collectingCountry name="Costa Rica">Costa Rica</collectingCountry>
,
<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
:
<collectingMunicipality>Monteverde</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:477D70860D1E55ED819E0194624AC020:870C56DDBF6B00317EEE4E7B9FC97326" country="Costa Rica" latitude="10.286389" longLatPrecision="21" longitude="-84.80139" municipality="Monteverde" name="San Luis" stateProvince="Puntarenas">San Luis</location>
,
<geoCoordinate degrees="10" direction="north" minutes="17" orientation="latitude" precision="15" seconds="11" value="10.286389">10°17'11&quot;N</geoCoordinate>
,
<geoCoordinate degrees="84" direction="west" minutes="48" orientation="longitude" precision="15" seconds="5" value="-84.80139">84°48'5&quot;W</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="distribution">
<paragraph pageId="0" pageNumber="157">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="157">
<collectingCountry name="Costa Rica">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="discussion">
<paragraph pageId="0" pageNumber="157">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="157">This species is easily identifiable by the dorsal coloration, yellow with black spots.</paragraph>
</subSubSection>
</treatment>
</document>